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<document id="5891C586425586FE898593EC8F052C40" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E5406CC7BFF06FC2EFA44FD00" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium Gravenhorst 1802" docType="key" docVersion="1" lastPageNumber="187" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="186" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="131218224C8A159D6F81FF9C3F8A266E">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<treatment id="038DBF6E5406CC7BFF06FC2EFA44FD00" LSID="urn:lsid:plazi:treatment:038DBF6E5406CC7BFF06FC2EFA44FD00" httpUri="http://treatment.plazi.org/id/038DBF6E5406CC7BFF06FC2EFA44FD00" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186">
<subSubSection id="C33E5DF35406CC7CFF06FC2EFE46FB93" box="[151,414,1014,1041]" pageId="35" pageNumber="186" type="nomenclature">
<paragraph id="8B9B0E785406CC7CFF06FC2EFE46FB93" blockId="35.[151,414,1014,1041]" box="[151,414,1014,1041]" pageId="35" pageNumber="186">
<heading id="D0D3B9145406CC7CFF06FC2EFE46FB93" bold="true" box="[151,414,1014,1041]" fontSize="11" level="1" pageId="35" pageNumber="186" reason="1">
<emphasis id="B950D26A5406CC7CFF06FC2EFE46FB93" bold="true" box="[151,414,1014,1041]" pageId="35" pageNumber="186">
3.2.5.
<taxonomicName id="4C2475FB5406CC7CFF70FC2EFE96FB92" ID-CoL="8VRX7" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[225,334,1014,1040]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="genus">Rivulare</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35406CC7CFF06FBE6FD12FBFF" pageId="35" pageNumber="186" type="diagnosis">
<paragraph id="8B9B0E785406CC7CFF06FBE6FD12FBFF" blockId="35.[151,1437,1086,1509]" pageId="35" pageNumber="186">
<emphasis id="B950D26A5406CC7CFF06FBE6FECAFBDA" bold="true" box="[151,274,1086,1112]" pageId="35" pageNumber="186">Diagnosis.</emphasis>
Median lobe narrow, in middle part significantly narrower than basal portion; parameres complicated, broadened apically, usually with long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35406CC7CFF56FB5EFBEDFB47" pageId="35" pageNumber="186" type="taxon_list">
<paragraph id="8B9B0E785406CC7CFF56FB5EFBEDFB47" blockId="35.[151,1437,1086,1509]" pageId="35" pageNumber="186">
<emphasis id="B950D26A5406CC7CFF56FB5EFE4EFB22" bold="true" box="[199,406,1158,1184]" pageId="35" pageNumber="186">Species included:</emphasis>
<taxonomicName id="4C2475FB5406CC7CFE0CFB5FFDBEFB22" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[413,614,1159,1185]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="angustissimum" status="sp. n.">
<emphasis id="B950D26A5406CC7CFE0CFB5FFDBEFB22" box="[413,614,1159,1185]" italics="true" pageId="35" pageNumber="186">O. angustissimum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5406CC7CFDFFFB5FFD76FB23" bold="true" box="[622,686,1159,1185]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="A2636F115406CC7CFDFFFB5FFD76FB23" box="[622,686,1159,1185]" pageId="35" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2475FB5406CC7CFD2DFB5FFC27FB23" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[700,1023,1159,1185]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5406CC7CFD2DFB5FFC9FFB22" box="[700,839,1159,1185]" italics="true" pageId="35" pageNumber="186">O. cariosum</emphasis>
<bibRefCitation id="EFB573895406CC7CFCDFFB5FFC27FB23" author="Cameron, M." box="[846,1023,1159,1185]" pageId="35" pageNumber="186" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2475FB5406CC7CFB9BFB5FFB0DFB23" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1034,1237,1159,1185]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
<emphasis id="B950D26A5406CC7CFB9BFB5FFB0DFB23" box="[1034,1237,1159,1185]" italics="true" pageId="35" pageNumber="186">O. flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5406CC7CFB4DFB5FFAC4FB23" bold="true" box="[1244,1308,1159,1185]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="A2636F115406CC7CFB4DFB5FFAC4FB23" box="[1244,1308,1159,1185]" pageId="35" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2475FB5406CC7CFABBFB5FFA44FB23" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1322,1436,1159,1185]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="kumari" status="sp. n.">
<emphasis id="B950D26A5406CC7CFABBFB5FFA44FB23" box="[1322,1436,1159,1185]" italics="true" pageId="35" pageNumber="186">O. kumari</emphasis>
</taxonomicName>
<emphasis id="B950D26A5406CC7CFF06FB73FF0FFB47" bold="true" box="[151,215,1195,1221]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="A2636F115406CC7CFF06FB73FF0FFB47" box="[151,215,1195,1221]" pageId="35" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2475FB5406CC7CFF75FB73FEA3FB47" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[228,379,1195,1221]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="latocavum" status="sp. n.">
<emphasis id="B950D26A5406CC7CFF75FB73FEA3FB47" box="[228,379,1195,1221]" italics="true" pageId="35" pageNumber="186">O. latocavum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5406CC7CFE13FB73FE1AFB47" bold="true" box="[386,450,1195,1221]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="A2636F115406CC7CFE13FB73FE1AFB47" box="[386,450,1195,1221]" pageId="35" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2475FB5406CC7CFE5EFB73FD31FB47" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[463,745,1195,1221]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B950D26A5406CC7CFE5EFB73FD9AFB46" box="[463,578,1195,1221]" italics="true" pageId="35" pageNumber="186">O. nigrum</emphasis>
<bibRefCitation id="EFB573895406CC7CFDD8FB73FD31FB47" author="Coiffait, H." box="[585,745,1195,1221]" pageId="35" pageNumber="186" pagination="21 - 179" refId="ref32915" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179. [1981]" type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2475FB5406CC7CFD65FB73FBE9FB47" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[756,1073,1195,1221]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="secretum">
<emphasis id="B950D26A5406CC7CFD65FB73FCA2FB46" box="[756,890,1195,1221]" italics="true" pageId="35" pageNumber="186">O. secretum</emphasis>
<bibRefCitation id="EFB573895406CC7CFC10FB73FBE9FB47" author="Cameron, M." box="[897,1073,1195,1221]" pageId="35" pageNumber="186" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35406CC7CFF56FB17FF38FA67" pageId="35" pageNumber="186" type="discussion">
<paragraph id="8B9B0E785406CC7CFF56FB17FF38FA67" blockId="35.[151,1437,1086,1509]" pageId="35" pageNumber="186">
<emphasis id="B950D26A5406CC7CFF56FB17FEE3FB6B" bold="true" box="[199,315,1231,1257]" pageId="35" pageNumber="186">Remarks.</emphasis>
The Rivulare group was originally erected for eight Western Palaearctic species by
<bibRefCitation id="EFB573895406CC7CFB62FB17FA40FB6B" author="Zanetti, A." box="[1267,1432,1231,1257]" pageId="35" pageNumber="186" pagination="1 - 472" refId="ref33896" refString="Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472." type="book chapter" year="1987">Zanetti (1987)</bibRefCitation>
. Later, the same author(Zanetti 2022)added to this group several additional species from
<collectingCountry id="F3334EE85406CC7CFBC0FB2BFB7DFA8F" box="[1105,1189,1267,1293]" name="Cyprus" pageId="35" pageNumber="186">Cyprus</collectingCountry>
,
<collectingCountry id="F3334EE85406CC7CFB3DFB2BFB24FA8F" box="[1196,1276,1267,1293]" name="Turkey" pageId="35" pageNumber="186">Turkey</collectingCountry>
and Caucasus. The Himalayan species have deeper and broader anteocellar foveae usually distinctly divergent lateroapicad about as that in species of the Oxyacanthae group (see above). Based on the general shape of the aedeagus, species of the Rivulare group differs from other groups by the distinctly narrower median lobe. Two species, originally described from
<collectingCountry id="F3334EE85406CC7CFF47FA5BFEC1FA1F" box="[214,281,1411,1437]" name="Japan" pageId="35" pageNumber="186">Japan</collectingCountry>
,
<taxonomicName id="4C2475FB5406CC7CFEB4FA5BFD97FA1F" authority="Sharp, 1874" authorityName="Sharp" authorityYear="1874" box="[293,591,1411,1437]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis id="B950D26A5406CC7CFEB4FA5BFEE1FA1F" box="[293,313,1411,1437]" italics="true" pageId="35" pageNumber="186">O</emphasis>
.
<emphasis id="B950D26A5406CC7CFED9FA5BFE66FA1E" box="[328,446,1411,1436]" italics="true" pageId="35" pageNumber="186">japonicum</emphasis>
Sharp, 1874
</taxonomicName>
(this species is also known from
<collectingCountry id="F3334EE85406CC7CFC5CFA5BFBC9FA1F" box="[973,1041,1411,1437]" name="South Korea" pageId="35" pageNumber="186">Korea</collectingCountry>
and north-western
<collectingCountry id="F3334EE85406CC7CFB7FFA5BFAE9FA1F" box="[1262,1329,1411,1437]" name="China" pageId="35" pageNumber="186">China</collectingCountry>
(
<bibRefCitation id="EFB573895406CC7CFAD1FA5BFE81FA43" author="Schulke, M. &amp; Smetana, A." pageId="35" pageNumber="186" refId="ref33396" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015</bibRefCitation>
)) and
<taxonomicName id="4C2475FB5406CC7CFE36FA7FFD08FA43" authority="Sharp, 1889" authorityName="Sharp" authorityYear="1889" box="[423,720,1447,1473]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="niponense">
<emphasis id="B950D26A5406CC7CFE36FA7FFE63FA43" box="[423,443,1447,1473]" italics="true" pageId="35" pageNumber="186">O</emphasis>
.
<emphasis id="B950D26A5406CC7CFE5AFA7FFDE5FA42" box="[459,573,1447,1472]" italics="true" pageId="35" pageNumber="186">niponense</emphasis>
Sharp, 1889
</taxonomicName>
belongs to this group (see figures of the aedeagi in
<bibRefCitation id="EFB573895406CC7CFABEFA7FFF0BFA67" author="Watanabe, Y." pageId="35" pageNumber="186" pagination="59 - 391" refId="ref33860" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35406CC7CFF06F9EEFC9DF9D2" box="[151,837,1590,1617]" pageId="35" pageNumber="186" type="multiple">
<paragraph id="8B9B0E785406CC7CFF06F9EEFC9DF9D2" blockId="35.[151,837,1590,1617]" box="[151,837,1590,1617]" pageId="35" pageNumber="186">
<heading id="D0D3B9145406CC7CFF06F9EEFC9DF9D2" bold="true" box="[151,837,1590,1617]" fontSize="11" level="1" pageId="35" pageNumber="186" reason="1">
<emphasis id="B950D26A5406CC7CFF06F9EEFC9DF9D2" bold="true" box="[151,837,1590,1617]" pageId="35" pageNumber="186">3.2.5.1. Key to Rivulare group of the Himalayan region</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35406CC7BFF06F9A4FA44FD00" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186" type="key">
<key id="FFB559495406CC7BFF06F9A4FA44FD00" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186">
<keyStep id="30D016DD5406CC7CFF06F9A4FA44F96B" pageId="35" pageNumber="186">
<paragraph id="8B9B0E785406CC7CFF06F9A4FA44F92C" blockId="35.[151,1437,1660,2034]" pageId="35" pageNumber="186">
<keyLead id="30D5AD4D5406CC7CFF06F9A4FA44F92C" pageId="35" pageNumber="186">1 Temples short, about twice as long or shorter than longitudinal length of eyes, gradually narrowed toward nuchal constriction ................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B9B0E785406CC7CFF06F96FFA44F96B" blockId="35.[151,1437,1660,2034]" pageId="35" pageNumber="186">
<keyLead id="30D5AD4D5406CC7CFF06F96FFA44F96B" pageId="35" pageNumber="186">- Temples moderately long, distinctly less than twice as long as longitudinal length of eyes, with convex and widely rounded temples............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D016DD5406CC7CFF06F929FA44F851" pageId="35" pageNumber="186">
<paragraph id="8B9B0E785406CC7CFF06F929FA44F8DC" blockId="35.[151,1437,1660,2034]" pageId="35" pageNumber="186">
<keyLead id="30D5AD4D5406CC7CFF06F929FA44F8DC" pageId="35" pageNumber="186">
2 Temples very short, distinctly more than twice shorter than longitudinal length of eyes. Pronotum widest in middle. Elytra about as long as broad. Median lobe moderately short and narrow; parameres short and very wide (
<figureCitation id="131F12FD5406CC7CFBDFF8D6FB4DF8A1" box="[1102,1173,1806,1828]" captionStart="FIGURES 73–75" captionStartId="40.[151,264,1430,1454]" captionTargetBox="[169,1433,187,1403]" captionTargetId="figure-22@40.[151,1436,181,1406]" captionTargetPageId="40" captionText="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="35" pageNumber="186">Fig. 73</figureCitation>
); apical part of median lobe forming subacute tooth if see laterally as in
<figureCitation id="131F12FD5406CC7CFD14F8F4FD14F8C3" box="[645,716,1835,1858]" captionStart="FIGURES 73–75" captionStartId="40.[151,264,1430,1454]" captionTargetBox="[169,1433,187,1403]" captionTargetId="figure-22@40.[151,1436,181,1406]" captionTargetPageId="40" captionText="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="35" pageNumber="186">Fig. 75</figureCitation>
. Female accessory sclerite as in
<figureCitation id="131F12FD5406CC7CFB90F8F4FB90F8C0" box="[1025,1096,1836,1858]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="35" pageNumber="186">Fig. 76</figureCitation>
. Body reddish-brown, with yellow-brown elytra. Habitus as in
<figureCitation id="131F12FD5406CC7CFE7AF891FDEAF8DC" box="[491,562,1865,1887]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="35" pageNumber="186">Fig. 70</figureCitation>
. Body length:
<quantity id="4CDCA39D5406CC7CFD2FF891FC94F8DC" box="[702,844,1865,1887]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.15" metricValueMin="2.85" pageId="35" pageNumber="186" unit="mm" value="3.0" valueMax="3.15" valueMin="2.85">2.85–3.15 mm</quantity>
.
<collectingCountry id="F3334EE85406CC7CFCC4F891FC5EF8DD" box="[853,902,1865,1887]" name="India" pageId="35" pageNumber="186">India</collectingCountry>
.....................
<taxonomicName id="4C2475FB5406CC7CFB1DF891FA44F8DC" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[1164,1436,1865,1887]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5406CC7CFB1DF891FADAF8DD" box="[1164,1282,1865,1887]" italics="true" pageId="35" pageNumber="186">O. cariosum</emphasis>
<bibRefCitation id="EFB573895406CC7CFA99F891FA44F8DC" author="Cameron, M." box="[1288,1436,1865,1886]" pageId="35" pageNumber="186" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9B0E785406CC7CFF06F8BFFA44F851" blockId="35.[151,1437,1660,2034]" pageId="35" pageNumber="186">
<keyLead id="30D5AD4D5406CC7CFF06F8BFFA44F851" pageId="35" pageNumber="186">
- Temples slightly longer, about twice shorter than longitudinal length of eyes. Pronotum widest in apical third. Elytra slightly broader than long. Median lobe elongate and moderately wide; parameres longer, distinctly narrowed in preapical portion (
<figureCitation id="131F12FD5406CC7CFAE4F85CFF25F834" captionStart="FIGURES 84–86" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 84–86. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="35" pageNumber="186">Fig. 84</figureCitation>
); apical part of median lobe without tooth if see laterally as in
<figureCitation id="131F12FD5406CC7CFCC1F879FC4EF835" box="[848,918,1953,1975]" captionStart="FIGURES 84–86" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 84–86. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="35" pageNumber="186">Fig. 86</figureCitation>
. Female accessory sclerite as in
<figureCitation id="131F12FD5406CC7CFB5BF879FAC9F834" box="[1226,1297,1953,1975]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="35" pageNumber="186">Fig. 77</figureCitation>
. Body yellow-brown, with darker head. Habitus as in
<figureCitation id="131F12FD5406CC7CFDCDF866FD7BF851" box="[604,675,1982,2004]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="35" pageNumber="186">Fig. 71</figureCitation>
. Body length:
<quantity id="4CDCA39D5406CC7CFCBFF866FC64F856" box="[814,956,1982,2004]" metricMagnitude="-3" metricUnit="m" metricValue="3.02" metricValueMax="3.55" metricValueMin="2.49" pageId="35" pageNumber="186" unit="mm" value="3.02" valueMax="3.55" valueMin="2.49">2.49–3.55 mm</quantity>
.
<collectingCountry id="F3334EE85406CC7CFC57F866FC22F856" box="[966,1018,1982,2004]" name="India" pageId="35" pageNumber="186">India</collectingCountry>
,
<collectingCountry id="F3334EE85406CC7CFB92F866FBE4F856" box="[1027,1084,1982,2004]" name="Nepal" pageId="35" pageNumber="186">Nepal</collectingCountry>
..........
<taxonomicName id="4C2475FB5406CC7CFB22F867FAB8F851" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1203,1376,1982,2004]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
<emphasis id="B950D26A5406CC7CFB22F867FAB8F851" box="[1203,1376,1982,2004]" italics="true" pageId="35" pageNumber="186">O. flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5406CC7CFAF7F866FA44F851" bold="true" box="[1382,1436,1982,2003]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="A2636F115406CC7CFAF7F866FA44F851" box="[1382,1436,1982,2003]" pageId="35" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D016DD5406CC7BFF06F804FA44FF65" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186">
<paragraph id="8B9B0E785406CC7BFF06F804FA44FF48" blockId="35.[151,1437,1660,2034]" lastBlockId="36.[151,1437,151,643]" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186">
<keyLead id="30D5AD4D5406CC7BFF06F804FA44FF48" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186">
3 Pronotum widest in apical third. Median lobe with distinctly widened portion in middle, from middle strongly narrowed apicad and from preapical part slightly broadened toward apex (
<figureCitation id="131F12FD5401CC7BFD6EFF4FFC9EFF2F" box="[767,838,151,173]" captionStart="FIGURES 87–89" captionStartId="44.[151,264,1413,1437]" captionTargetBox="[157,1433,190,1385]" captionTargetId="figure-22@44.[151,1436,181,1389]" captionTargetPageId="44" captionText="FIGURES 87–89. Aedeagus of Omalium kumari (holotype): 87—aedeagus, parameral view, 88—aedeagus, lateral view, 89— median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146228" httpUri="https://zenodo.org/record/10146228/files/figure.png" pageId="36" pageNumber="187">Fig. 87</figureCitation>
). Female accessory sclerite as in
<figureCitation id="131F12FD5401CC7BFB13FF4FFB10FF2F" box="[1154,1224,151,173]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="36" pageNumber="187">Fig. 79</figureCitation>
. Body reddish-brown. Habitus as in
<figureCitation id="131F12FD5401CC7BFEF4FF6DFE74FF48" box="[357,428,181,203]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="36" pageNumber="187">Fig. 72</figureCitation>
. Body smaller: 2.40–3.00 mm.
<collectingCountry id="F3334EE85401CC7BFD4BFF6DFCCBFF49" box="[730,787,181,203]" name="Nepal" pageId="36" pageNumber="187">Nepal</collectingCountry>
.........................................
<taxonomicName id="4C2475FB5401CC7BFB6EFF6DFAB8FF48" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1279,1376,181,202]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="kumari" status="sp. n.">
<emphasis id="B950D26A5401CC7BFB6EFF6DFAB8FF48" box="[1279,1376,181,202]" italics="true" pageId="36" pageNumber="187">O. kumari</emphasis>
</taxonomicName>
<emphasis id="B950D26A5401CC7BFAF7FF6DFA44FF48" bold="true" box="[1382,1436,181,202]" pageId="36" pageNumber="187">
<taxonomicNameLabel id="A2636F115401CC7BFAF7FF6DFA44FF48" box="[1382,1436,181,202]" pageId="36" pageNumber="187" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9B0E785401CC7BFF06FF0BFA44FF65" blockId="36.[151,1437,151,643]" box="[151,1436,210,232]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FF0BFA44FF65" box="[151,1436,210,232]" pageId="36" pageNumber="187">- Pronotum widest in or slightly above middle. Median lobe different............................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D016DD5401CC7BFF06FF28FA44FEA1" pageId="36" pageNumber="187">
<paragraph id="8B9B0E785401CC7BFF06FF28FA44FE87" blockId="36.[151,1437,151,643]" box="[151,1436,239,261]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FF28FA44FE87" box="[151,1436,239,261]" pageId="36" pageNumber="187">4 Pronotum gradually narrowed both apicad and posteriad...................................................... 5</keyLead>
</paragraph>
<paragraph id="8B9B0E785401CC7BFF06FED5FA44FEA1" blockId="36.[151,1437,151,643]" box="[151,1436,269,291]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FED5FA44FEA1" box="[151,1436,269,291]" pageId="36" pageNumber="187">- Pronotum more narrowed posteriad than apicad............................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D016DD5401CC7BFF06FEF2FA44FD8F" pageId="36" pageNumber="187">
<paragraph id="8B9B0E785401CC7BFF06FEF2FA44FE15" blockId="36.[151,1437,151,643]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FEF2FA44FE15" pageId="36" pageNumber="187">
5 Elytra slightly less than twice as long as pronotum. Punctation of pronotum and elytra very dense and coarse. Median lobe spear-shaped, slightly broadened in preapical portion; apical portions of parameres very wide (
<figureCitation id="131F12FD5401CC7BFB11FE9FFB11FEDF" box="[1152,1225,327,349]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="36" pageNumber="187">Fig. 93</figureCitation>
); preapical portion of median lobe with distinct obtuse hook, if see laterally as in
<figureCitation id="131F12FD5401CC7BFCA3FEBDFCA3FEF9" box="[818,891,357,379]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="36" pageNumber="187">Fig. 95</figureCitation>
. Female accessory sclerite as in
<figureCitation id="131F12FD5401CC7BFB2FFEBDFADEFEF8" box="[1214,1286,357,379]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="36" pageNumber="187">Fig. 81</figureCitation>
. Body reddish-brown. Habitus as in
<figureCitation id="131F12FD5401CC7BFE3FFE5AFE2EFE15" box="[430,502,386,408]" captionStart="FIGURES 90–92" captionStartId="45.[151,264,1333,1357]" captionTargetBox="[164,1418,397,1298]" captionTargetId="figure-104@45.[151,1435,381,1310]" captionTargetPageId="45" captionText="FIGURES 90–92. Habitus of Omalium: 90—O. latocavum (paratype), 91—O. nigrum (holotype), 92—O. secretum (Nepal: Parbat). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146230" httpUri="https://zenodo.org/record/10146230/files/figure.png" pageId="36" pageNumber="187">Fig. 90</figureCitation>
. Body larger:
<quantity id="4CDCA39D5401CC7BFDECFE5AFCD3FE1A" box="[637,779,386,408]" metricMagnitude="-3" metricUnit="m" metricValue="3.17" metricValueMax="3.28" metricValueMin="3.06" pageId="36" pageNumber="187" unit="mm" value="3.17" valueMax="3.28" valueMin="3.06">3.06–3.28 mm</quantity>
. Nepa.................................
<taxonomicName id="4C2475FB5401CC7BFB4EFE5AFAB8FE15" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1247,1376,386,407]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="latocavum" status="sp. n.">
<emphasis id="B950D26A5401CC7BFB4EFE5AFAB8FE15" box="[1247,1376,386,407]" italics="true" pageId="36" pageNumber="187">O. latocavum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5401CC7BFAF7FE5AFA44FE15" bold="true" box="[1382,1436,386,407]" pageId="36" pageNumber="187">
<taxonomicNameLabel id="A2636F115401CC7BFAF7FE5AFA44FE15" box="[1382,1436,386,407]" pageId="36" pageNumber="187" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9B0E785401CC7BFF06FE78FA44FD8F" blockId="36.[151,1437,151,643]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FE78FA44FD8F" pageId="36" pageNumber="187">
- Elytra about twice as long as pronotum. Punctation of pronotum and elytra sparse and fine. Median lobe narrow, from widest median part gradually narrowed apically; preapical portions of parameres narrow (
<figureCitation id="131F12FD5401CC7BFC6DFE65FB88FE50" box="[1020,1104,445,467]" captionStart="FIGURES 100–102" captionStartId="52.[151,264,1447,1471]" captionTargetBox="[154,1433,185,1420]" captionTargetId="figure-22@52.[151,1436,181,1423]" captionTargetPageId="52" captionText="FIGURES 100–102. Aedeagus of Omalium secretum (Nepal, Parbat): 100—aedeagus, parameral view, 101—aedeagus, lateral view, 102—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146238" httpUri="https://zenodo.org/record/10146238/files/figure.png" pageId="36" pageNumber="187">Fig. 100</figureCitation>
); preapical portion of median lobe without hook, if see laterally as in
<figureCitation id="131F12FD5401CC7BFDAAFE02FD48FE6D" box="[571,656,474,496]" captionStart="FIGURES 100–102" captionStartId="52.[151,264,1447,1471]" captionTargetBox="[154,1433,185,1420]" captionTargetId="figure-22@52.[151,1436,181,1423]" captionTargetPageId="52" captionText="FIGURES 100–102. Aedeagus of Omalium secretum (Nepal, Parbat): 100—aedeagus, parameral view, 101—aedeagus, lateral view, 102—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146238" httpUri="https://zenodo.org/record/10146238/files/figure.png" pageId="36" pageNumber="187">Fig. 102</figureCitation>
. Female accessory sclerite as in
<figureCitation id="131F12FD5401CC7BFC43FE02FBC2FE6D" box="[978,1050,474,496]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="36" pageNumber="187">Fig. 82</figureCitation>
. Body yellow-brown to reddish-brown. Habitus as in
<figureCitation id="131F12FD5401CC7BFEF4FE2FFE74FD8F" box="[357,428,503,525]" captionStart="FIGURES 90–92" captionStartId="45.[151,264,1333,1357]" captionTargetBox="[164,1418,397,1298]" captionTargetId="figure-104@45.[151,1435,381,1310]" captionTargetPageId="45" captionText="FIGURES 90–92. Habitus of Omalium: 90—O. latocavum (paratype), 91—O. nigrum (holotype), 92—O. secretum (Nepal: Parbat). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146230" httpUri="https://zenodo.org/record/10146230/files/figure.png" pageId="36" pageNumber="187">Fig. 92</figureCitation>
. Body length:
<quantity id="4CDCA39D5401CC7BFDA9FE2FFD1EFD8F" box="[568,710,503,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.815" metricValueMax="3.28" metricValueMin="2.35" pageId="36" pageNumber="187" unit="mm" value="2.815" valueMax="3.28" valueMin="2.35">2.35–3.28 mm</quantity>
.
<collectingCountry id="F3334EE85401CC7BFD5EFE2FFCDBFD8F" box="[719,771,503,525]" name="India" pageId="36" pageNumber="187">India</collectingCountry>
,
<collectingCountry id="F3334EE85401CC7BFC9CFE2FFC9EFD8F" box="[781,838,503,525]" name="Nepal" pageId="36" pageNumber="187">Nepal</collectingCountry>
........................................
<taxonomicName id="4C2475FB5401CC7BFAB8FE20FA44FD8F" authorityName="Cameron" authorityYear="1924" box="[1321,1436,504,525]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="secretum">
<emphasis id="B950D26A5401CC7BFAB8FE20FA44FD8F" box="[1321,1436,504,525]" italics="true" pageId="36" pageNumber="187">O. secretum</emphasis>
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</keyStep>
<keyStep id="30D016DD5401CC7BFF06FDCDFA44FD00" pageId="36" pageNumber="187">
<paragraph id="8B9B0E785401CC7BFF06FDCDFA44FDC5" blockId="36.[151,1437,151,643]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FDCDFA44FDC5" pageId="36" pageNumber="187">
6 Apical angles of pronotum not protruded anteriad. Parameres wide (
<figureCitation id="131F12FD5401CC7BFCEAFDCDFC1CFDA9" box="[891,964,533,555]" captionStart="FIGURES 66–68" captionStartId="37.[151,264,1311,1335]" captionTargetBox="[154,1433,184,1284]" captionTargetId="figure-16@37.[151,1436,181,1287]" captionTargetPageId="37" captionText="FIGURES 66–68. Aedeagus of Omalium angustissimum (holotype): 66—aedeagus, parameral view, 67—aedeagus, lateral view, 68—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146216" httpUri="https://zenodo.org/record/10146216/files/figure.png" pageId="36" pageNumber="187">Fig. 66</figureCitation>
). Body dark-brown. Habitus as in
<figureCitation id="131F12FD5401CC7BFA87FDCDFA86FDA9" box="[1302,1374,533,555]" captionStart="FIGURES 56–58" captionStartId="30.[151,264,1283,1307]" captionTargetBox="[163,1420,278,1245]" captionTargetId="figure-52@30.[159,1428,273,1260]" captionTargetPageId="30" captionText="FIGURES 56–58. Habitus of Omalium: 56—O. marginatum (holotype), 57—O. scapulare (Nepal, Bagmati), 58—O. angustissimum (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146208" httpUri="https://zenodo.org/record/10146208/files/figure.png" pageId="36" pageNumber="187">Fig. 58</figureCitation>
. Body length:
<quantity id="4CDCA39D5401CC7BFEBAFDEAFE5BFDCA" box="[299,387,562,584]" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="36" pageNumber="187" unit="mm" value="2.76">2.76 mm</quantity>
.
<collectingCountry id="F3334EE85401CC7BFE1DFDEAFE1DFDCA" box="[396,453,562,584]" name="Nepal" pageId="36" pageNumber="187">Nepal</collectingCountry>
..............................................................
<taxonomicName id="4C2475FB5401CC7BFB25FDEAFAB8FDCA" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1204,1376,562,584]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="angustissimum" status="sp. n.">
<emphasis id="B950D26A5401CC7BFB25FDEAFAB8FDCA" box="[1204,1376,562,584]" italics="true" pageId="36" pageNumber="187">O. angustissimum</emphasis>
</taxonomicName>
<emphasis id="B950D26A5401CC7BFAF7FDEAFA44FDC5" bold="true" box="[1382,1436,562,583]" pageId="36" pageNumber="187">
<taxonomicNameLabel id="A2636F115401CC7BFAF7FDEAFA44FDC5" box="[1382,1436,562,583]" pageId="36" pageNumber="187" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9B0E785401CC7BFF06FD88FA44FD00" blockId="36.[151,1437,151,643]" pageId="36" pageNumber="187">
<keyLead id="30D5AD4D5401CC7BFF06FD88FA44FD00" pageId="36" pageNumber="187">
- Apical angles of pronotum slightly protruded anteriad. Parameres moderately narrow (
<figureCitation id="131F12FD5401CC7BFBA1FD97FBA1FDE7" box="[1072,1145,591,613]" captionStart="FIGURES 96–98" captionStartId="49.[151,264,1529,1553]" captionTargetBox="[161,1433,387,1503]" captionTargetId="figure-101@49.[151,1436,345,1506]" captionTargetPageId="49" captionText="FIGURES 96–98. Aedeagus of Omalium nigrum (Nepal: Mahakali): 96—aedeagus, parameral view, 97—aedeagus, lateral view, 98—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146234" httpUri="https://zenodo.org/record/10146234/files/figure.png" pageId="36" pageNumber="187">Fig. 96</figureCitation>
). Body brown to dark-brown. Habitus as in
<figureCitation id="131F12FD5401CC7BFEF4FDB5FE74FD00" box="[357,428,621,643]" captionStart="FIGURES 90–92" captionStartId="45.[151,264,1333,1357]" captionTargetBox="[164,1418,397,1298]" captionTargetId="figure-104@45.[151,1435,381,1310]" captionTargetPageId="45" captionText="FIGURES 90–92. Habitus of Omalium: 90—O. latocavum (paratype), 91—O. nigrum (holotype), 92—O. secretum (Nepal: Parbat). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146230" httpUri="https://zenodo.org/record/10146230/files/figure.png" pageId="36" pageNumber="187">Fig. 91</figureCitation>
. Body length: 2.80–3.00 mm.
<collectingCountry id="F3334EE85401CC7BFD5EFDB5FCD0FD01" box="[719,776,621,643]" name="Nepal" pageId="36" pageNumber="187">Nepal</collectingCountry>
..............................................
<taxonomicName id="4C2475FB5401CC7BFAABFDB5FA44FD00" authorityName="Coiffait" authorityYear="1982" box="[1338,1436,621,642]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B950D26A5401CC7BFAABFDB5FA44FD00" box="[1338,1436,621,642]" italics="true" pageId="36" pageNumber="187">O. nigrum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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