treatments-xml/data/03/86/C9/0386C943FFFDFFDCD6DBF9519955FA85.xml
2024-06-21 12:22:17 +02:00

325 lines
46 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2903D980A1FF3ACBC638F773D7D8339D" ID-DOI="10.11646/zootaxa.4021.1.10" ID-GBIF-Dataset="149eeecc-8edc-4bc8-9184-d4cc3d532ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="242639" ID-ZooBank="ED10FD99-B9B3-432F-BFD7-14D9A1B80989" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461273865916" checkinUser="plazi" docAuthor="Perreau, Michel &amp; Faille, Arnaud" docDate="2015" docId="0386C943FFFDFFDCD6DBF9519955FA85" docLanguage="en" docName="zt04021p200.pdf" docOrigin="Zootaxa 4021 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptomaphaminus granophilus Perreau &amp; Faille, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="FFBFB13BFFFDFFD8D64CFFD09C7FFFA2" masterDocTitle="One new species of Ptomaphaminus Perreau, 2000 (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from a granitic subterranean environment in Vietnam" masterLastPageNumber="200" masterPageNumber="195" pageNumber="195" updateTime="1698612285074" updateUser="plazi">
<mods:mods id="317F492B2B6B3B8DCB3A84941A4F1A2E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EA76BCA6CA4D4073A0ED6CEEE1765DB7">
<mods:title id="33894009DA1E3507BC5279B5020853E0">One new species of Ptomaphaminus Perreau, 2000 (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from a granitic subterranean environment in Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="C42428C07ECEB7ED41AD4B74E0BDC8E5" type="personal">
<mods:role id="B2B6A3D136477A584C93BEAB7E719F70">
<mods:roleTerm id="135273A264EFECCE0492121390AD4A8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1DFB5B1C825EAEE9874708862588009F">Perreau, Michel</mods:namePart>
</mods:name>
<mods:name id="67E0CE4938C87C0B3FDCBED092E3CE37" type="personal">
<mods:role id="FE51A2D9DEC7DB387B4CF3ED9A6DC06F">
<mods:roleTerm id="9D4CA5243098AC85E7DD1070BCFD3600">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEC8289D12EECAD135BA501DB11A072E">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="7B2EE4C076D85CB03D940EEBD1DB9C6F">text</mods:typeOfResource>
<mods:relatedItem id="56A4B1EE2B891BCFE9B6610624663C38" type="host">
<mods:titleInfo id="A21E03D987DACE2A9CFCCB7843B0D074">
<mods:title id="EA57515A5AE168DAFA8129921B602481">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CA19F3D28003D861C0138946DBE124B9">
<mods:date id="38866A87C83836AEDCAF0F07FB02F8E7">2015</mods:date>
<mods:detail id="974D588AA7974B3F0B91BFD765C5EB2F" type="volume">
<mods:number id="87EA061B3B3A8E2603A2669D412EED42">4021</mods:number>
</mods:detail>
<mods:detail id="8A67A89CC91A8ACFBA53A7FBB92E35E7" type="issue">
<mods:number id="8B3C3689D0896BC75029BE236609E13A">1</mods:number>
</mods:detail>
<mods:extent id="D14A16FF1F42ACEFFC7F4475C1CAB624" unit="page">
<mods:start id="30048EC9245B6DD2B55DF0DD00A533E4">195</mods:start>
<mods:end id="FD42670AE77DA2065F531740605AD2BF">200</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0D63EBE9A02D69DC08B35973ABFEF320">journal article</mods:classification>
<mods:identifier id="E3A30BDFE61331B912B3F47942B64EC3" type="DOI">10.11646/zootaxa.4021.1.10</mods:identifier>
<mods:identifier id="57D4128DD886D2E53237B216694EDFB8" type="GBIF-Dataset">149eeecc-8edc-4bc8-9184-d4cc3d532ed9</mods:identifier>
<mods:identifier id="2DE5343768CDD4A6E15FA3EE0FB11951" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7EDAE14B27D318F6320973B97CB5FF4C" type="Zenodo-Dep">242639</mods:identifier>
<mods:identifier id="5E2AA3939543B426F0EB424C83C21FCD" type="ZooBank">ED10FD99-B9B3-432F-BFD7-14D9A1B80989</mods:identifier>
</mods:mods>
<treatment id="0386C943FFFDFFDCD6DBF9519955FA85" ID-DOI="http://doi.org/10.5281/zenodo.6101407" ID-GBIF-Taxon="119639149" ID-Zenodo-Dep="6101407" LSID="urn:lsid:plazi:treatment:0386C943FFFDFFDCD6DBF9519955FA85" httpUri="http://treatment.plazi.org/id/0386C943FFFDFFDCD6DBF9519955FA85" lastPageId="4" lastPageNumber="199" pageId="0" pageNumber="195">
<subSubSection id="C3352BDEFFFDFFD8D6DBF9519C98F919" pageId="0" pageNumber="195" type="nomenclature">
<paragraph id="8B907855FFFDFFD8D6DBF9519E3EF93E" blockId="0.[151,577,1665,1723]" box="[151,577,1665,1692]" pageId="0" pageNumber="195">
<heading id="D0D8CF39FFFDFFD8D6DBF9519E3EF93E" bold="true" box="[151,577,1665,1692]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
<emphasis id="B95BA447FFFDFFD8D6DBF9519E3EF93E" bold="true" box="[151,577,1665,1692]" pageId="0" pageNumber="195">
<taxonomicName id="4C2F03D6FFFDFFD8D6DBF9519D87F939" ID-CoL="4Q9KD" box="[151,504,1665,1691]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFFDFFD8D6DBF9519D87F939" bold="true" box="[151,504,1665,1691]" italics="true" pageId="0" pageNumber="195">Ptomaphaminus granophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268193CFFFDFFD8D7B3F9529E3EF93E" box="[511,577,1666,1692]" pageId="0" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B907855FFFDFFD8D6DBF9749C98F919" blockId="0.[151,577,1665,1723]" box="[151,231,1700,1723]" pageId="0" pageNumber="195">
(
<figureCitation id="131464D0FFFDFFD8D6D3F9749CA1F919" box="[159,222,1700,1723]" captionStart="FIGURES 1 3" captionStartId="1.[151,258,1978,2000]" captionTargetBox="[173,1405,194,1948]" captionTargetId="figure@1.[173,1406,194,1950]" captionTargetPageId="1" captionText="FIGURES 1 3. Ptomaphaminus granophilus n. sp. 1 - 2: biotope: the granitic blockfield. 3: habitus." httpUri="https://zenodo.org/record/242640/files/figure.png" pageId="0" pageNumber="195">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3352BDEFFFDFFD8D6DBF9379FA3F883" pageId="0" pageNumber="195" type="materials_examined">
<paragraph id="8B907855FFFDFFD8D6DBF9379FA3F883" blockId="0.[151,1436,1767,2033]" pageId="0" pageNumber="195">
<emphasis id="B95BA447FFFDFFD8D6DBF9379D7AF95C" bold="true" box="[151,261,1767,1790]" pageId="0" pageNumber="195">
<typeStatus id="5494C6F7FFFDFFD8D6DBF9379D7EF95C" box="[151,257,1767,1790]" pageId="0" pageNumber="195" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D29B3DCFFFDFFD8D742F9379D24F95C" box="[270,347,1767,1790]" pageId="0" pageNumber="195" type="male">1 male</specimenCount>
,
<collectingCountry id="F33838C5FFFDFFD8D724F9379DBAF95C" box="[360,453,1767,1790]" name="Vietnam" pageId="0" pageNumber="195">Vietnam</collectingCountry>
, granitic blockfield, Hon Ba, Suoi cat,
<quantity id="4CD7D5B0FFFDFFD8D53AF9379FC1F95C" box="[886,958,1767,1790]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="0" pageNumber="195" unit="m" value="650.0">650 m</quantity>
, pitfall baited with vinegar,
<date id="FF915E95FFFDFFD8D2BAF937990BF95C" box="[1270,1396,1767,1790]" pageId="0" pageNumber="195" value="2013-09-20">20.IX 2013</date>
, A. Faille leg., deposited in Museum national d'Histoire naturelle de Paris,
<collectingCountry id="F33838C5FFFDFFD8D5C0F8DA9FA7F883" box="[908,984,1802,1825]" name="France" pageId="0" pageNumber="195">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3352BDEFFFDFFDCD68BF8FC9DF8FDAB" lastPageId="4" lastPageNumber="199" pageId="0" pageNumber="195" type="description">
<paragraph id="8B907855FFFDFFD8D68BF8FC9800F8E1" blockId="0.[151,1436,1767,2033]" box="[199,1151,1836,1859]" pageId="0" pageNumber="195">
<emphasis id="B95BA447FFFDFFD8D68BF8FC9D2DF8E1" bold="true" box="[199,338,1836,1859]" pageId="0" pageNumber="195">Description.</emphasis>
Length:
<quantity id="4CD7D5B0FFFDFFD8D7FEF8FC9E78F8E1" box="[434,519,1836,1859]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="195" unit="mm" value="1.6">1.6 mm</quantity>
. General color light brown, antenna and protarsi yellowish.
</paragraph>
<paragraph id="8B907855FFFDFFD8D68BF89F9E40F809" blockId="0.[151,1436,1767,2033]" pageId="0" pageNumber="195">Head. Eyes reduced, composed of approximately 40 ommatidia. Antennae slender, all antennomeres longer than wide except the eighth. The length of antennomeres (divided by the length of the first one) are: 1.00; 0.80; 0.35; 0.33; 0.30; 0.32; 0.45; 0.27; 0.50; 0.40; 0.87.</paragraph>
<paragraph id="8B907855FFFDFFD8D68BF8679D96F853" blockId="0.[151,1436,1767,2033]" pageId="0" pageNumber="195">Pronotum transverse, 1.6 times wider than long, largest width near the base, sides parallel near the base. Surface covered with transverse strigae.</paragraph>
<caption id="DF5028DDFFFCFFD9D6DBF86A9828F872" box="[151,1111,1978,2000]" httpUri="https://zenodo.org/record/242640/files/figure.png" pageId="1" pageNumber="196" targetBox="[173,1405,194,1948]" targetPageId="1">
<paragraph id="8B907855FFFCFFD9D6DBF86A9828F872" blockId="1.[151,1111,1978,2000]" box="[151,1111,1978,2000]" pageId="1" pageNumber="196">
<emphasis id="B95BA447FFFCFFD9D6DBF86A9D4DF872" bold="true" box="[151,306,1978,2000]" pageId="1" pageNumber="196">FIGURES 13.</emphasis>
<taxonomicName id="4C2F03D6FFFCFFD9D77BF86B9E32F872" box="[311,589,1979,2000]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFFCFFD9D77BF86B9E32F872" box="[311,589,1979,2000]" italics="true" pageId="1" pageNumber="196">Ptomaphaminus granophilus</emphasis>
</taxonomicName>
<emphasis id="B95BA447FFFCFFD9D41FF86A9EC9F872" bold="true" box="[595,694,1978,2000]" pageId="1" pageNumber="196">
<taxonomicNameLabel id="A268193CFFFCFFD9D41FF86A9EF5F872" box="[595,650,1978,2000]" pageId="1" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
1-2:
</emphasis>
biotope: the granitic blockfield.
<emphasis id="B95BA447FFFCFFD9D5BDF86A987BF872" bold="true" box="[1009,1028,1978,2000]" pageId="1" pageNumber="196">3:</emphasis>
habitus.
</paragraph>
</caption>
<caption id="DF5028DDFFFFFFDAD6DBF94F9852F8EA" httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="2" pageNumber="197" targetBox="[166,1414,227,1631]" targetPageId="2">
<paragraph id="8B907855FFFFFFDAD6DBF94F9852F8EA" blockId="2.[151,1437,1695,1864]" pageId="2" pageNumber="197">
<emphasis id="B95BA447FFFFFFDAD6DBF94F9D42F917" bold="true" box="[151,317,1695,1717]" pageId="2" pageNumber="197">FIGURES 416.</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD708F9709E50F917" authority="Perreau" authorityName="Perreau" box="[324,559,1696,1717]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA447FFFFFFDAD708F9709D9FF917" box="[324,480,1696,1717]" italics="true" pageId="2" pageNumber="197">Ptomaphaminus</emphasis>
Perreau
</taxonomicName>
from Vietnam and Laos.
<emphasis id="B95BA447FFFFFFDAD56BF94F9F45F917" bold="true" box="[807,826,1695,1717]" pageId="2" pageNumber="197">4:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD50DF9709FB2F917" box="[833,973,1696,1717]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFFFFFDAD50DF9709FB2F917" box="[833,973,1696,1717]" italics="true" pageId="2" pageNumber="197">P. granophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268193CFFFFFFDAD598F94F9874F917" box="[980,1035,1695,1717]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B95BA447FFFFFFDAD598F94F9874F917" bold="true" box="[980,1035,1695,1717]" pageId="2" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
, aedeagus, dorsal view.
<emphasis id="B95BA447FFFFFFDAD2B9F94F9975F917" bold="true" box="[1269,1290,1695,1717]" pageId="2" pageNumber="197">5:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD35CF97099E3F917" box="[1296,1436,1696,1717]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFFFFFDAD35CF97099E3F917" box="[1296,1436,1696,1717]" italics="true" pageId="2" pageNumber="197">P. granophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268193CFFFFFFDAD6DBF96D9CB1F971" box="[151,206,1725,1747]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B95BA447FFFFFFDAD6DBF96D9CB1F971" bold="true" box="[151,206,1725,1747]" pageId="2" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
, male urite IX.
<emphasis id="B95BA447FFFFFFDAD72BF96D9D05F971" bold="true" box="[359,378,1725,1747]" pageId="2" pageNumber="197">6:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD7CDF96D9E72F970" box="[385,525,1725,1746]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFFFFFDAD7CDF96D9E72F970" box="[385,525,1725,1746]" italics="true" pageId="2" pageNumber="197">P. granophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268193CFFFFFFDAD45FF96D9E35F971" box="[531,586,1725,1747]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B95BA447FFFFFFDAD45FF96D9E35F971" bold="true" box="[531,586,1725,1747]" pageId="2" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
, aedeagus, lateral view.
<emphasis id="B95BA447FFFFFFDAD57AF96D9F35F971" bold="true" box="[822,842,1725,1747]" pageId="2" pageNumber="197">7:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD51DF96D98FEF970" authority="Szymczakowski" authorityName="Szymczakowski" box="[849,1153,1724,1746]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="bihamatus">
<emphasis id="B95BA447FFFFFFDAD51DF96D9FB2F970" box="[849,973,1725,1746]" italics="true" pageId="2" pageNumber="197">P. bihamatus</emphasis>
(Szymczakowski)
</taxonomicName>
, aedeagus, dorsal view (from Szymczakowski, 1972).
<emphasis id="B95BA447FFFFFFDAD7C4F90A9DE4F952" bold="true" box="[392,411,1754,1776]" pageId="2" pageNumber="197">8:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD7E8F90B9EA5F952" authority="Szymczakowski" authorityName="Szymczakowski" box="[420,730,1754,1776]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="bihamatus">
<emphasis id="B95BA447FFFFFFDAD7E8F90B9E5CF952" box="[420,547,1755,1776]" italics="true" pageId="2" pageNumber="197">P. bihamatus</emphasis>
(Szymczakowski)
</taxonomicName>
, aedeagus, lateral view (from Szymczakowski, 1972).
<emphasis id="B95BA447FFFFFFDAD2B5F90A9971F952" bold="true" box="[1273,1294,1754,1776]" pageId="2" pageNumber="197">9:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD35AF90B9C9CF8AF" authority="Perreau" authorityName="Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B95BA447FFFFFFDAD35AF90B99E3F952" box="[1302,1436,1755,1776]" italics="true" pageId="2" pageNumber="197">P. deharvengi</emphasis>
Perreau
</taxonomicName>
, aedeagus, dorsal view.
<emphasis id="B95BA447FFFFFFDAD784F9279D97F8AF" bold="true" box="[456,488,1783,1805]" pageId="2" pageNumber="197">10:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD7A2F9289EBCF8AF" authority="Perreau" authorityName="Perreau" box="[494,707,1784,1805]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B95BA447FFFFFFDAD7A2F9289E0EF8AF" box="[494,625,1784,1805]" italics="true" pageId="2" pageNumber="197">P. deharvengi</emphasis>
Perreau
</taxonomicName>
, aedeagus lateral view.
<emphasis id="B95BA447FFFFFFDAD5E8F9279FBEF8AF" bold="true" box="[932,961,1783,1805]" pageId="2" pageNumber="197">11:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD584F9289800F8AF" authority="Perreau" authorityName="Perreau" box="[968,1151,1784,1805]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B95BA447FFFFFFDAD584F9289852F8AF" box="[968,1069,1784,1805]" italics="true" pageId="2" pageNumber="197">P. bedosae</emphasis>
Perreau
</taxonomicName>
, aedeagus, dorsal view.
<emphasis id="B95BA447FFFFFFDAD328F92799FBF8AF" bold="true" box="[1380,1412,1783,1805]" pageId="2" pageNumber="197">12:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD3C6F9289D48F888" authority="Perreau" authorityName="Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B95BA447FFFFFFDAD3C6F9289C9AF888" italics="true" pageId="2" pageNumber="197">P. bedosae</emphasis>
Perreau
</taxonomicName>
, aedeagus, lateral view.
<emphasis id="B95BA447FFFFFFDAD452F8C59E41F889" bold="true" box="[542,574,1813,1835]" pageId="2" pageNumber="197">13:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD408F8C59E86F888" authority="Jarrige" authorityName="Jarrige" box="[580,761,1812,1834]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="boutini">
<emphasis id="B95BA447FFFFFFDAD408F8C59EDFF888" box="[580,672,1813,1834]" italics="true" pageId="2" pageNumber="197">P. boutini</emphasis>
(Jarrige)
</taxonomicName>
, aedeagus, dorsal view.
<emphasis id="B95BA447FFFFFFDAD593F8C59F80F889" bold="true" box="[991,1023,1813,1835]" pageId="2" pageNumber="197">14:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD249F8C598C5F888" authority="Jarrige" authorityName="Jarrige" box="[1029,1210,1812,1834]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="boutini">
<emphasis id="B95BA447FFFFFFDAD249F8C5981EF888" box="[1029,1121,1813,1834]" italics="true" pageId="2" pageNumber="197">P. boutini</emphasis>
(Jarrige)
</taxonomicName>
, aedeagus, lateral view.
<emphasis id="B95BA447FFFFFFDAD6DBF8E29CC8F8EA" bold="true" box="[151,183,1842,1864]" pageId="2" pageNumber="197">15:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD6F1F8E39D02F8EA" authority="Perreau" authorityName="Perreau" box="[189,381,1842,1864]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="leclerci">
<emphasis id="B95BA447FFFFFFDAD6F1F8E39D64F8EA" box="[189,283,1843,1864]" italics="true" pageId="2" pageNumber="197">P. leclerci</emphasis>
(Perreau)
</taxonomicName>
, aedeagus, dorsal view.
<emphasis id="B95BA447FFFFFFDAD428F8E29EFBF8EA" bold="true" box="[612,644,1842,1864]" pageId="2" pageNumber="197">16:</emphasis>
<taxonomicName id="4C2F03D6FFFFFFDAD4C6F8E39F35F8EA" authority="Perreau" authorityName="Perreau" box="[650,842,1842,1864]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="leclerci">
<emphasis id="B95BA447FFFFFFDAD4C6F8E39E97F8EA" box="[650,744,1843,1864]" italics="true" pageId="2" pageNumber="197">P. leclerci</emphasis>
(Perreau)
</taxonomicName>
, aedeagus, lateral view.
</paragraph>
</caption>
<caption id="DF5028DDFFFEFFDBD6DBF85C9F8FF800" box="[151,1008,1932,1954]" httpUri="https://zenodo.org/record/242642/files/figure.png" pageId="3" pageNumber="198" targetBox="[150,1435,191,1905]" targetPageId="3">
<paragraph id="8B907855FFFEFFDBD6DBF85C9F8FF800" blockId="3.[151,1008,1932,1954]" box="[151,1008,1932,1954]" pageId="3" pageNumber="198">
<emphasis id="B95BA447FFFEFFDBD6DBF85C9D67F800" bold="true" box="[151,280,1932,1954]" pageId="3" pageNumber="198">FIGURE 17.</emphasis>
Distribution map of the genus
<taxonomicName id="4C2F03D6FFFEFFDBD40EF85D9EA1F800" box="[578,734,1933,1954]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA447FFFEFFDBD40EF85D9EA1F800" box="[578,734,1933,1954]" italics="true" pageId="3" pageNumber="198">Ptomaphaminus</emphasis>
</taxonomicName>
from Indo-China Peninsula.
</paragraph>
</caption>
<paragraph id="8B907855FFF9FFDCD68BFF489D6BFF56" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">Elytra 1.35 times longer than wide together, as wide as the pronotum, largest width near the base. Surface covered with oblique (not strictly transversal) strigae, without longitudinal striae except the sutural one. Metasternal sutures convergent.</paragraph>
<paragraph id="8B907855FFF9FFDCD68BFED09DCFFEFE" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">Protibiae with a longitudinal row of equal spines along outer margin, and with ventral spines randomly dispersed. Mesotibiae and metatibiae with a circular row of equal spines at apex. Tarsi pentamerous. Male protarsi dilated, 0.75 times as wide as protibias.</paragraph>
<paragraph id="8B907855FFF9FFDCD68BFEB89913FE45" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">
Urite IX with a long and narrow spiculum gastrale, protruding beyond the anterior edge of epipleurite IX by the apical half of its length which is imperceptibly widened (
<figureCitation id="131464D0FFF9FFDCD54AFE5A9F36FE03" box="[774,841,394,417]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Fig. 5</figureCitation>
). Aedeagus long and rather narrow, with parallel sides which are narrowly convergent at apex. Lateral apical expansions of the median lobe asymmetrical (
<figureCitation id="131464D0FFF9FFDCD2A9FE7D9956FE66" box="[1253,1321,429,452]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Fig. 4</figureCitation>
), the apex with a hook in lateral view (
<figureCitation id="131464D0FFF9FFDCD78EFE009E7BFE45" box="[450,516,464,487]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Fig. 6</figureCitation>
). Internal stylus of the endophallus thin, helically winded in a single turn (
<figureCitation id="131464D0FFF9FFDCD352FE009920FE45" box="[1310,1375,464,487]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B907855FFF9FFDCD68BFE229DF8FDAB" blockId="4.[151,1436,152,1319]" box="[199,391,498,521]" pageId="4" pageNumber="199">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3352BDEFFF9FFDCD68BFDC59CA2FDED" pageId="4" pageNumber="199" type="etymology">
<paragraph id="8B907855FFF9FFDCD68BFDC59CA2FDED" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">
<emphasis id="B95BA447FFF9FFDCD68BFDC59D3FFD8E" bold="true" box="[199,320,533,556]" pageId="4" pageNumber="199">Etymology</emphasis>
. The epithet is from
<emphasis id="B95BA447FFF9FFDCD47DFDC59EFBFD8E" box="[561,644,533,556]" italics="true" pageId="4" pageNumber="199">granum</emphasis>
, latin origin of &quot;granit&quot;, refering to the biotope where this species was found.
</paragraph>
</subSubSection>
<subSubSection id="C3352BDEFFF9FFDCD68BFD8A9955FA85" pageId="4" pageNumber="199" type="discussion">
<paragraph id="8B907855FFF9FFDCD68BFD8A9D1FFC25" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">
<emphasis id="B95BA447FFF9FFDCD68BFD8A9D40FDD3" bold="true" box="[199,319,602,625]" pageId="4" pageNumber="199">Discussion</emphasis>
. The general morphology of this species, either external or internal, is typical of the genus
<taxonomicName id="4C2F03D6FFF9FFDCD6DBFDAC9DDFFD36" authority="Perreau" authorityName="Perreau" box="[151,416,636,660]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA447FFF9FFDCD6DBFDAC9D3BFD36" box="[151,324,636,660]" italics="true" pageId="4" pageNumber="199">Ptomaphaminus</emphasis>
Perreau
</taxonomicName>
: median lobe of the aedeagus long, parallel, with the lateral apical expansions asymmetrical: the left one (which appears on the right side on pictures traditionally represented in reversed orientation) with a hook ventrally deflexed in lateral view. But it is evidently different from other species: aedeagus longer and narrower than
<taxonomicName id="4C2F03D6FFF9FFDCD3C4FD129E6CFD5E" authority="Szymczakowski, 1972" authorityName="Szymczakowski" authorityYear="1972" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="bihamatus">
<emphasis id="B95BA447FFF9FFDCD3C4FD129D79FD5E" italics="true" pageId="4" pageNumber="199">P. bihamatus</emphasis>
(
<bibRefCitation id="EFBE05A4FFF9FFDCD75AFD359E73FD5E" author="Szymczakowski" box="[278,524,741,764]" pageId="4" pageNumber="199" refString="Szymczakowski, W. (1972) Catopidae et Colonidae (Coleoptera) de Ceylan (resultats du voyage entomologique du Museum d'histoire naturelle de Geneve en 1970). Acta Zoologica cracoviensia, 17 (7), 163 - 192." type="journal article" year="1972">Szymczakowski, 1972</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131464D0FFF9FFDCD468FD359E0CFD5E" box="[548,627,741,764]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Figs. 7</figureCitation>
; 8),
<taxonomicName id="4C2F03D6FFF9FFDCD4EBFD359FC5FD5E" authority="Perreau, 2009" authorityName="Perreau" authorityYear="2009" box="[679,954,740,764]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B95BA447FFF9FFDCD4EBFD359F64FD5E" box="[679,795,740,764]" italics="true" pageId="4" pageNumber="199">P. bedosae</emphasis>
<bibRefCitation id="EFBE05A4FFF9FFDCD568FD359FC5FD5E" author="Perreau" box="[804,954,741,764]" pageId="4" pageNumber="199" refString="Perreau, M. (2009) Nouveaux Ptomaphagini et Anemadini souterrains, dont la premiere espece anophtalme d' Anemadus (Coleoptera: Leiodidae Cholevinae). Annales de la Societe entomologique de France, 45, 1 - 10. http: // dx. doi. org / 10.1080 / 00379271.2009.10697587" type="journal article" year="2009">Perreau, 2009</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131464D0FFF9FFDCD580FD35982EFD5E" box="[972,1105,741,764]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Figs. 11, 12</figureCitation>
),
<taxonomicName id="4C2F03D6FFF9FFDCD22BFD3599E3FD5E" authority="Perreau, 2009" authorityName="Perreau" authorityYear="2009" box="[1127,1436,740,764]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B95BA447FFF9FFDCD22BFD359883FD5E" box="[1127,1276,740,764]" italics="true" pageId="4" pageNumber="199">P. deharvengi</emphasis>
<bibRefCitation id="EFBE05A4FFF9FFDCD349FD3599E3FD5E" author="Perreau" box="[1285,1436,741,764]" pageId="4" pageNumber="199" refString="Perreau, M. (2009) Nouveaux Ptomaphagini et Anemadini souterrains, dont la premiere espece anophtalme d' Anemadus (Coleoptera: Leiodidae Cholevinae). Annales de la Societe entomologique de France, 45, 1 - 10. http: // dx. doi. org / 10.1080 / 00379271.2009.10697587" type="journal article" year="2009">Perreau, 2009</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131464D0FFF9FFDCD6D3FCD89C91FCBD" box="[159,238,776,799]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Figs. 9</figureCitation>
; 10),
<taxonomicName id="4C2F03D6FFF9FFDCD77DFCD89E3BFCBD" authority="Jarrige, 1969" authorityName="Jarrige" authorityYear="1969" box="[305,580,775,799]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="boutini">
<emphasis id="B95BA447FFF9FFDCD77DFCD89DE6FCBD" box="[305,409,775,799]" italics="true" pageId="4" pageNumber="199">P. boutini</emphasis>
(
<bibRefCitation id="EFBE05A4FFF9FFDCD7E7FCD89E44FCBD" author="Jarrige" box="[427,571,776,799]" pageId="4" pageNumber="199" refString="Jarrige, J. (1969) Un Catopide nouveau du Cambodge (Col.). Bulletin de la Societe entomologique de France, 74, 234 - 235." type="journal article" year="1969">Jarrige, 1969</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131464D0FFF9FFDCD419FCD89ECDFCBD" box="[597,690,776,799]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Figs. 13</figureCitation>
; 14) and
<taxonomicName id="4C2F03D6FFF9FFDCD552FCD89842FCBD" authority="Perreau, 1993" authorityName="Perreau" authorityYear="1993" box="[798,1085,775,799]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="leclerci">
<emphasis id="B95BA447FFF9FFDCD552FCD89FF4FCBD" box="[798,907,775,799]" italics="true" pageId="4" pageNumber="199">P. leclerci</emphasis>
(
<bibRefCitation id="EFBE05A4FFF9FFDCD5D7FCD8984AFCBD" author="Perreau" box="[923,1077,776,799]" pageId="4" pageNumber="199" refString="Perreau, M. (1993) Nouvelles especes de Ptomaphaginus de Thailande et de Taiwan (Coleoptera, Cholevidae). Bulletin de la Societe entomologique de France, 97 (4), 343 - 347. [1992]" type="journal article" year="1993">Perreau, 1993</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131464D0FFF9FFDCD203FCD898D4FCBD" box="[1103,1195,776,799]" captionStart="FIGURES 4 16" captionStartId="2.[151,258,1695,1717]" captionTargetBox="[166,1414,227,1631]" captionTargetId="figure@2.[151,1436,217,1648]" captionTargetPageId="2" captionText="FIGURES 4 16. Ptomaphaminus Perreau from Vietnam and Laos. 4: P. granophilus n. sp., aedeagus, dorsal view. 5: P. granophilus n. sp., male urite IX. 6: P. granophilus n. sp., aedeagus, lateral view. 7: P. bihamatus (Szymczakowski), aedeagus, dorsal view (from Szymczakowski, 1972). 8: P. bihamatus (Szymczakowski), aedeagus, lateral view (from Szymczakowski, 1972). 9: P. deharvengi Perreau, aedeagus, dorsal view. 10: P. deharvengi Perreau, aedeagus lateral view. 11: P. bedosae Perreau, aedeagus, dorsal view. 12: P. bedosae Perreau, aedeagus, lateral view. 13: P. boutini (Jarrige), aedeagus, dorsal view. 14: P. boutini (Jarrige), aedeagus, lateral view. 15: P. leclerci (Perreau), aedeagus, dorsal view. 16: P. leclerci (Perreau), aedeagus, lateral view." httpUri="https://zenodo.org/record/242641/files/figure.png" pageId="4" pageNumber="199">Figs. 15</figureCitation>
; 16), and the apex of median lobe with a different conformation. These five species are distributed not far from the
<typeStatus id="5494C6F7FFF9FFDCD2D5FCFA98B8FCE3" box="[1177,1223,810,833]" pageId="4" pageNumber="199">type</typeStatus>
locality of the new species: the first two are known from
<collectingCountry id="F33838C5FFF9FFDCD405FC9D9ED9FCC6" box="[585,678,845,868]" name="Vietnam" pageId="4" pageNumber="199">Vietnam</collectingCountry>
, the third from
<collectingCountry id="F33838C5FFF9FFDCD510FC9D9FEDFCC6" box="[860,914,845,868]" name="Laos" pageId="4" pageNumber="199">Laos</collectingCountry>
, the fourth from
<collectingCountry id="F33838C5FFF9FFDCD215FC9D98B9FCC6" box="[1113,1222,845,868]" name="Cambodia" pageId="4" pageNumber="199">Cambodia</collectingCountry>
and the fifth from
<collectingCountry id="F33838C5FFF9FFDCD6DBFCA09C89FC25" box="[151,246,880,903]" name="Thailand" pageId="4" pageNumber="199">Thailand</collectingCountry>
(
<figureCitation id="131464D0FFF9FFDCD748FCA09D2CFC25" box="[260,339,880,903]" captionStart="FIGURE 17" captionStartId="3.[151,244,1932,1954]" captionTargetBox="[150,1435,191,1905]" captionTargetId="figure@3.[150,1436,191,1911]" captionTargetPageId="3" captionText="FIGURE 17. Distribution map of the genus Ptomaphaminus from Indo-China Peninsula." httpUri="https://zenodo.org/record/242642/files/figure.png" pageId="4" pageNumber="199">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8B907855FFF9FFDCD68BFC42997AFBDB" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">
The biotope where the new species was discovered is remarkable.
<taxonomicName id="4C2F03D6FFF9FFDCD5DDFC4198B9FC0B" box="[913,1222,913,937]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="granophilus" status="sp. nov.">
<emphasis id="B95BA447FFF9FFDCD5DDFC4198B9FC0B" box="[913,1222,913,937]" italics="true" pageId="4" pageNumber="199">Ptomaphaminus granophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268193CFFF9FFDCD281FC429975FC0B" box="[1229,1290,914,937]" pageId="4" pageNumber="199" rank="species">
<emphasis id="B95BA447FFF9FFDCD281FC429975FC0B" bold="true" box="[1229,1290,914,937]" pageId="4" pageNumber="199">n. sp.</emphasis>
</taxonomicNameLabel>
was found in a granitic blockfield made of huge, piled up blocks which are forming more or less important void spaces (
<figureCitation id="131464D0FFF9FFDCD367FC6599ECFC6E" box="[1323,1427,949,972]" captionStart="FIGURES 1 3" captionStartId="1.[151,258,1978,2000]" captionTargetBox="[173,1405,194,1948]" captionTargetId="figure@1.[173,1406,194,1950]" captionTargetPageId="1" captionText="FIGURES 1 3. Ptomaphaminus granophilus n. sp. 1 - 2: biotope: the granitic blockfield. 3: habitus." httpUri="https://zenodo.org/record/242640/files/figure.png" pageId="4" pageNumber="199">Figs. 1, 2</figureCitation>
) allowing to get through the screes and to go down several meters. The bottom is extremely wet and completely dark (
<figureCitation id="131464D0FFF9FFDCD33CFC089CD8FBB3" captionStart="FIGURES 1 3" captionStartId="1.[151,258,1978,2000]" captionTargetBox="[173,1405,194,1948]" captionTargetId="figure@1.[173,1406,194,1950]" captionTargetPageId="1" captionText="FIGURES 1 3. Ptomaphaminus granophilus n. sp. 1 - 2: biotope: the granitic blockfield. 3: habitus." httpUri="https://zenodo.org/record/242640/files/figure.png" pageId="4" pageNumber="199">Fig. 2</figureCitation>
). During our visit, some water flowed there, forming a stream difficult to reach but clearly audible. In this peculiar biotope we also found most of the Arthropod groups common in the caves of limestones areas of the region:
<taxonomicName id="4C2F03D6FFF9FFDCD36DFBCD99E6FB96" box="[1313,1433,1053,1076]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
:
<taxonomicName id="4C2F03D6FFF9FFDCD6DBFB909D20FBF5" box="[151,351,1088,1111]" class="Insecta" family="Rhaphidophoridae" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
,
<taxonomicName id="4C2F03D6FFF9FFDCD73CFB909DB2FBF5" box="[368,461,1088,1111]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="199" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
:
<taxonomicName id="4C2F03D6FFF9FFDCD791FB909E21FBF5" box="[477,606,1088,1111]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="199" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
,
<taxonomicName id="4C2F03D6FFF9FFDCD423FB909E9BFBF5" box="[623,740,1088,1111]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
: Fulgoroidea of the families
<taxonomicName id="4C2F03D6FFF9FFDCD27BFB9098EEFBF5" box="[1079,1169,1088,1111]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
and
<taxonomicName id="4C2F03D6FFF9FFDCD29FFB90991DFBF5" box="[1235,1378,1088,1111]" class="Insecta" family="Meenoplidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="family">Meenoplidae</taxonomicName>
. The description of one of the species collected on hanging roots in the same blockfield is currently in progress.
</paragraph>
<paragraph id="8B907855FFF9FFDCD68BFB559955FA85" blockId="4.[151,1436,152,1319]" pageId="4" pageNumber="199">The specimen was collected in the deepest reachable area of the blockfield, by means of a pitfall trap baited with vinegar. It is not excluded that the species will be found in the deep ground of the area. A second blockfield was localized at an altitude of 1500 meters behind the house of the bacteriologist A. Yersin who lived there until his death in 1943, close to the top of the massif. The new species was not found there, but these remarkable geological structures, forming a real subterranean environment in granitic area, would deserve to be inventoried and systematically sampled.</paragraph>
</subSubSection>
</treatment>
</document>