treatments-xml/data/03/86/87/038687B6697A8F7FFF1BFA62FF5B859E.xml
2024-06-21 12:22:17 +02:00

319 lines
53 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A63BDA0C68819BF90DB95B734D299543" ID-CLB-Dataset="112021" ID-DOI="10.11646/zootaxa.5143.1.1" ID-GBIF-Dataset="9d7cad94-f472-47eb-a988-5569e479204e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6601404" ID-ZooBank="F71553B2-7D58-4E61-A883-546B2A0124D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654063338489" checkinUser="plazi" docAuthor="Burington, Zelia L." docDate="2022" docId="038687B6697A8F7FFF1BFA62FF5B859E" docLanguage="en" docName="zootaxa.5143.1.1.pdf" docOrigin="Zootaxa 5143 (1)" docSource="http://zoobank.org/f71553b2-7d58-4e61-a883-546b2a0124d5" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eucelatoria gustavogutierrezi Burington 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="FFBFFFCE694A8F4DFF8CFFE0FFA28416" masterDocTitle="A taxonomic revision of the Eucelatoria ferox species group (Diptera: Tachinidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="49" updateTime="1699338509909" updateUser="ExternalLinkService">
<mods:mods id="87385FC13C2CC8045FD78B7884670CB5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F7404BD32E26CD1753D6F2739214257">
<mods:title id="A2E6216811FBCC8DAC8AA69C2A589685">A taxonomic revision of the Eucelatoria ferox species group (Diptera: Tachinidae)</mods:title>
</mods:titleInfo>
<mods:name id="F1695778154CED48533CB4A90B058FA4" type="personal">
<mods:role id="35B8A792D102E62A23CB64BD276D485E">
<mods:roleTerm id="21C2C969EBDFE28CF5671FC4C6567180">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D769A52023ADCF0EF7380A7BF657D924">Burington, Zelia L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FF8074567A4D89EDF3EAEC86D39CEF42">text</mods:typeOfResource>
<mods:relatedItem id="7E2421A9B2FAAD3736E464FFD79B2B99" type="host">
<mods:titleInfo id="95880A418133EC0288A6627E2D3C34D2">
<mods:title id="C14F5CDD4F02D85A0C939304D76A75C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B0BEEC6F1FBAE6914894A9882E30562A">
<mods:date id="8C5535A8E478ADCF8925940BCADEEB35">2022</mods:date>
<mods:detail id="017028142FFAB919C6C06287D7EA3C02" type="pubDate">
<mods:number id="9D32044437AB7B105BF2B54F8FC67AD7">2022-06-01</mods:number>
</mods:detail>
<mods:detail id="6580E46FC1020994AA1F7724D8C753F7" type="volume">
<mods:number id="F46224AD2F1EE51B7846EC9D52F33826">5143</mods:number>
</mods:detail>
<mods:detail id="F9274E7DC4E3E1E41BD6B8C5AEDA978C" type="issue">
<mods:number id="B03B6EAEFE87C419E83D35EB10F60503">1</mods:number>
</mods:detail>
<mods:extent id="0F009A57993E1DD3720114F0E48B43F2" unit="page">
<mods:start id="B87505663834A193194B24D01B233D6C">1</mods:start>
<mods:end id="01A4666CA401C3EAE0DACF1BE7496C4C">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CD3A2B8168F5CC0EF3A2009D5DCDA1AA">
<mods:url id="AFE06DA9B127B9FFF4FCCDACCE687B7D">http://zoobank.org/f71553b2-7d58-4e61-a883-546b2a0124d5</mods:url>
</mods:location>
<mods:classification id="F055CD4285ED2816F233BF80D2C8A54A">journal article</mods:classification>
<mods:identifier id="7914B40ACEC7E1689A0E1B49681CB033" type="CLB-Dataset">112021</mods:identifier>
<mods:identifier id="595D28A5036FE0AD3624703B8F1C9894" type="DOI">10.11646/zootaxa.5143.1.1</mods:identifier>
<mods:identifier id="D7A5368A6F0D46914E265460A0507089" type="GBIF-Dataset">9d7cad94-f472-47eb-a988-5569e479204e</mods:identifier>
<mods:identifier id="7703E40D9E99D88D1BC00E0A3BDEA5A3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="16FF880E7AB67BB9CA6CD92CF8301137" type="Zenodo-Dep">6601404</mods:identifier>
<mods:identifier id="62733D43B861A1D51CACBADE42C56F29" type="ZooBank">F71553B2-7D58-4E61-A883-546B2A0124D5</mods:identifier>
</mods:mods>
<treatment id="038687B6697A8F7FFF1BFA62FF5B859E" ID-DOI="http://doi.org/10.5281/zenodo.6958424" ID-GBIF-Taxon="198574809" ID-Zenodo-Dep="6958424" LSID="urn:lsid:plazi:treatment:038687B6697A8F7FFF1BFA62FF5B859E" httpUri="http://treatment.plazi.org/id/038687B6697A8F7FFF1BFA62FF5B859E" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49">
<subSubSection id="C335652B697A8F7DFF1BFA62FDC0818B" box="[151,610,1410,1437]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8B9036A0697A8F7DFF1BFA62FDC0818B" blockId="48.[151,763,1410,1474]" box="[151,610,1410,1437]" pageId="48" pageNumber="49">
<heading id="D0D881CC697A8F7DFF1BFA62FDC0818B" bold="true" box="[151,610,1410,1437]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
<emphasis id="B95BEAB2697A8F7DFF1BFA62FDC0818B" bold="true" box="[151,610,1410,1437]" pageId="48" pageNumber="49">
<taxonomicName id="4C2F4D23697A8F7DFF1BFA62FE5F818B" ID-CoL="9J36B" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[151,509,1410,1437]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="gustavogutierrezi" status="sp. nov.">
<emphasis id="B95BEAB2697A8F7DFF1BFA62FE5F818B" bold="true" box="[151,509,1410,1437]" italics="true" pageId="48" pageNumber="49">Eucelatoria gustavogutierrezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857C9697A8F7DFD88FA63FDC0818B" box="[516,610,1411,1437]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C335652B697A8F7DFF1BFA47FD5981D4" box="[151,763,1446,1474]" pageId="48" pageNumber="49" type="description">
<paragraph id="8B9036A0697A8F7DFF1BFA47FD5981D4" blockId="48.[151,763,1410,1474]" box="[151,763,1446,1474]" pageId="48" pageNumber="49">
(
<figureCitation id="13142A25697A8F7DFF13FA47FF5E81D7" box="[159,252,1447,1473]" captionStart="FIGURE 1" captionStartId="9.[151,250,1766,1790]" captionTargetBox="[164,1421,431,1729]" captionTargetId="figure-49@9.[153,1434,417,1743]" captionTargetPageId="9" captionText="FIGURE 1. CO1-based neighbor-joining tree of 12 species of the Eucelatoria ferox species group, with inset illustrations of mesonota. The outgroup is Eucelatoria armigera (Coquillett). Node labels indicate support values over 60. Scale bar = 2% genetic difference. The E. charapensis (Townsend) sequence with the axial break is of poor quality but is included to show unity of El Bigal specimens with the rest of the metapopulation. Mesonota are illustrated with setal annuli only for clarity. A. E. gustavogutierrezi sp. nov. B. E. kopis sp. nov. C. E. ritavargasae sp. nov. D. E. ferox (Townsend). E. E. huitepecensis sp. nov. F. E. auriceps (Aldrich). G. E. gladiatrix (Townsend). H. E. borealis sp. nov. I. E. yanayacu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601406" httpUri="https://zenodo.org/record/6601406/files/figure.png" pageId="48" pageNumber="49">Figs 1A</figureCitation>
,
<figureCitation id="13142A25697A8F7DFE84FA47FE8481D7" box="[264,294,1447,1473]" captionStart="FIGURES 1821" captionStartId="66.[151,264,1943,1967]" captionTargetBox="[171,1416,181,1920]" captionTargetId="figure-31@66.[170,1416,181,1920]" captionTargetPageId="66" captionText="FIGURES 1821. Female habitus photographs of Eucelatoria ferox group spp., dorsal view. 18. E. texana (Reinhard), Holly Springs (ZLB_E.Ferox 00074) (USNM). 19. E. woodorum sp. nov., Chiapas (ZLB_E.Ferox 00286) (DMW). 20. E. yanayacu sp. nov., allotype (ZLB_E.Ferox 00184) (USNM). 21. E. gustavogutierrezi sp. nov., holotype (ZLB_E.Ferox 00006) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601416" httpUri="https://zenodo.org/record/6601416/files/figure.png" pageId="48" pageNumber="49">21</figureCitation>
,
<figureCitation id="13142A25697A8F7DFEBEFA47FEF281D7" box="[306,336,1447,1473]" captionStart="FIGURES 3841" captionStartId="71.[151,264,1908,1932]" captionTargetBox="[184,1404,182,1885]" captionTargetId="figure-15@71.[184,1404,181,1885]" captionTargetPageId="71" captionText="FIGURES 3841. Male habitus photographs of Eucelatoria ferox group spp., dorsal view. 38. E. texana (Reinhard), College Station (ZLB_E.Ferox 00049) (CNC). 39. E. woodorum sp. nov., holotype (ZLB_E.Ferox 00288) (DMW). 40. E. yanayacu sp. nov., holotype (ZLB_E.Ferox 00191) (USNM). 41. E. gustavogutierrezi sp. nov., Guanacaste Province (ZLB_E.Ferox 00008) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601426" httpUri="https://zenodo.org/record/6601426/files/figure.png" pageId="48" pageNumber="49">41</figureCitation>
,
<figureCitation id="13142A25697A8F7DFED0FA47FED881D7" box="[348,378,1447,1473]" captionStart="FIGURES 6667" captionStartId="83.[151,264,1943,1967]" captionTargetBox="[197,1390,181,1920]" captionTargetId="figure-15@83.[197,1390,181,1920]" captionTargetPageId="83" captionText="FIGURES 6667. Female habitus photographs of Eucelatoria ferox group spp., lateral view. 66. E. gustavogutierrezi sp. nov., holotype (ZLB_E.Ferox 00006) (CNC). 67. E. kopis sp. nov., holotype (ZLB_E.Ferox 00002) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601452" httpUri="https://zenodo.org/record/6601452/files/figure.png" pageId="48" pageNumber="49">66</figureCitation>
,
<figureCitation id="13142A25697A8F7DFE0AFA47FE0681D7" box="[390,420,1447,1473]" captionStart="FIGURES 8687" captionStartId="93.[151,264,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-15@93.[198,1390,181,1920]" captionTargetPageId="93" captionText="FIGURES 8687. Male habitus photographs of Eucelatoria ferox group spp., lateral view. 86. E. gustavogutierrezi sp. nov., Guanacaste Province (ZLB_E.Ferox 00008) (CNC). 87. E. hafelei sp. nov., Chapare Province (ZLB_E.Ferox 00386) (DMW)." figureDoi="http://doi.org/10.5281/zenodo.6601484" httpUri="https://zenodo.org/record/6601484/files/figure.png" pageId="48" pageNumber="49">86</figureCitation>
,
<figureCitation id="13142A25697A8F7DFE3CFA47FE7E81D7" box="[432,476,1447,1473]" captionStart="FIGURES 94108" captionStartId="97.[151,264,1704,1728]" captionTargetBox="[264,1316,193,1668]" captionTargetId="figure-15@97.[251,1337,181,1680]" captionTargetPageId="97" captionText="FIGURES 94108. Female seventh abdominal segment of Eucelatoria ferox group spp., lateral view. Numbers are “ZLB_ E.Ferox” specimen IDs. 94. E. ferox (Townsend), Santa Cruz (00398) (USNM). 95. E. aurata (Townsend), Nova Teutonia (00250) (DMW). 96. E. auriceps (Aldrich), Crane Hollow (00410) (JOSC). 97. E. borealis sp. nov., Sheffeld (00041) (DMW). 98. E. charapensis (Townsend), Chapare Province (00393) (DMW). 99. E. falcata sp. nov., Nova Teutonia (00274) (CNC). 100. E. fordlandia sp. nov., Fordlandia (00311) (USNM). 101. E. gladiatrix (Townsend), Guanacaste Province (00376) (CNC). 102. E. sabroskyi sp. nov., Herb Martyr Campground (00084) (CNC). 103. E. strigata (Wulp), Trinidad (00138) (CNC). 104. E. texana (Reinhard), Madera Greaterville Rd. (00048) (JOSC). 105. E. yanayacu sp. nov., Yanayacu (no “E.Ferox” number, labeled “DIP328”) (USNM). 106. E. gustavogutierrezi sp. nov., Guanacaste Province (00007) (CNC). 107. E. kopis sp. nov., Avispas (00217) (CNC). 108. E. sica sp. nov., Yanayacu (00211) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601500" httpUri="https://zenodo.org/record/6601500/files/figure.png" pageId="48" pageNumber="49">106</figureCitation>
,
<figureCitation id="13142A25697A8F7DFE64FA47FDB681D7" box="[488,532,1447,1473]" captionStart="FIGURES 109123" captionStartId="98.[151,264,1695,1719]" captionTargetBox="[221,1365,193,1657]" captionTargetId="figure-31@98.[209,1378,181,1672]" captionTargetPageId="98" captionText="FIGURES 109123. Female abdominal segments eight through ten of Eucelatoria ferox group spp., dorsal view. Numbers are “ZLB_E.Ferox” specimen IDs. 119. E. ferox (Townsend), Santa Cruz (00398) (USNM). 110. E. aurata (Townsend), Nova Teutonia (00250) (DMW). 111. E. auriceps (Aldrich), Crane Hollow (00410) (JOSC). 112. E. borealis sp. nov., Sheffeld (00041) (DMW). 113. E. charapensis (Townsend), Chapare Province (00393) (DMW). 114. E. falcata sp. nov., Nova Teutonia (00274) (CNC). 115. E. fordlandia sp. nov., Fordlandia (00311) (USNM). 116. E. gladiatrix (Townsend), Guanacaste Province (ZLB_ E.Ferox 00379) (CNC). 117. E. sabroskyi sp. nov., Herb Martyr Campground (00084) (CNC). 118. E. strigata (Wulp), Trinidad (00138) (CNC). 119. E. texana (Reinhard), Madera Greaterville Rd. (00048) (JOSC). 120. E. yanayacu sp. nov., Yanayacu (no “E.Ferox” number, labeled “DIP328”) (USNM). 121. E. gustavogutierrezi sp. nov., Guanacaste Province (00007) (CNC). 122. E. kopis sp. nov., Avispas (00217) (CNC). 123. E. sica sp. nov., Yanayacu (00211) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601502" httpUri="https://zenodo.org/record/6601502/files/figure.png" pageId="48" pageNumber="49">121</figureCitation>
,
<figureCitation id="13142A25697A8F7DFDACFA47FD2681D7" box="[544,644,1446,1473]" captionStart="FIGURES 136155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="48" pageNumber="49">136, 152</figureCitation>
,
<figureCitation id="13142A25697A8F7DFD1CFA47FD1E81D7" box="[656,700,1447,1473]" captionStart="FIGURES 156173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="48" pageNumber="49">169</figureCitation>
,
<figureCitation id="13142A25697A8F7DFD44FA47FD5181D4" box="[712,755,1447,1474]" captionStart="FIGURES 174179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="48" pageNumber="49">176</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C335652B697A8F7CFF1BFA0FFC2F84C3" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8B9036A0697A8F7DFF1BFA0FFEE68263" blockId="48.[151,1437,1518,2013]" pageId="48" pageNumber="49">
<emphasis id="B95BEAB2697A8F7DFF1BFA0FFEEB821E" bold="true" box="[151,329,1518,1545]" pageId="48" pageNumber="49">Type material.</emphasis>
<materialsCitation id="3B473CFD697A8F7DFEDFFA0FFEE68263" ID-GBIF-Occurrence="3878846602" collectionCode="CNC" collectorName="Z. L. Burington" country="Costa Rica" location="Area de Conservacion" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54948802697A8F7DFEDFFA0FFE19821F" box="[339,443,1519,1545]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
<specimenCount id="9D29FD29697A8F7DFE4AFA0FFE79821F" box="[454,475,1519,1545]" pageId="48" pageNumber="49" type="female"></specimenCount>
, labeled “DHJPAR0034387”, “Voucher:
<collectorName id="26DA5376697A8F7DFC3BFA0FFBE6821F" box="[951,1092,1519,1545]" pageId="48" pageNumber="49">D.H. Janzen</collectorName>
&amp;
<collectorName id="26DA5376697A8F7DFBE2FA0FFAAF821F" box="[1134,1293,1519,1545]" pageId="48" pageNumber="49">W. Hallwachs</collectorName>
/ DB: http:// janzen.sas.upen.edu/
<location id="8EF0607B697A8F7DFE09F9F3FD23823B" LSID="urn:lsid:plazi:treatment:038687B6697A8F7FFF1BFA62FF5B859E:8EF0607B697A8F7DFE09F9F3FD23823B" box="[389,641,1555,1581]" country="Costa Rica" name="Area de Conservacion" pageId="48" pageNumber="49" stateProvince="Guanacaste">Area de Conservacion</location>
<collectingRegion id="49EBF842697A8F7DFD05F9F3FCAE823B" box="[649,780,1555,1581]" country="Costa Rica" name="Guanacaste" pageId="48" pageNumber="49">Guanacaste</collectingRegion>
,/
<collectingCountry id="F3387630697A8F7DFCADF9F3FC6A823B" box="[801,968,1555,1581]" name="Costa Rica" pageId="48" pageNumber="49">COSTA RICA</collectingCountry>
./ 09 -SRNP- 35146”, “
<typeStatus id="54948802697A8F7DFB41F9F3FAC1823B" box="[1229,1379,1555,1581]" pageId="48" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C2F4D23697A8F7DFAF8F9F3FF558247" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ gustavogutierrezi/
<collectorName id="26DA5376697A8F7DFE56F9D7FD268247" box="[474,644,1591,1617]" pageId="48" pageNumber="49">Z.L. Burington</collectorName>
[red label]”, “LEGS AWAY FOR DNA”, “Janzen06”, “ZLB_E.Ferox 00006” (
<collectionCode id="ED3EAE65697A8F7DFF75F9BBFE9A8263" box="[249,312,1627,1653]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="48" pageNumber="49">CNC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9036A0697A8F7DFF4BF99FFAF083CB" blockId="48.[151,1437,1518,2013]" pageId="48" pageNumber="49">
<materialsCitation id="3B473CFD697A8F7DFF4BF99FFCAF828F" ID-GBIF-Occurrence="3878846552" box="[199,781,1663,1689]" country="Costa Rica" location="Guanacaste" pageId="48" pageNumber="49" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54948802697A8F7DFF4BF99FFE95828F" box="[199,311,1663,1689]" pageId="48" pageNumber="49" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D29FD29697A8F7DFECFF99FFECF828F" box="[323,365,1663,1689]" pageId="48" pageNumber="49" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D29FD29697A8F7DFE29F99FFE44828F" box="[421,486,1663,1689]" pageId="48" pageNumber="49" type="female">4 ♀♀</specimenCount>
.
<collectingCountry id="F3387630697A8F7DFE7FF99FFDCF828F" box="[499,621,1663,1689]" name="Costa Rica" pageId="48" pageNumber="49">Costa Rica</collectingCountry>
<emphasis id="B95BEAB2697A8F7DFDE2F99FFD25828E" box="[622,647,1663,1688]" italics="true" pageId="48" pageNumber="49"></emphasis>
<collectingRegion id="49EBF842697A8F7DFD0BF99FFCAF828F" box="[647,781,1663,1689]" country="Costa Rica" name="Guanacaste" pageId="48" pageNumber="49">Guanacaste</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B473CFD697A8F7DFC9BF99FFBA18313" ID-GBIF-Occurrence="3878846558" collectionCode="CNC" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" location="Area de Conservacion" municipality="Voucher" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D29FD29697A8F7DFC9BF99FFCE0828F" box="[791,834,1663,1689]" pageId="48" pageNumber="49" type="male">1 ♂</specimenCount>
, labeled “DHJPAR0050613”, “
<collectingMunicipality id="6BF4ACDA697A8F7DFB2FF99FFAA5828F" box="[1187,1287,1663,1689]" pageId="48" pageNumber="49">Voucher</collectingMunicipality>
:
<collectorName id="26DA5376697A8F7DFA9EF99FFA3E828F" box="[1298,1436,1663,1689]" pageId="48" pageNumber="49">D.H. Janzen</collectorName>
&amp;
<collectorName id="26DA5376697A8F7DFF39F943FEF382AB" box="[181,337,1699,1725]" pageId="48" pageNumber="49">W. Hallwachs</collectorName>
/ DB: http://janzen.sas.upen.edu/
<location id="8EF0607B697A8F7DFD45F943FC6782AB" LSID="urn:lsid:plazi:treatment:038687B6697A8F7FFF1BFA62FF5B859E:8EF0607B697A8F7DFD45F943FC6782AB" box="[713,965,1699,1725]" country="Costa Rica" municipality="Voucher" name="Area de Conservacion" pageId="48" pageNumber="49" stateProvince="Guanacaste">Area de Conservacion</location>
<collectingRegion id="49EBF842697A8F7DFC41F943FBF282AB" box="[973,1104,1699,1725]" country="Costa Rica" name="Guanacaste" pageId="48" pageNumber="49">Guanacaste</collectingRegion>
,/
<collectingCountry id="F3387630697A8F7DFBE9F943FAA982AB" box="[1125,1291,1699,1725]" name="Costa Rica" pageId="48" pageNumber="49">COSTA RICA</collectingCountry>
./ 13 -SRNP-69094”, “
<typeStatus id="54948802697A8F7DFE8CF927FE2C82F7" box="[256,398,1735,1761]" pageId="48" pageNumber="49" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C2F4D23697A8F7DFE11F927FDBE82F7" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[413,540,1735,1761]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ gustavogutierrezi/
<collectorName id="26DA5376697A8F7DFD70F927FC0782F7" box="[764,933,1735,1761]" pageId="48" pageNumber="49">Z.L. Burington</collectorName>
[yellow label]”, “LEGS AWAY FOR DNA”, “Janzen06”, “ZLB_E.Ferox 00008” [genitalia in vial under specimen] (
<collectionCode id="ED3EAE65697A8F7DFC30F90BFC598313" box="[956,1019,1771,1797]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="48" pageNumber="49">CNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B473CFD697A8F7DFB83F90BFAC5835B" ID-GBIF-Occurrence="3878846430" collectionCode="CNC" country="Costa Rica" pageId="48" pageNumber="49" specimenCount="2" specimenCount-female="2" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D29FD29697A8F7DFB83F90BFBED8313" box="[1039,1103,1771,1797]" pageId="48" pageNumber="49" type="female">2 ♀♀</specimenCount>
, as previous except DHJPAR numbers “DHJPAR0049524” and “DHJPAR0053214”, SRNP numbers “12 -SRNP- 81030” and “13 -SRNP-35825”, and specimen identifiers “00007” and “00009” [first with terminalia in vial under specimen] (
<collectionCode id="ED3EAE65697A8F7DFA93F8D3FAFC835B" box="[1311,1374,1843,1869]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="48" pageNumber="49">CNC</collectionCode>
)
</materialsCitation>
.
<collectingRegion id="49EBF842697A8F7DFAFEF8D3FF718367" country="Costa Rica" name="Heredia" pageId="48" pageNumber="49">Heredia</collectingRegion>
:
<materialsCitation id="3B473CFD697A8F7DFF6DF8B7FF4D83AF" ID-GBIF-Occurrence="3878846578" collectingDate="2015-01-02" collectionCode="JOSC" collectorName="A. Eckhardt" country="Costa Rica" elevation="58" latitude="-10.425333" location="La Selva Biol Sta." longLatPrecision="13" longitude="-84.004" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="9D29FD29697A8F7DFF6DF8B7FEAF8367" box="[225,269,1879,1905]" pageId="48" pageNumber="49" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F3387630697A8F7DFEABF8B7FE718367" box="[295,467,1879,1905]" name="Costa Rica" pageId="48" pageNumber="49">COSTA RICA</collectingCountry>
:
<collectingRegion id="49EBF842697A8F7DFE6CF8B7FDDF8367" box="[480,637,1879,1905]" country="Costa Rica" name="Heredia" pageId="48" pageNumber="49">Heredia Prov.</collectingRegion>
/
<location id="8EF0607B697A8F7DFD02F8B7FCC08367" LSID="urn:lsid:plazi:treatment:038687B6697A8F7FFF1BFA62FF5B859E:8EF0607B697A8F7DFD02F8B7FCC08367" box="[654,866,1879,1905]" country="Costa Rica" latitude="-10.425333" longLatPrecision="13" longitude="-84.004" name="La Selva Biol Sta." pageId="48" pageNumber="49" stateProvince="Heredia">La Selva Biol Sta.</location>
<quantity id="4CD79B45697A8F7DFCE7F8B6FC048367" box="[875,934,1878,1905]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="48" pageNumber="49" unit="m" value="58.0">
<elevation id="0002D193697A8F7DFCE7F8B6FC048367" box="[875,934,1878,1905]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="48" pageNumber="49" unit="m" value="58.0">58 m</elevation>
</quantity>
/
<geoCoordinate id="EE1B5067697A8F7DFC34F8B8FB968365" box="[952,1076,1880,1907]" degrees="10" direction="south" minutes="25.52" orientation="latitude" pageId="48" pageNumber="49" precision="9" value="-10.425333">10°25.52S</geoCoordinate>
[sic; N]
<geoCoordinate id="EE1B5067697A8F7DFB13F8B8FA868365" box="[1183,1316,1880,1907]" degrees="84" direction="west" minutes="00.24" orientation="longitude" pageId="48" pageNumber="49" precision="9" value="-84.004">84°00.24W</geoCoordinate>
/
<date id="FF911060697A8F7DFABBF8B6FA318367" box="[1335,1427,1878,1905]" pageId="48" pageNumber="49" value="2015-01-02">
<collectingDate id="EFD5E988697A8F7DFABBF8B6FA318367" box="[1335,1427,1878,1905]" pageId="48" pageNumber="49" value="2015-01-02">2.I.2015</collectingDate>
</date>
/ J.O. Stireman,
<collectorName id="26DA5376697A8F7DFEB3F89BFE6F8383" box="[319,461,1915,1941]" pageId="48" pageNumber="49">A. Eckhardt</collectorName>
”, “JOS115.20”, [pink label], “
<taxonomicName id="4C2F4D23697A8F7DFCADF89BFC018383" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[801,931,1915,1941]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ (
<taxonomicName id="4C2F4D23697A8F7DFC36F89BFB988383" box="[954,1082,1915,1941]" class="Insecta" family="Tachinidae" genus="Xiphomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Xiphomyia</taxonomicName>
grp?)”, “ZLB_E.Ferox 00405” (
<collectionCode id="ED3EAE65697A8F7DFF13F87FFF4483AF" box="[159,230,1951,1977]" pageId="48" pageNumber="49">JOSC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B473CFD697A8F7DFF76F87FFAEC83CB" ID-GBIF-Occurrence="3878846496" collectionCode="JOSC" collectorName="L. Dyer" country="Costa Rica" location="La Selva Biological Station" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="9D29FD29697A8F7DFF76F87FFE8783AF" box="[250,293,1951,1977]" pageId="48" pageNumber="49" type="female">1 ♀</specimenCount>
, “WCCTR/ 7119”, “
<taxonomicName id="4C2F4D23697A8F7DFD83F87FFD3583AF" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[527,663,1951,1977]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
”, “
<collectingCountry id="F3387630697A8F7DFD39F87FFCFD83AF" box="[693,863,1951,1977]" name="Costa Rica" pageId="48" pageNumber="49">COSTA RICA</collectingCountry>
:
<collectingRegion id="49EBF842697A8F7DFCE7F87FFBA583AF" box="[875,1031,1951,1977]" country="Costa Rica" name="Heredia" pageId="48" pageNumber="49">Heredia Prov.</collectingRegion>
,/
<location id="8EF0607B697A8F7DFB97F87FFAF683AF" LSID="urn:lsid:plazi:treatment:038687B6697A8F7FFF1BFA62FF5B859E:8EF0607B697A8F7DFB97F87FFAF683AF" box="[1051,1364,1951,1977]" country="Costa Rica" name="La Selva Biological Station" pageId="48" pageNumber="49" stateProvince="Heredia">La Selva Biological Station</location>
,/ STR 1650, 3[/]22[/]2000/
<collectorName id="26DA5376697A8F7DFE0FF823FE7E83CB" box="[387,476,1987,2013]" pageId="48" pageNumber="49">L. Dyer</collectorName>
, Coll.”, “Reared:
<taxonomicName id="4C2F4D23697A8F7DFD2FF823FC8C83CB" authorityName="Lederer" authorityYear="1863" box="[675,814,1987,2013]" class="Insecta" family="Crambidae" genus="Pantographa" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="50" phylum="Arthropoda" rank="genus">Pantographa</taxonomicName>
/ suffusalis/ ex. Hampia/ appendiculata” (
<collectionCode id="ED3EAE65697A8F7DFB72F823FAE783CB" box="[1278,1349,1987,2013]" pageId="48" pageNumber="49">JOSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BFF76FC2F84C3" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">
<emphasis id="B95BEAB2697B8F7CFF4BFF76FEFF84A6" bold="true" box="[199,349,150,176]" pageId="49" pageNumber="50">Recognition.</emphasis>
This species is similar to other
<taxonomicName id="4C2F4D23697B8F7CFD4CFF77FCBA84A6" authorityName=": Wood" authorityYear="1985" box="[704,792,150,176]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B95BEAB2697B8F7CFD4CFF77FCBA84A6" box="[704,792,150,176]" italics="true" pageId="49" pageNumber="50">E. ferox</emphasis>
</taxonomicName>
group species with a single postsutural vitta, but is easily recognized by the large yellow to tan palpi and absent ocellar setae.
</paragraph>
</subSubSection>
<subSubSection id="C335652B697B8F7CFF4BFF3EFD2B8556" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8B9036A0697B8F7CFF4BFF3EFD2B8556" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">
<emphasis id="B95BEAB2697B8F7CFF4BFF3EFEEC84EE" bold="true" box="[199,334,222,248]" pageId="49" pageNumber="50">Etymology.</emphasis>
By recommendation of Dr. Daniel Janzen, this species “is named in recognition of Dr. Gustavo Gutierrez Espeleta for his full support of the new collaborations between Costa Ricas Area de Conservación
<collectingRegion id="49EBF842697B8F7CFAE8FEE2FF4B8557" country="Costa Rica" name="Guanacaste" pageId="49" pageNumber="50">Guanacaste</collectingRegion>
and the Universidad de
<collectingCountry id="F3387630697B8F7CFE77FEC6FDD88556" box="[507,634,294,320]" name="Costa Rica" pageId="49" pageNumber="50">Costa Rica</collectingCountry>
”.
</paragraph>
</subSubSection>
<subSubSection id="C335652B697B8F7CFF4BFEAAFCD0835B" pageId="49" pageNumber="50" type="description">
<paragraph id="8B9036A0697B8F7CFF4BFEAAFB268573" blockId="49.[151,1437,150,2013]" box="[199,1156,330,357]" pageId="49" pageNumber="50">
<emphasis id="B95BEAB2697B8F7CFF4BFEAAFEFB8572" bold="true" box="[199,345,330,356]" pageId="49" pageNumber="50">Description.</emphasis>
[Described from
<specimenCount id="9D29FD29697B8F7CFD93FEAAFDFC8573" box="[543,606,330,357]" pageId="49" pageNumber="50" type="female">4 ♀♀</specimenCount>
and
<specimenCount id="9D29FD29697B8F7CFD18FEAAFD608573" box="[660,706,330,357]" pageId="49" pageNumber="50" type="male">1 ♂</specimenCount>
.] Length 4.0
<quantity id="4CD79B45697B8F7CFCD2FEAAFC118573" box="[862,947,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="49" pageNumber="50" unit="mm" value="5.5">5.5 mm</quantity>
(mean = 5.0 mm).
</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BFE8EFB828796" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">Head. Fronto-orbital plate, vertex, upper half of post-ocular plate, and vibrissal angle tomentum yellow, appearing dark gold. Genal dilation, lower half of post-ocular plate, postgena, and occiput tomentum ash-gray. Parafacial tomentum yellow on upper three-fourths, merging to ash-gray near ventral corner, occasionally entirely yellow. Pale occipital setae ash-gray, usually merging to yellow on upper half of occiput. Ocellar setae minute or apparently absent, not distinguishable from setulae of ocellar triangle, rarely with a single seta. Gena with 4 setae, subvibrissal ridge with 1 seta. Facial ridge with setulae on lower one-third to one-half. Postocellar setae one-third to two-thirds length of posterior reclinate orbital seta. Paravertical seta one-half to subequal to length of postocellar setae. Outer vertical seta undifferentiated in male, one-third length of inner vertical seta in female. Reclinate orbital setae 3. Frontal setae 45, last frontal seta level with arista base. Eye with short setulae, no more than 12 facets in length. Eye height to head height ratio 0.860.90 (mean = 0.88). Postpedicel length 0.410.44 (mean = 0.42) height of head. Facial ridge length 0.540.58 (mean = 0.56) height of head. Parafacial width 0.030.07 (mean = 0.04) lateral length of head. Pedicel 0.250.33 (mean = 0.30) length of postpedicel. Postpedicel 4.07.0 (mean = 5.4) times width of parafacial in lateral view. Vertex 0.220.23 width of head in dorsal view. Palpus tan to yellow; subcylindrical and slightly dilated at tip in male; greatly dilated and flattened at apex in female; with short dark setae apically, longer ventrally, shorter dorsally in female; with several thin black setae mediolaterally.</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BFC6AFAF580D2" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">Thorax. Dorsomedial length 1.3 times width of thorax. Lateral tomentum ash-gray, merging to yellow on upper part of anepisternum. Dorsal tomentum yellow, appearing dark gold, occasionally gray medially. Presutural outer vitta subquadrate, width two times that of inner vitta, connected to postsutural vittae. Presutural inner thoracic vittae bar-shaped, narrowly separated from outer vittae, rarely fused to outer vittae creating two presutural vittae. Postsutural vittae apparently fused into a single vitta covering most of area; tomentum visible on subquadrate area posteromedially and adjacent to supra-alar and intra-alar setae. Scutellar dorsal tomentum diffuse, yellow to gray, covering apical third to one-half of dorsal area. Postpronotum with 3 setae. Presutural area with 2 supra-alar setae. Postsutural area with 4 dorsocentral setae. Scutellum with 1 pair discal setae; the long clothing setulae occasionally seeming as a false second pair. Fore tibia with a single posterodorsal seta. Wing hyaline. Calypters tan to cream colored.</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BFB2EFC788143" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">Abdomen. Cuticle and setulae entirely black. Dorsal tomentum bands diffuse gray to yellow-gray, appearing gold in some individuals; covering one-third to one-half area of T35, narrower on T3 than T5; with distinct median vitta. Ventral tomentum bands ash-gray merging to yellow dorsally; covering three-fourths to entirety of T35. T4 with 2 pairs of median marginal setae and 2 pairs of lateral marginal setae.</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BFABEFB6E8262" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">
Male terminalia (
<figureCitation id="13142A25697B8F7CFE00FABEFD8F816F" box="[396,557,1374,1401]" captionStart="FIGURES 136155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="49" pageNumber="50">Figs 136, 152</figureCitation>
,
<figureCitation id="13142A25697B8F7CFDB7FABEFDCB816F" box="[571,617,1374,1401]" captionStart="FIGURES 156173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="49" pageNumber="50">169</figureCitation>
). Sternite 5 basal plate with median teeth separated 0.53 width of basal notch; apical lobes subequal to length of basal plate. Postgonite broadly rounded on posterior margin, slightly narrowed from base to digitiform apex, the anterior emargination subequal in depth to width of postgonite at mid length. Surstylus paddle shaped, 2.5 times as long as wide, gradually curved on margins to blunt apex; the basal lobe small with distinct angle on margin. Cercus in lateral view slightly dilated at base, then gradually narrowed to parallel sided, with blunt apex; in caudal view gradually narrowed to mid length, then subparallel to apex. Upper lobe subquadrate, inner margin curved; 0.29 length of cercus. Median section 0.41 length of cercus. Apical cleft parallel sided, 0.28 length of cercus. Syncercus apex width in caudal view 0.42 width of syncercus base.
</paragraph>
<paragraph id="8B9036A0697B8F7CFF4BF99EFCD0835B" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">
Female terminalia (
<figureCitation id="13142A25697B8F7CFE2EF99EFDA5828E" box="[418,519,1662,1688]" captionStart="FIGURES 94108" captionStartId="97.[151,264,1704,1728]" captionTargetBox="[264,1316,193,1668]" captionTargetId="figure-15@97.[251,1337,181,1680]" captionTargetPageId="97" captionText="FIGURES 94108. Female seventh abdominal segment of Eucelatoria ferox group spp., lateral view. Numbers are “ZLB_ E.Ferox” specimen IDs. 94. E. ferox (Townsend), Santa Cruz (00398) (USNM). 95. E. aurata (Townsend), Nova Teutonia (00250) (DMW). 96. E. auriceps (Aldrich), Crane Hollow (00410) (JOSC). 97. E. borealis sp. nov., Sheffeld (00041) (DMW). 98. E. charapensis (Townsend), Chapare Province (00393) (DMW). 99. E. falcata sp. nov., Nova Teutonia (00274) (CNC). 100. E. fordlandia sp. nov., Fordlandia (00311) (USNM). 101. E. gladiatrix (Townsend), Guanacaste Province (00376) (CNC). 102. E. sabroskyi sp. nov., Herb Martyr Campground (00084) (CNC). 103. E. strigata (Wulp), Trinidad (00138) (CNC). 104. E. texana (Reinhard), Madera Greaterville Rd. (00048) (JOSC). 105. E. yanayacu sp. nov., Yanayacu (no “E.Ferox” number, labeled “DIP328”) (USNM). 106. E. gustavogutierrezi sp. nov., Guanacaste Province (00007) (CNC). 107. E. kopis sp. nov., Avispas (00217) (CNC). 108. E. sica sp. nov., Yanayacu (00211) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601500" httpUri="https://zenodo.org/record/6601500/files/figure.png" pageId="49" pageNumber="50">Figs 106</figureCitation>
,
<figureCitation id="13142A25697B8F7CFD9FF99EFDE2828E" box="[531,576,1662,1688]" captionStart="FIGURES 109123" captionStartId="98.[151,264,1695,1719]" captionTargetBox="[221,1365,193,1657]" captionTargetId="figure-31@98.[209,1378,181,1672]" captionTargetPageId="98" captionText="FIGURES 109123. Female abdominal segments eight through ten of Eucelatoria ferox group spp., dorsal view. Numbers are “ZLB_E.Ferox” specimen IDs. 119. E. ferox (Townsend), Santa Cruz (00398) (USNM). 110. E. aurata (Townsend), Nova Teutonia (00250) (DMW). 111. E. auriceps (Aldrich), Crane Hollow (00410) (JOSC). 112. E. borealis sp. nov., Sheffeld (00041) (DMW). 113. E. charapensis (Townsend), Chapare Province (00393) (DMW). 114. E. falcata sp. nov., Nova Teutonia (00274) (CNC). 115. E. fordlandia sp. nov., Fordlandia (00311) (USNM). 116. E. gladiatrix (Townsend), Guanacaste Province (ZLB_ E.Ferox 00379) (CNC). 117. E. sabroskyi sp. nov., Herb Martyr Campground (00084) (CNC). 118. E. strigata (Wulp), Trinidad (00138) (CNC). 119. E. texana (Reinhard), Madera Greaterville Rd. (00048) (JOSC). 120. E. yanayacu sp. nov., Yanayacu (no “E.Ferox” number, labeled “DIP328”) (USNM). 121. E. gustavogutierrezi sp. nov., Guanacaste Province (00007) (CNC). 122. E. kopis sp. nov., Avispas (00217) (CNC). 123. E. sica sp. nov., Yanayacu (00211) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601502" httpUri="https://zenodo.org/record/6601502/files/figure.png" pageId="49" pageNumber="50">121</figureCitation>
). Piercer generally extending to base of T3 or beyond, often hidden between tergite edges, in lateral view sharply curved anterad near base, then gradually tapered to slightly bent apex; flat or concave anteriorally with strong ridge between anterior and lateral faces; in posterior view parallel sided on basal half, then gradually narrowed to tip. Aculeate lobe 2.5 times height of segment 7 base. End tergite halves irregularly fused medially, the unit more than four times as long as wide. Cercus with 6 setae, the ventral elongation subequal in length to width of cercus body. Postgenital plate with 10 setae.
</paragraph>
</subSubSection>
<subSubSection id="C335652B697B8F7FFF4BF8B6FBE884C3" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="biology_ecology">
<paragraph id="8B9036A0697B8F7CFF4BF8B6FE7483CA" blockId="49.[151,1437,150,2013]" pageId="49" pageNumber="50">
<emphasis id="B95BEAB2697B8F7CFF4BF8B6FEB38366" bold="true" box="[199,273,1878,1904]" pageId="49" pageNumber="50">Hosts.</emphasis>
Thirteen individuals have been reared from caterpillars of
<taxonomicName id="4C2F4D23697B8F7CFC4CF8B7FABA8367" box="[960,1304,1878,1905]" class="Magnoliopsida" family="Fabaceae" genus="Inga" kingdom="Plantae" order="Fabales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B95BEAB2697B8F7CFC4CF8B7FC518366" box="[960,1011,1879,1904]" italics="true" pageId="49" pageNumber="50">Inga</emphasis>
Mill. sp. (Oecophoridae)
</taxonomicName>
feeding on
<taxonomicName id="4C2F4D23697B8F7CFF1BF89BFD158383" authority="Donn. Sm." authorityName="Donn. Sm." box="[151,695,1914,1941]" class="Magnoliopsida" family="Nyctaginaceae" genus="Neea" kingdom="Plantae" order="Caryophyllales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="psychotrioides">
<emphasis id="B95BEAB2697B8F7CFF1BF89BFEDF8382" box="[151,381,1914,1940]" italics="true" pageId="49" pageNumber="50">Neea psychotrioides</emphasis>
Donn.Sm. (Nyctaginaceae)
</taxonomicName>
(
<bibRefCitation id="EFBE4B51697B8F7CFD44F89AFC5A8382" author="Janzen, D. H. &amp; Hallwachs, W." box="[712,1016,1914,1940]" pageId="49" pageNumber="50" refId="ref67919" refString="Janzen, D. H. &amp; Hallwachs, W. (2008) Event-based database of caterpillars, their food plants, and their parasitoids in Area de Conservacion Guanacaste, Northwestern Costa Rica. Available from: http: // janzen. sas. upenn. edu (accessed 11 February 2015)" type="url" year="2008">Janzen &amp; Hallwachs 2008</bibRefCitation>
), and one individual has been reared from
<taxonomicName id="4C2F4D23697B8F7CFF5AF87EFD1C83AF" authority="Druce" authorityName="Druce" box="[214,702,1950,1977]" class="Insecta" family="Crambidae" genus="Pantographa" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="suffusalis">
<emphasis id="B95BEAB2697B8F7CFF5AF87EFE7E83AE" box="[214,476,1950,1976]" italics="true" pageId="49" pageNumber="50">Pantographa suffusalis</emphasis>
Druce (Crambidae)
</taxonomicName>
feeding on
<taxonomicName id="4C2F4D23697B8F7CFCCBF87FFF7183CB" authority="(Donn. Sm.) Standl." authorityName="Standl." baseAuthorityName="Donn. Sm." class="Magnoliopsida" family="Malvaceae" genus="Hampea" kingdom="Plantae" order="Malvales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="appendiculata">
<emphasis id="B95BEAB2697B8F7CFCCBF87FFBEC83AE" box="[839,1102,1950,1976]" italics="true" pageId="49" pageNumber="50">Hampea appendiculata</emphasis>
(Donn. Sm.) Standl. (Malvaceae)
</taxonomicName>
(
<bibRefCitation id="EFBE4B51697B8F7CFF6DF822FE6983CA" author="Stireman, J. O. III &amp; Dyer, L. A. &amp; Greeney, H. F." box="[225,459,1986,2012]" pageId="49" pageNumber="50" pagination="367 - 384" refId="ref68830" refString="Stireman, J. O. III, Dyer, L. A. &amp; Greeney, H. F. (2017) Specialised generalists? Food web structure of a tropical tachinid-caterpillar community. Insect Conservation and Diversity, 10 (5), 367 - 384. https: // doi. org / 10.1111 / icad. 12238" type="journal article" year="2017">
Stireman
<emphasis id="B95BEAB2697B8F7CFEC3F823FE2583CA" box="[335,391,1986,2012]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9036A069788F7FFF4BFF76FBE884C3" blockId="50.[151,1437,150,392]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF4BFF76FD7C84A6" bold="true" box="[199,734,150,176]" pageId="50" pageNumber="51">Geographic extent and seasonal occurrence.</emphasis>
Individuals of
<taxonomicName id="4C2F4D2369788F7FFC1CFF77FBD784A6" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[912,1141,151,176]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="gustavogutierrezi" status="sp. nov.">
<emphasis id="B95BEAB269788F7FFC1CFF77FBD784A6" box="[912,1141,151,176]" italics="true" pageId="50" pageNumber="51">E. gustavogutierrezi</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269788F7FFBF2FF76FB7984A6" bold="true" box="[1150,1243,150,176]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A26857C969788F7FFBF2FF76FB7984A6" box="[1150,1243,150,176]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are known from montane and moist lowland forests in
<collectingCountry id="F338763069788F7FFDCFFF5AFD1F84C2" box="[579,701,186,212]" name="Costa Rica" pageId="50" pageNumber="51">Costa Rica</collectingCountry>
at
<quantity id="4CD79B4569788F7FFD53FF5AFCFC84C3" box="[735,862,186,213]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="12.0" metricValueMin="0.5" pageId="50" pageNumber="51" unit="m" value="625.0" valueMax="1200.0" valueMin="50.0">501200 m</quantity>
elevation (
<figureCitation id="13142A2569788F7FFC57FF5AFB9F84C3" box="[987,1085,186,213]" captionStart="FIGURES 174179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="50" pageNumber="51">Fig. 176</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335652B69788F7FFF4BFF3EFF5B859E" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8B9036A069788F7FFF4BFF3EFF5B859E" blockId="50.[151,1437,150,392]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF4BFF3EFEEE84EE" bold="true" box="[199,332,222,248]" pageId="50" pageNumber="51">Discussion.</emphasis>
This species includes “Janzen06” in the Area de Conservación
<collectingRegion id="49EBF84269788F7FFBBEFF3EFB1184EE" box="[1074,1203,222,248]" country="Costa Rica" name="Guanacaste" pageId="50" pageNumber="51">Guanacaste</collectingRegion>
database (
<bibRefCitation id="EFBE4B5169788F7FFAA2FF3EFEF0850A" author="Janzen, D. H. &amp; Hallwachs, W." pageId="50" pageNumber="51" refId="ref67919" refString="Janzen, D. H. &amp; Hallwachs, W. (2008) Event-based database of caterpillars, their food plants, and their parasitoids in Area de Conservacion Guanacaste, Northwestern Costa Rica. Available from: http: // janzen. sas. upenn. edu (accessed 11 February 2015)" type="url" year="2008">Janzen &amp; Hallwachs 2008</bibRefCitation>
). Genetic data show no population structure; one individual from La Selva (ZLB_E.Ferox 00405) does not differ genetically from those in
<collectingRegion id="49EBF84269788F7FFDD2FEC6FCEA8556" box="[606,840,294,320]" country="Costa Rica" name="Guanacaste" pageId="50" pageNumber="51">Guanacaste province</collectingRegion>
(
<figureCitation id="13142A2569788F7FFCDBFEC6FC3A8556" box="[855,920,294,320]" captionStart="FIGURE 1" captionStartId="9.[151,250,1766,1790]" captionTargetBox="[164,1421,431,1729]" captionTargetId="figure-49@9.[153,1434,417,1743]" captionTargetPageId="9" captionText="FIGURE 1. CO1-based neighbor-joining tree of 12 species of the Eucelatoria ferox species group, with inset illustrations of mesonota. The outgroup is Eucelatoria armigera (Coquillett). Node labels indicate support values over 60. Scale bar = 2% genetic difference. The E. charapensis (Townsend) sequence with the axial break is of poor quality but is included to show unity of El Bigal specimens with the rest of the metapopulation. Mesonota are illustrated with setal annuli only for clarity. A. E. gustavogutierrezi sp. nov. B. E. kopis sp. nov. C. E. ritavargasae sp. nov. D. E. ferox (Townsend). E. E. huitepecensis sp. nov. F. E. auriceps (Aldrich). G. E. gladiatrix (Townsend). H. E. borealis sp. nov. I. E. yanayacu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601406" httpUri="https://zenodo.org/record/6601406/files/figure.png" pageId="50" pageNumber="51">Fig. 1</figureCitation>
). The extent of yellow tomentum on the parafacial seems to vary within this species, as does the shape of the presutural thoracic vittae, but other characters are uniform.
</paragraph>
</subSubSection>
</treatment>
</document>