331 lines
36 KiB
XML
331 lines
36 KiB
XML
<document id="0A4BBBCC1330A8E73EC9FDA394A0C6C3" ID-CLB-Dataset="5605" ID-DOI="10.11646/zootaxa.3795.5.2" ID-GBIF-Dataset="87d17125-337d-4679-bdcd-102f6b022ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915083" ID-ZooBank="9394634C-9836-4973-868B-BDEE414E4EA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623198152603" checkinUser="felipe" docAuthor="Petzold, Alice, Vargas-Ramírez, Mario, Kehlmaier, Christian, Vamberger, Melita, Branch, William R., Preez, Louis Du, Hofmeyr, Margaretha D., Meyer, Leon, Schleicher, Alfred, Široký, Pavel & Fritz, Uwe" docDate="2014" docId="CB02879FF923FFD4FF74FE47FED1F905" docLanguage="en" docName="zt03795p548.pdf" docOrigin="Zootaxa 3795 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelomedusa variabilis Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="543" masterDocId="373BFFE7F937FFC0FFE3FFB8FFC9FFBF" masterDocTitle="A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species" masterLastPageNumber="548" masterPageNumber="523" pageNumber="543" updateTime="1699006367747" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="F7F81D7A13261D3BDC8110411F38FEAD" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9CD97F62D5CEBD6C0DD13E0D73337095">
|
||
<mods:title id="BA15608A3EECA11776897C333342D7F7">A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2C439E029397BB23E991FCD556204C6C" type="personal">
|
||
<mods:role id="A90E2927912C57569597794E0F0449F7">
|
||
<mods:roleTerm id="D32F0C9366CAC402A2CFFB90F22280F8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A9100F4537B1D80200B3AC5FC7349144">Petzold, Alice</mods:namePart>
|
||
<mods:affiliation id="D193F3B77E2FFDB155E97E2B0152100A">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="71D686F03FD72B9F5FC6E6C3C084BC79" type="personal">
|
||
<mods:role id="05F671C8853EBC1592073C3696CC935D">
|
||
<mods:roleTerm id="2E1E04F1F07B64C272B3B38407AD861F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="806B066E9F18A8200150720DDE5BF5DF">Vargas-Ramírez, Mario</mods:namePart>
|
||
<mods:affiliation id="3767DD99BD40FA5D73D69CD20412B798">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="043B58DC51507BE6B704D2EB7550527E" type="personal">
|
||
<mods:role id="62CAAEB4D0209BD96B070AE651EF5E0F">
|
||
<mods:roleTerm id="8D4A49B6FB922CF788995CD6C952617A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9422659A665E77EF7605DD49D1245A69">Kehlmaier, Christian</mods:namePart>
|
||
<mods:affiliation id="2E238193B9129DFB16BB4F7D2A5D1D3A">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="8C4E326A7DBF2F484B591F7935921541" type="personal">
|
||
<mods:role id="DBC9ED2CC8D730906B53C6EE7CBD87A5">
|
||
<mods:roleTerm id="27BDF9648F21EBDF7CAF79FAA6055FDF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FC2FD825D9D9D0735171BB3AB82FB901">Vamberger, Melita</mods:namePart>
|
||
<mods:affiliation id="821D1368535D56273DA340C436C0CB3B">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="CC669780D7F150E5CC3565F2A109A154" type="personal">
|
||
<mods:role id="B44A6E9ECF8860F7B3FC40194C8DFDFF">
|
||
<mods:roleTerm id="F389F4B7169C355751A74E946883254A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="216C603B257B25361662A8C0356FD28F">Branch, William R.</mods:namePart>
|
||
<mods:affiliation id="D74BCA71F6973E60AE847ED221057F9E">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="8846B86DE5BE464F1E85BCA083DD4613" type="personal">
|
||
<mods:role id="874D83E4634A53ECB8039FC433981006">
|
||
<mods:roleTerm id="4DBFE96CDFB4986993B81CBFF9998189">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="54196FD5C509A5B5308400F6435D00A9">Preez, Louis Du</mods:namePart>
|
||
<mods:affiliation id="B59C52B0AD320160F301D9AD0F1785D9">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="66D6183D45271B04843ECA672C256B0F" type="personal">
|
||
<mods:role id="FE6E01447654CD83799AA749647843E8">
|
||
<mods:roleTerm id="EEB30F9780E46AAC42DADA5E0911E59F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="95B67647AAB5F11816ECD4ECA91B29B8">Hofmeyr, Margaretha D.</mods:namePart>
|
||
<mods:affiliation id="B902E4C9082374527B0610297053A29A">Chelonian Biodiversity and Conservation, Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X 17, Bellville 7535, South Africa</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="D9364489ACF3EFA10AF9EB00C6C06D86" type="personal">
|
||
<mods:role id="49082EBA4501B3F65CFA04C450F3899F">
|
||
<mods:roleTerm id="1A829652AD8D61E836380EF05D409FF0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C481E087A937E8BCC8F0725D1FB8F4AD">Meyer, Leon</mods:namePart>
|
||
<mods:affiliation id="D9BBEC46320F53CF5FEC8E84DBD74E16">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="49DB575D3E504800AE92CBC0401CF9B0" type="personal">
|
||
<mods:role id="6C80A6740596F2DCF7ED339CC50ABCCD">
|
||
<mods:roleTerm id="851FD2F5F5BCD5A9625DC4A5D652698D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="861828868FA7469A18DC4152EA1205CB">Schleicher, Alfred</mods:namePart>
|
||
<mods:affiliation id="989149BA55ACBAC34DF4CBE76F6BEDB2">P. O. Box 30566, Windhoek, Namibia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="55C9AA15CE59EBA8CFDC3B80AAEB1FDA" type="personal">
|
||
<mods:role id="AE09C98C5FC8B35AA068B8952C7FED30">
|
||
<mods:roleTerm id="3067041E70D30AA0EEABD9A787EB9B11">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C5F5D62BCEBAEC99E37B2AF05DFC224F">Široký, Pavel</mods:namePart>
|
||
<mods:affiliation id="FA190CA3CF2303B9ED42C63E91F71A0F">Department of Biology and Wildlife Diseases, Faculty of Veterinary Hygiene and Ecology, University of Veterinary and Pharmaceutical Sciences, Palackého tř. 1 / 3, 612 42 Brno, Czech Republic</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="991D76440787EC349ACA624EBBF809B2" type="personal">
|
||
<mods:role id="C54A5168BC6088279FDE0B628640D13C">
|
||
<mods:roleTerm id="A209E0275BC35A0B51FB88D5A8A493F1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5BDA42D770271A4DC6794DEBA2A19509">Fritz, Uwe</mods:namePart>
|
||
<mods:affiliation id="A330F7187993C519BECCB62929CAC2C4">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A777200904692DA7BAC937E77F88403B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3ECA0A4C2689BCA5DC2929F486671AAD" type="host">
|
||
<mods:titleInfo id="2E3034F2448F5F80C19CB3737C757B36">
|
||
<mods:title id="755C8686A24FD6B523287F53B5521222">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="18403BB2CAF08165C4B9D7008C102D4B">
|
||
<mods:date id="1FFD8B594E297CC94703DA3009F9E73E">2014</mods:date>
|
||
<mods:detail id="7400F205A99B6E607A31A3F66CD0210C" type="pubDate">
|
||
<mods:number id="BB45950F764597D418FD4F70CC3584FE">2014-05-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AD1E33E21B79E3FC3E1EF8BE9B35A4A5" type="volume">
|
||
<mods:number id="17F9AC3459ED7480793F0916A83060F0">3795</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6BF92C2EFABCD7FF23DA572E8A9C2F37" type="issue">
|
||
<mods:number id="C666CBC0CF7A78B179AEE73E235C7A79">5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="523C6CA6F9A703F703B1605DF19E5436" unit="page">
|
||
<mods:start id="51706EED212B834C17AE9F008384E493">523</mods:start>
|
||
<mods:end id="49A3FDDCDAA99F8289F726D2370B8DCE">548</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="9E5A77ABFFC90F4831D79838D4EC8313">journal article</mods:classification>
|
||
<mods:identifier id="59FFB86124CF825E8CA4B8254C73A74A" type="CLB-Dataset">5605</mods:identifier>
|
||
<mods:identifier id="0639DEBCA81E72F99EAB0B31FC86E471" type="DOI">10.11646/zootaxa.3795.5.2</mods:identifier>
|
||
<mods:identifier id="3B647811118027382C547EF5834BD082" type="GBIF-Dataset">87d17125-337d-4679-bdcd-102f6b022ad2</mods:identifier>
|
||
<mods:identifier id="FE27DE709843D4AA892E1F34033F6119" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1991FA85B099484985C85E4E299CA346" type="Zenodo-Dep">4915083</mods:identifier>
|
||
<mods:identifier id="631D890771C0AD7278840AFC075BE93B" type="ZooBank">9394634C-9836-4973-868B-BDEE414E4EA8</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CB02879FF923FFD4FF74FE47FED1F905" ID-DOI="http://doi.org/10.5281/zenodo.4915103" ID-GBIF-Taxon="183403863" ID-Zenodo-Dep="4915103" LSID="urn:lsid:plazi:treatment:CB02879FF923FFD4FF74FE47FED1F905" httpUri="http://treatment.plazi.org/id/CB02879FF923FFD4FF74FE47FED1F905" lastPageNumber="543" pageId="20" pageNumber="543">
|
||
<subSubSection id="0BB16502F923FFD4FF74FE47FDCFFDA5" box="[151,518,511,538]" pageId="20" pageNumber="543" type="nomenclature">
|
||
<paragraph id="43143689F923FFD4FF74FE47FDCFFDA5" blockId="20.[151,518,511,538]" box="[151,518,511,538]" pageId="20" pageNumber="543">
|
||
<heading id="185C81E5F923FFD4FF74FE47FDCFFDA5" bold="true" box="[151,518,511,538]" fontSize="11" level="1" pageId="20" pageNumber="543" reason="1">
|
||
<emphasis id="71DFEA9BF923FFD4FF74FE47FDCFFDA5" bold="true" box="[151,518,511,538]" pageId="20" pageNumber="543">
|
||
<taxonomicName id="84AB4D0AF923FFD4FF74FE47FE68FDA6" authority="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz, 2014" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[151,417,511,537]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis" status="sp. nov.">
|
||
<emphasis id="71DFEA9BF923FFD4FF74FE47FE68FDA6" bold="true" box="[151,417,511,537]" italics="true" pageId="20" pageNumber="543">Pelomedusa variabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F923FFD4FE4BFDB8FDCFFDA5" box="[424,518,512,538]" pageId="20" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF74FDFEFDBDFCE4" pageId="20" pageNumber="543" type="diagnosis">
|
||
<paragraph id="43143689F923FFD4FF74FDFEFDBDFCE4" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<emphasis id="71DFEA9BF923FFD4FF74FDFEFEDAFDE0" bold="true" box="[151,275,582,607]" pageId="20" pageNumber="543">Diagnosis:</emphasis>
|
||
Medium- to large-sized helmeted terrapins with a known maximum straight carapacial length of
|
||
<quantity id="84539B6CF923FFD4FA88FDFEFF74FD3D" metricMagnitude="-1" metricUnit="m" metricValue="2.48" pageId="20" pageNumber="543" unit="cm" value="24.8">24.8 cm</quantity>
|
||
. Pectoral scutes mostly triangular without midseam contact, but terrapins with pectorals having narrow or wide midseam contact occur. One large undivided temporal head scale. Two small barbels under chin. Colouration extremely variable; very dark individuals are known, as well as specimens with light horn-coloured carapace and completely yellow plastron. Soft parts ventrally lighter than dorsally.
|
||
<taxonomicName id="84AB4D0AF923FFD4FC48FD6EFB6EFD50" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[939,1191,726,751]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="71DFEA9BF923FFD4FC48FD6EFB6EFD50" box="[939,1191,726,751]" italics="true" pageId="20" pageNumber="543">Pelomedusa variabilis</emphasis>
|
||
</taxonomicName>
|
||
differs from all other
|
||
<taxonomicName id="84AB4D0AF923FFD4FF74FD42FEB0FCAC" box="[151,377,762,787]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF923FFD4FF74FD42FEE9FCAC" box="[151,288,762,787]" italics="true" pageId="20" pageNumber="543">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the presence of adenine (A) instead of cytosine (C) or thymine (T) at position 189 and by the presence of adenine (A) instead of guanine (G) at position 322 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF24FCDFFD94FC1C" pageId="20" pageNumber="543" type="materials_examined">
|
||
<paragraph id="43143689F923FFD4FF24FCDFFD94FC1C" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<materialsCitation id="F3C33CD4F923FFD4FF24FCDFFD94FC1C" ID-GBIF-Occurrence="3319794316" accessionNumber="SMF 58075" collectingDate="1957-02-18" collectionCode="SMF" collectorName="H. Lang" country="Ghana" location="Senckenberg-Museum" municipality="Frankfurt am Main" pageId="20" pageNumber="543" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FCDFFEF4FCC0" bold="true" box="[199,317,871,895]" pageId="20" pageNumber="543">
|
||
<typeStatus id="9C10882BF923FFD4FF24FCDFFEF1FCC0" box="[199,312,871,895]" pageId="20" pageNumber="543" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<location id="46746052F923FFD4FEA0FCDEFD8BFCC0" LSID="urn:lsid:plazi:treatment:CB02879FF923FFD4FF74FE47FED1F905:46746052F923FFD4FEA0FCDEFD8BFCC0" box="[323,578,870,895]" country="Ghana" municipality="Frankfurt am Main" name="Senckenberg-Museum" pageId="20" pageNumber="543">Senckenberg-Museum</location>
|
||
,
|
||
<collectingMunicipality id="A370ACF3F923FFD4FDAEFCDEFCE8FCC0" box="[589,801,870,895]" pageId="20" pageNumber="543">Frankfurt am Main</collectingMunicipality>
|
||
(
|
||
<collectionCode id="25BAAE4CF923FFD4FCCCFCDEFCA0FCC0" box="[815,873,870,895]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="20" pageNumber="543">SMF</collectionCode>
|
||
58075, subadult male,
|
||
<collectingCountry id="3BBC7619F923FFD4FB8EFCDEFB26FCC0" box="[1133,1263,870,895]" name="Ghana" pageId="20" pageNumber="543">Gold Coast</collectingCountry>
|
||
,
|
||
<collectingCountry id="3BBC7619F923FFD4FB1AFCDEFA8FFCC0" box="[1273,1350,870,895]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
; leg.
|
||
<collectorName id="EE5E535FF923FFD4FA62FCDFFF1AFC1C" pageId="20" pageNumber="543">H. Lang</collectorName>
|
||
,
|
||
<date id="37151049F923FFD4FF3DFC32FE61FC1C" box="[222,424,906,931]" pageId="20" pageNumber="543" value="1957-02-18">
|
||
<collectingDate id="2751E9A1F923FFD4FF3DFC32FE61FC1C" box="[222,424,906,931]" pageId="20" pageNumber="543" value="1957-02-18">18 February 1957</collectingDate>
|
||
</date>
|
||
;
|
||
<figureCitation id="DB902A0CF923FFD4FE57FC32FE3CFC1C" box="[436,501,906,931]" captionStart="FIGURE 5" captionStartId="18.[151,250,1918,1940]" captionTargetBox="[226,1382,244,1853]" captionTargetId="figure-68@18.[151,1436,145,1884]" captionTargetPageId="18" captionText="FIGURE 5. Dorsal and ventral aspects of the holotype of Pelomedusa somalica sp. nov. (BMNH 1970.1481, adult, Borama district, Awdal, Somaliland/Somalia; top) and of Pelomedusa variabilis sp. nov. (SMF 58075, subadult male, Gold Coast, Ghana; bottom). Scale bars, 3 cm. Photos: A. Petzold and E. Morawa." figureDoi="http://doi.org/10.5281/zenodo.4915095" httpUri="https://zenodo.org/record/4915095/files/figure.png" pageId="20" pageNumber="543">Fig. 5</figureCitation>
|
||
bottom).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF24FC16FC23FB7C" pageId="20" pageNumber="543" type="description">
|
||
<paragraph id="43143689F923FFD4FF24FC16FBA4FB8C" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FC16FDEAFC78" bold="true" box="[199,547,942,967]" pageId="20" pageNumber="543">
|
||
Description of the
|
||
<typeStatus id="9C10882BF923FFD4FE57FC17FDD6FC78" box="[436,543,943,967]" pageId="20" pageNumber="543" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Straight carapacial length
|
||
<quantity id="84539B6CF923FFD4FC86FC16FC01FC79" box="[869,968,942,967]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="20" pageNumber="543" unit="cm" value="10.3">10.3 cm</quantity>
|
||
, plastral length
|
||
<quantity id="84539B6CF923FFD4FB69FC16FB16FC79" box="[1162,1247,942,967]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="20" pageNumber="543" unit="cm" value="8.7">8.7 cm</quantity>
|
||
. Pectoral scutes triangular, widely separated, without midseam contact. Temporal scales undivided; injury on the right side of head. Two small barbels under chin. Carapace covered by algae; plastron dark brown, seams distinctly lighter. Soft parts dorsally dark grey; throat cream-coloured, ventral side of legs greyish brown. Tail long.
|
||
</paragraph>
|
||
<paragraph id="43143689F923FFD4FF24FB87FC23FB7C" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<materialsCitation id="F3C33CD4F923FFD4FF24FB87FC2FFB7C" ID-GBIF-Occurrence="3319794310" accessionNumber="T 35041" collectionCode="MTD, T, BMHN" country="Ghana" county="The Natural History Museum" location="Zoologisches Forschungsmuseum Alexander Koenig" municipality="London" pageId="20" pageNumber="543" specimenCode="T 33818-33819, BMNH 1863.3, ZFMK 47521-47523" specimenCount="1" stateProvince="Greater Accra" typeStatus="paratype">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FB87FE81FBE8" bold="true" box="[199,328,1087,1111]" pageId="20" pageNumber="543">
|
||
<typeStatus id="9C10882BF923FFD4FF24FB87FE8AFBE8" box="[199,323,1087,1111]" pageId="20" pageNumber="543" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Museum für Tierkunde, Senckenberg Dresden (
|
||
<collectionCode id="25BAAE4CF923FFD4FC93FB86FC79FBE8" box="[880,944,1086,1111]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="20" pageNumber="543">MTD</collectionCode>
|
||
<specimenCode id="130D9EF2F923FFD4FC5AFB87FBB9FBE8" box="[953,1136,1086,1111]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="543">T 33818–33819</specimenCode>
|
||
, male and female,
|
||
<collectingCountry id="3BBC7619F923FFD4FAA8FB86FA51FBE8" box="[1355,1432,1086,1111]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
;
|
||
<collectionCode id="25BAAE4CF923FFD4FF74FBDBFF1CFBC5" box="[151,213,1123,1146]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="20" pageNumber="543">MTD</collectionCode>
|
||
<collectionCode id="25BAAE4CF923FFD4FF01FBDBFF3AFBC5" box="[226,243,1123,1146]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="543">T</collectionCode>
|
||
35041, juvenile,
|
||
<collectingCountry id="3BBC7619F923FFD4FE25FBDAFDDDFBC4" box="[454,532,1122,1147]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
);
|
||
<collectingCounty id="AA754E05F923FFD4FDCEFBDAFC42FBC5" box="[557,907,1122,1147]" pageId="20" pageNumber="543">The Natural History Museum</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="A370ACF3F923FFD4FC78FBDAFC3BFBC4" box="[923,1010,1122,1147]" pageId="20" pageNumber="543">London</collectingMunicipality>
|
||
(
|
||
<specimenCode id="130D9EF2F923FFD4FBE5FBDAFB73FBC4" box="[1030,1210,1122,1147]" collectionCode="BMNH" pageId="20" pageNumber="543">BMNH 1863.3</specimenCode>
|
||
.27.1, male,
|
||
<collectingCountry id="3BBC7619F923FFD4FAA8FBDAFA51FBC4" box="[1355,1432,1122,1147]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
;
|
||
<collectionCode id="25BAAE4CF923FFD4FF74FB3FFF22FB21" box="[151,235,1159,1182]" pageId="20" pageNumber="543">BMHN</collectionCode>
|
||
1927.8.27.240–242, juveniles,
|
||
<collectingCountry id="3BBC7619F923FFD4FD88FB3EFD70FB20" box="[619,697,1158,1183]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
);
|
||
<location id="46746052F923FFD4FD34FB3EFA83FB20" LSID="urn:lsid:plazi:treatment:CB02879FF923FFD4FF74FE47FED1F905:46746052F923FFD4FD34FB3EFA83FB20" box="[727,1354,1158,1183]" country="Ghana" county="The Natural History Museum" municipality="London" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="543" stateProvince="Greater Accra">Zoologisches Forschungsmuseum Alexander Koenig</location>
|
||
,
|
||
<location id="46746052F923FFD4FA83FB3FFA55FB21" LSID="urn:lsid:plazi:treatment:CB02879FF923FFD4FF74FE47FED1F905:46746052F923FFD4FA83FB3FFA55FB21" box="[1376,1436,1159,1182]" country="Ghana" county="The Natural History Museum" municipality="London" name="Bonn" pageId="20" pageNumber="543" stateProvince="Greater Accra">Bonn</location>
|
||
(
|
||
<specimenCode id="130D9EF2F923FFD4FF7DFB12FE5BFB7C" box="[158,402,1194,1219]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="543" type="Museum">ZFMK 47521–47523</specimenCode>
|
||
, juveniles,
|
||
<location id="46746052F923FFD4FDF3FB12FD48FB7C" LSID="urn:lsid:plazi:treatment:CB02879FF923FFD4FF74FE47FED1F905:46746052F923FFD4FDF3FB12FD48FB7C" box="[528,641,1194,1219]" country="Ghana" county="The Natural History Museum" municipality="London" name="Achimota" pageId="20" pageNumber="543" stateProvince="Greater Accra">Achimota</location>
|
||
,
|
||
<collectingRegion id="816FF86BF923FFD4FD6EFB12FC4CFB7C" box="[653,901,1194,1219]" country="Ghana" name="Greater Accra" pageId="20" pageNumber="543">Greater Accra Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F923FFD4FC73FB12FC17FB7C" box="[912,990,1194,1219]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF24FB76FE34FAB4" pageId="20" pageNumber="543" type="etymology">
|
||
<paragraph id="43143689F923FFD4FF24FB76FE34FAB4" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FB76FE63FB58" bold="true" box="[199,426,1230,1255]" pageId="20" pageNumber="543">Derivatio nominis:</emphasis>
|
||
The species name
|
||
<taxonomicName id="84AB4D0AF923FFD4FD6EFB76FD31FB58" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[653,760,1230,1255]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="71DFEA9BF923FFD4FD6EFB76FD31FB58" box="[653,760,1230,1255]" italics="true" pageId="20" pageNumber="543">variabilis</emphasis>
|
||
</taxonomicName>
|
||
is an adjective in female gender, referring to the variable morphology of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF24FAAEFCBAFAEC" pageId="20" pageNumber="543" type="distribution">
|
||
<paragraph id="43143689F923FFD4FF24FAAEFCBAFAEC" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FAAEFEABFA90" bold="true" box="[199,354,1302,1327]" pageId="20" pageNumber="543">Distribution:</emphasis>
|
||
Genetically verified records for
|
||
<taxonomicName id="84AB4D0AF923FFD4FD01FAAFFCA4FA90" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[738,877,1302,1327]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="71DFEA9BF923FFD4FD01FAAFFCA4FA90" box="[738,877,1302,1327]" italics="true" pageId="20" pageNumber="543">P. variabilis</emphasis>
|
||
</taxonomicName>
|
||
are only known for
|
||
<collectingCountry id="3BBC7619F923FFD4FB81FAAEFB62FA90" box="[1122,1195,1302,1327]" name="Ghana" pageId="20" pageNumber="543">Ghana</collectingCountry>
|
||
and the
|
||
<collectingCountry id="3BBC7619F923FFD4FAF6FAAFFA55FA90" box="[1301,1436,1302,1327]" name="Ivory Coast" pageId="20" pageNumber="543">Ivory Coast</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F923FFD4FF7DFA82FE1CFAEC" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[158,469,1338,1363]" pageId="20" pageNumber="543" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF923FFD4FEB8FA83FE5AFAEC" box="[347,403,1338,1363]" italics="true" pageId="20" pageNumber="543">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="273A4B78F923FFD4FE02FA83FD6DFAEC" author="Wong, R. A. & Fong, J. J. & Papenfuss, T. J." box="[481,676,1338,1363]" pageId="20" pageNumber="543" pagination="575 - 585" refId="ref19006" refString="Wong, R. A., Fong, J. J. & Papenfuss, T. J. (2010) Phylogeography of the African helmeted terrapin, Pelomedusa subrufa: genetic structure, dispersal, and human introduction. Proceedings of the California Academy of Sciences, 61, 575 - 585." type="journal article" year="2010">
|
||
Wong
|
||
<emphasis id="71DFEA9BF923FFD4FDC9FA83FDABFAEC" box="[554,610,1338,1363]" italics="true" pageId="20" pageNumber="543">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="273A4B78F923FFD4FD53FA82FCA1FAEC" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[688,872,1338,1363]" pageId="20" pageNumber="543" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF923FFD4FD0FFA83FCEDFAEC" box="[748,804,1338,1363]" italics="true" pageId="20" pageNumber="543">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F923FFD4FF24FAE7FED1F905" pageId="20" pageNumber="543" type="discussion">
|
||
<paragraph id="43143689F923FFD4FF24FAE7FAF4FA24" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FAE7FEF4FAC8" bold="true" box="[199,317,1375,1399]" pageId="20" pageNumber="543">Remarks:</emphasis>
|
||
<taxonomicName id="84AB4D0AF923FFD4FEA0FAE6FDF7FAC8" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[323,574,1374,1399]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="71DFEA9BF923FFD4FEA0FAE6FDF7FAC8" box="[323,574,1374,1399]" italics="true" pageId="20" pageNumber="543">Pelomedusa variabilis</emphasis>
|
||
</taxonomicName>
|
||
corresponds to mtDNA lineage II of
|
||
<bibRefCitation id="273A4B78F923FFD4FC3EFAE6FAEAFAC8" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[989,1315,1374,1399]" pageId="20" pageNumber="543" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF923FFD4FB74FAE7FB06FAC8" box="[1175,1231,1374,1399]" italics="true" pageId="20" pageNumber="543">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
. It belongs to the northern clade of
|
||
<taxonomicName id="84AB4D0AF923FFD4FE40FA3AFDE5FA24" authorityName="Wagler" authorityYear="1830" box="[419,556,1410,1435]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="genus">
|
||
<emphasis id="71DFEA9BF923FFD4FE40FA3AFDE5FA24" box="[419,556,1410,1435]" italics="true" pageId="20" pageNumber="543">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
and is according to our mtDNA analyses sister to
|
||
<taxonomicName id="84AB4D0AF923FFD4FB83FA3BFB12FA24" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[1120,1243,1410,1435]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="olivacea">
|
||
<emphasis id="71DFEA9BF923FFD4FB83FA3BFB12FA24" box="[1120,1243,1410,1435]" italics="true" pageId="20" pageNumber="543">P. olivacea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DB902A0CF923FFD4FB09FA3AFAE2FA24" box="[1258,1323,1410,1435]" captionStart="FIGURE 1" captionStartId="3.[151,250,999,1021]" captionTargetBox="[167,1426,219,969]" captionTargetId="figure-480@3.[151,1436,199,975]" captionTargetPageId="3" captionText="FIGURE 1. Maximum Likelihood tree for helmeted terrapins (Pelomedusa spp.) using 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. Terminal clades collapsed to cartoons. The topology of the Bayesian 50% majority rule consensus tree was identical. Numbers along branches indicate bootstrap support and clade support under BI (posterior probabilities) greater than 50 or 0.95, respectively. Asterisks represent maximum support under both methods. Colours and symbol correspond to the map (Fig. 2). On the right, proposed species names indicated; in brackets, mtDNA lineages. For clade membership of individual samples, see Table S1. The questionable sample MTD T 5484 from Swellendam District (Western Cape), South Africa, is highlighted in red (see text). The shown terrapins are Pelomedusa neumanni (top; Kakamega, Kenya) and P. galeata (bottom; Port Elizabeth, South Africa—photos: H. Prokop and W.R. Branch)." figureDoi="http://doi.org/10.5281/zenodo.4915085" httpUri="https://zenodo.org/record/4915085/files/figure.png" pageId="20" pageNumber="543">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="43143689F923FFD4FF24FA1EFE50F9CD" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
<materialsCitation id="F3C33CD4F923FFD4FF24FA1EFE5CF9CD" ID-GBIF-Occurrence="3319794302" accessionNumber="D 33819" collectionCode="MTD" collectorName="M. - O. Rodel" country="Ivory Coast" location="Ivory Coast" pageId="20" pageNumber="543" specimenCode="BMNH 1863.3" specimenCount="11" typeStatus="paratype">
|
||
Male helmeted terrapins from the
|
||
<collectingCountry id="3BBC7619F923FFD4FDADFA1FFD1DFA00" box="[590,724,1446,1471]" name="Ivory Coast" pageId="20" pageNumber="543">Ivory Coast</collectingCountry>
|
||
were reported to have red-spotted nearly white heads (
|
||
<bibRefCitation id="273A4B78F923FFD4FAAFFA1EFEEAFA5C" author="Bull, J. J. & Legler, J. M." pageId="20" pageNumber="543" pagination="828 - 841" refId="ref15257" refString="Bull, J. J. & Legler, J. M. (1980) Karyotypes of side-necked turtles (Testudines: Pleurodira). Canadian Journal of Zoology, 58, 828 - 841. http: // dx. doi. org / 10.1139 / z 80 - 115" type="journal article" year="1980">Bull & Legler 1980</bibRefCitation>
|
||
). However, specimens with such colouration were never seen during fieldwork by
|
||
<collectorName id="EE5E535FF923FFD4FB24FA72FA9FFA5C" box="[1223,1366,1482,1507]" pageId="20" pageNumber="543">M.-O. Rödel</collectorName>
|
||
(pers. comm.). Among the
|
||
<specimenCount id="55ADFD00F923FFD4FE9DFA56FDDCF9B8" box="[382,533,1518,1543]" pageId="20" pageNumber="543" type="generic">11 specimens</specimenCount>
|
||
of the type series, there are nine terrapins with triangular pectoral scutes without midseam contact; in another specimen (
|
||
<collectionCode id="25BAAE4CF923FFD4FD89F9AAFD63F994" box="[618,682,1554,1579]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="20" pageNumber="543">MTD</collectionCode>
|
||
D 33819) the tip of the left pectoral just reaches the midseam, whereas the right pectoral does not, and in the
|
||
<typeStatus id="9C10882BF923FFD4FD7FF98FFD34F9F0" box="[668,765,1591,1615]" pageId="20" pageNumber="543" type="paratype">paratype</typeStatus>
|
||
<specimenCode id="130D9EF2F923FFD4FCE0F98FFC67F9F0" box="[771,942,1590,1615]" collectionCode="BMNH" pageId="20" pageNumber="543">BMNH 1863.3</specimenCode>
|
||
.27.1 the pectoral scutes are rectangular with wide midseam contact
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="43143689F923FFD4FF24F9C7FED1F905" blockId="20.[151,1437,582,1723]" pageId="20" pageNumber="543">
|
||
For the monogenean flatworm
|
||
<taxonomicName id="84AB4D0AF923FFD4FDC4F9C6FCE2F928" authorityName="Kulo" authorityYear="1980" box="[551,811,1662,1687]" class="Monogenea" family="Polystomatidae" genus="Polystomoides" kingdom="Animalia" order="Polystomatidea" pageId="20" pageNumber="543" phylum="Platyhelminthes" rank="species" species="nabedei">
|
||
<emphasis id="71DFEA9BF923FFD4FDC4F9C6FCE2F928" box="[551,811,1662,1687]" italics="true" pageId="20" pageNumber="543">Polystomoides nabedei</emphasis>
|
||
</taxonomicName>
|
||
possibly parasitizing
|
||
<taxonomicName id="84AB4D0AF923FFD4FBC7F9C6FAE9F928" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1060,1312,1662,1687]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="71DFEA9BF923FFD4FBC7F9C6FAE9F928" box="[1060,1312,1662,1687]" italics="true" pageId="20" pageNumber="543">Pelomedusa variabilis</emphasis>
|
||
</taxonomicName>
|
||
, see under
|
||
<taxonomicName id="84AB4D0AF923FFD4FF74F91BFED8F904" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[151,273,1698,1723]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="olivacea">
|
||
<emphasis id="71DFEA9BF923FFD4FF74F91BFED8F904" box="[151,273,1698,1723]" italics="true" pageId="20" pageNumber="543">P. olivacea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |