581 lines
103 KiB
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581 lines
103 KiB
XML
<document id="348C0AE8B5984ED92A63873AC4BC3D92" ID-CLB-Dataset="92411" ID-DOI="10.1080/00222933.2022.2049389" ID-GBIF-Dataset="4cebde67-ce4c-402d-b8d5-901a5a528fab" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1656311063267" checkinUser="plazi" docAuthor="Leever, Ellen M., Daniels, Savel R., Soma, Julia B. & Cumberlidge, Neil" docDate="2022" docId="9529DB19FFEDFFAA3950FE07FECB8F60" docLanguage="en" docName="JNATHIST.56.1-4.241-263.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2022.2049389" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Vahatra Leever & Daniels & Soma & Cumberlidge 2022, gen. nov." docType="treatment" docVersion="9" lastPageNumber="248" masterDocId="6910A361FFE9FFA23B75FFBAFFF98911" masterDocTitle="Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)" masterLastPageNumber="263" masterPageNumber="241" pageNumber="245" updateTime="1699350272557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4B8C0C4151F4469C8BDF7237D382A2E1">Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)</mods:title>
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<mods:namePart id="5F2C94710A69DFF2DBE8895768E80503">Leever, Ellen M.</mods:namePart>
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<mods:namePart id="CACDF0D803F5221D311A76C0BD555FE1">Daniels, Savel R.</mods:namePart>
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<mods:namePart id="5FF949FF51D543F3381FAE001D664F08">Soma, Julia B.</mods:namePart>
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<mods:namePart id="011474D9EE58BE8E982BF5718D64E450">Cumberlidge, Neil</mods:namePart>
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<mods:title id="9DDF421760305D38F5A0113631BB8C67">Journal of Natural History</mods:title>
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<mods:part id="113F001FC394E1AE780CE6EEC8F79ADF">
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<mods:date id="FAFC2EB7D84B9EEFD97CEB8B2C080BE9">2022</mods:date>
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<mods:number id="98F5828870C58A50EDAF7332D8680058">2022-05-24</mods:number>
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<mods:number id="2A925433EB89514557817F5DC6B061A8">56</mods:number>
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<mods:number id="16554070738959398AFC7001236BC419">1 - 4</mods:number>
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<treatment id="9529DB19FFEDFFAA3950FE07FECB8F60" ID-DOI="http://doi.org/10.5281/zenodo.6762503" ID-GBIF-Taxon="197158399" ID-Zenodo-Dep="6762503" LSID="urn:lsid:plazi:treatment:9529DB19FFEDFFAA3950FE07FECB8F60" httpUri="http://treatment.plazi.org/id/9529DB19FFEDFFAA3950FE07FECB8F60" lastPageId="8" lastPageNumber="248" pageId="4" pageNumber="245">
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<subSubSection id="559A3984FFEDFFA63950FE07FD0788C6" box="[549,766,445,471]" pageId="4" pageNumber="245" type="nomenclature">
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<paragraph id="1D3F6A0FFFEDFFA63950FE07FD0788C6" blockId="4.[297,1019,445,506]" box="[549,766,445,471]" pageId="4" pageNumber="245">
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<heading id="4677DD63FFEDFFA63950FE07FD0788C6" box="[549,766,445,471]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="245" reason="2">
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<taxonomicName id="DA80118CFFEDFFA63950FE07FD7088C6" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[549,649,445,471]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2FF4B61DFFEDFFA63950FE07FD7088C6" bold="true" box="[549,649,445,471]" italics="true" pageId="4" pageNumber="245">Vahatra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34C70B66FFEDFFA639E4FE07FD0788C6" box="[657,766,445,471]" pageId="4" pageNumber="245" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="559A3984FFEDFFA63A5CFE5AFD2A8B41" pageId="4" pageNumber="245" type="description">
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<paragraph id="1D3F6A0FFFEDFFA63A5CFE5AFC0288EB" blockId="4.[297,1019,445,506]" box="[297,1019,480,506]" pageId="4" pageNumber="245">
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(
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<figureCitation id="85BB768AFFEDFFA63A44FE5AFE6E88EB" box="[305,407,480,506]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="4" pageNumber="245">Figures 1</figureCitation>
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||
(a,c,e), 2(a,c,e), 3(a), 4(a,b,e,f), 5(a,b,c), 6;
|
||
<tableCitation id="50025FB4FFEDFFA63800FE5AFC0B88EB" box="[885,1010,480,506]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="3.[224,281,142,165]" captionStartId-1="17.[160,216,142,165]" captionStartId-2="18.[160,216,142,165]" captionStartId-3="18.[160,216,534,557]" captionTargetBox-0="[235,821,235,1031]" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="18" captionText-0="Table 1. Updated list of all known Malagasy freshwater crabs (Family Deckeniidae Ortmann, 1897, Subfamily Hydrothelphusinae Bott, 1955) from 1853 to July 2021. The" captionText-1="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." captionText-2="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." captionText-3="Table 4. Gonopod 1 (G1) measurements (taken on the ventral side) showing lengths of the G1 terminal article (TA) and the G1 subterminal article (SA) in mm for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The relative length of the G1 TA and the G1 SA is also shown, along with the size classes for the G1 TA." httpUri-0="http://table.plazi.org/id/49FF3A87FFEAFFA13B95FF34FC1A89F1" httpUri-1="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" httpUri-2="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="4" pageNumber="245" tableUuid-0="49FF3A87FFEAFFA13B95FF34FC1A89F1" tableUuid-1="49FF3A87FFF8FFB33BD5FF34FC3A89EC" tableUuid-2="49FF3A87FFFBFFB03BD5FF34FE20880B">Tables 1–4</tableCitation>
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||
)
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||
</paragraph>
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||
<paragraph id="1D3F6A0FFFEDFFA63BB6FDAFFD2A8B41" blockId="4.[195,1152,533,592]" pageId="4" pageNumber="245">
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<emphasis id="2FF4B61DFFEDFFA63BB6FDAFFE208B3F" box="[195,473,533,558]" italics="true" pageId="4" pageNumber="245">Nomenclatural statement</emphasis>
|
||
: A life science identifier (LSID) number was obtained for the new genus:
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="559A3984FFEDFFA63BD5FD3FFCA58B8F" box="[160,860,645,670]" pageId="4" pageNumber="245" type="reference_group">
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<paragraph id="1D3F6A0FFFEDFFA63BD5FD3FFCA58B8F" blockId="4.[160,860,645,670]" box="[160,860,645,670]" pageId="4" pageNumber="245">
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<treatmentCitationGroup id="3D904D21FFEDFFA63BD5FD3FFCA58B8F" box="[160,860,645,670]" pageId="4" pageNumber="245">
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<emphasis id="2FF4B61DFFEDFFA63BD5FD3FFDEF8B8F" bold="true" box="[160,534,645,670]" pageId="4" pageNumber="245">
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||
<emphasis id="2FF4B61DFFEDFFA63BD5FD3FFF158B8F" bold="true" box="[160,236,645,670]" italics="true" pageId="4" pageNumber="245">
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||
<taxonomicName id="DA80118CFFEDFFA63BD5FD3FFF2E8B8F" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[160,215,645,670]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="genus">Foza</taxonomicName>
|
||
–
|
||
</emphasis>
|
||
<treatmentCitation id="9C214C1EFFEDFFA63B81FD3FFDEF8B8F" ID-DOI="http://doi.org/10.5281/zenodo.3791535" ID-Zenodo-Dep="3791535" author="Cumberlidge N & Meyer KS" box="[244,534,645,670]" httpUri="http://treatment.plazi.org/id/03EF536EFFCE4662FF173B18FBFFFEF1" page="79" pageId="4" pageNumber="245" year="2009">
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||
<bibRefCitation id="791117FEFFEDFFA63B81FD3FFDEF8B8F" author="Cumberlidge N & Meyer KS" box="[244,534,645,670]" pageId="4" pageNumber="245" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer</bibRefCitation>
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||
</treatmentCitation>
|
||
</emphasis>
|
||
2009: 79–84, figs 1–3, tab. 1.
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="559A3984FFEDFFA63BD5FD77FC398A18" pageId="4" pageNumber="245" type="type_taxon">
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||
<paragraph id="1D3F6A0FFFEDFFA63BD5FD77FECA8BF6" blockId="4.[160,960,717,777]" box="[160,307,717,743]" pageId="4" pageNumber="245">
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||
<emphasis id="2FF4B61DFFEDFFA63BD5FD77FECA8BF6" bold="true" box="[160,307,717,743]" italics="true" pageId="4" pageNumber="245">
|
||
<typeStatus id="C23BD4ADFFEDFFA63BD5FD77FF218BF6" box="[160,216,717,743]" pageId="4" pageNumber="245">Type</typeStatus>
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||
species
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FD4AFC398A18" blockId="4.[160,960,717,777]" box="[160,960,752,777]" pageId="4" pageNumber="245">
|
||
<taxonomicName id="DA80118CFFEDFFA63BD5FD4AFD508A18" authority="Cumberlidge and Meyer, 2009" authorityName="Cumberlidge and Meyer" authorityYear="2009" box="[160,681,752,777]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5FD4AFEB48A18" box="[160,333,752,777]" italics="true" pageId="4" pageNumber="245">Foza ambohitra</emphasis>
|
||
<bibRefCitation id="791117FEFFEDFFA63A20FD4AFD508A18" author="Cumberlidge N & Meyer KS" box="[341,681,752,777]" pageId="4" pageNumber="245" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="559A3984FFEDFFA63BD5FC83FED48DE3" pageId="4" pageNumber="245" type="diagnosis">
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FC83FEED8A42" blockId="4.[160,1156,825,1266]" box="[160,276,825,851]" pageId="4" pageNumber="245">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5FC83FEED8A42" bold="true" box="[160,276,825,851]" italics="true" pageId="4" pageNumber="245">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FCE6FED48DE3" blockId="4.[160,1156,825,1266]" pageId="4" pageNumber="245">
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||
Entire carapace surface smooth; anterolateral margin smooth; epibranchial tooth small, acute, close to exorbital tooth, positioned in line with postorbital margin (
|
||
<figureCitation id="85BB768AFFEDFFA63889FCC5FBAC8A89" box="[1020,1109,895,920]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="4" pageNumber="245">Figure 1</figureCitation>
|
||
(a)); shallow notch between exorbital, epibranchial teeth; lateral carapace margin smooth; suborbital, subhepatic regions of branchiostegite smooth, pterygostomial region smooth except for small field of granules at junction of longitudinal, vertical sutures; large anterior lobe on terminal article of mandibular palp; ambulatory legs (P2–5) stout, not elongated; cheliped ischium margin smooth, rounded; S4/5 thoracic sternal sulcus meeting pleon at PL5/6 suture; PL6 relatively narrow with convex lateral margins; telson lateral margins gently sinuous, apex rounded; third maxilliped exopod with short flagellum (~ 0.5× merus length). G1 TA dorsal side glabrous, broadest proximally, distal half curving upward, tip broad (
|
||
<figureCitation id="85BB768AFFEDFFA63B85FB0DFEB58DC1" box="[240,332,1207,1232]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="4" pageNumber="245">Figure 5</figureCitation>
|
||
(a,b)); G2 TA long (G2 TA/SA = 0.6), flagellum-like, with distinct distal curve (
|
||
<figureCitation id="85BB768AFFEDFFA63BDDFB63FEFD8DE3" box="[168,260,1241,1266]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="4" pageNumber="245">Figure 5</figureCitation>
|
||
(c)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFEDFFA63BD5FA9BFCB08CB2" pageId="4" pageNumber="245" type="distribution">
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FA9BFED58C2A" blockId="4.[160,1156,1313,1443]" box="[160,300,1313,1339]" pageId="4" pageNumber="245">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5FA9BFED58C2A" bold="true" box="[160,300,1313,1339]" italics="true" pageId="4" pageNumber="245">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FAFFFCB08CB2" blockId="4.[160,1156,1313,1443]" pageId="4" pageNumber="245">
|
||
Endemic to
|
||
<collectingCountry id="65972A9FFFEDFFA63A5EFAFFFE4D8C4F" box="[299,436,1349,1374]" name="Madagascar" pageId="4" pageNumber="245">Madagascar</collectingCountry>
|
||
.
|
||
<taxonomicName id="DA80118CFFEDFFA63AB5FAFFFDEE8C4F" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[448,535,1349,1374]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEDFFA63AB5FAFFFDEE8C4F" box="[448,535,1349,1374]" italics="true" pageId="4" pageNumber="245">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEDFFA63954FAFFFD728C4F" box="[545,651,1349,1374]" pageId="4" pageNumber="245" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is known from three locations in the Diana Region of
|
||
<collectingRegion id="DF44A4EDFFEDFFA63A6FFAD2FDF28C90" box="[282,523,1384,1409]" country="Madagascar" name="Antsiranana" pageId="4" pageNumber="245">Antsiranana Province</collectingRegion>
|
||
in northern
|
||
<collectingCountry id="65972A9FFFEDFFA639D7FAD2FCD28C90" box="[674,811,1384,1409]" name="Madagascar" pageId="4" pageNumber="245">Madagascar</collectingCountry>
|
||
: Ambohitra, the Analamerana Special Reserve and the Ankarana Special Reserve (
|
||
<figureCitation id="85BB768AFFEDFFA63995FA30FCC48CB2" box="[736,829,1418,1443]" captionStart="Figure 6" captionStartId="16.[160,225,1033,1056]" captionTargetBox="[165,1150,144,998]" captionTargetId="figure-69@16.[163,1153,142,1001]" captionTargetPageId="16" captionText="Figure 6. Map of Madagascar showing the distributional range of Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov. (black circles, not shown to scale), Toamasina clarki sp. nov. (white triangle in black square, not shown to scale), Foza raimundi Reed and Cumberlidge, 2006 (white asterisk with black margin), and Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015 (white square with black margin). Scale bar = 184 km." figureDoi="http://doi.org/10.5281/zenodo.6758305" httpUri="https://zenodo.org/record/6758305/files/figure.png" pageId="4" pageNumber="245">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFEDFFA63BD5FA69FC398F1E" pageId="4" pageNumber="245" type="type_taxon">
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FA69FECA8CFC" blockId="4.[160,960,1491,1551]" box="[160,307,1491,1517]" pageId="4" pageNumber="245">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5FA69FECA8CFC" bold="true" box="[160,307,1491,1517]" italics="true" pageId="4" pageNumber="245">
|
||
<typeStatus id="C23BD4ADFFEDFFA63BD5FA69FF218CFC" box="[160,216,1491,1517]" pageId="4" pageNumber="245">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5FA4CFC398F1E" blockId="4.[160,960,1491,1551]" box="[160,960,1526,1551]" pageId="4" pageNumber="245">
|
||
<taxonomicName id="DA80118CFFEDFFA63BD5FA4CFD508F1E" authority="Cumberlidge and Meyer, 2009" authorityName="Cumberlidge and Meyer" authorityYear="2009" box="[160,681,1526,1551]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5FA4CFEB48F1E" box="[160,333,1526,1551]" italics="true" pageId="4" pageNumber="245">Foza ambohitra</emphasis>
|
||
<bibRefCitation id="791117FEFFEDFFA63A20FA4CFD508F1E" author="Cumberlidge N & Meyer KS" box="[341,681,1526,1551]" pageId="4" pageNumber="245" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFEDFFA73BD5F984FC988844" lastPageId="5" lastPageNumber="246" pageId="4" pageNumber="245" type="etymology">
|
||
<paragraph id="1D3F6A0FFFEDFFA63BD5F984FEE48F49" blockId="4.[160,1156,1598,1728]" box="[160,285,1598,1624]" pageId="4" pageNumber="245">
|
||
<emphasis id="2FF4B61DFFEDFFA63BD5F984FEE48F49" bold="true" box="[160,285,1598,1624]" italics="true" pageId="4" pageNumber="245">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFEDFFA73BD5F9D8FB8989FC" blockId="4.[160,1156,1598,1728]" lastBlockId="5.[160,1155,142,237]" lastPageId="5" lastPageNumber="246" pageId="4" pageNumber="245">
|
||
The genus name recognises the contributions of the
|
||
<taxonomicName id="DA80118CFFEDFFA63983F9D8FC2A8F6A" authority="Association" authorityName="Association" box="[758,979,1634,1659]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="245" phylum="Arthropoda" rank="genus">Vahatra Association</taxonomicName>
|
||
of
|
||
<collectingCountry id="65972A9FFFEDFFA6388DF9D8FB788F6A" box="[1016,1153,1634,1659]" name="Madagascar" pageId="4" pageNumber="245">Madagascar</collectingCountry>
|
||
, whose objective is to train Malagasy field scientists in conservation biology in order to protect the unique biotic resources of that island nation. Most of the specimens of
|
||
<taxonomicName id="DA80118CFFEDFFA73F59F91DFD3C89B6" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="246" order="Decapoda" pageId="4" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFEDFFA73F59F91DFEEB89B6" italics="true" lastPageId="5" lastPageNumber="246" pageId="4" pageNumber="245">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFECFFA73A6AFF34FD8889B6" author="Cumberlidge N & Meyer KS" box="[287,625,142,167]" pageId="5" pageNumber="246" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFECFFA7390AFF34FD3C89B6" box="[639,709,142,167]" pageId="5" pageNumber="246" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov. in the present work were collected by Dr Steve M. Goodman, a founding member of the
|
||
<taxonomicName id="DA80118CFFECFFA7387AFF0BFC0C89DB" authority="Association." authorityName="Association." box="[783,1013,177,202]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">Vahatra Association.</taxonomicName>
|
||
The generic name,
|
||
<taxonomicName id="DA80118CFFECFFA73B99FF6EFEBA89FC" box="[236,323,212,237]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73B99FF6EFEBA89FC" box="[236,323,212,237]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
, is used as a Latin noun in nominative singular and treated as feminine.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA73BD5FEA2FE9A8823" blockId="5.[160,865,280,341]" box="[160,355,280,306]" pageId="5" pageNumber="246">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FEA2FE9A8823" bold="true" box="[160,355,280,306]" italics="true" pageId="5" pageNumber="246">Species included</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA73BD5FE86FC988844" blockId="5.[160,865,280,341]" box="[160,865,316,341]" pageId="5" pageNumber="246">
|
||
<taxonomicName id="DA80118CFFECFFA73BD5FE86FCD38844" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[160,810,316,341]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FE86FE888844" box="[160,369,316,341]" italics="true" pageId="5" pageNumber="246">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFECFFA73A0BFE86FD2C8844" author="Cumberlidge N & Meyer KS" box="[382,725,316,341]" pageId="5" pageNumber="246" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73996FE86FCD38844" box="[739,810,316,341]" pageId="5" pageNumber="246" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFECFFAA3BD5FE3BFECB8F60" lastPageId="8" lastPageNumber="249" pageId="5" pageNumber="246" type="discussion">
|
||
<paragraph id="1D3F6A0FFFECFFA73BD5FE3BFEFF888A" blockId="5.[160,1156,385,1728]" box="[160,262,385,411]" pageId="5" pageNumber="246">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FE3BFEFF888A" bold="true" box="[160,262,385,411]" italics="true" pageId="5" pageNumber="246">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA73BD5FE1FFD278B7A" blockId="5.[160,1156,385,1728]" pageId="5" pageNumber="246">
|
||
The classification used here for the Afrotropical freshwater crab fauna follows that proposed recently by
|
||
<bibRefCitation id="791117FEFFECFFA73AE8FE7DFCF788F1" author="Cumberlidge N & Daniels SR" box="[413,782,455,480]" pageId="5" pageNumber="246" pagination="1268 - 1311" refId="ref11238" refString="Cumberlidge N, Daniels SR. 2022. A new multilocus phylogeny reveals overlooked diversity in African freshwater crabs (Potamonautidae Bott, 1970): a major revision with new higher taxa and genera. Zool J Linn Soc. 194: 1268 - 1311. https: // zlab 082, % 26 # x 00 A 0; https: // doi. org / 10.1093 / zoolinnean / zlab 082." type="journal article" year="2022">Cumberlidge and Daniels (2022)</bibRefCitation>
|
||
, who recognised two families in the region: the
|
||
<taxonomicName id="DA80118CFFECFFA73A2BFE50FD668B12" authority="Bott, 1970" authorityName="Bott" authorityYear="1970" box="[350,671,490,515]" class="Malacostraca" family="Potamonautidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="family">Potamonautidae Bott, 1970</taxonomicName>
|
||
(endemic to continental Africa) and the
|
||
<taxonomicName id="DA80118CFFECFFA73BD5FDB7FE088B37" authority="Ortmann, 1897" authorityName="Ortmann" authorityYear="1897" box="[160,497,525,550]" class="Malacostraca" family="Deckeniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="family">Deckeniidae Ortmann, 1897</taxonomicName>
|
||
(found in West and East Africa, The
|
||
<collectingCountry id="65972A9FFFECFFA738A4FDB7FBB18B37" box="[977,1096,525,550]" name="Seychelles" pageId="5" pageNumber="246">Seychelles</collectingCountry>
|
||
and
|
||
<collectingCountry id="65972A9FFFECFFA73BD5FD95FED28B59" box="[160,299,559,584]" name="Madagascar" pageId="5" pageNumber="246">Madagascar</collectingCountry>
|
||
). Those authors included the entire Malagasy freshwater crab fauna in the deckeniid subfamily
|
||
<taxonomicName id="DA80118CFFECFFA73AFDFDE8FD218B7A" authority="Bott, 1955" authorityName="Bott" authorityYear="1955" box="[392,728,594,619]" class="Malacostraca" family="Potamonautidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="subFamily" subFamily="Hydrothelphusinae">
|
||
Hydrothelphusinae
|
||
<bibRefCitation id="791117FEFFECFFA73910FDE8FD218B7A" author="Bott R" box="[613,728,594,619]" pageId="5" pageNumber="246" pagination="209 - 352" refId="ref11054" refString="Bott R. 1955. Die Susswasserkrabben von Afrika und ihre Stammesgeschichte. Ann Mus Congo Belge. 1 (3): 209 - 352. Taf. 1 - 30, Abb. 103." type="journal article" year="1955">Bott, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA73BB5FDCFFB7D8BE4" blockId="5.[160,1156,385,1728]" pageId="5" pageNumber="246">
|
||
The new genus is established based on morphological and phylogenetic evidence. The molecular phylogeny in
|
||
<bibRefCitation id="791117FEFFECFFA73AB4FD2DFD1C8BA1" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[449,741,663,688]" pageId="5" pageNumber="246" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
|
||
, fig. 1) indicates that
|
||
<taxonomicName id="DA80118CFFECFFA73885FD2DFDF18BC2" authority="(Cumberlidge and Meyer, 2009)" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="ambohitra" status="comb. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73885FD2DFB7D8BA1" box="[1008,1156,663,688]" italics="true" pageId="5" pageNumber="246">V. ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFECFFA73BD3FD00FDF88BC2" author="Cumberlidge N & Meyer KS" box="[166,513,698,723]" pageId="5" pageNumber="246" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73964FD00FD698BC2" box="[529,656,698,723]" pageId="5" pageNumber="246" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is part of a unique clade that is genetically separated from
|
||
<taxonomicName id="DA80118CFFECFFA73A24FD66FE328BE4" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[337,459,732,757]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFECFFA73A24FD66FE328BE4" box="[337,459,732,757]" italics="true" pageId="5" pageNumber="246">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="C23BD4ADFFECFFA73970FD66FDCE8BE4" box="[517,567,732,757]" pageId="5" pageNumber="246">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="DA80118CFFECFFA739C4FD66FD1D8BE4" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[689,740,732,757]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA739C4FD66FD1D8BE4" box="[689,740,732,757]" italics="true" pageId="5" pageNumber="246">Foza</emphasis>
|
||
</taxonomicName>
|
||
) and from all other Malagasy genera.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA73BB5FD45FC5B8D87" blockId="5.[160,1156,385,1728]" pageId="5" pageNumber="246">
|
||
<taxonomicName id="DA80118CFFECFFA73BB5FD45FEEE8A09" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[192,279,767,792]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73BB5FD45FEEE8A09" box="[192,279,767,792]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73A6BFD45FE7E8A09" box="[286,391,767,792]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is part of a morphological group with seven other Malagasy genera (
|
||
<emphasis id="2FF4B61DFFECFFA73BDDFC98FF088A2A" box="[168,241,802,827]" italics="true" pageId="5" pageNumber="246">
|
||
<taxonomicName id="DA80118CFFECFFA73BDDFC98FF148A2A" authorityName="Cumberlidge, Soma, Leever and Daniels" authorityYear="2020" box="[168,237,802,827]" class="Malacostraca" family="Potamonautidae" genus="Agora" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">Agora</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="DA80118CFFECFFA73A70FC98FBC98A2A" authority="Cumberlidge, Soma, Leever, Clark and Daniels, 2021" authorityName="Cumberlidge, Soma, Leever, Clark and Daniels" authorityYear="2021" box="[261,1072,802,827]" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73A70FC98FE798A2A" box="[261,384,802,827]" italics="true" pageId="5" pageNumber="246">Crosnautes</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73AE6FC98FBC98A2A" author="Cumberlidge N & Soma J & Leever EM & Clark PF & Daniels SR" box="[403,1072,802,827]" pageId="5" pageNumber="246" pagination="1 - 10" refId="ref11550" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: brachyura: potamonautidae) from southeastern Madagascar. Journal of Crustacean Biology. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge, Soma, Leever, Clark and Daniels, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="2FF4B61DFFECFFA73F3CFC98FB7A8A2A" box="[1097,1155,802,827]" italics="true" pageId="5" pageNumber="246">
|
||
<taxonomicName id="DA80118CFFECFFA73F3CFC98FB868A2A" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[1097,1151,802,827]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">Foza</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="DA80118CFFECFFA73BD5FCFEFCC68A4C" authority="Meyer, Cumberlidge and Koppin, 2014" authorityName="Meyer, Cumberlidge, and Koppin" authorityYear="2014" box="[160,831,836,861]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FCFEFEAB8A4C" box="[160,338,836,861]" italics="true" pageId="5" pageNumber="246">Glabrithelphusa</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73A17FCFEFCC68A4C" author="Meyer KS & Cumberlidge N & Koppin JC" box="[354,831,836,861]" pageId="5" pageNumber="246" pagination="65 - 72" refId="ref11796" refString="Meyer KS, Cumberlidge N, Koppin JC. 2014. A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae). Potamoidea, Potamonautidae). Zootaxa. 3884 (1): 65 - 72. doi: 10.11646 / zootaxa. 3884.1.5." type="journal article" year="2014">Meyer, Cumberlidge and Koppin, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DA80118CFFECFFA73820FCFEFEAE8A91" authority="A. MilneEdwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73820FCFEFBFF8A4C" box="[853,1030,836,861]" italics="true" pageId="5" pageNumber="246">Hydrothelphusa</emphasis>
|
||
A. MilneEdwards, 1872
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DA80118CFFECFFA73A00FCDDFC158A91" authority="Cumberlidge, Boyko, and Harvey, 2002" authorityName="Cumberlidge, Boyko, and Harvey" authorityYear="2002" box="[373,1004,871,896]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73A00FCDDFE2C8A91" box="[373,469,871,896]" italics="true" pageId="5" pageNumber="246">Marojejy</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73A98FCDDFC158A91" author="Cumberlidge N & Boyko CB & Harvey AW" box="[493,1004,871,896]" pageId="5" pageNumber="246" pagination="65 - 77" refId="ref11178" refString="Cumberlidge N, Boyko CB, Harvey AW. 2002. A new genus and species of freshwater crab (Crustacea, Decapoda, Potamoidea) from northern Madagascar, and a second new species associated with Pandanus leaf axils. J Nat Hist. 36 (1): 65 - 77. doi: 10.1080 / 00222930010003800." type="journal article" year="2002">Cumberlidge, Boyko, and Harvey, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DA80118CFFECFFA73F7FFCDDFDD98AB2" authority="Cumberlidge and Sternberg, 2002" authorityName="Cumberlidge and von Sternberg" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73F7FFCDDFB7A8A91" box="[1034,1155,871,896]" italics="true" pageId="5" pageNumber="246">Malagasya</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73BD5FC30FDD98AB2" author="Cumberlidge N & Sternberg R" box="[160,544,906,931]" pageId="5" pageNumber="246" pagination="41 - 79" refId="ref11694" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DA80118CFFECFFA73915FC30FD218AB2" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[608,728,906,931]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73915FC30FD218AB2" box="[608,728,906,931]" italics="true" pageId="5" pageNumber="246">Toamasina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73995FC30FCB38AB2" box="[736,842,906,931]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
) that all have a mandibular palp with a clearly bilobed terminal article, whose anterior lobe is conspicuous and medium-sized (MPAL/MPTA = 0.4–0.6) (see
|
||
<bibRefCitation id="791117FEFFECFFA739F8FC75FBFE8AF9" author="Cumberlidge N & Sternberg R" box="[653,1031,975,1000]" pageId="5" pageNumber="246" pagination="41 - 79" refId="ref11694" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 4a–g). On the other hand, the size of the anterior lobe on the terminal article of the mandibular palp of
|
||
<taxonomicName id="DA80118CFFECFFA73B83FBAFFEB48D3F" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[246,333,1045,1070]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73B83FBAFFEB48D3F" box="[246,333,1045,1070]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73A21FBAFFE458D3F" box="[340,444,1045,1070]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(MPAL/MPTA = 0.4) (
|
||
<figureCitation id="85BB768AFFECFFA739DCFBAFFCFC8D3F" box="[681,773,1045,1070]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="5" pageNumber="246">Figure 2</figureCitation>
|
||
(a,c);
|
||
<tableCitation id="50025FB4FFECFFA7384BFBAFFC698D3F" box="[830,912,1045,1070]" captionStart="Table 2" captionStartId="17.[160,216,142,165]" captionTargetPageId="17" captionText="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." httpUri="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" pageId="5" pageNumber="246" tableUuid="49FF3A87FFF8FFB33BD5FF34FC3A89EC">Table 2</tableCitation>
|
||
) distinguishes it from
|
||
<taxonomicName id="DA80118CFFECFFA73BD5FB8DFDC48D41" authority="(Cumberlidge, 2010)" baseAuthorityName="Cumberlidge" baseAuthorityYear="2010" box="[160,573,1079,1104]" class="Malacostraca" family="Potamonautidae" genus="Boreathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FB8DFEB08D41" box="[160,329,1079,1104]" italics="true" pageId="5" pageNumber="246">Boreathelphusa</emphasis>
|
||
(Cumberlidge, 2010)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DA80118CFFECFFA7393FFB8DFC7E8D41" authority="Bott, 1955" authorityName="Bott" authorityYear="1955" box="[586,903,1079,1104]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA7393FFB8DFCF18D41" box="[586,776,1079,1104]" italics="true" pageId="5" pageNumber="246">Madagapotamon</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73867FB8DFC7E8D41" author="Bott R" box="[786,903,1079,1104]" pageId="5" pageNumber="246" pagination="209 - 352" refId="ref11054" refString="Bott R. 1955. Die Susswasserkrabben von Afrika und ihre Stammesgeschichte. Ann Mus Congo Belge. 1 (3): 209 - 352. Taf. 1 - 30, Abb. 103." type="journal article" year="1955">Bott, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA80118CFFECFFA738B3FB8DFE9A8D62" authority="Ng and Takeda, 1994" authorityName="Ng and Takeda" authorityYear="1994" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA738B3FB8DFBA08D41" box="[966,1113,1079,1104]" italics="true" pageId="5" pageNumber="246">Skelosophusa</emphasis>
|
||
<bibRefCitation id="791117FEFFECFFA73F16FB8DFE9A8D62" author="Ng PKL & Takeda M" pageId="5" pageNumber="246" pagination="161 - 172" refId="ref11918" refString="Ng PKL, Takeda M. 1994. Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bull Nat Sci Mus Ser A (Zool). 20: 161 - 172." type="journal article" year="1994">Ng and Takeda, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
, whose mandibular palp has a small but distinct ledge-like anterior lobe (MPAL/MPTA = 0.2) (see
|
||
<bibRefCitation id="791117FEFFECFFA73AC3FBC7FCD78D87" author="Cumberlidge N & Sternberg R" box="[438,814,1149,1174]" pageId="5" pageNumber="246" pagination="41 - 79" refId="ref11694" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 4h–l).
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFECFFA53BB5FB25FCFE8D73" blockId="5.[160,1156,385,1728]" lastBlockId="7.[160,1157,993,1712]" lastPageId="7" lastPageNumber="248" pageId="5" pageNumber="246">
|
||
<taxonomicName id="DA80118CFFECFFA73BB5FB25FEE08DA9" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[192,281,1183,1208]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73BB5FB25FEE08DA9" box="[192,281,1183,1208]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73A50FB25FE6D8DA9" box="[293,404,1183,1208]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFECFFA73991FB25FCDF8DA9" authorityName="Cumberlidge, Soma, Leever and Daniels" authorityYear="2020" box="[740,806,1183,1208]" class="Malacostraca" family="Potamonautidae" genus="Agora" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73991FB25FCDF8DA9" box="[740,806,1183,1208]" italics="true" pageId="5" pageNumber="246">Agora</emphasis>
|
||
</taxonomicName>
|
||
by the incomplete and faint postfrontal crest that does not traverse the entire carapace (
|
||
<figureCitation id="85BB768AFFECFFA738F1FB78FC0C8DCA" box="[900,1013,1218,1243]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="5" pageNumber="246">Figure 1a</figureCitation>
|
||
), vs a welldefined postfrontal crest that completely traverses the carapace (see
|
||
<bibRefCitation id="791117FEFFECFFA738C6FB5FFF2E8C31" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" pageId="5" pageNumber="246" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
|
||
).
|
||
<taxonomicName id="DA80118CFFECFFA73B99FABDFEBA8C31" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[236,323,1287,1312]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73B99FABDFEBA8C31" box="[236,323,1287,1312]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73A3EFABDFE4C8C31" box="[331,437,1287,1312]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFECFFA7399BFABDFC9F8C31" authorityName="Cumberlidge, Soma, Leever, Clark and Daniels" authorityYear="2021" box="[750,870,1287,1312]" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA7399BFABDFC9F8C31" box="[750,870,1287,1312]" italics="true" pageId="5" pageNumber="246">Crosnautes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA80118CFFECFFA738D4FABDFC2D8C31" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[929,980,1287,1312]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA738D4FABDFC2D8C31" box="[929,980,1287,1312]" italics="true" pageId="5" pageNumber="246">Foza</emphasis>
|
||
</taxonomicName>
|
||
by the smooth anterolateral corners of the carapace surface (
|
||
<figureCitation id="85BB768AFFECFFA739D0FA90FD058C52" box="[677,764,1322,1347]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="5" pageNumber="246">Figure 1</figureCitation>
|
||
(a)), vs anterolateral corners of the carapace with fields of well-defined carinae in
|
||
<taxonomicName id="DA80118CFFECFFA739EBFAF7FCEF8C77" authorityName="Cumberlidge, Soma, Leever, Clark and Daniels" authorityYear="2021" box="[670,790,1357,1382]" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA739EBFAF7FCEF8C77" box="[670,790,1357,1382]" italics="true" pageId="5" pageNumber="246">Crosnautes</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="791117FEFFECFFA7383BFAF7FBAF8C77" author="Cumberlidge N & Soma J & Leever EM & Clark PF & Daniels SR" box="[846,1110,1357,1382]" pageId="5" pageNumber="246" pagination="1 - 10" refId="ref11550" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: brachyura: potamonautidae) from southeastern Madagascar. Journal of Crustacean Biology. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. 2021</bibRefCitation>
|
||
, fig. 1a,b) and
|
||
<taxonomicName id="DA80118CFFECFFA73A78FAD5FEB98C99" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[269,320,1391,1416]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73A78FAD5FEB98C99" box="[269,320,1391,1416]" italics="true" pageId="5" pageNumber="246">Foza</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="85BB768AFFECFFA73A38FAD5FE518C99" box="[333,424,1391,1416]" captionStart="Figure 7" captionStartId="19.[160,225,1121,1144]" captionTargetBox="[163,1153,142,1089]" captionTargetId="figure-84@19.[163,1153,142,1090]" captionTargetPageId="19" captionText="Figure 7. Foza raimundi Reed and Cumberlidge, 2006, adult ♂, CW 18.7 mm, FMNH 15336, from Marojejy National Park, Madagascar: a, dorsal view of cephalothorax; b, frontal view of cephalothorax; c, ventral view of cephalothorax; g, ventral view of left G1; h, dorsal view of left G1; i, ventral view of left G2. Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015, holotype, adult ♂, CW 43.07 mm, ZSM A20145003, from Ankarana Special Reserve, Madagascar: d, dorsal view of cephalothorax; e, frontal view of cephalothorax; f, ventral view of cephalothorax; j, ventral view of left G1; k, dorsal view of left G1; l, ventral view of left G2. Scale bar:a–c = 6.6 mm; d–f = 10.0 mm; g, h = 2.0 mm; i = 2.5 mm; j, k = 3.3 mm; l = 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758309" httpUri="https://zenodo.org/record/6758309/files/figure.png" pageId="5" pageNumber="246">Figure 7</figureCitation>
|
||
(a,c)).
|
||
<taxonomicName id="DA80118CFFECFFA73A92FAD5FDC78C99" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[487,574,1391,1416]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73A92FAD5FDC78C99" box="[487,574,1391,1416]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73931FAD5FD528C99" box="[580,683,1391,1416]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFECFFA738A2FAD5FB7A8C99" authorityName="Meyer, Cumberlidge, and Koppin" authorityYear="2014" box="[983,1155,1391,1416]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA738A2FAD5FB7A8C99" box="[983,1155,1391,1416]" italics="true" pageId="5" pageNumber="246">Glabrithelphusa</emphasis>
|
||
</taxonomicName>
|
||
by the G1 TA which is broad basally, tapers evenly and curves upward distally in
|
||
<taxonomicName id="DA80118CFFECFFA73F59FA28FB7A8CBA" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[1068,1155,1426,1451]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73F59FA28FB7A8CBA" box="[1068,1155,1426,1451]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73BD5FA0FFEF18CDF" box="[160,264,1461,1486]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFECFFA73A60FA0FFE888CDF" box="[277,369,1461,1486]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="5" pageNumber="246">Figure 5</figureCitation>
|
||
(a,b)), vs a G1 TA that is broadest at the mid-point and is straight along its length (
|
||
<bibRefCitation id="791117FEFFECFFA73A6AFA6DFE178CE1" author="Meyer KS & Cumberlidge N & Koppin JC" box="[287,494,1495,1520]" pageId="5" pageNumber="246" pagination="65 - 72" refId="ref11796" refString="Meyer KS, Cumberlidge N, Koppin JC. 2014. A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae). Potamoidea, Potamonautidae). Zootaxa. 3884 (1): 65 - 72. doi: 10.11646 / zootaxa. 3884.1.5." type="journal article" year="2014">Meyer et al. 2014</bibRefCitation>
|
||
, figs 1a, 3a). The new genus can be distinguished from
|
||
<taxonomicName id="DA80118CFFECFFA73BD5FA40FEB58F02" authorityName="A. Milne-Edwards" authorityYear="1872" box="[160,332,1530,1555]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5FA40FEB58F02" box="[160,332,1530,1555]" italics="true" pageId="5" pageNumber="246">Hydrothelphusa</emphasis>
|
||
</taxonomicName>
|
||
by the shallow notch between the exorbital and epibranchial teeth in
|
||
<taxonomicName id="DA80118CFFECFFA73BD5F9A7FF0E8F27" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,247,1565,1590]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73BD5F9A7FF0E8F27" box="[160,247,1565,1590]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA73B88F9A7FE9C8F27" box="[253,357,1565,1590]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFECFFA73A04F9A7FE3E8F27" box="[369,455,1565,1590]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="5" pageNumber="246">Figure 1</figureCitation>
|
||
(a)), vs a deep cleft between these teeth (see
|
||
<bibRefCitation id="791117FEFFECFFA738B4F9A7FEB68F49" author="Cumberlidge N & Sternberg R" pageId="5" pageNumber="246" pagination="41 - 79" refId="ref11694" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 1a–c;
|
||
<bibRefCitation id="791117FEFFECFFA73ABCF985FD238F49" author="Cumberlidge N & Marijnissen SAE & Thompson J" box="[457,730,1599,1624]" pageId="5" pageNumber="246" pagination="61 - 68" refId="ref11366" refString="Cumberlidge N, Marijnissen SAE, Thompson J. 2007. Hydrothelphusa vencesi, a new species of freshwater crab (Brachyura: potamoidea: potamonautidae) from southeastern Madagascar. Zootaxa. 1524 (1): 61 - 68. doi: 10.11646 / zootaxa. 1524.1.6." type="journal article" year="2007">Cumberlidge et al. 2007</bibRefCitation>
|
||
, figs 1, 16).
|
||
<taxonomicName id="DA80118CFFECFFA73810F985FC458F49" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[869,956,1599,1624]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73810F985FC458F49" box="[869,956,1599,1624]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA738B0F985FBD68F49" box="[965,1071,1599,1624]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFECFFA73AFFF9D8FDFA8F6A" authorityName="Cumberlidge and von Sternberg" authorityYear="2002" box="[394,515,1634,1659]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFECFFA73AFFF9D8FDFA8F6A" box="[394,515,1634,1659]" italics="true" pageId="5" pageNumber="246">Malagasya</emphasis>
|
||
</taxonomicName>
|
||
by the smooth anterolateral margin of the carapace (
|
||
<figureCitation id="85BB768AFFECFFA73BDDF93FFEEE8F8F" box="[168,279,1669,1694]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="5" pageNumber="246">Figure 1a</figureCitation>
|
||
), vs small or conspicuous teeth lining this margin (see
|
||
<bibRefCitation id="791117FEFFECFFA738CDF93FFEB78FD1" author="Cumberlidge N & Sternberg R" pageId="5" pageNumber="246" pagination="41 - 79" refId="ref11694" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 1e,f;
|
||
<bibRefCitation id="791117FEFFECFFA73AC9F91DFD348FD1" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[444,717,1703,1728]" pageId="5" pageNumber="246" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
|
||
, fig. 10a,c,e).
|
||
<taxonomicName id="DA80118CFFECFFA73813F91DFC448FD1" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[870,957,1703,1728]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFECFFA73813F91DFC448FD1" box="[870,957,1703,1728]" italics="true" pageId="5" pageNumber="246">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFECFFA738B3F91DFBD68FD1" box="[966,1071,1703,1728]" pageId="5" pageNumber="246" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFEEFFA53A0BFC5BFE228AEB" authorityName="Cumberlidge, Boyko, and Harvey" authorityYear="2002" box="[382,475,993,1018]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFEEFFA53A0BFC5BFE228AEB" box="[382,475,993,1018]" italics="true" pageId="7" pageNumber="248">Marojejy</emphasis>
|
||
</taxonomicName>
|
||
by its short ambulatory legs (ΣP2–5/CW = 5.7) (
|
||
<figureCitation id="85BB768AFFEEFFA53F71FC5BFB9B8AEB" box="[1028,1122,993,1018]" captionStart="Figure 3" captionStartId="7.[160,225,797,820]" captionTargetBox="[163,1153,142,765]" captionTargetId="figure-326@7.[163,1153,142,765]" captionTargetPageId="7" captionText="Figure 3. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., paratype, adult ♀, CW 35.0 mm, FMNH 11046, from Analamerana Special Reserve, Madagascar, right ambulatory legs (a). Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, right cheliped and ambulatory legs (b). Scale bar = 6.6 mm." figureDoi="http://doi.org/10.5281/zenodo.6758297" httpUri="https://zenodo.org/record/6758297/files/figure.png" pageId="7" pageNumber="248">Figure 3</figureCitation>
|
||
(a);
|
||
<tableCitation id="50025FB4FFEEFFA53BD5FBBEFF0F8D0C" box="[160,246,1028,1053]" captionStart="Table 3" captionStartId="18.[160,216,142,165]" captionTargetPageId="18" captionText="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." httpUri="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="7" pageNumber="248" tableUuid="49FF3A87FFFBFFB03BD5FF34FE20880B">Table 3</tableCitation>
|
||
) and normal-length eyestalks and corneas (
|
||
<figureCitation id="85BB768AFFEEFFA5398EFBBEFCAF8D0C" box="[763,854,1028,1053]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(a)), vs long, slender, and elongated ambulatory legs (ΣP2–5/CW = 7.0) and eyestalks that taper distally and have a reduced cornea (see Cumberlidge et al. 2002, fig. 2f).
|
||
</paragraph>
|
||
<caption id="49FF3A87FFEFFFA43BD5F9EAFBD78FEB" ID-DOI="http://doi.org/10.5281/zenodo.6758295" ID-Zenodo-Dep="6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="6" pageNumber="247" startId="6.[160,225,1616,1639]" targetBox="[163,1153,142,1584]" targetPageId="6">
|
||
<paragraph id="1D3F6A0FFFEFFFA43BD5F9EAFBD78FEB" blockId="6.[159,1156,1616,1786]" pageId="6" pageNumber="247">
|
||
<emphasis id="2FF4B61DFFEFFFA43BD5F9EAFF038F76" bold="true" box="[160,250,1616,1639]" pageId="6" pageNumber="247">Figure 2.</emphasis>
|
||
<taxonomicName id="DA80118CFFEFFFA43A74F9EAFCE78F76" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[257,798,1616,1639]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFEFFFA43A74F9EAFE568F76" box="[257,431,1616,1639]" italics="true" pageId="6" pageNumber="247">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFEFFFA43ACCF9EAFD2E8F76" author="Cumberlidge N & Meyer KS" box="[441,727,1616,1639]" pageId="6" pageNumber="247" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFEFFFA43997F9EAFCE78F76" box="[738,798,1616,1639]" pageId="6" pageNumber="247" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar:
|
||
<emphasis id="2FF4B61DFFEFFFA43910F9D7FD8B8F95" bold="true" box="[613,626,1645,1668]" pageId="6" pageNumber="247">a</emphasis>
|
||
, frontal view of mandible;
|
||
<emphasis id="2FF4B61DFFEFFFA43810F9D7FC968F95" bold="true" box="[869,879,1645,1668]" pageId="6" pageNumber="247">c</emphasis>
|
||
, superior view of mandible;
|
||
<emphasis id="2FF4B61DFFEFFFA43F04F9D7FB878F95" bold="true" box="[1137,1150,1645,1668]" pageId="6" pageNumber="247">e</emphasis>
|
||
, outer view of left third maxilliped.
|
||
<taxonomicName id="DA80118CFFEFFFA43A96F931FD848FB3" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[483,637,1675,1698]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="247" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
|
||
<emphasis id="2FF4B61DFFEFFFA43A96F931FD848FB3" box="[483,637,1675,1698]" italics="true" pageId="6" pageNumber="247">Toamasina clarki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEFFFA439F6F931FD378FB3" box="[643,718,1675,1698]" pageId="6" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003, from
|
||
<taxonomicName id="DA80118CFFEFFFA43A40F912FE638FAE" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[309,410,1704,1727]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province, Madagascar:b, dorsal view of mandible;
|
||
<emphasis id="2FF4B61DFFEFFFA43806F912FC788FAE" bold="true" box="[883,897,1704,1727]" pageId="6" pageNumber="247">d</emphasis>
|
||
, superior view of mandible;
|
||
<emphasis id="2FF4B61DFFEFFFA43BEAF97FFF5E8FCD" bold="true" box="[159,167,1733,1756]" pageId="6" pageNumber="247">f</emphasis>
|
||
, outer view of left third maxilliped.Scale bar:
|
||
<emphasis id="2FF4B61DFFEFFFA43932F97FFD978FCD" bold="true" box="[583,622,1733,1756]" pageId="6" pageNumber="247">a–d</emphasis>
|
||
= 1.0 mm;
|
||
<emphasis id="2FF4B61DFFEFFFA439A1F97FFD188FCD" bold="true" box="[724,737,1733,1756]" pageId="6" pageNumber="247">e</emphasis>
|
||
,
|
||
<emphasis id="2FF4B61DFFEFFFA4399CF97FFD088FCD" bold="true" box="[745,753,1733,1756]" pageId="6" pageNumber="247">f</emphasis>
|
||
= 2.2 mm. The anterior lobe of the palp has been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="49FF3A87FFEEFFA53BD5FCA7FC2F8A9D" ID-DOI="http://doi.org/10.5281/zenodo.6758297" ID-Zenodo-Dep="6758297" httpUri="https://zenodo.org/record/6758297/files/figure.png" pageId="7" pageNumber="248" startId="7.[160,225,797,820]" targetBox="[163,1153,142,765]" targetPageId="7">
|
||
<paragraph id="1D3F6A0FFFEEFFA53BD5FCA7FC2F8A9D" blockId="7.[160,1155,797,908]" pageId="7" pageNumber="248">
|
||
<emphasis id="2FF4B61DFFEEFFA53BD5FCA7FF078A25" bold="true" box="[160,254,797,820]" pageId="7" pageNumber="248">Figure 3.</emphasis>
|
||
<taxonomicName id="DA80118CFFEEFFA53A7DFCA7FCA38A25" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[264,858,797,820]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFEEFFA53A7DFCA7FE3A8A25" box="[264,451,797,820]" italics="true" pageId="7" pageNumber="248">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFEEFFA53AA6FCA7FCF38A25" author="Cumberlidge N & Meyer KS" box="[467,778,797,820]" pageId="7" pageNumber="248" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53869FCA7FCA38A25" box="[796,858,797,820]" pageId="7" pageNumber="248" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov., paratype, adult ♀, CW 35.0 mm, FMNH 11046, from Analamerana Special Reserve, Madagascar, right ambulatory legs (
|
||
<emphasis id="2FF4B61DFFEEFFA53F1EFC80FB818A40" bold="true" box="[1131,1144,826,849]" pageId="7" pageNumber="248">a</emphasis>
|
||
).
|
||
<taxonomicName id="DA80118CFFEEFFA53BD5FCEDFEBD8A7F" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,324,855,878]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53BD5FCEDFEBD8A7F" box="[160,324,855,878]" italics="true" pageId="7" pageNumber="248">Toamasina clarki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53A39FCEDFE648A7F" box="[332,413,855,878]" pageId="7" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from
|
||
<taxonomicName id="DA80118CFFEEFFA53F6FFCEDFB7A8A7F" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[1050,1155,855,878]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province, Madagascar, right cheliped and ambulatory legs (
|
||
<emphasis id="2FF4B61DFFEEFFA53998FCCFFD028A9D" bold="true" box="[749,763,885,908]" pageId="7" pageNumber="248">b</emphasis>
|
||
). Scale bar = 6.6 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1D3F6A0FFFEEFFAA3BB5FBD6FECB8F60" blockId="7.[160,1157,993,1712]" lastBlockId="8.[160,1156,1520,1649]" lastPageId="8" lastPageNumber="249" pageId="7" pageNumber="248">
|
||
Finally,
|
||
<taxonomicName id="DA80118CFFEEFFA53A54FBD6FE828D94" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[289,379,1132,1157]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53A54FBD6FE828D94" box="[289,379,1132,1157]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53AFFFBD6FE048D94" box="[394,509,1132,1157]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFEEFFA53815FBD6FC258D94" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[864,988,1132,1157]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53815FBD6FC258D94" box="[864,988,1132,1157]" italics="true" pageId="7" pageNumber="248">Toamasina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA5389EFBD6FBA78D94" box="[1003,1118,1132,1157]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
as follows. The margin of the cheliped ischium of
|
||
<taxonomicName id="DA80118CFFEEFFA53985FB35FCB38DB9" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[752,842,1167,1192]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53985FB35FCB38DB9" box="[752,842,1167,1192]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53822FB35FC318DB9" box="[855,968,1167,1192]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is smooth and rounded (
|
||
<figureCitation id="85BB768AFFEEFFA53A6AFB0BFE748DDB" box="[287,397,1201,1226]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="7" pageNumber="248">Figure 4</figureCitation>
|
||
(e,f)), vs a cheliped ischium margin lined with small teeth (
|
||
<figureCitation id="85BB768AFFEEFFA53BDDFB6EFEEB8DFC" box="[168,274,1236,1261]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="7" pageNumber="248">Figure 4</figureCitation>
|
||
(g,h)); the S4/5 thoracic sternal sulcus of
|
||
<taxonomicName id="DA80118CFFEEFFA5386BFB6EFC818DFC" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[798,888,1236,1261]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA5386BFB6EFC818DFC" box="[798,888,1236,1261]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA538F2FB6EFC038DFC" box="[903,1018,1236,1261]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
meets the pleon at the PL5/6 suture (
|
||
<figureCitation id="85BB768AFFEEFFA53A9EFB4DFDB08C01" box="[491,585,1271,1296]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(e)), vs an S4/5 sulcus that meets the pleon at the PL/6 suture (
|
||
<figureCitation id="85BB768AFFEEFFA53A0BFAA3FE1C8C23" box="[382,485,1305,1330]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(f)); the PL6 is relatively narrow with convex lateral margins in
|
||
<taxonomicName id="DA80118CFFEEFFA53A5BFA86FE718C44" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[302,392,1340,1365]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53A5BFA86FE718C44" box="[302,392,1340,1365]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53AE3FA86FDF18C44" box="[406,520,1340,1365]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFEEFFA53969FA86FD858C44" box="[540,636,1340,1365]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(e)), vs a relatively broad PL6 with gently concave lateral margins (
|
||
<figureCitation id="85BB768AFFEEFFA53ABAFAE5FDD68C69" box="[463,559,1375,1400]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(f)); the telson has gently sinuous lateral margins with a rounded apex in
|
||
<taxonomicName id="DA80118CFFEEFFA53AA5FA3BFDD38C8B" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[464,554,1409,1434]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53AA5FA3BFDD38C8B" box="[464,554,1409,1434]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53942FA3BFD518C8B" box="[567,680,1409,1434]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFEEFFA539CEFA3BFCE28C8B" box="[699,795,1409,1434]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(e)), vs a telson with straight lateral margins with a rounded apex (
|
||
<figureCitation id="85BB768AFFEEFFA5390FFA1EFD248CAC" box="[634,733,1444,1469]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="7" pageNumber="248">Figure 1</figureCitation>
|
||
(f)); the exopod of the third maxilliped has a short flagellum (less than the merus length) in
|
||
<taxonomicName id="DA80118CFFEEFFA538DDFA7DFBFB8CF1" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[936,1026,1479,1504]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA538DDFA7DFBFB8CF1" box="[936,1026,1479,1504]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53F64FA7DFB7D8CF1" box="[1041,1156,1479,1504]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFEEFFA53BDDFA53FEE98F13" box="[168,272,1513,1538]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="7" pageNumber="248">Figure 2</figureCitation>
|
||
(e)), vs a third maxilliped exopod with a long flagellum (equal to the merus length) (
|
||
<figureCitation id="85BB768AFFEEFFA53A2FF9B6FE478F34" box="[346,446,1548,1573]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="7" pageNumber="248">Figure 2</figureCitation>
|
||
(f)); the ambulatory legs (P2–5) are short (ΣP2–5/CW = 5.7) in
|
||
<taxonomicName id="DA80118CFFEEFFA53BCAF995FEE08F59" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[191,281,1583,1608]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53BCAF995FEE08F59" box="[191,281,1583,1608]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53A56F995FE688F59" box="[291,401,1583,1608]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<tableCitation id="50025FB4FFEEFFA53AD4F995FE018F59" box="[417,504,1583,1608]" captionStart="Table 3" captionStartId="18.[160,216,142,165]" captionTargetPageId="18" captionText="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." httpUri="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="7" pageNumber="248" tableUuid="49FF3A87FFFBFFB03BD5FF34FE20880B">Table 3</tableCitation>
|
||
), vs ambulatory legs (P2–5) that are of medium length (ΣP2–5/CW = 6.4) in
|
||
<taxonomicName id="DA80118CFFEEFFA53AEDF9EBFDFF8F7B" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[408,518,1617,1642]" class="Lingulata" family="Obolidae" genus="Tomasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="7" pageNumber="248" phylum="Brachiopoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA53AEDF9EBFDFF8F7B" box="[408,518,1617,1642]" italics="true" pageId="7" pageNumber="248">Tomasina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA5397AF9EBFD848F7B" box="[527,637,1617,1642]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<tableCitation id="50025FB4FFEEFFA539FBF9EBFD1C8F7B" box="[654,741,1617,1642]" captionStart="Table 3" captionStartId="18.[160,216,142,165]" captionTargetPageId="18" captionText="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." httpUri="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="7" pageNumber="248" tableUuid="49FF3A87FFFBFFB03BD5FF34FE20880B">Table 3</tableCitation>
|
||
); the G1 TA is broadest proximally with a distal half that curves upward and ends in a broad tip in
|
||
<taxonomicName id="DA80118CFFEEFFA538C4F9CEFBF28F9C" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[945,1035,1652,1677]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="248" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFEEFFA538C4F9CEFBF28F9C" box="[945,1035,1652,1677]" italics="true" pageId="7" pageNumber="248">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFEEFFA53F60F9CEFB7D8F9C" box="[1045,1156,1652,1677]" pageId="7" pageNumber="248" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFEEFFA53BDDF92DFEF48FA1" box="[168,269,1687,1712]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="7" pageNumber="248">Figure 5</figureCitation>
|
||
(a,b)), vs a G1 TA that is widest at the midpoint, with a distal half that is straight and ends in a pointed tip (
|
||
<figureCitation id="85BB768AFFE1FFAA392BFA4AFD3D8F18" box="[606,708,1520,1545]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="8" pageNumber="249">Figure 5</figureCitation>
|
||
(d,e)); and a G2 TA that is long and flagellum-like and ends in a distinct distal curve in
|
||
<taxonomicName id="DA80118CFFE1FFAA3865F9A9FC938F3D" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[784,874,1555,1580]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="249" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFE1FFAA3865F9A9FC938F3D" box="[784,874,1555,1580]" italics="true" pageId="8" pageNumber="249">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE1FFAA3800F9A9FC1D8F3D" box="[885,996,1555,1580]" pageId="8" pageNumber="249" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="85BB768AFFE1FFAA3883F9A9FBA08F3D" box="[1014,1113,1555,1580]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="8" pageNumber="249">Figure 5</figureCitation>
|
||
(c)), vs a G2 TA that is long and flagellum-like but is straight and not curved distally (
|
||
<figureCitation id="85BB768AFFE1FFAA3BDDF9E2FEF58F60" box="[168,268,1624,1649]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="8" pageNumber="249">Figure 5</figureCitation>
|
||
(f)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |