323 lines
59 KiB
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323 lines
59 KiB
XML
<document id="53E0C05992AC8089EB8D869F6A29D830" ID-CLB-Dataset="92411" ID-DOI="10.1080/00222933.2022.2049389" ID-GBIF-Dataset="4cebde67-ce4c-402d-b8d5-901a5a528fab" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1656311063267" checkinUser="plazi" docAuthor="Leever, Ellen M., Daniels, Savel R., Soma, Julia B. & Cumberlidge, Neil" docDate="2022" docId="9529DB19FFE4FFAC3963FE21FCD98C03" docLanguage="en" docName="JNATHIST.56.1-4.241-263.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2022.2049389" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Toamasina Leever & Daniels & Soma & Cumberlidge 2022, gen. nov." docType="treatment" docVersion="9" lastPageNumber="255" masterDocId="6910A361FFE9FFA23B75FFBAFFF98911" masterDocTitle="Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)" masterLastPageNumber="263" masterPageNumber="241" pageNumber="254" updateTime="1699350272557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="984D34F3704AD5524FFBD924B7024438">Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)</mods:title>
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<mods:namePart id="D9FFC53C67AC6DB9F89B4A3C1EC7592E">Leever, Ellen M.</mods:namePart>
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<mods:namePart id="B49A45D6DEC69C2BC901DEB4AD7899C5">Daniels, Savel R.</mods:namePart>
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<mods:namePart id="D39DBBD464B6D0B48A36169594D90B3A">Soma, Julia B.</mods:namePart>
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<mods:title id="18600082862667D1532A38F8486EB964">Journal of Natural History</mods:title>
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<mods:date id="7B32553CB1ABDFC5397D6A451FF48A50">2022</mods:date>
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<treatment id="9529DB19FFE4FFAC3963FE21FCD98C03" ID-DOI="http://doi.org/10.5281/zenodo.6762507" ID-GBIF-Taxon="197158402" ID-Zenodo-Dep="6762507" LSID="urn:lsid:plazi:treatment:9529DB19FFE4FFAC3963FE21FCD98C03" httpUri="http://treatment.plazi.org/id/9529DB19FFE4FFAC3963FE21FCD98C03" lastPageId="14" lastPageNumber="255" pageId="13" pageNumber="254">
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<subSubSection id="559A3984FFE4FFAF3963FE21FCF788A4" box="[534,782,411,437]" pageId="13" pageNumber="254" type="nomenclature">
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<paragraph id="1D3F6A0FFFE4FFAF3963FE21FCF788A4" blockId="13.[534,782,411,437]" box="[534,782,411,437]" pageId="13" pageNumber="254">
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<heading id="4677DD63FFE4FFAF3963FE21FCF788A4" box="[534,782,411,437]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="254" reason="5">
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<emphasis id="2FF4B61DFFE4FFAF3963FE21FCF788A4" box="[534,782,411,437]" italics="true" pageId="13" pageNumber="254">
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<taxonomicName id="DA80118CFFE4FFAF3963FE21FD6488A4" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[534,669,411,437]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="254" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2FF4B61DFFE4FFAF3963FE21FD6488A4" bold="true" box="[534,669,411,437]" italics="true" pageId="13" pageNumber="254">Toamasina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34C70B66FFE4FFAF39D0FE21FCF788A4" box="[677,782,411,437]" pageId="13" pageNumber="254" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="559A3984FFE4FFAF3A22FE6AFF128B55" pageId="13" pageNumber="254" type="description">
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<paragraph id="1D3F6A0FFFE4FFAF3A22FE6AFC1588F8" blockId="13.[343,1004,464,489]" box="[343,1004,464,489]" pageId="13" pageNumber="254">
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(
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<figureCitation id="85BB768AFFE4FFAF3A2AFE6AFE3888F8" box="[351,449,464,489]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="13" pageNumber="254">Figures 1</figureCitation>
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(b,d,f), 2(b,d,f), 3(b), 4(c,d,g,h), 5(d), 6;
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<tableCitation id="50025FB4FFE4FFAF3818FE6AFC1D88F8" box="[877,996,464,489]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="3.[224,281,142,165]" captionStartId-1="17.[160,216,142,165]" captionStartId-2="18.[160,216,142,165]" captionStartId-3="18.[160,216,534,557]" captionTargetBox-0="[235,821,235,1031]" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="18" captionText-0="Table 1. Updated list of all known Malagasy freshwater crabs (Family Deckeniidae Ortmann, 1897, Subfamily Hydrothelphusinae Bott, 1955) from 1853 to July 2021. The" captionText-1="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." captionText-2="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." captionText-3="Table 4. Gonopod 1 (G1) measurements (taken on the ventral side) showing lengths of the G1 terminal article (TA) and the G1 subterminal article (SA) in mm for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The relative length of the G1 TA and the G1 SA is also shown, along with the size classes for the G1 TA." httpUri-0="http://table.plazi.org/id/49FF3A87FFEAFFA13B95FF34FC1A89F1" httpUri-1="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" httpUri-2="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="13" pageNumber="254" tableUuid-0="49FF3A87FFEAFFA13B95FF34FC1A89F1" tableUuid-1="49FF3A87FFF8FFB33BD5FF34FC3A89EC" tableUuid-2="49FF3A87FFFBFFB03BD5FF34FE20880B">Tables 1–4</tableCitation>
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)
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</paragraph>
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<paragraph id="1D3F6A0FFFE4FFAF3BD5FDB2FF128B55" blockId="13.[160,1155,520,580]" pageId="13" pageNumber="254">
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<emphasis id="2FF4B61DFFE4FFAF3BD5FDB2FE4D8B30" box="[160,436,520,545]" italics="true" pageId="13" pageNumber="254">Nomenclatural statement</emphasis>
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: A life science identifier (LSID) number was obtained for the new genus:
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</paragraph>
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</subSubSection>
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<subSubSection id="559A3984FFE4FFAF3BD5FDD9FE1E8B8F" pageId="13" pageNumber="254" type="reference_group">
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<paragraph id="1D3F6A0FFFE4FFAF3BD5FDD9FE1E8B8F" blockId="13.[160,1155,611,670]" pageId="13" pageNumber="254">
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<treatmentCitationGroup id="3D904D21FFE4FFAF3BD5FDD9FE1E8B8F" pageId="13" pageNumber="254">
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<treatmentCitation id="9C214C1EFFE4FFAF3BD5FDD9FD978B6D" ID-DOI="http://doi.org/10.5281/zenodo.3791535" ID-Zenodo-Dep="3791535" author="Cumberlidge N & Meyer KS" box="[160,622,611,636]" httpUri="http://treatment.plazi.org/id/03EF536EFFCE4662FF173B18FBFFFEF1" page="81" pageId="13" pageNumber="254" year="2009">
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<taxonomicName id="DA80118CFFE4FFAF3BD5FDD9FD978B6D" authority="Cumberlidge and Meyer, 2009: 81" authorityName="Cumberlidge and Meyer" authorityPageNumber="81" authorityYear="2009" box="[160,622,611,636]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="2FF4B61DFFE4FFAF3BD5FDD9FF2D8B6D" box="[160,212,611,636]" italics="true" pageId="13" pageNumber="254">Foza</emphasis>
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<bibRefCitation id="791117FEFFE4FFAF3BAAFDD9FD978B6D" author="Cumberlidge N & Meyer KS" box="[223,622,611,636]" pageId="13" pageNumber="254" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009: 81</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(partim): (not
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<taxonomicName id="DA80118CFFE4FFAF386BFDD9FED08B8F" authority="Cumberlidge and Meyer, 2009" authorityName="Cumberlidge and Meyer" authorityYear="2009" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="254" phylum="Arthropoda" rank="species" species="ambohitra">
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<emphasis id="2FF4B61DFFE4FFAF386BFDD9FC568B6D" box="[798,943,611,636]" italics="true" pageId="13" pageNumber="254">F. ambohitra</emphasis>
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<bibRefCitation id="791117FEFFE4FFAF38CFFDD9FED08B8F" author="Cumberlidge N & Meyer KS" pageId="13" pageNumber="254" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
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</taxonomicName>
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, figs 1–3, tab. 1).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="559A3984FFE4FFAF3BD5FD69FD368A1E" pageId="13" pageNumber="254" type="type_taxon">
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<paragraph id="1D3F6A0FFFE4FFAF3BD5FD69FECA8BFC" blockId="13.[160,719,723,783]" box="[160,307,723,749]" pageId="13" pageNumber="254">
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<emphasis id="2FF4B61DFFE4FFAF3BD5FD69FECA8BFC" bold="true" box="[160,307,723,749]" italics="true" pageId="13" pageNumber="254">
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<typeStatus id="C23BD4ADFFE4FFAF3BD5FD69FF218BFC" box="[160,216,723,749]" pageId="13" pageNumber="254">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="1D3F6A0FFFE4FFAF3BD5FD4CFD368A1E" blockId="13.[160,719,723,783]" box="[160,719,758,783]" pageId="13" pageNumber="254">
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<taxonomicName id="DA80118CFFE4FFAF3BD5FD4CFEA38A1E" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,346,758,783]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="254" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
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<emphasis id="2FF4B61DFFE4FFAF3BD5FD4CFEA38A1E" box="[160,346,758,783]" italics="true" pageId="13" pageNumber="254">Toamasina clarki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34C70B66FFE4FFAF3A14FD4CFE428A1E" box="[353,443,758,783]" pageId="13" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="559A3984FFE4FFAC3BD5FCFEFED18866" lastPageId="14" lastPageNumber="255" pageId="13" pageNumber="254" type="diagnosis">
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<paragraph id="1D3F6A0FFFE4FFAF3BD5FCFEFEED8A4F" blockId="13.[160,1156,836,1728]" box="[160,276,836,862]" pageId="13" pageNumber="254">
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<emphasis id="2FF4B61DFFE4FFAF3BD5FCFEFEED8A4F" bold="true" box="[160,276,836,862]" italics="true" pageId="13" pageNumber="254">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="1D3F6A0FFFE4FFAC3BD5FCDDFED18866" blockId="13.[160,1156,836,1728]" lastBlockId="14.[160,1156,142,375]" lastPageId="14" lastPageNumber="255" pageId="13" pageNumber="254">
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Entire carapace surface smooth except for faint carinae on anterolateral corners; front narrow (FW/CW = 0.27); epigastric crests, epibranchial teeth in advanced position in line with the postorbital margin; postorbital crests faint, ending before meeting epibranchial teeth; cardiac, urogastric sulci, semicircular sulcus, cervical sulci faint (
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<figureCitation id="85BB768AFFE4FFAF3886FC75FBAB8AF9" box="[1011,1106,975,1000]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="13" pageNumber="254">Figure 1</figureCitation>
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(b)). Branchiostegite suborbital region smooth except for horizontal line of granules across middle, subhepatic region smooth, pterygostomial region smooth except for small field of granules near third maxilliped (
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<figureCitation id="85BB768AFFE4FFAF3964FB8DFD948D41" box="[529,621,1079,1104]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="13" pageNumber="254">Figure 1</figureCitation>
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||
(d)). Mandibular palp terminal article bilobed, anterior lobe on terminal article conspicuous, medium sized (MPAL/MPTA = 0.5) (
|
||
<figureCitation id="85BB768AFFE4FFAF3F5DFBE0FB7A8D62" box="[1064,1155,1114,1139]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="13" pageNumber="254">Figure 2</figureCitation>
|
||
(b,d);
|
||
<tableCitation id="50025FB4FFE4FFAF3B97FBC7FECF8D87" box="[226,310,1149,1174]" captionStart="Table 2" captionStartId="17.[160,216,142,165]" captionTargetPageId="17" captionText="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." httpUri="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" pageId="13" pageNumber="254" tableUuid="49FF3A87FFF8FFB33BD5FF34FC3A89EC">Table 2</tableCitation>
|
||
). Exopod of third maxilliped with long flagellum (equal to merus length); ischium with deep vertical sulcus angled diagonally, located medially (
|
||
<figureCitation id="85BB768AFFE4FFAF388FFB25FBA68DA9" box="[1018,1119,1183,1208]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="13" pageNumber="254">Figure 2</figureCitation>
|
||
(f)). Thoracic sternal suture S2/3 deep, completely traversing sternum, S3/4 broadly U-shaped, deep at edges, not meeting anterior margin of subpleonal cavity; anterior subpleonal cavity lacking setae; S4/5 meeting pleon at PL5/6 suture; S6/7 meeting submedial part of PL6 (
|
||
<figureCitation id="85BB768AFFE4FFAF3AD2FA90FDF98C52" box="[423,512,1322,1347]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="13" pageNumber="254">Figure 1</figureCitation>
|
||
(f)). Left chela larger, cutting edge of fixed finger (pollex) with four large molars on proximal third, followed by medium and small teeth distally, cutting edge of dactylus with four large teeth proximally followed by small teeth (
|
||
<figureCitation id="85BB768AFFE4FFAF3F5CFAD5FB7A8C99" box="[1065,1155,1391,1416]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(d)); dactylus of left chela distinctly arched (
|
||
<figureCitation id="85BB768AFFE4FFAF39DAFA28FCEB8CBA" box="[687,786,1426,1451]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(d)); left chela propodus longer (
|
||
<quantity id="DA78C7EAFFE4FFAF3BD3FA0FFF0F8CDF" box="[166,246,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="254" unit="mm" value="29.0">29 mm</quantity>
|
||
vs
|
||
<quantity id="DA78C7EAFFE4FFAF3A68FA0FFE948CDF" box="[285,365,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="254" unit="mm" value="24.0">24 mm</quantity>
|
||
) and higher (
|
||
<quantity id="DA78C7EAFFE4FFAF3971FA0FFDAD8CDF" box="[516,596,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="254" unit="mm" value="16.0">16 mm</quantity>
|
||
vs
|
||
<quantity id="DA78C7EAFFE4FFAF390EFA0FFD448CDF" box="[635,701,1461,1486]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="254" unit="mm" value="8.0">8 mm</quantity>
|
||
) (
|
||
<figureCitation id="85BB768AFFE4FFAF39A6FA0FFCC98CDF" box="[723,816,1461,1486]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(d)) than right chela propodus (
|
||
<figureCitation id="85BB768AFFE4FFAF3BDDFA6DFEFD8CE1" box="[168,260,1495,1520]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(c)); cutting edges of dactylus, pollex of right chela lined with small, even, sharp teeth (
|
||
<figureCitation id="85BB768AFFE4FFAF3B9FFA40FEBF8F02" box="[234,326,1530,1555]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(c)); dactylus of right chela straight, not arched (
|
||
<figureCitation id="85BB768AFFE4FFAF3817FA40FC478F02" box="[866,958,1530,1555]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(c)); lower margin of propodus of left chela strongly indented medially (
|
||
<figureCitation id="85BB768AFFE4FFAF3870F9A7FC9A8F27" box="[773,867,1565,1590]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(d)). Distal tooth on inner margin of cheliped carpus large, acute; proximal tooth smaller, pointed, followed by small granules (
|
||
<figureCitation id="85BB768AFFE4FFAF3A7BF9D8FE938F6A" box="[270,362,1634,1659]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(g)). Inferior margins of cheliped merus lined with small, rounded teeth, distal tooth largest; inferior margins of cheliped ischium lined with small, rounded teeth (
|
||
<figureCitation id="85BB768AFFE4FFAF3BDDF91DFEF28FD1" box="[168,267,1703,1728]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="13" pageNumber="254">Figure 4</figureCitation>
|
||
(h)). Ambulatory legs (P2–5) medium length (ΣP2–5/CW = 6.4) (
|
||
<figureCitation id="85BB768AFFE4FFAF388BF91DFB998FD1" box="[1022,1120,1703,1728]" captionStart="Figure 3" captionStartId="7.[160,225,797,820]" captionTargetBox="[163,1153,142,765]" captionTargetId="figure-326@7.[163,1153,142,765]" captionTargetPageId="7" captionText="Figure 3. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., paratype, adult ♀, CW 35.0 mm, FMNH 11046, from Analamerana Special Reserve, Madagascar, right ambulatory legs (a). Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, right cheliped and ambulatory legs (b). Scale bar = 6.6 mm." figureDoi="http://doi.org/10.5281/zenodo.6758297" httpUri="https://zenodo.org/record/6758297/files/figure.png" pageId="13" pageNumber="254">Figure 3</figureCitation>
|
||
(b);
|
||
<tableCitation id="50025FB4FFE7FFAC3BD5FF34FF0D89B6" box="[160,244,142,167]" captionStart="Table 3" captionStartId="18.[160,216,142,165]" captionTargetPageId="18" captionText="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." httpUri="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="14" pageNumber="255" tableUuid="49FF3A87FFFBFFB03BD5FF34FE20880B">Table 3</tableCitation>
|
||
). Male pleon plus telson triangular, tapered distally, widest at PL3, narrowest at telson; PL6 relatively broad, with gently concave lateral margins; telson with straight lateral margins, apex rounded (
|
||
<figureCitation id="85BB768AFFE7FFAC3973FF6EFD9989FC" box="[518,608,212,237]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="14" pageNumber="255">Figure 1</figureCitation>
|
||
(f)). G1 TA medium length (G1 TA/G1 SA = 0.3), slim, widened by distinct hump at midpoint, distal half straight, tip pointed; G1 SA with distinct raised rounded shoulder on external margin near G1 TA-SA junction (
|
||
<figureCitation id="85BB768AFFE7FFAC3886FEA3FBB78823" box="[1011,1102,281,306]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="14" pageNumber="255">Figure 5</figureCitation>
|
||
(d,e);
|
||
<tableCitation id="50025FB4FFE7FFAC3BD5FE86FF018844" box="[160,248,316,341]" captionStart="Table 4" captionStartId="18.[160,216,534,557]" captionTargetPageId="18" captionText="Table 4. Gonopod 1 (G1) measurements (taken on the ventral side) showing lengths of the G1 terminal article (TA) and the G1 subterminal article (SA) in mm for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The relative length of the G1 TA and the G1 SA is also shown, along with the size classes for the G1 TA." pageId="14" pageNumber="255">Table 4</tableCitation>
|
||
). G2 TA long (G2 TA/SA = 0.62), flagellum-like, straight, not curved distally (
|
||
<figureCitation id="85BB768AFFE7FFAC3BDDFEE4FEFC8866" box="[168,261,350,375]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="14" pageNumber="255">Figure 5</figureCitation>
|
||
(f)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE7FFAC3BD5FE0BFC6A8B22" pageId="14" pageNumber="255" type="distribution">
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FE0BFED588DA" blockId="14.[160,1156,433,563]" box="[160,300,433,459]" pageId="14" pageNumber="255">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BD5FE0BFED588DA" bold="true" box="[160,300,433,459]" italics="true" pageId="14" pageNumber="255">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FE6EFC6A8B22" blockId="14.[160,1156,433,563]" pageId="14" pageNumber="255">
|
||
Endemic to
|
||
<collectingCountry id="65972A9FFFE7FFAC3A46FE6EFE4688FC" box="[307,447,468,493]" name="Madagascar" pageId="14" pageNumber="255">Madagascar</collectingCountry>
|
||
;
|
||
<taxonomicName id="DA80118CFFE7FFAC3AB8FE6EFDBE88FC" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[461,583,468,493]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3AB8FE6EFDBE88FC" box="[461,583,468,493]" italics="true" pageId="14" pageNumber="255">Toamasina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3926FE6EFD3B88FC" box="[595,706,468,493]" pageId="14" pageNumber="255" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
<materialsCitation id="ADE86052FFE7FFAC39B8FE6EFC768B22" ID-GBIF-Occurrence="3861175304" collectionCode="ASL" elevation="600" latitude="-15.2883" location="Analanjirofo Region" longLatPrecision="7" longitude="49.5483" municipality="Akirindro Mountain" pageId="14" pageNumber="255" specimenCount="1" stateProvince="Toamasina">
|
||
is known only from a single locality,
|
||
<collectingMunicipality id="FD5BF075FFE7FFAC3BD5FE4DFE868B01" box="[160,383,503,528]" pageId="14" pageNumber="255">Akirindro Mountain</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="0C8D732DFFE7FFAC3AFEFE4DFD2C8B01" box="[395,725,503,528]" location="Ambinantelo" pageId="14" pageNumber="255">
|
||
<quantity id="DA78C7EAFFE7FFAC3AFEFE4DFE218B01" box="[395,472,503,528]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="255" unit="km" value="7.6">7.6 km</quantity>
|
||
NNW of Ambinantelo
|
||
</locationDeviation>
|
||
(
|
||
<geoCoordinate id="78B40CC8FFE7FFAC3996FE4DFCAC8B01" box="[739,853,503,528]" degrees="15.2883" direction="south" orientation="latitude" pageId="14" pageNumber="255" precision="5" value="-15.2883">15.2883°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78B40CC8FFE7FFAC3814FE4DFC2A8B01" box="[865,979,503,528]" degrees="49.5483" direction="east" orientation="longitude" pageId="14" pageNumber="255" precision="5" value="49.5483">49.5483°E</geoCoordinate>
|
||
),
|
||
<quantity id="DA78C7EAFFE7FFAC3891FE4DFBD58B01" box="[996,1068,503,528]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="255" unit="m" value="600.0">
|
||
<elevation id="96AD8D3CFFE7FFAC3891FE4DFBD58B01" box="[996,1068,503,528]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="255" unit="m" value="600.0">600 m</elevation>
|
||
</quantity>
|
||
<collectionCode id="7B91F2CAFFE7FFAC3F40FE4DFB9B8B01" box="[1077,1122,503,528]" pageId="14" pageNumber="255">ASL</collectionCode>
|
||
, in the
|
||
<location id="185F3CD4FFE7FFAC3BB9FDA0FE538B22" LSID="urn:lsid:plazi:treatment:9529DB19FFE4FFAC3963FE21FCD98C03:185F3CD4FFE7FFAC3BB9FDA0FE538B22" box="[204,426,538,563]" latitude="-15.2883" longLatPrecision="7" longitude="49.5483" municipality="Akirindro Mountain" name="Analanjirofo Region" pageId="14" pageNumber="255" stateProvince="Toamasina">Analanjirofo Region</location>
|
||
of northern
|
||
<collectingRegion id="DF44A4EDFFE7FFAC394FFDA0FCE28B22" box="[570,795,538,563]" country="Madagascar" name="Toamasina" pageId="14" pageNumber="255">
|
||
<taxonomicName id="DA80118CFFE7FFAC394FFDA0FD4B8B22" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[570,690,538,563]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province
|
||
</collectingRegion>
|
||
(
|
||
<figureCitation id="85BB768AFFE7FFAC385FFDA0FC7F8B22" box="[810,902,538,563]" captionStart="Figure 6" captionStartId="16.[160,225,1033,1056]" captionTargetBox="[165,1150,144,998]" captionTargetId="figure-69@16.[163,1153,142,1001]" captionTargetPageId="16" captionText="Figure 6. Map of Madagascar showing the distributional range of Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov. (black circles, not shown to scale), Toamasina clarki sp. nov. (white triangle in black square, not shown to scale), Foza raimundi Reed and Cumberlidge, 2006 (white asterisk with black margin), and Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015 (white square with black margin). Scale bar = 184 km." figureDoi="http://doi.org/10.5281/zenodo.6758305" httpUri="https://zenodo.org/record/6758305/files/figure.png" pageId="14" pageNumber="255">Figure 6</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE7FFAC3BD5FDD6FE408A50" pageId="14" pageNumber="255" type="etymology">
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FDD6FEE48B97" blockId="14.[160,1155,620,715]" box="[160,285,620,646]" pageId="14" pageNumber="255">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BD5FDD6FEE48B97" bold="true" box="[160,285,620,646]" italics="true" pageId="14" pageNumber="255">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FD2AFC518BDA" blockId="14.[160,1155,620,715]" pageId="14" pageNumber="255">
|
||
The genus name refers to
|
||
<collectingRegion id="DF44A4EDFFE7FFAC3AB1FD2AFD5D8BB8" box="[452,676,656,681]" country="Madagascar" name="Toamasina" pageId="14" pageNumber="255">
|
||
<taxonomicName id="DA80118CFFE7FFAC3AB1FD2AFDC58BB8" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[452,572,656,681]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province
|
||
</collectingRegion>
|
||
where the crab was collected. The name is used as a Latin noun in nominative singular and treated as feminine.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FCBFFE9A8A0E" blockId="14.[160,441,773,833]" box="[160,355,773,799]" pageId="14" pageNumber="255">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BD5FCBFFE9A8A0E" bold="true" box="[160,355,773,799]" italics="true" pageId="14" pageNumber="255">Species included</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FC92FE408A50" blockId="14.[160,441,773,833]" box="[160,441,808,833]" pageId="14" pageNumber="255">
|
||
<taxonomicName id="DA80118CFFE7FFAC3BD5FC92FEA38A50" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,346,808,833]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BD5FC92FEA38A50" box="[160,346,808,833]" italics="true" pageId="14" pageNumber="255">Toamasina clarki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3A14FC92FE408A50" box="[353,441,808,833]" pageId="14" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE7FFAC3BD5FCC0FCD98C03" pageId="14" pageNumber="255" type="discussion">
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FCC0FEFF8A85" blockId="14.[160,1156,890,1298]" box="[160,262,890,916]" pageId="14" pageNumber="255">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BD5FCC0FEFF8A85" bold="true" box="[160,262,890,916]" italics="true" pageId="14" pageNumber="255">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE7FFAC3BD5FC24FCD98C03" blockId="14.[160,1156,890,1298]" pageId="14" pageNumber="255">
|
||
The new genus is established based on morphological and phylogenetic evidence. The phylogeny in
|
||
<bibRefCitation id="791117FEFFE7FFAC3A48FC7BFDAF8ACB" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[317,598,961,986]" pageId="14" pageNumber="255" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
|
||
, fig. 1) indicates that
|
||
<taxonomicName id="DA80118CFFE7FFAC383AFC7BFBF38ACB" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[847,1034,961,986]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC383AFC7BFBF38ACB" box="[847,1034,961,986]" italics="true" pageId="14" pageNumber="255">Toamasina clarki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3F67FC7BFB928ACB" box="[1042,1131,961,986]" pageId="14" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is genetically separate from
|
||
<taxonomicName id="DA80118CFFE7FFAC3ABCFC59FC338AED" authority="(Cumberlidge and Meyer, 2009)" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[457,970,995,1020]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="ambohitra" status="comb. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3ABCFC59FDA18AED" box="[457,600,995,1020]" italics="true" pageId="14" pageNumber="255">V. ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE7FFAC391DFC59FC3B8AED" author="Cumberlidge N & Meyer KS" box="[616,962,995,1020]" pageId="14" pageNumber="255" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC38A6FC59FBAB8AED" box="[979,1106,995,1020]" pageId="14" pageNumber="255" rank="species">comb. nov.</taxonomicNameLabel>
|
||
The morphological studies of
|
||
<bibRefCitation id="791117FEFFE7FFAC3AB1FBBCFCDF8D0E" author="Cumberlidge N & Meyer KS" box="[452,806,1030,1055]" pageId="14" pageNumber="255" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer (2009)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="791117FEFFE7FFAC3817FBBCFB7D8D0E" author="Cumberlidge N & Klaus S & Meyer KS & Koppin JC" box="[866,1156,1030,1055]" pageId="14" pageNumber="255" pagination="1 - 15" refId="ref11313" refString="Cumberlidge N, Klaus S, Meyer KS, Koppin JC. 2015. New collections of freshwater crabs from northern Madagascar, with the description of a new species of Foza Reed and Cumberlidge, 2006 (Brachyura, Potamonautidae). Euro J Tax. 109: 1 - 15." type="journal article" year="2015">Cumberlidge et al. (2015)</bibRefCitation>
|
||
included the specimen of
|
||
<taxonomicName id="DA80118CFFE7FFAC3A90FB93FD538D53" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[485,682,1065,1090]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3A90FB93FD538D53" box="[485,682,1065,1090]" italics="true" pageId="14" pageNumber="255">Toamasina clarki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC39CDFB93FCEE8D53" box="[696,791,1065,1090]" pageId="14" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NMU PN 17–
|
||
<date id="693E4CCFFFE7FFAC38B0FB93FBC08D53" box="[965,1081,1065,1090]" pageId="14" pageNumber="255" value="2003-03-21">21.3.2003</date>
|
||
) from
|
||
<collectingRegion id="DF44A4EDFFE7FFAC3BD5FBF1FE728D75" box="[160,395,1099,1124]" country="Madagascar" name="Toamasina" pageId="14" pageNumber="255">
|
||
<taxonomicName id="DA80118CFFE7FFAC3BD5FBF1FEE28D75" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,283,1099,1124]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province
|
||
</collectingRegion>
|
||
in
|
||
<taxonomicName id="DA80118CFFE7FFAC3ACCFBF1FDB58D75" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[441,588,1099,1124]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE7FFAC3ACCFBF1FDB58D75" box="[441,588,1099,1124]" italics="true" pageId="14" pageNumber="255">F. ambohitra</emphasis>
|
||
</taxonomicName>
|
||
; however, phylogenetically (
|
||
<bibRefCitation id="791117FEFFE7FFAC38D1FBF1FF2F8D96" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" pageId="14" pageNumber="255" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
|
||
, fig. 1) this specimen is positioned on a related, but clearly separate, lineage from the
|
||
<typeStatus id="C23BD4ADFFE7FFAC3BB8FB2BFEF08DBB" box="[205,265,1169,1194]" pageId="14" pageNumber="255">types</typeStatus>
|
||
of both
|
||
<taxonomicName id="DA80118CFFE7FFAC3A1BFB2BFE038DBB" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[366,506,1169,1194]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE7FFAC3A1BFB2BFE038DBB" box="[366,506,1169,1194]" italics="true" pageId="14" pageNumber="255">F. ambohitra</emphasis>
|
||
</taxonomicName>
|
||
(now
|
||
<taxonomicName id="DA80118CFFE7FFAC3931FB2BFF1E8DDD" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3931FB2BFCEC8DBB" box="[580,789,1169,1194]" italics="true" pageId="14" pageNumber="255">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE7FFAC3856FB2BFB858DBB" author="Cumberlidge N & Meyer KS" box="[803,1148,1169,1194]" pageId="14" pageNumber="255" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3BD5FB09FF1E8DDD" box="[160,231,1203,1228]" pageId="14" pageNumber="255" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov.) and
|
||
<taxonomicName id="DA80118CFFE7FFAC3A2EFB09FE2F8DDD" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[347,470,1203,1228]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFE7FFAC3A2EFB09FE2F8DDD" box="[347,470,1203,1228]" italics="true" pageId="14" pageNumber="255">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
. Distinguishing characters that separate
|
||
<taxonomicName id="DA80118CFFE7FFAC38EBFB09FBEF8DDD" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[926,1046,1203,1228]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC38EBFB09FBEF8DDD" box="[926,1046,1203,1228]" italics="true" pageId="14" pageNumber="255">Toamasina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3F69FB09FB7D8DDD" box="[1052,1156,1203,1228]" pageId="14" pageNumber="255" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="DA80118CFFE7FFAC3BABFB6CFECC8DFE" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[222,309,1238,1263]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3BABFB6CFECC8DFE" box="[222,309,1238,1263]" italics="true" pageId="14" pageNumber="255">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3A48FB6CFE518DFE" box="[317,424,1238,1263]" pageId="14" pageNumber="255" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, and both of these new genera from the other Malagasy genera are provided in the remarks for
|
||
<taxonomicName id="DA80118CFFE7FFAC3972FB43FDA78C03" authority="Leever & Daniels & Soma & Cumberlidge, 2022" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[519,606,1273,1298]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="255" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FF4B61DFFE7FFAC3972FB43FDA78C03" box="[519,606,1273,1298]" italics="true" pageId="14" pageNumber="255">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE7FFAC3913FB43FD378C03" box="[614,718,1273,1298]" pageId="14" pageNumber="255" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |