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<document ID-DOI="10.1206/832.1" ID-GBIF-Dataset="85bd2c66-f9f0-4172-8d82-2e8841cd354a" ID-ISSN="0003-0090" ID-Zenodo-Dep="4611863" checkinTime="1615991711531" checkinUser="felipe" docAuthor="LeCroy, Mary" docDate="2013" docId="2B4687A09E52FF9BFD09762CFEC113C1" docLanguage="en" docName="BulAmeMusNatHis.2013.381.1-155.pdf" docOrigin="Bulletin of the American Museum of Natural History 2013 (381)" docSource="http://www.bioone.org/doi/abs/10.1206/832.1" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Pyrenestes ostrinus subsp. rothschildi Neumann" docType="treatment" docVersion="9" lastPageNumber="87" masterDocId="D77FFFD89E04FFCCFFB3741AFFF6164D" masterDocTitle="Type Specimens Of Birds In The American Museum Of Natural History Part 11. Passeriformes: Parulidae, Drepanididae, Vireonidae, Icteridae, Fringillinae, Carduelinae, Estrildidae, And Viduinae" masterLastPageNumber="155" masterPageNumber="1" pageNumber="86" updateTime="1674063294785" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Type Specimens Of Birds In The American Museum Of Natural History Part 11. Passeriformes: Parulidae, Drepanididae, Vireonidae, Icteridae, Fringillinae, Carduelinae, Estrildidae, And Viduinae</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>LeCroy, Mary</mods:namePart>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-09-26</mods:number>
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<mods:detail type="volume">
<mods:number>2013</mods:number>
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<mods:detail type="issue">
<mods:number>381</mods:number>
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<mods:identifier type="DOI">10.1206/832.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">85bd2c66-f9f0-4172-8d82-2e8841cd354a</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4627660" ID-GBIF-Taxon="180662663" ID-Zenodo-Dep="4627660" LSID="urn:lsid:plazi:treatment:2B4687A09E52FF9BFD09762CFEC113C1" httpUri="http://treatment.plazi.org/id/2B4687A09E52FF9BFD09762CFEC113C1" lastPageId="87" lastPageNumber="87" pageId="86" pageNumber="86">
<subSubSection box="[698,1162,565,588]" pageId="86" pageNumber="86" type="nomenclature">
<paragraph blockId="86.[698,1162,565,588]" box="[698,1162,565,588]" pageId="86" pageNumber="86">
<heading box="[698,1162,565,588]" centered="true" fontSize="9" level="2" pageId="86" pageNumber="86" reason="2">
<taxonomicName authority="Neumann" authorityName="Neumann" box="[698,1162,565,588]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="86" pageNumber="86" phylum="Chordata" rank="subSpecies" species="ostrinus" subSpecies="rothschildi">
<emphasis bold="true" box="[698,1162,565,588]" pageId="86" pageNumber="86">
<emphasis bold="true" box="[698,1045,565,588]" italics="true" pageId="86" pageNumber="86">Pyrenestes ostrinus rothschildi</emphasis>
Neumann
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="86" pageNumber="86" type="reference_group">
<paragraph blockId="86.[667,1194,611,793]" pageId="86" pageNumber="86">
<treatmentCitationGroup pageId="86" pageNumber="86">
<treatmentCitation author="Neumann, O." page="528" pageId="86" pageNumber="86" year="1910">
<taxonomicName authority="Neumann, 1910: 528 (Warri)" authorityName="Neumann" authorityYear="1910" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="86" pageNumber="86" phylum="Chordata" rank="subSpecies" species="ostrinus" subSpecies="rothschildi">
<emphasis box="[667,973,611,631]" italics="true" pageId="86" pageNumber="86">Pyrenestes ostrinus rothschildi</emphasis>
<bibRefCitation author="Neumann, O." box="[979,1193,611,631]" pageId="86" pageNumber="86" pagination="522 - 530" refId="ref124086" refString="Neumann, O. 1910. Revision der genera Spermospiza and Pyrenestes. Journal fur Ornithologie 58: 522 - 530." type="journal article" year="1910">Neumann, 1910: 528</bibRefCitation>
(Warri)
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="86.[667,1194,611,793]" pageId="86" pageNumber="86">
<treatmentCitationGroup pageId="86" pageNumber="86">
Now
<taxonomicName ID-CoL="4QRFM" authority="(Vieillot, 1805)" baseAuthorityName="Vieillot" baseAuthorityYear="1805" box="[738,1135,666,685]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ostrinus">
<emphasis box="[738,945,667,685]" italics="true" pageId="86" pageNumber="86">Pyrenestes ostrinus</emphasis>
(Vieillot, 1805)
</taxonomicName>
. See
<treatmentCitation author="Hartert, E." box="[693,909,693,712]" page="144" pageId="86" pageNumber="86" year="1919">
<bibRefCitation author="Hartert, E." box="[693,909,693,712]" pageId="86" pageNumber="86" pagination="123 - 178" refId="ref119004" refString="Hartert, E. 1919 a. Types of birds in the Tring Museum. B. Types in the general collection. Novitates Zoologicae 26: 123 - 178." type="journal article" year="1919">Hartert, 1919a: 144</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Chapin, J. P." box="[925,1190,693,712]" page="490" pageId="86" pageNumber="86" year="1954">
<bibRefCitation author="Chapin, J. P." box="[925,1190,693,712]" pageId="86" pageNumber="86" pagination="1 - 846" refId="ref114020" refString="Chapin, J. P. 1954 b. The birds of the Belgian Congo, Part 4. Bulletin of the American Museum of Natural History 75 B: i-ix, 1 - 846, 27 pls." type="book chapter" year="1954">Chapin, 1954b: 490494</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Mayr, E. &amp; R. A. Paynter, Jr. &amp; M. A. Traylor &amp; African" box="[693,987,720,739]" page="319" pageId="86" pageNumber="86" year="1968">
<bibRefCitation author="Mayr, E. &amp; R. A. Paynter, Jr. &amp; M. A. Traylor &amp; African" box="[693,987,720,739]" pageId="86" pageNumber="86" pagination="306 - 390" refId="ref123101" refString="Mayr, E. [Australo-Papuan]., R. A. Paynter, Jr. [Oriental], and M. A. Traylor [African]. 1968. Family Estrildidae. In R. A. Paynter, Jr. (editor), Check-list of birds of the world, vol. 14: 306 - 390. Cambridge, MA: Museum of Comparative Zoology, 433 pp." type="book chapter" year="1968">Mayr et al., 1968: 319320</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Smith, T. B." page="381" pageId="86" pageNumber="86" year="1990">
<bibRefCitation author="Smith, T. B." pageId="86" pageNumber="86" pagination="381 - 414" refId="ref128462" refString="Smith, T. B. 1990. Patterns of morphological and geographic variation in trophic bill morphs of the African finch Pyrenestes. Biological Journal of the Linnean Society 41: 381 - 414." type="journal article" year="1990">Smith, 1990: 381 414</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Dickinson, E. C." box="[743,964,747,766]" page="728" pageId="86" pageNumber="86" year="2003">
<bibRefCitation author="Dickinson, E. C." box="[743,964,747,766]" pageId="86" pageNumber="86" refId="ref116121" refString="Dickinson, E. C. (editor). 2003. The Howard and Moore complete checklist of the birds of the world, 3 rd ed. London: Christopher Helm, 1039 pp." type="book" year="2003">Dickinson, 2003: 728</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Fry, C. H. &amp; S. Keith" page="317" pageId="86" pageNumber="86" year="2004">
<bibRefCitation author="Fry, C. H. &amp; S. Keith" pageId="86" pageNumber="86" refId="ref116661" refString="Fry, C. H., and S. Keith (editors). 2004. The birds of Africa, vol. 7. Princeton, NJ: Princeton University Press, xxi + 666 pp., 36 pls." type="book" year="2004">Fry and Keith, 2004: 317321</bibRefCitation>
</treatmentCitation>
; and
<treatmentCitation author="Payne, R. B." box="[840,1022,774,793]" page="322" pageId="86" pageNumber="86" year="2010">
<bibRefCitation author="Payne, R. B." box="[840,1022,774,793]" pageId="86" pageNumber="86" pagination="299 - 377" refId="ref124981" refString="Payne, R. B. 2010. Family Estrildidae (waxbills), species accounts. In J. del Hoyo, A. Elliott, and D. Christie (editors), Handbook of the birds of the world, vol. 15, weavers to New World warblers: 299 - 377. Barcelona: Lynx Edicions, 879 pp., 60 pls., photographs." type="book chapter" year="2010">Payne, 2010: 322</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="86.[667,1193,816,1759]" pageId="86" pageNumber="86">
<materialsCitation ID-GBIF-Occurrence="3059738996" collectingDate="1897-05-11" collectionCode="AMNH" country="Niger" latitude="5.36" location="Warri" longLatPrecision="783" longitude="5.5" pageId="86" pageNumber="86" specimenCode="AMNH 728281" specimenCount="1" stateProvince="Rivers" typeStatus="lectotype">
<typeStatus box="[696,833,816,840]" pageId="86" pageNumber="86">LECTOTYPE</typeStatus>
:
<emphasis bold="true" box="[853,1038,818,839]" pageId="86" pageNumber="86">
<specimenCode box="[853,1038,818,839]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="86" pageNumber="86">AMNH 728281</specimenCode>
</emphasis>
, adult male, collected at
<location LSID="urn:lsid:plazi:treatment:2B4687A09E52FF9BFD09762CFEC113C1:A630606D9E52FF9AFC9C774AFC81152B" box="[815,887,848,870]" country="Niger" latitude="5.36" longLatPrecision="783" longitude="5.5" name="Warri" pageId="86" pageNumber="86" stateProvince="Rivers">Warri</location>
,
<geoCoordinate box="[903,987,848,870]" degrees="05.36" direction="north" orientation="latitude" pageId="86" pageNumber="86" precision="555" value="5.36">05.36N</geoCoordinate>
<geoCoordinate box="[999,1079,848,870]" degrees="05.50" direction="east" orientation="longitude" pageId="86" pageNumber="86" precision="555" value="5.5">05.50E</geoCoordinate>
(
<bibRefCitation author="Chapin, J. P." pageId="86" pageNumber="86" pagination="638 - 738" refId="ref113973" refString="Chapin, J. P. 1954 a. Gazetteer of African localities mentioned in Parts 1 to 4 [of '' The birds of the Belgian Congo' ']. In Bulletin of the American Museum of Natural History 75 B: 638 - 738." type="book chapter" year="1954">Chapin, 1954a</bibRefCitation>
), lower
<collectingCountry box="[833,899,877,899]" name="Niger" pageId="86" pageNumber="86">Niger</collectingCountry>
<collectingRegion box="[908,976,877,899]" country="Nigeria" name="Rivers" pageId="86" pageNumber="86">River</collectingRegion>
, southern
<collectingCountry box="[1099,1189,877,899]" name="Nigeria" pageId="86" pageNumber="86">Nigeria</collectingCountry>
, on
<date box="[709,872,907,929]" pageId="86" pageNumber="86" value="1897-05-11">
<collectingDate box="[709,872,907,929]" pageId="86" pageNumber="86" value="1897-05-11">11 May 1897</collectingDate>
</date>
, by Felix Roth. From the Rothschild Collection.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="87" lastPageNumber="87" pageId="86" pageNumber="86" type="discussion">
<paragraph blockId="86.[667,1193,816,1759]" pageId="86" pageNumber="86">
COMMENTS: In the original description, Neumann designated as the type a male collected at Warri on
<date box="[921,1073,1026,1048]" pageId="86" pageNumber="86" value="1897-05-11">11 May 1897</date>
and listed his type series: in the Rothschild Collection, three males, two females from Warri; two males, four females, one juvenile from Buguma and Degama,
<collectingCountry box="[909,975,1144,1166]" name="Niger" pageId="86" pageNumber="86">Niger</collectingCountry>
Delta; one female from Abutshi. Also in his type series were a male and a female from
<collectingCountry box="[983,1106,1204,1225]" name="Cameroon" pageId="86" pageNumber="86">Cameroon</collectingCountry>
in the Berlin Museum. All three of the males from Warri were collected on the same date;
<materialsCitation ID-GBIF-Occurrence="3059738464" collectionCode="AMNH" pageId="86" pageNumber="86" specimenCode="AMNH 728281" specimenCount="1">
one of them is not the type as it is molting into adult plumage, but the other two males cannot be differentiated on the basis of the type description nor from
<bibRefCitation author="Hartert, E." box="[832,1085,1381,1403]" pageId="86" pageNumber="86" pagination="123 - 178" refId="ref119004" refString="Hartert, E. 1919 a. Types of birds in the Tring Museum. B. Types in the general collection. Novitates Zoologicae 26: 123 - 178." type="journal article" year="1919">Harterts (1919a: 144)</bibRefCitation>
listing of the type.
<specimenCode box="[793,984,1411,1432]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="86" pageNumber="86">AMNH 728281</specimenCode>
was Neumanns intended type as the Rothschild Museum label is annotated by Neumann with the new name and Typus, and it bears a Rothschild type label. This specimen was cataloged as the type when the Rothschild Collection came to
<collectionCode box="[667,760,1589,1610]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="86" pageNumber="86">AMNH</collectionCode>
and has always been so considered
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3059738409" collectionCode="AMNH" pageId="86" pageNumber="86" specimenCode="AMNH 728281" specimenCount="1" typeStatus="lectotype">
in order to remove the ambiguity, I hereby designate
<specimenCode box="[790,981,1648,1669]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="86" pageNumber="86">AMNH 728281</specimenCode>
the
<typeStatus box="[1048,1152,1648,1670]" pageId="86" pageNumber="86">lectotype</typeStatus>
of
<taxonomicName box="[667,1011,1676,1699]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="86" pageNumber="86" phylum="Chordata" rank="subSpecies" species="ostrinus" subSpecies="rothschildi">
<emphasis box="[667,1011,1676,1699]" italics="true" pageId="86" pageNumber="86">Pyrenestes ostrinus rothschildi</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="86.[667,1193,816,1759]" lastBlockId="87.[128,654,213,1420]" lastPageId="87" lastPageNumber="87" pageId="86" pageNumber="86">
<typeStatus box="[696,865,1707,1729]" pageId="86" pageNumber="86">Paralectotypes</typeStatus>
in AMNH are difficult to list with certainty. The two additional males and two females collected by Roth at Warri are definitely
<typeStatus box="[285,457,242,264]" pageId="87" pageNumber="87">paralectotypes</typeStatus>
:
<emphasis bold="true" box="[467,646,242,263]" pageId="87" pageNumber="87">AMNH 728282</emphasis>
, male;
<emphasis bold="true" box="[199,379,272,293]" pageId="87" pageNumber="87">AMNH 728283</emphasis>
, sexed as a male but in female plumage;
<emphasis bold="true" box="[331,514,301,322]" pageId="87" pageNumber="87">AMNH 728284</emphasis>
, sexed as a female but in male plumage with the only sign of immaturity a brown crown (these latter two specimens may have had the labels put on the wrong bird), collected on
<date pageId="87" pageNumber="87" value="1897-05-11">11 May 1897</date>
;
<emphasis bold="true" box="[196,376,449,470]" pageId="87" pageNumber="87">AMNH 728285</emphasis>
, female, collected on
<date pageId="87" pageNumber="87" value="1897-05-15">15 May 1897</date>
. Also a definite
<typeStatus box="[489,651,479,501]" pageId="87" pageNumber="87">paralectotype</typeStatus>
:
<emphasis bold="true" box="[128,309,509,530]" pageId="87" pageNumber="87">AMNH 728286</emphasis>
, female, collected at Abutshi by Kemp in
<date box="[306,505,538,560]" pageId="87" pageNumber="87" value="1901-09">September 1901</date>
. The seven specimens listed by Neumann from Buguma and Degama collected by Ansorge, all in 1902, are more problematic. There are nine specimens in AMNH from those localities, some of which bear both the names
<taxonomicName baseAuthorityName="J.F.Gmelin" baseAuthorityYear="1789" box="[547,653,687,708]" class="Aves" family="Fringillidae" genus="Loxops" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="coccineus">
<emphasis box="[547,653,687,708]" italics="true" pageId="87" pageNumber="87">coccineus</emphasis>
</taxonomicName>
and
<emphasis box="[186,304,716,739]" italics="true" pageId="87" pageNumber="87">rothschildi</emphasis>
. In some cases
<taxonomicName baseAuthorityName="J.F.Gmelin" baseAuthorityYear="1789" box="[515,621,717,738]" class="Aves" family="Fringillidae" genus="Loxops" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="coccineus">
<emphasis box="[515,621,717,738]" italics="true" pageId="87" pageNumber="87">coccineus</emphasis>
</taxonomicName>
is marked out, in other cases not. I have included as
<typeStatus box="[273,441,775,797]" pageId="87" pageNumber="87">paralectotypes</typeStatus>
only those specimens which bear the name
<emphasis box="[445,564,804,827]" italics="true" pageId="87" pageNumber="87">rothschildi</emphasis>
written by Neumann, whether or not the name
<taxonomicName baseAuthorityName="J.F.Gmelin" baseAuthorityYear="1789" box="[128,234,865,886]" class="Aves" family="Fringillidae" genus="Loxops" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="coccineus">
<emphasis box="[128,234,865,886]" italics="true" pageId="87" pageNumber="87">coccineus</emphasis>
</taxonomicName>
appears: Buguma,
<emphasis bold="true" box="[465,646,864,885]" pageId="87" pageNumber="87">AMNH 728287</emphasis>
, male, 5 June;
<emphasis bold="true" box="[314,503,894,915]" pageId="87" pageNumber="87">AMNH 728289</emphasis>
, female, 16 July;
<emphasis bold="true" box="[207,403,923,945]" pageId="87" pageNumber="87">AMNH 728290</emphasis>
, female, 21 May;
<emphasis bold="true" box="[128,317,953,974]" pageId="87" pageNumber="87">AMNH 728291</emphasis>
, female, 21 May;
<emphasis bold="true" pageId="87" pageNumber="87">AMNH 728292</emphasis>
, female, 9 May. Degama,
<emphasis bold="true" pageId="87" pageNumber="87">AMNH 728294</emphasis>
, male, 15 March;
<emphasis bold="true" box="[458,646,1012,1033]" pageId="87" pageNumber="87">AMNH 728295</emphasis>
, female juvenile, 9 May. AMNH 728288, male, 16 July, from Buguma, and AMNH 728293, male, 3 August, from Degama have only the name
<taxonomicName baseAuthorityName="J.F.Gmelin" baseAuthorityYear="1789" box="[328,434,1132,1153]" class="Aves" family="Fringillidae" genus="Loxops" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="coccineus">
<emphasis box="[328,434,1132,1153]" italics="true" pageId="87" pageNumber="87">coccineus</emphasis>
</taxonomicName>
written by Neumann, and are not considered
<typeStatus box="[478,650,1161,1183]" pageId="87" pageNumber="87">paralectotypes</typeStatus>
.
</paragraph>
<paragraph blockId="87.[128,654,213,1420]" pageId="87" pageNumber="87">
Species and subspecies recognition has varied greatly over time and the concensus of opinion most recently is that three species comprising a superspecies should be recognized:
<taxonomicName authorityName="Shelley" authorityYear="1894" box="[208,408,1310,1331]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="minor">
<emphasis box="[208,408,1310,1331]" italics="true" pageId="87" pageNumber="87">Pyrenestes minor</emphasis>
</taxonomicName>
,
<emphasis box="[427,560,1310,1331]" italics="true" pageId="87" pageNumber="87">
<taxonomicName baseAuthorityName="Vieillot" baseAuthorityYear="1805" box="[427,556,1310,1331]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="ostrinus">P. ostrinus</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Swainson" authorityYear="1837" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="sanguineus">
<emphasis italics="true" pageId="87" pageNumber="87">P. sanguineus</emphasis>
</taxonomicName>
, with
<taxonomicName baseAuthorityName="Vieillot" baseAuthorityYear="1805" box="[333,460,1340,1361]" class="Aves" family="Estrildidae" genus="Pyrenestes" kingdom="Animalia" order="Passeriformes" pageId="87" pageNumber="87" phylum="Chordata" rank="species" species="ostrinus">
<emphasis box="[333,460,1340,1361]" italics="true" pageId="87" pageNumber="87">P. ostrinus</emphasis>
</taxonomicName>
monotypic (see
<bibRefCitation author="Fry, C. H. &amp; S. Keith" box="[128,501,1369,1391]" pageId="87" pageNumber="87" refId="ref116661" refString="Fry, C. H., and S. Keith (editors). 2004. The birds of Africa, vol. 7. Princeton, NJ: Princeton University Press, xxi + 666 pp., 36 pls." type="book" year="2004">Fry and Keith, 2004: 317318</bibRefCitation>
, and
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).
</paragraph>
</subSubSection>
</treatment>
</document>