127 lines
20 KiB
XML
127 lines
20 KiB
XML
<document id="2067E0232633882243AFF61C117280E4" ID-DOI="10.11646/zootaxa.3846.1.1" ID-GBIF-Dataset="01a9d979-7b10-4c36-b1af-4acc2c0cb9a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286792" ID-ZooBank="97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460464243854" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2014" docId="113287AD9B58FFDEFF13FB49FCE0FEEE" docLanguage="en" docName="zt03846p041.pdf" docOrigin="Zootaxa 3846 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvarium Pic 1918" docType="treatment" docVersion="6" lastPageNumber="2" masterDocId="ED0BFFD59B59FFDCFF84FFF7FFB5FFF3" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer" masterLastPageNumber="41" masterPageNumber="1" pageNumber="2" updateTime="1698304595582" updateUser="plazi">
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<mods:titleInfo id="181C0747D13E8053DD3DAD1387CF7560">
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<mods:title id="483532130F83141A19B110BC538ADA7B">Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer</mods:title>
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<mods:name id="68061EECAD11985257BDB504E72457C3" type="personal">
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<mods:namePart id="A0A9DA662C51EA70BBDCA1B02CDC631D">Zwick, Peter</mods:namePart>
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<mods:typeOfResource id="7EFB6EF590F9C58852DFFC23ECDD4D50">text</mods:typeOfResource>
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<mods:title id="DBCFDE89DBD991081DFBC14A23FE8BB6">Zootaxa</mods:title>
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<mods:part id="5979D5C71CDA80B2A423B480D048CD9B">
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<mods:date id="D12649DE67B8C4BF2D0C6EED6A43F71B">2014</mods:date>
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<mods:detail id="39FB44B8171B67A223D7C94A3508AFB7" type="volume">
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<mods:number id="5DBCD64DAD2B60029C813F3CC2D204DB">3846</mods:number>
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<mods:detail id="549B28DA1EC754BC834CC8F911E4BFCE" type="issue">
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<mods:number id="EBB15CFFFB5B0BE5F61F6B452B40F665">1</mods:number>
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<mods:classification id="1BE23D006969CB990410BD20AD158DD5">journal article</mods:classification>
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<mods:identifier id="D2FF14964017140EE2A17B7F106CE907" type="DOI">10.11646/zootaxa.3846.1.1</mods:identifier>
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<treatment id="113287AD9B58FFDEFF13FB49FCE0FEEE" ID-DOI="http://doi.org/10.5281/zenodo.6126797" ID-GBIF-Taxon="127669609" ID-Zenodo-Dep="6126797" LSID="urn:lsid:plazi:treatment:113287AD9B58FFDEFF13FB49FCE0FEEE" httpUri="http://treatment.plazi.org/id/113287AD9B58FFDEFF13FB49FCE0FEEE" lastPageId="2" lastPageNumber="2" pageId="1" pageNumber="2">
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<subSubSection id="D18165309B58FFDEFF13FB49FCE0FEEE" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="992436BB9B58FFDDFF13FB49FE52FB2B" blockId="1.[151,487,1214,1240]" box="[151,487,1214,1240]" pageId="1" pageNumber="2">
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<heading id="C26C81D79B58FFDDFF13FB49FE52FB2B" bold="true" box="[151,487,1214,1240]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="ABEFEAA99B58FFDDFF13FB49FE52FB2B" bold="true" box="[151,487,1214,1240]" pageId="1" pageNumber="2">
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Genus
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<taxonomicName id="5E9B4D389B58FFDDFF6AFB49FE52FB2B" authority="Pic, 1918" authorityName="Pic" authorityYear="1918" box="[238,487,1214,1240]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="ABEFEAA99B58FFDDFF6AFB49FEDAFB2B" bold="true" box="[238,367,1214,1240]" italics="true" pageId="1" pageNumber="2">Calvarium</emphasis>
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Pic, 1918
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF13FAF2FEF6FA79" blockId="1.[151,1436,1285,1490]" pageId="1" pageNumber="2">
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The genus
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<emphasis id="ABEFEAA99B58FFDDFE93FAF2FE1EFAEE" box="[279,427,1285,1309]" italics="true" pageId="1" pageNumber="2">
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<taxonomicName id="5E9B4D389B58FFDDFE93FAF2FE3AFAEE" box="[279,399,1285,1309]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Calvarium</taxonomicName>
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w
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</emphasis>
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as proposed for non-saltatorial African
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<taxonomicName id="5E9B4D389B58FFDDFCF5FAF2FC60FAED" box="[881,981,1285,1310]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
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with unmodified hind legs. The poorly known genus was redescribed by Ruta (2010). He transferred several species first described in genus
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<taxonomicName id="5E9B4D389B58FFDDFAA2FADEFAC8FAB2" box="[1318,1405,1321,1345]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="ABEFEAA99B58FFDDFAA2FADEFAC8FAB2" box="[1318,1405,1321,1345]" italics="true" pageId="1" pageNumber="2">Cyphon</emphasis>
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</taxonomicName>
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to
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<taxonomicName id="5E9B4D389B58FFDDFF13FABAFEBAFA96" box="[151,271,1357,1381]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="ABEFEAA99B58FFDDFF13FABAFEBAFA96" box="[151,271,1357,1381]" italics="true" pageId="1" pageNumber="2">Calvarium</emphasis>
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</taxonomicName>
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and provided a list of the named species, many of them Asian. The genus is for the first time recorded from
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<collectingCountry id="E18C762B9B58FFDDFF50FA86FE8BFA79" box="[212,318,1393,1418]" name="Australia" pageId="1" pageNumber="2">Australia</collectingCountry>
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.
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF43FA62FEA4FA21" blockId="1.[151,1436,1285,1490]" pageId="1" pageNumber="2">
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The six new Australian species agree with the redescription of
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<taxonomicName id="5E9B4D389B58FFDDFC00FA62FB3CFA5D" authority="Ruta 2010" authorityName="Ruta" authorityYear="2010" box="[900,1161,1429,1454]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="ABEFEAA99B58FFDDFC00FA62FC49FA5E" box="[900,1020,1429,1453]" italics="true" pageId="1" pageNumber="2">Calvarium</emphasis>
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(Ruta 2010)
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</taxonomicName>
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in the following derived characters:
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF13F9F5FF64F9CD" blockId="1.[151,1437,1537,2033]" pageId="1" pageNumber="2">
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• A wide short head with a narrow oral area, the labrum corresponding to only
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<emphasis id="ABEFEAA99B58FFDDFBA2F9F5FBF4F9EA" box="[1062,1089,1538,1561]" italics="true" pageId="1" pageNumber="2">ca</emphasis>
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20% of total head width (
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<figureCitation id="01A02A3E9B58FFDDFAE0F9F6FF76F9CD" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="1" pageNumber="2">Figs. 1, 6</figureCitation>
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).
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF13F9BDFD99F991" blockId="1.[151,1437,1537,2033]" box="[151,556,1609,1634]" pageId="1" pageNumber="2">
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• A deep antennal sulcus (
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<figureCitation id="01A02A3E9B58FFDDFE5DF9BEFDABF991" box="[473,542,1609,1634]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
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).
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF13F999FBFBF907" blockId="1.[151,1437,1537,2033]" pageId="1" pageNumber="2">
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• Antennal scape wide, with sharp anterior edge. At rest it lies horizontally in the space between the antennal insertion and the eye, filling it completely. Pedicel inserted on the lower face of the scape, near its distal edge. The pedicel and the base of the flagellum lie deep in the antennal sulcus when the pedicel stands at right angle to the scape. The tip of the long 2nd flagellomere projects caudally from the sulcus (
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<figureCitation id="01A02A3E9B58FFDDFC78F92BFBF5F907" box="[1020,1088,1756,1781]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
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).
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</paragraph>
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<paragraph id="992436BB9B58FFDDFF13F8F6FEEEF85B" blockId="1.[151,1437,1537,2033]" pageId="1" pageNumber="2">
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• Prosternal process with anterior portion parallel-sided, posterior portion wide, lanceolate, pilose (
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<figureCitation id="01A02A3E9B58FFDDFA8CF8F7FAECF8EB" box="[1288,1369,1792,1817]" captionStart="FIGURES 18 − 20" captionStartId="7.[151,264,1825,1848]" captionTargetBox="[248,1335,1100,1793]" captionTargetId="figure@7.[244,1342,1095,1801]" captionTargetPageId="7" captionText="FIGURES 18 − 20. Calvarium (Calvariellum) hamifer, n. sp., male: 18, contour in dorsal view; 19, antenna; 20, central portion of thorax, ventral side. After photographs. Fig. 20 not to scale, diagrammatic. Abbreviations: cx I, fore coxa; cx II, middle coxa; g, groove receiving prosternal process (psp); msp, process of mesoventrite; mtp, process of metaventrite." httpUri="https://zenodo.org/record/286796/files/figure.png" pageId="1" pageNumber="2">Fig. 20</figureCitation>
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). • Groove on mesoventrite receiving the prosternal process wide and short, bordered by ridges (
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<figureCitation id="01A02A3E9B58FFDDFB5FF8D3FA9AF8CF" box="[1243,1327,1828,1853]" captionStart="FIGURES 18 − 20" captionStartId="7.[151,264,1825,1848]" captionTargetBox="[248,1335,1100,1793]" captionTargetId="figure@7.[244,1342,1095,1801]" captionTargetPageId="7" captionText="FIGURES 18 − 20. Calvarium (Calvariellum) hamifer, n. sp., male: 18, contour in dorsal view; 19, antenna; 20, central portion of thorax, ventral side. After photographs. Fig. 20 not to scale, diagrammatic. Abbreviations: cx I, fore coxa; cx II, middle coxa; g, groove receiving prosternal process (psp); msp, process of mesoventrite; mtp, process of metaventrite." httpUri="https://zenodo.org/record/286796/files/figure.png" pageId="1" pageNumber="2">Fig. 20</figureCitation>
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). • Mesoventrite wide and short, barely longer than distance between middle coxae (
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<figureCitation id="01A02A3E9B58FFDDFBD0F8BFFB1DF893" box="[1108,1192,1864,1889]" captionStart="FIGURES 18 − 20" captionStartId="7.[151,264,1825,1848]" captionTargetBox="[248,1335,1100,1793]" captionTargetId="figure@7.[244,1342,1095,1801]" captionTargetPageId="7" captionText="FIGURES 18 − 20. Calvarium (Calvariellum) hamifer, n. sp., male: 18, contour in dorsal view; 19, antenna; 20, central portion of thorax, ventral side. After photographs. Fig. 20 not to scale, diagrammatic. Abbreviations: cx I, fore coxa; cx II, middle coxa; g, groove receiving prosternal process (psp); msp, process of mesoventrite; mtp, process of metaventrite." httpUri="https://zenodo.org/record/286796/files/figure.png" pageId="1" pageNumber="2">Fig. 20</figureCitation>
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). • Coarse, groove-like punctures laterally from mesoventral groove and behind front margin of metaventral process (
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<figureCitation id="01A02A3E9B58FFDDFF7EF867FEFBF85B" box="[250,334,1936,1961]" captionStart="FIGURES 18 − 20" captionStartId="7.[151,264,1825,1848]" captionTargetBox="[248,1335,1100,1793]" captionTargetId="figure@7.[244,1342,1095,1801]" captionTargetPageId="7" captionText="FIGURES 18 − 20. Calvarium (Calvariellum) hamifer, n. sp., male: 18, contour in dorsal view; 19, antenna; 20, central portion of thorax, ventral side. After photographs. Fig. 20 not to scale, diagrammatic. Abbreviations: cx I, fore coxa; cx II, middle coxa; g, groove receiving prosternal process (psp); msp, process of mesoventrite; mtp, process of metaventrite." httpUri="https://zenodo.org/record/286796/files/figure.png" pageId="1" pageNumber="2">Fig. 20</figureCitation>
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).
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</paragraph>
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<paragraph id="992436BB9B58FFDEFF43F843FCE0FEEE" blockId="1.[151,1437,1537,2033]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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The characteristic structure of the described male genitalia of
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<taxonomicName id="5E9B4D389B58FFDDFC0BF843FBB2F83F" box="[911,1031,1972,1996]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="ABEFEAA99B58FFDDFC0BF843FBB2F83F" box="[911,1031,1972,1996]" italics="true" pageId="1" pageNumber="2">Calvarium</emphasis>
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</taxonomicName>
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species, including membranous S8 and S9 lacking sclerites (Klausnitzer 2006a, b, Ruta 2007, 2010,Yoshitomi 1998, 2007, Yoshitomi & Sato 2004) is shared by only two of the new Australian species, from the Northern Territory (
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<figureCitation id="01A02A3E9B5BFFDEFBB7FF60FB0BFF43" box="[1075,1214,151,176]" captionStart-0="FIGURES 10 − 13" captionStart-1="FIGURES 14 − 17" captionStartId-0="4.[151,264,1270,1293]" captionStartId-1="5.[151,264,1608,1631]" captionTargetBox-0="[236,1346,204,1245]" captionTargetBox-1="[168,1417,462,1568]" captionTargetId-0="figure@4.[232,1355,195,1247]" captionTargetId-1="figure@5.[151,1436,447,1584]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 10 − 13. Calvarium (Calvarium) australiense, n. sp., male: 10, T 8; 11, T 9; 12, penis; 13, tegmen and parameres. All to the same scale." captionText-1="FIGURES 14 − 17. Calvarium (Calvarium) superbum, n. sp., male: 14, superimposed T 8 and T 9; 15, tegmen and parameres; 16, penis; 14 – 16 are dorsal views, to the same scale. 17, slightly oblique lateral view of tegmen and parameres overlying the penis; after a photograph of the cleared, artificially everted genitalia, not to scale." httpUri-0="https://zenodo.org/record/286794/files/figure.png" httpUri-1="https://zenodo.org/record/286795/files/figure.png" pageId="2" pageNumber="3">Figs. 10–17</figureCitation>
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). Four new species from North Queensland differ in male genitalia (
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<figureCitation id="01A02A3E9B5BFFDEFD35FF4BFC8DFF26" box="[689,824,188,213]" captionStart="FIGURES 21 − 25" captionStartId="8.[151,264,1336,1359]" captionTargetBox="[249,1337,203,1303]" captionTargetId="figure@8.[242,1345,195,1312]" captionTargetPageId="8" captionText="FIGURES 21 − 25. Calvarium (Calvariellum) hamifer, n. sp., male: 21, T 8, ventral; 22, S 8; 23, S 9 and the apodemes representing T 9, superimposed; 24, penis; 25 tegmen (25 holotype, the other paratypes from Mt. Tozer; all to the same scale)." httpUri="https://zenodo.org/record/286797/files/figure.png" pageId="2" pageNumber="3">Figs. 21–25</figureCitation>
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,
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<figureCitation id="01A02A3E9B5BFFDEFCC7FF4BFC38FF27" box="[835,909,188,212]" captionStart-0="FIGURES 29 − 37" captionStart-1="FIGURES 38 − 42" captionStartId-0="10.[151,264,1707,1730]" captionStartId-1="12.[151,264,1706,1729]" captionTargetBox-0="[246,1345,203,1671]" captionTargetBox-1="[244,1342,846,1680]" captionTargetId-0="figure@10.[228,1358,197,1686]" captionTargetId-1="figure@12.[237,1349,842,1683]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." captionText-1="FIGURES 38 − 42. Calvarium (Calvariellum) lancifer, n. sp., male: 38, T 8; 39, detail of microtrichial armature of same; 40, one of the struts representing T 9; 41, S 9 and tegmen (tn) with parameres (p) superimposed; 42, penis. All figures except 39 to the same scale." httpUri-0="https://zenodo.org/record/286799/files/figure.png" httpUri-1="https://zenodo.org/record/286800/files/figure.png" pageId="2" pageNumber="3">29–42</figureCitation>
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). The sutures on the lower face of the head also differ somewhat (
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<figureCitation id="01A02A3E9B5BFFDEFEE6FF28FE67FF0B" box="[354,466,223,248]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 6</figureCitation>
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). Therefore, a new subgenus,
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<emphasis id="ABEFEAA99B5BFFDEFCB5FF17FC70FF0B" box="[817,965,224,248]" italics="true" pageId="2" pageNumber="3">Calvariellum</emphasis>
|
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<emphasis id="ABEFEAA99B5BFFDEFC4AFF28FBF6FF0B" bold="true" box="[974,1091,223,248]" pageId="2" pageNumber="3">n. subgen</emphasis>
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., is proposed separating them from the nominate subgenus in which all other species remain.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |