treatments-xml/data/03/B1/16/03B11679A46DFFC9FF48F9B8FB20FD21.xml
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<document id="E1A9AFF2DC2205B57D5D91C8327B9A32" ID-CLB-Dataset="6486" ID-DOI="10.11646/zootaxa.4966.5.1" ID-GBIF-Dataset="3e79baaa-3ab0-4829-9ed9-2867ea361655" ID-ISSN="1175-5326" ID-Zenodo-Dep="4745182" ID-ZooBank="E708D37D-D3C5-41AC-A87B-EC9153167D38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620632671783" checkinUser="plazi" docAuthor="Krupitsky, Anatoly V." docDate="2021" docId="03B11679A46DFFC9FF48F9B8FB20FD21" docLanguage="en" docName="zootaxa.4966.5.1.pdf" docOrigin="Zootaxa 4966 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolycaena (Rhymnaria) davidi subsp. tannuola Krupitsky 2021, ssp. n." docType="treatment" docVersion="7" lastPageNumber="514" masterDocId="FF886E01A466FFC4FFDFFF89FFF7FFA4" masterDocTitle="Taxonomy of the Neolycaena (Rhymnaria) davidi (Oberthür, 1881) species group with the description of two new taxa from Russia and China (Lepidoptera, Lycaenidae)" masterLastPageNumber="518" masterPageNumber="501" pageNumber="512" updateTime="1698938401448" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="47E928F3369D4384E6D42C74880ECA77">Taxonomy of the Neolycaena (Rhymnaria) davidi (Oberthür, 1881) species group with the description of two new taxa from Russia and China (Lepidoptera, Lycaenidae)</mods:title>
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<mods:name id="2A9662A860B0505E39567E033D672005" type="personal">
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<mods:namePart id="13CE7EFFEFBF5C03848DB67A649FE350">Krupitsky, Anatoly V.</mods:namePart>
<mods:nameIdentifier id="6C68952CE48F1BC4E338D25E06EE1D3B" type="ORCID">0000-0002-6095-3570</mods:nameIdentifier>
<mods:affiliation id="D4FED7951B94ED4FF0C8618CA9A1C2D1">Department of Entomology, Biological Faculty, Lomonosov Moscow State University, Leninskie gory, GSP- 1, korp. 12, Moscow 119991, Russia. &amp; Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect, 33, Moscow, 119071, Russia. nephrurus @ yandex. ru; https: // orcid. org / 0000 - 0002 - 6095 - 3570</mods:affiliation>
<mods:nameIdentifier id="EB90F39E344BD0EAE384F46B60582C21" type="email">nephrurus@yandex.ru</mods:nameIdentifier>
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<mods:date id="88CEAFCF1E53D4E6A24A2EDC25A72641">2021</mods:date>
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<treatment id="03B11679A46DFFC9FF48F9B8FB20FD21" ID-DOI="http://doi.org/10.5281/zenodo.4792235" ID-GBIF-Taxon="182388148" ID-Zenodo-Dep="4792235" LSID="urn:lsid:plazi:treatment:03B11679A46DFFC9FF48F9B8FB20FD21" httpUri="http://treatment.plazi.org/id/03B11679A46DFFC9FF48F9B8FB20FD21" lastPageId="13" lastPageNumber="514" pageId="11" pageNumber="512">
<subSubSection id="C302F4E4A46DFFCFFF48F9B8FC86F9EF" box="[151,881,1585,1612]" pageId="11" pageNumber="512" type="nomenclature">
<paragraph id="8BA7A76FA46DFFCFFF48F9B8FC86F9EF" blockId="11.[151,881,1585,1647]" box="[151,881,1585,1612]" pageId="11" pageNumber="512">
<heading id="D0EF1003A46DFFCFFF48F9B8FC86F9EF" bold="true" box="[151,881,1585,1612]" fontSize="11" level="1" pageId="11" pageNumber="512" reason="1">
<emphasis id="B96C7B7DA46DFFCFFF48F9B8FC86F9EF" bold="true" box="[151,881,1585,1612]" pageId="11" pageNumber="512">
<taxonomicName id="4C18DCECA46DFFCFFF48F9B8FCE1F9EF" authority="Krupitsky" authorityName="Krupitsky" authorityYear="2021" box="[151,790,1585,1612]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="subSpecies" species="davidi" status="subsp. nov." subGenus="Rhymnaria" subSpecies="tannuola">
<emphasis id="B96C7B7DA46DFFCFFF48F9B8FED2F9E8" bold="true" box="[151,293,1585,1612]" italics="true" pageId="11" pageNumber="512">Neolycaena</emphasis>
(
<emphasis id="B96C7B7DA46DFFCFFEE9F9B8FE48F9E8" bold="true" box="[310,447,1585,1612]" italics="true" pageId="11" pageNumber="512">Rhymnaria</emphasis>
)
<emphasis id="B96C7B7DA46DFFCFFE0FF9B8FD79F9E8" bold="true" box="[464,654,1585,1612]" italics="true" pageId="11" pageNumber="512">davidi tannuola</emphasis>
Krupitsky
</taxonomicName>
,
<taxonomicNameLabel id="A25FC606A46DFFCFFCFCF9B8FC86F9EF" box="[803,881,1585,1611]" pageId="11" pageNumber="512" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46DFFCFFF48F9DCFE13F9CB" box="[151,484,1621,1647]" pageId="11" pageNumber="512" type="description">
<paragraph id="8BA7A76FA46DFFCFFF48F9DCFE13F9CB" blockId="11.[151,881,1585,1647]" box="[151,484,1621,1647]" pageId="11" pageNumber="512">
(
<figureCitation id="1323BBEAA46DFFCFFF7EF9DCFF19F9CB" box="[161,238,1621,1647]" captionStart="FIGURE 8" captionStartId="5.[151,250,1206,1230]" captionTargetBox="[159,1427,188,1170]" captionTargetId="figure-15@5.[151,1436,181,1182]" captionTargetPageId="5" captionText="FIGURE 8. N. (R.) d. tannuola ssp. n., holotype, ♂, dorsal side and ventral side, Russia, southern Tuva, southern slopes of Tannuola Mts., Samagaltai v., 1400 m, 26.V.05.VI.2004 (larvae), ex pupa 2630.VI.2004, S. Vaschenko leg. (ZMMU); FIGURE 9. N. (R.) d. tannuola ssp. n., paratype, ♀, dorsal side and ventral side, same label as in holotype; FIGURE 10. N. (R.) d. victoriae Churkin, 2004, ♂, dorsal side and ventral side, Russia, C Tuva, S slopes of Ujuksky Mts., near Seserlig vill., 10001200 m, 29.VI.10.VII.2002 (AKM); FIGURE 11. N. (R.) d. victoriae Churkin, 2004, ♀, dorsal side and ventral side, Russia, Tuva, Ujuksky Mts., Seserlig vill., 1600 m, 20.VI.2002 (AKM); FIGURE 12. N. (R.) d. irkuta Zhdanko, 1996, ♂, holotype, “Irkut // Leder” (ZISP); FIGURE 13. Id., label of the holotype; FIGURE 14. N. (R.) tangutica (Grum-Grshimailo, 1891), ♂, dorsal side and ventral side, China, Gansu Prov., Gannan Prefecture, Xiahe County, Xiahe (H河), Daxia River (±H河) valley SE slope, N 35°1050” E 102°3034”, 30003300 m, 26.VII.2016, A.V. Krupitsky leg. (AKM); FIGURE 15. N. (R.) tangutica (Grum-Grshimailo, 1891), ♀, dorsal side and ventral side, China, Gansu Prov., Gannan Prefecture, Xiahe County, Xiahe (H河), Daxia river (±H河) valley SE slope, N 35°1050” E 102°3034”, 30003300 m, 26.VII.2016, A.V. Krupitsky leg. (AKM);" figureDoi="http://doi.org/10.5281/zenodo.4745188" httpUri="https://zenodo.org/record/4745188/files/figure.png" pageId="11" pageNumber="512">Figs. 8</figureCitation>
, 9, 19, 23, 31, 34, 39)
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46DFFCFFF48F913FCEAF974" pageId="11" pageNumber="512" type="reference_group">
<paragraph id="8BA7A76FA46DFFCFFF48F913FD6FF916" blockId="11.[151,797,1690,1745]" box="[151,664,1690,1714]" pageId="11" pageNumber="512">
<treatmentCitationGroup id="AB088041A46DFFCFFF48F913FD6FF916" box="[151,664,1690,1714]" pageId="11" pageNumber="512">
<taxonomicName id="4C18DCECA46DFFCFFF48F913FEA6F916" authorityName="Zhdanko" authorityYear="1996" box="[151,337,1690,1714]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="irkuta">
<emphasis id="B96C7B7DA46DFFCFFF48F913FEA6F916" box="[151,337,1690,1714]" italics="true" pageId="11" pageNumber="512">Neolycaena irkuta</emphasis>
</taxonomicName>
<treatmentCitation id="0AB9817EA46DFFCFFEB3F913FDCFF916" author="Zhdanko, A. B." box="[364,568,1690,1714]" page="658" pageId="11" pageNumber="512" year="1998">
<bibRefCitation id="EF89DA9EA46DFFCFFEB3F913FDCFF916" author="Zhdanko, A. B." box="[364,568,1690,1714]" pageId="11" pageNumber="512" pagination="639 - 662" refId="ref14451" refString="Zhdanko, A. B. (1998) A review of the genus Neolycaena de Niceville, 1890 (Lepidoptera, Lycaenidae) with descriptions of the new subspecies. Entomologicheskoe obozreniye, 77 (4), 639 - 662. [in Russian]" type="journal article" year="1998">Zhdanko, 1998: 658</bibRefCitation>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA7A76FA46DFFCFFF48F931FCEAF974" blockId="11.[151,797,1690,1745]" box="[151,797,1720,1745]" pageId="11" pageNumber="512">
<treatmentCitationGroup id="AB088041A46DFFCFFF48F931FCEAF974" box="[151,797,1720,1745]" pageId="11" pageNumber="512">
<taxonomicName id="4C18DCECA46DFFCFFF48F931FE14F974" box="[151,483,1720,1744]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="subSpecies" species="davidi" subSpecies="namkhaidorji">
<emphasis id="B96C7B7DA46DFFCFFF48F931FE14F974" box="[151,483,1720,1744]" italics="true" pageId="11" pageNumber="512">Neolycaena davidi namkhaidorji</emphasis>
</taxonomicName>
<treatmentCitation id="0AB9817EA46DFFCFFE22F931FD4BF975" author="Churkin, S." box="[509,700,1720,1745]" page="182" pageId="11" pageNumber="512" year="2004">
<bibRefCitation id="EF89DA9EA46DFFCFFE22F931FD4BF975" author="Churkin, S." box="[509,700,1720,1745]" pageId="11" pageNumber="512" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin 2004: 182</bibRefCitation>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46DFFCFFF48F977FABEF869" pageId="11" pageNumber="512" type="materials_examined">
<paragraph id="8BA7A76FA46DFFCFFF48F977FABEF869" blockId="11.[151,1437,1790,2033]" pageId="11" pageNumber="512">
<emphasis id="B96C7B7DA46DFFCFFF48F977FEF1F8BC" bold="true" box="[151,262,1790,1816]" pageId="11" pageNumber="512">Material.</emphasis>
<materialsCitation id="3B70AD32A46DFFCFFED0F976FABEF869" ID-GBIF-Occurrence="3118564309" collectingDate="2001-06-18" collectingDateMax="2004-06-30" collectingDateMin="2001-06-18" collectionCode="ZMMU, IYuM, RYaB" collectorName="S. Vaschenko &amp; R. Yakovlev" country="Russia" county="Tanuola Mts." elevation="1400" location="Tanuola Mts." municipality="Samagaltai" pageId="11" pageNumber="512" specimenCount="85" specimenCount-female="32" specimenCount-male="53" stateProvince="Tyva" typeStatus="holotype">
<typeStatus id="54A319CDA46DFFCFFED0F976FE8BF8BD" box="[271,380,1791,1817]" pageId="11" pageNumber="512" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1E6CE6A46DFFCFFE58F976FE43F8BD" box="[391,436,1791,1817]" pageId="11" pageNumber="512" type="male">1 ♂</specimenCount>
,
<collectingCountry id="F30FE7FFA46DFFCFFE1DF976FDF8F8BD" box="[450,527,1791,1817]" name="Russia" pageId="11" pageNumber="512">Russia</collectingCountry>
, southern
<collectingRegion id="49DC698DA46DFFCFFD5BF976FD48F8BD" box="[644,703,1791,1817]" country="Russia" name="Tyva" pageId="11" pageNumber="512">Tuva</collectingRegion>
, southern slopes of
<collectingCounty id="62C6DFE3A46DFFCFFC7DF976FBCDF8BD" box="[930,1082,1791,1817]" pageId="11" pageNumber="512">Tanuola Mts.</collectingCounty>
,
<collectingMunicipality id="6BC33D15A46DFFCFFB98F976FB34F8BD" box="[1095,1219,1791,1817]" pageId="11" pageNumber="512">Samagaltai</collectingMunicipality>
v.,
<quantity id="4CE00A8AA46DFFCFFB31F976FAB1F8BD" box="[1262,1350,1791,1817]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="512" unit="m" value="1400.0">
<elevation id="0035405CA46DFFCFFB31F976FAB1F8BD" box="[1262,1350,1791,1817]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="512" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FFA681AFA46DFFCFFA8CF976FEE0F899" pageId="11" pageNumber="512" value="2004-05-26" valueMax="2004-06-05" valueMin="2004-05-26">
<collectingDate id="EFE27847A46DFFCFFA8CF976FEE0F899" pageId="11" pageNumber="512" value="2004-05-26" valueMax="2004-06-05" valueMin="2004-05-26">26.V. 05.VI.2004</collectingDate>
</date>
(larvae), ex pupa
<date id="FFA681AFA46DFFCFFE31F8AAFD6EF899" box="[494,665,1827,1853]" pageId="11" pageNumber="512" value="2004-06-26" valueMax="2004-06-30" valueMin="2004-06-26">
<collectingDate id="EFE27847A46DFFCFFE31F8AAFD6EF899" box="[494,665,1827,1853]" pageId="11" pageNumber="512" value="2004-06-26" valueMax="2004-06-30" valueMin="2004-06-26">2630.VI.2004</collectingDate>
</date>
,
<collectorName id="26EDC2B9A46DFFCFFD77F8AAFCB6F899" box="[680,833,1827,1853]" pageId="11" pageNumber="512">S. Vaschenko</collectorName>
leg. (
<collectionCode id="ED093FAAA46DFFCFFC59F8AAFC13F899" box="[902,996,1827,1853]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="512" type="Museum">ZMMU</collectionCode>
);
<typeStatus id="54A319CDA46DFFCFFC25F8AAFB9DF899" box="[1018,1130,1827,1853]" pageId="11" pageNumber="512" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D1E6CE6A46DFFCFFBA8F8AAFB51F899" box="[1143,1190,1827,1853]" pageId="11" pageNumber="512" type="male">7 ♂</specimenCount>
,
<specimenCount id="9D1E6CE6A46DFFCFFB6AF8AAFB07F899" box="[1205,1264,1827,1853]" pageId="11" pageNumber="512" type="female">10 ♀</specimenCount>
, same label as in the
<typeStatus id="54A319CDA46DFFCFFF3FF8CEFEB5F8C5" box="[224,322,1863,1889]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
(SChM),
<specimenCount id="9D1E6CE6A46DFFCFFE6AF8CEFE17F8C5" box="[437,480,1863,1889]" pageId="11" pageNumber="512" type="female">1 ♀</specimenCount>
, same label (
<collectionCode id="ED093FAAA46DFFCFFDA7F8CEFD37F8C5" box="[632,704,1863,1889]" pageId="11" pageNumber="512">IYuM</collectionCode>
);
<specimenCount id="9D1E6CE6A46DFFCFFD0BF8CEFD08F8C5" box="[724,767,1863,1889]" pageId="11" pageNumber="512" type="male">1 ♂</specimenCount>
(dissected),
<specimenCount id="9D1E6CE6A46DFFCFFC4FF8CEFC4CF8C5" box="[912,955,1863,1889]" pageId="11" pageNumber="512" type="female">1 ♀</specimenCount>
(dissected),
<collectingCountry id="F30FE7FFA46DFFCFFB93F8CEFB6EF8C5" box="[1100,1177,1863,1889]" name="Russia" pageId="11" pageNumber="512">Russia</collectingCountry>
,
<collectingRegion id="49DC698DA46DFFCFFB79F8CEFB16F8C5" box="[1190,1249,1863,1889]" country="Russia" name="Tyva" pageId="11" pageNumber="512">Tuva</collectingRegion>
, southern slopes of
<location id="8EC7F1B4A46DFFCFFF69F8E2FEB9F821" LSID="urn:lsid:plazi:treatment:03B11679A46DFFC9FF48F9B8FB20FD21:8EC7F1B4A46DFFCFFF69F8E2FEB9F821" box="[182,334,1899,1925]" country="Russia" county="Tanuola Mts." municipality="Samagaltai" name="Tanuola Mts." pageId="11" pageNumber="512" stateProvince="Tyva">Tanuola Mts.</location>
,
<locationDeviation id="9A15BE4DA46DFFCFFE85F8E2FDA9F821" box="[346,606,1899,1925]" location="Samagaltai" pageId="11" pageNumber="512">
<quantity id="4CE00A8AA46DFFCFFE85F8E2FE54F821" box="[346,419,1899,1925]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="512" unit="km" value="30.0">30 km</quantity>
NW Samagaltai
</locationDeviation>
v.,
<quantity id="4CE00A8AA46DFFCFFD56F8E3FD16F821" box="[649,737,1898,1925]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="512" unit="m" value="1500.0">
<elevation id="0035405CA46DFFCFFD56F8E3FD16F821" box="[649,737,1898,1925]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="512" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFA681AFA46DFFCFFD31F8E3FC86F821" box="[750,881,1898,1925]" pageId="11" pageNumber="512" value="2001-06-25">
<collectingDate id="EFE27847A46DFFCFFD31F8E3FC86F821" box="[750,881,1898,1925]" pageId="11" pageNumber="512" value="2001-06-25">25.VI.2001</collectingDate>
</date>
,
<collectorName id="26EDC2B9A46DFFCFFCA2F8E2FBFFF821" box="[893,1032,1899,1925]" pageId="11" pageNumber="512">R. Yakovlev</collectorName>
leg. (SChM);
<specimenCount id="9D1E6CE6A46DFFCFFB71F8E2FB2BF821" box="[1198,1244,1899,1925]" pageId="11" pageNumber="512" type="male">4 ♂</specimenCount>
,
<specimenCount id="9D1E6CE6A46DFFCFFB36F8E2FAE3F821" box="[1257,1300,1899,1925]" pageId="11" pageNumber="512" type="female">2 ♀</specimenCount>
, same label (
<collectionCode id="ED093FAAA46DFFCFFF40F806FF10F80D" box="[159,231,1935,1961]" pageId="11" pageNumber="512">RYaB</collectionCode>
);
<specimenCount id="9D1E6CE6A46DFFCFFF26F806FED5F80D" box="[249,290,1935,1961]" pageId="11" pageNumber="512" type="male">7 ♂</specimenCount>
<collectingCountry id="F30FE7FFA46DFFCFFEF7F806FE82F80D" box="[296,373,1935,1961]" name="Russia" pageId="11" pageNumber="512">Russia</collectingCountry>
, southern
<collectingRegion id="49DC698DA46DFFCFFE39F806FDD6F80D" box="[486,545,1935,1961]" country="Russia" name="Tyva" pageId="11" pageNumber="512">Tuva</collectingRegion>
,
<locationDeviation id="9A15BE4DA46DFFCFFDF4F807FD49F80D" box="[555,702,1934,1961]" location="Erzin" pageId="11" pageNumber="512">
<quantity id="4CE00A8AA46DFFCFFDF4F807FD94F80D" box="[555,611,1934,1961]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="512" unit="km" value="5.0">5 km</quantity>
S Erzin
</locationDeviation>
, semidesert with
<taxonomicName id="4C18DCECA46DFFCFFCA1F806FBBCF80C" box="[894,1099,1935,1961]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="512" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis id="B96C7B7DA46DFFCFFCA1F806FBBCF80C" box="[894,1099,1935,1961]" italics="true" pageId="11" pageNumber="512">Caragana spinosa</emphasis>
</taxonomicName>
,
<date id="FFA681AFA46DFFCFFB87F806FB2DF80D" box="[1112,1242,1935,1961]" pageId="11" pageNumber="512" value="2001-06-18">
<collectingDate id="EFE27847A46DFFCFFB87F806FB2DF80D" box="[1112,1242,1935,1961]" pageId="11" pageNumber="512" value="2001-06-18">18.VI.2001</collectingDate>
</date>
,
<collectorName id="26EDC2B9A46DFFCFFB3BF806FA9AF80D" box="[1252,1389,1935,1961]" pageId="11" pageNumber="512">R. Yakovlev</collectorName>
leg. (SChM);
<specimenCount id="9D1E6CE6A46DFFCFFEDDF83AFECBF869" box="[258,316,1971,1997]" pageId="11" pageNumber="512" type="male">32 ♂</specimenCount>
,
<specimenCount id="9D1E6CE6A46DFFCFFE97F83AFE77F869" box="[328,384,1971,1997]" pageId="11" pageNumber="512" type="female">18 ♀</specimenCount>
, same label (
<collectionCode id="ED093FAAA46DFFCFFDCCF83AFDACF869" box="[531,603,1971,1997]" pageId="11" pageNumber="512">RYaB</collectionCode>
);
<specimenCount id="9D1E6CE6A46DFFCFFDB0F83AFD6CF869" box="[623,667,1971,1997]" pageId="11" pageNumber="512" type="male">1 ♂</specimenCount>
, southern
<collectingRegion id="49DC698DA46DFFCFFCD1F83AFCBEF869" box="[782,841,1971,1997]" country="Russia" name="Tyva" pageId="11" pageNumber="512">Tuva</collectingRegion>
,
<location id="8EC7F1B4A46DFFCFFC8BF83AFC64F869" LSID="urn:lsid:plazi:treatment:03B11679A46DFFC9FF48F9B8FB20FD21:8EC7F1B4A46DFFCFFC8BF83AFC64F869" box="[852,915,1971,1997]" country="Russia" county="Tanuola Mts." municipality="Samagaltai" name="Erzin" pageId="11" pageNumber="512" stateProvince="Tyva">Erzin</location>
,
<date id="FFA681AFA46DFFCFFC40F83AFBD6F869" box="[927,1057,1971,1997]" pageId="11" pageNumber="512" value="2001-06-21">
<collectingDate id="EFE27847A46DFFCFFC40F83AFBD6F869" box="[927,1057,1971,1997]" pageId="11" pageNumber="512" value="2001-06-21">21.VI.2001</collectingDate>
</date>
,
<collectorName id="26EDC2B9A46DFFCFFBF3F83AFB41F869" box="[1068,1206,1971,1997]" pageId="11" pageNumber="512">R. Yakovlev</collectorName>
leg. (
<collectionCode id="ED093FAAA46DFFCFFB2AF83AFACAF869" box="[1269,1341,1971,1997]" pageId="11" pageNumber="512">IYuM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46DFFC8FF18F85FFB95FA95" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512" type="description">
<paragraph id="8BA7A76FA46DFFCFFF18F85FFDD4F855" blockId="11.[151,1437,1790,2033]" box="[199,547,2006,2033]" pageId="11" pageNumber="512">
<emphasis id="B96C7B7DA46DFFCFFF18F85FFEAEF854" bold="true" box="[199,345,2006,2032]" pageId="11" pageNumber="512">Description.</emphasis>
Male (figs. 8, 19)
</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FF1EFCA4FEB9" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Head: antenna black, white-ringed at bases of segments, club dark with brown tip. Eye surrounded by white stripe, brown with short sparse brownish hairs. Frons black with white hairs closer to vertex. Vertex covered with black scales. Occiput surrounded by whitish hairs. Palpi: segment 2 black dorsally, with white base and white sides ventrally; segment 3 black with white scales on base ventrally.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FEAEFE3BFEC1" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Thorax: dorsal side brown with grey hairs, ventral side brown, densely covered with whitish hairs. Legs brown with white scales and hairs.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FEE6FD1FFE2D" blockId="12.[151,1437,151,2013]" box="[199,744,367,393]" pageId="12" pageNumber="513">Abdomen: dorsal side brown, ventral side white.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FE1AFCE5FDC5" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Forewing: dorsal side brown, base of wing covered with long light brown scales.Androconial patch on forewing poorly developed, dark brown, narrow, rather small. Outer margin dark brown. Fringe white distally, brown mesially, dark-brown basally. Ventral side brown, with light brown submarginal area and upper half of cell, darker at postdiscal and submarginal pattern. Margin with creamy scales near base of wing. Transverse vein marked with white stroke. Spaces R1—M2 with white postdiscal strokes suffused by dark brown scales basally. Submarginal area and apex whitish between veins, with two rows of darkened spots.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FDE2FBBBFD0D" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Hindwing: rounded, with small anal lobe marked with long whitish hairs. Dorsal side greyish brown, outer margin black. Fringe dirty white distally, brown with admixture of white proximally.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FD3AFC27FCF9" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Ventral side greyish brown with bluish scales and hairs in basal area and light brown scales in discal and postdiscal area. Transverse vein marked by bracket-like white stroke. Postdiscal row consisting of separate white strokes with strong dark shading from inner side tending to form single band. Submarginal pattern somewhat reduced, separated by veins consisting of two rows of black spots connected by orange scales and surrounded by white and pale scales, with clear-cut inner row of white V-shaped strokes.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FCEEFC6BFC25" blockId="12.[151,1437,151,2013]" box="[199,924,871,897]" pageId="12" pageNumber="513">
Forewing length of
<typeStatus id="54A319CDA46AFFC8FE7AFCEEFDF0FC25" box="[421,519,871,897]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
14.0 mm,
<typeStatus id="54A319CDA46AFFC8FDA0FCEEFD1DFC25" box="[639,746,871,897]" pageId="12" pageNumber="513" type="paratype">paratypes</typeStatus>
13.016.0 mm.
</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FC02FA98FB91" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Male genitalia (fig. 31, 32). Falx long, rather thick, pointed on tip; lobes of uncus with well-developed pointed processes; valva narrow, elongate, reaching or nearly reaching lobe of uncus, with ovoid basal part and somewhat shorter distal part slightly broadened to tip and bearing small thorn; vinculum inwardly with large lateral projections nearly reaching valvae; saccus very short, rounded. Aedeagus slightly longer than genital capsule, with arcuate upper cornutus, short, nearly straight under cornutus and with nearly straight sclerotized keel with rounded tip.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FBB6FF3DFB05" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Female (figs. 9, 22). Similar to male in appearance. Dorsal side stronger suffused by bluish and greyish scales, dark suffusion of postdiscal pattern less developed. Abdomen with black anal tuft. Forewing length 12.015.0 mm.</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FB22FE1CFB61" blockId="12.[151,1437,151,2013]" box="[199,491,1195,1221]" pageId="12" pageNumber="513">Female genitalia (fig. 38).</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18FB46FB95FA95" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">Distal part of ductus bursae very broad, rhomboid, with long triangular lamella postvaginalis divided on two parts by membrane and funnel-shaped antrum turning in very short and thin ductus bursae broadened to base; bursa membranous, with very small bidentate signa, second tooth of signa poorly developed.</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46AFFC9FF18FAB3FACAFE09" lastPageId="13" lastPageNumber="514" pageId="12" pageNumber="513" type="diagnosis">
<paragraph id="8BA7A76FA46AFFC8FF18FAB3FCACF8A1" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">
<emphasis id="B96C7B7DA46AFFC8FF18FAB3FEB5FAF0" bold="true" box="[199,322,1338,1364]" pageId="12" pageNumber="513">Diagnosis.</emphasis>
In general, the male and female genitalia of
<emphasis id="B96C7B7DA46AFFC8FCE4FAB2FCB9FAF0" box="[827,846,1339,1364]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FC83FAB2FC23FAF1" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[860,980,1339,1365]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FCB9FAB2FC80FAF0" box="[870,887,1339,1364]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FC51FAB2FC23FAF1" box="[910,980,1339,1365]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
<emphasis id="B96C7B7DA46AFFC8FC03FAB2FBB7FAF1" box="[988,1088,1339,1365]" italics="true" pageId="12" pageNumber="513">tannuola</emphasis>
<emphasis id="B96C7B7DA46AFFC8FB98FAB2FB65FAF1" bold="true" box="[1095,1170,1339,1365]" pageId="12" pageNumber="513">ssp. n.</emphasis>
share all the diagnostic characters of the wings and genitalia structure with the other known subspecies of
<emphasis id="B96C7B7DA46AFFC8FB88FAD6FB9DFADC" box="[1111,1130,1375,1400]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FBA5FAD6FB03FADD" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[1146,1268,1375,1401]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FB5CFAD6FB63FADC" box="[1155,1172,1375,1400]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FB71FAD6FB03FADD" box="[1198,1268,1375,1401]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
as well as the colouration of the female abdomen, although the females of the new subspecies are characterised by the larger distal part of the ductus bursae. The pale ventral side of the wings makes the new subspecies similar to the geographically adjacent
<emphasis id="B96C7B7DA46AFFC8FF20FA42FEE5FA40" box="[255,274,1483,1508]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FEFCFA42FE69FA41" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[291,414,1483,1509]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FEF3FA42FECAFA40" box="[300,317,1483,1508]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FE87FA42FE69FA41" box="[344,414,1483,1509]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
<emphasis id="B96C7B7DA46AFFC8FE77FA42FDFBFA40" box="[424,524,1483,1508]" italics="true" pageId="12" pageNumber="513">victoriae</emphasis>
<bibRefCitation id="EF89DA9EA46AFFC8FDC9FA42FD48FA41" author="Churkin, S." box="[534,703,1483,1509]" pageId="12" pageNumber="513" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin, 2004</bibRefCitation>
. The unique diagnostic character of the new subspecies clearly distinguishing it from the other subspecies of
<emphasis id="B96C7B7DA46AFFC8FD41FA66FD46F9AC" box="[670,689,1519,1544]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FD60FA66FCCFF9AD" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[703,824,1519,1545]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FD16FA66FD2DF9AC" box="[713,730,1519,1544]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FD2DFA66FCCFF9AD" box="[754,824,1519,1545]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
is a peculiarity of the postdiscal pattern of the ventral side of the hindwing: white strokes of the postdiscal pattern are strongly shaded by dark brown scales, especially in the males, the postdiscal pattern tending to form a connected band (postdiscal pattern poorly shaded or brown scales absent, consisted of separate strokes in the other subspecies). The four strokes of the postdiscal band in the cells Sc+R1M2 form Z-shaped marking in the upper part of the hindwing in most of the studied specimens, especially in males. Additionally, the new subspecies is characterised by poorly developed orange elements of the submarginal pattern of the hindwing compared to the other subspecies of
<emphasis id="B96C7B7DA46AFFC8FCE1F94EFCA6F944" box="[830,849,1735,1760]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FC80F94EFC2CF945" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[863,987,1735,1761]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FCB7F94EFC8EF944" box="[872,889,1735,1760]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FC4EF94EFC2CF945" box="[913,987,1735,1761]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
<emphasis id="B96C7B7DA46AFFC8FC04F94EFC28F945" box="[987,991,1735,1761]" italics="true" pageId="12" pageNumber="513">.</emphasis>
This character somewhat recalls that of
<emphasis id="B96C7B7DA46AFFC8FF48F962FED9F8A1" box="[151,302,1771,1797]" italics="true" pageId="12" pageNumber="513">namkhaidorji</emphasis>
, see diagnosis above and in
<bibRefCitation id="EF89DA9EA46AFFC8FDB1F962FCAFF8A1" author="Churkin, S." box="[622,856,1771,1797]" pageId="12" pageNumber="513" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin (2004: 184)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA7A76FA46AFFC8FF18F887FB30F8E9" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="513">
<emphasis id="B96C7B7DA46AFFC8FF18F887FE48F88C" bold="true" box="[199,447,1806,1832]" pageId="12" pageNumber="513">Individual variation.</emphasis>
The intensity of the dark postdiscal shading varies between specimens. The shape of the male genital capsule slightly varies between specimens, as in other subspecies of
<emphasis id="B96C7B7DA46AFFC8FBF8F8BAFBCDF8E8" box="[1063,1082,1843,1868]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FB97F8BAFB37F8E9" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[1096,1216,1843,1869]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FB8EF8BAFB95F8E8" box="[1105,1122,1843,1868]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FBA5F8BAFB37F8E9" box="[1146,1216,1843,1869]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA7A76FA46AFFC9FF18F8DFFACAFE09" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1437,151,645]" lastPageId="13" lastPageNumber="514" pageId="12" pageNumber="513">
<emphasis id="B96C7B7DA46AFFC8FF18F8DFFE7BF8D5" bold="true" box="[199,396,1878,1905]" pageId="12" pageNumber="513">Taxonomic note.</emphasis>
Sergei Churkin (pers. comm.) kindly informed me that during the preparation of the review of the
<taxonomicName id="4C18DCECA46AFFC8FF3BF8F2FEDDF831" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[228,298,1915,1941]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B96C7B7DA46AFFC8FF3BF8F2FEDDF831" box="[228,298,1915,1941]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF89DA9EA46AFFC8FE00F8F2FD76F831" author="Churkin, S." box="[479,641,1915,1941]" pageId="12" pageNumber="513" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin 2004</bibRefCitation>
) he ascribed populations of
<emphasis id="B96C7B7DA46AFFC8FC16F8F2FC2BF830" box="[969,988,1915,1940]" italics="true" pageId="12" pageNumber="513">N</emphasis>
.
<taxonomicName id="4C18DCECA46AFFC8FC33F8F2FB91F831" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[1004,1126,1915,1941]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46AFFC8FC2AF8F2FBF1F830" box="[1013,1030,1915,1940]" italics="true" pageId="12" pageNumber="513">R</emphasis>
.)
<emphasis id="B96C7B7DA46AFFC8FBFFF8F2FB91F831" box="[1056,1126,1915,1941]" italics="true" pageId="12" pageNumber="513">davidi</emphasis>
</taxonomicName>
from southern Tuva to the subspecies
<emphasis id="B96C7B7DA46AFFC8FECAF816FE5BF81D" box="[277,428,1951,1977]" italics="true" pageId="12" pageNumber="513">namkhaidorji</emphasis>
based on some common characters such as poorly developed orange pattern and relatively small size. In the description of the latter taxon, the author united the diagnostic characters of both
<emphasis id="B96C7B7DA46AFFC8FADAF84AFA6BF879" box="[1285,1436,1987,2013]" italics="true" pageId="12" pageNumber="513">namkhaidorji</emphasis>
from the
<typeStatus id="54A319CDA46BFFC9FF22FF1EFEDAFF15" box="[253,301,151,177]" pageId="13" pageNumber="514">type</typeStatus>
locality in
<collectingCountry id="F30FE7FFA46BFFC9FE75FF1EFDE0FF15" box="[426,535,151,177]" name="Mongolia" pageId="13" pageNumber="514">Mongolia</collectingCountry>
and the so-called “northern race” of
<emphasis id="B96C7B7DA46BFFC9FC6DFF1EFBBEFF15" box="[946,1097,151,177]" italics="true" pageId="13" pageNumber="514">namkhaidorji</emphasis>
from southern Tuva. It caused an erroneous treatment of the taxon
<emphasis id="B96C7B7DA46BFFC9FDFBFF32FD4CFF71" box="[548,699,187,213]" italics="true" pageId="13" pageNumber="514">namkhaidorji</emphasis>
as a widely distributed subspecies of
<emphasis id="B96C7B7DA46BFFC9FB85FF32FB9AFF70" box="[1114,1133,187,212]" italics="true" pageId="13" pageNumber="514">N</emphasis>
.
<taxonomicName id="4C18DCECA46BFFC9FBA6FF32FB07FF71" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[1145,1264,187,213]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46BFFC9FB5DFF32FB64FF70" box="[1154,1171,187,212]" italics="true" pageId="13" pageNumber="514">R</emphasis>
.)
<emphasis id="B96C7B7DA46BFFC9FB75FF32FB07FF71" box="[1194,1264,187,213]" italics="true" pageId="13" pageNumber="514">davidi</emphasis>
</taxonomicName>
(with the range extending from southwestern
<collectingCountry id="F30FE7FFA46BFFC9FE3CFF56FDA7FF5D" box="[483,592,223,249]" name="Mongolia" pageId="13" pageNumber="514">Mongolia</collectingCountry>
to
<collectingRegion id="49DC698DA46BFFC9FDACFF56FD5BFF5D" box="[627,684,223,249]" country="Russia" name="Tyva" pageId="13" pageNumber="514">Tuva</collectingRegion>
in
<collectingCountry id="F30FE7FFA46BFFC9FD11FF56FCEDFF5E" box="[718,794,223,250]" name="Russia" pageId="13" pageNumber="514">Russia</collectingCountry>
) by subsequent authors (
<bibRefCitation id="EF89DA9EA46BFFC9FBF1FF56FABDFF5D" author="Weidenhoffer, Z. &amp; Bozano, G. C. &amp; Zhdanko, A. &amp; Churkin, S." box="[1070,1354,223,249]" pageId="13" pageNumber="514" refId="ref14142" refString="Weidenhoffer, Z., Bozano, G. C., Zhdanko, A. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae. Part II. Omnes Artes, Milano, 94 pp." type="book" year="2004">
Weidenhoffer
<emphasis id="B96C7B7DA46BFFC9FB0FFF56FAF0FF5D" box="[1232,1287,223,249]" italics="true" pageId="13" pageNumber="514">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="EF89DA9EA46BFFC9FA8BFF56FA66FF5E" author="Weidenhoffer, Z. &amp; Bozano, G. C. &amp; Zhdanko, A. &amp; Churkin, S." box="[1364,1425,223,250]" pageId="13" pageNumber="514" refId="ref14188" refString="Weidenhoffer, Z., Bozano, G. C., Zhdanko, A. &amp; Churkin, S. (2016) Guide to the butterflies of the Palearctic Region. Lycaenidae. Part II. 2 nd Edition. Omnes Artes, Milano, 106 pp." type="book" year="2016">2016</bibRefCitation>
). It is important to note that specimens from
<collectingRegion id="49DC698DA46BFFC9FDA9FE8AFD58FEB9" box="[630,687,259,285]" country="Russia" name="Tyva" pageId="13" pageNumber="514">Tuva</collectingRegion>
were not included in the
<typeStatus id="54A319CDA46BFFC9FC18FE8AFC00FEB9" box="[967,1015,259,285]" pageId="13" pageNumber="514">type</typeStatus>
series of
<emphasis id="B96C7B7DA46BFFC9FBBDFE8AFB0EFEB9" box="[1122,1273,259,285]" italics="true" pageId="13" pageNumber="514">namkhaidorji</emphasis>
from environs of Biger in southwestern
<collectingCountry id="F30FE7FFA46BFFC9FE6CFEAEFDD3FEE5" box="[435,548,295,321]" name="Mongolia" pageId="13" pageNumber="514">Mongolia</collectingCountry>
. The “northern race” was not described as a separate subspecies due to the lack of distinct external characters compared with
<emphasis id="B96C7B7DA46BFFC9FD45FEC2FCC6FEC1" box="[666,817,331,357]" italics="true" pageId="13" pageNumber="514">namkhaidorji</emphasis>
and the absence of comparative material from western
<collectingCountry id="F30FE7FFA46BFFC9FF48FEE6FEF3FE2D" box="[151,260,367,393]" name="Mongolia" pageId="13" pageNumber="514">Mongolia</collectingCountry>
(
<bibRefCitation id="EF89DA9EA46BFFC9FECDFEE6FE1CFE2D" author="Churkin, S." box="[274,491,366,393]" pageId="13" pageNumber="514" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin 2004: 185</bibRefCitation>
). The black anal abdominal tuft of the female and the genitalic characters mentioned above clarified the position of this isolated population and led to consider it a subspecies of
<emphasis id="B96C7B7DA46BFFC9FB42FE1AFB47FE08" box="[1181,1200,403,428]" italics="true" pageId="13" pageNumber="514">N</emphasis>
.
<taxonomicName id="4C18DCECA46BFFC9FB62FE1AFACEFE09" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[1213,1337,403,429]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46BFFC9FB18FE1AFB2FFE08" box="[1223,1240,403,428]" italics="true" pageId="13" pageNumber="514">R</emphasis>
.)
<emphasis id="B96C7B7DA46BFFC9FB30FE1AFACEFE09" box="[1263,1337,403,429]" italics="true" pageId="13" pageNumber="514">davidi</emphasis>
</taxonomicName>
<emphasis id="B96C7B7DA46BFFC9FAE6FE1AFACAFE09" box="[1337,1341,403,429]" italics="true" pageId="13" pageNumber="514">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46BFFC9FF18FE3EFA7EFE76" box="[199,1417,439,466]" pageId="13" pageNumber="514" type="etymology">
<paragraph id="8BA7A76FA46BFFC9FF18FE3EFA7EFE76" blockId="13.[151,1437,151,645]" box="[199,1417,439,466]" pageId="13" pageNumber="514">
<emphasis id="B96C7B7DA46BFFC9FF18FE3EFEB9FE75" bold="true" box="[199,334,439,465]" pageId="13" pageNumber="514">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54A319CDA46BFFC9FD3CFE3EFCE4FE75" box="[739,787,439,465]" pageId="13" pageNumber="514">type</typeStatus>
locality, Tannu-Ola Mountains (southern
<collectingRegion id="49DC698DA46BFFC9FB38FE3EFAD5FE75" box="[1255,1314,439,465]" country="Russia" name="Tyva" pageId="13" pageNumber="514">Tuva</collectingRegion>
,
<collectingCountry id="F30FE7FFA46BFFC9FAF1FE3EFA89FE76" box="[1326,1406,439,466]" name="Russia" pageId="13" pageNumber="514">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C302F4E4A46BFFC9FF18FE53FB20FD21" pageId="13" pageNumber="514" type="distribution">
<paragraph id="8BA7A76FA46BFFC9FF18FE53FB20FD21" blockId="13.[151,1437,151,645]" pageId="13" pageNumber="514">
<emphasis id="B96C7B7DA46BFFC9FF18FE53FE1AFE50" bold="true" box="[199,493,474,501]" pageId="13" pageNumber="514">Distribution and biology.</emphasis>
<emphasis id="B96C7B7DA46BFFC9FE2DFE52FDF2FE50" box="[498,517,475,500]" italics="true" pageId="13" pageNumber="514">N</emphasis>
.
<taxonomicName id="4C18DCECA46BFFC9FDCEFE52FD70FE51" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[529,647,475,501]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46BFFC9FDC5FE52FDDCFE50" box="[538,555,475,500]" italics="true" pageId="13" pageNumber="514">R</emphasis>
.)
<emphasis id="B96C7B7DA46BFFC9FD9EFE52FD70FE51" box="[577,647,475,501]" italics="true" pageId="13" pageNumber="514">davidi</emphasis>
</taxonomicName>
<emphasis id="B96C7B7DA46BFFC9FD54FE52FD18FE51" box="[651,751,475,501]" italics="true" pageId="13" pageNumber="514">tannuola</emphasis>
<emphasis id="B96C7B7DA46BFFC9FD2BFE52FCCBFE51" bold="true" box="[756,828,475,501]" pageId="13" pageNumber="514">ssp. n.</emphasis>
inhabits the Tannu-Ola Mountains in
<collectingRegion id="49DC698DA46BFFC9FB06FE52FAE3FE51" box="[1241,1300,475,501]" country="Russia" name="Tyva" pageId="13" pageNumber="514">Tuva</collectingRegion>
,
<collectingCountry id="F30FE7FFA46BFFC9FAC2FE52FA9DFE51" box="[1309,1386,475,501]" name="Russia" pageId="13" pageNumber="514">Russia</collectingCountry>
, and probably the Khan-Khokhi Mountains in the southwestern Khangai mountain system in
<collectingCountry id="F30FE7FFA46BFFC9FBB2FE76FB2DFDBD" box="[1133,1242,511,537]" name="Mongolia" pageId="13" pageNumber="514">Mongolia</collectingCountry>
(
<figureCitation id="1323BBEAA46BFFC9FB35FE76FACAFDBD" box="[1258,1341,511,537]" captionStart="FIGURE 42" captionStartId="13.[151,250,1567,1591]" captionTargetBox="[151,1436,669,1544]" captionTargetId="figure-275@13.[151,1436,669,1544]" captionTargetPageId="13" captionText="FIGURE 42. Distribution of the Neolycaena (Rhymnaria) davidi (Oberthür, 1881) species group based on the specimens examined and data from literature (Balint 1989; Churkin 2004; Kosterin 2009; Yakovlev 2012; Weidenhoffer et al. 2004, 2016). Squares refer to the type localities." figureDoi="http://doi.org/10.5281/zenodo.4745198" httpUri="https://zenodo.org/record/4745198/files/figure.png" pageId="13" pageNumber="514">Fig. 42</figureCitation>
); further distribution needs confirmation. Populations of the new subspecies are among the westernmost known populations of
<emphasis id="B96C7B7DA46BFFC9FF69FDCEFF3EFDC4" box="[182,201,583,608]" italics="true" pageId="13" pageNumber="514">N</emphasis>
.
<taxonomicName id="4C18DCECA46BFFC9FF09FDCEFEB9FDC5" baseAuthorityName="Oberthur" baseAuthorityYear="1881" box="[214,334,583,609]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
(
<emphasis id="B96C7B7DA46BFFC9FF3FFDCEFF06FDC4" box="[224,241,583,608]" italics="true" pageId="13" pageNumber="514">R</emphasis>
.)
<emphasis id="B96C7B7DA46BFFC9FED7FDCEFEB9FDC5" box="[264,334,583,609]" italics="true" pageId="13" pageNumber="514">davidi</emphasis>
</taxonomicName>
. The host plant reported in the literature for the populations of Tuva is
<taxonomicName id="4C18DCECA46BFFC9FBB4FDCEFF2BFD21" authority="(Churkin 2004)" baseAuthorityName="Churkin" baseAuthorityYear="2004" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="514" phylum="Tracheophyta" rank="species" species="bungei">
<emphasis id="B96C7B7DA46BFFC9FBB4FDCEFAC7FDC5" box="[1131,1328,583,609]" italics="true" pageId="13" pageNumber="514">Caragana bungei</emphasis>
(
<bibRefCitation id="EF89DA9EA46BFFC9FAE0FDCEFF23FD21" author="Churkin, S." pageId="13" pageNumber="514" pagination="153 - 218" refId="ref13308" refString="Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153 - 218." type="journal article" year="2004">Churkin 2004</bibRefCitation>
)
</taxonomicName>
. According to the data of labels, the flight period is from the end of June to the end of July.
</paragraph>
</subSubSection>
</treatment>
</document>