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<document id="559CB4D3169038DE8EC27DBD510F4E2A" ID-DOI="10.11646/phytotaxa.311.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13700563" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725574901492" checkinUser="felipe" docAuthor="Ranjbar, Massoud &amp; Ranjbar, Zahra" docDate="2017" docId="039787EDFF939C4DDEBFD155FBFCA542" docLanguage="en" docName="phytotaxa.311.2.1.pdf" docOrigin="Phytotaxa 311 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.311.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Cephalaria torbatejamensis Ranjbar &amp; Z. Ranjbar 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="134" masterDocId="FFAEFF95FF8C9C6CDE37D27DFF83A467" masterDocTitle="A taxonomic revision of Cephalaria subgen. Cephalaria sect. Atrocephalae (Caprifoliaceae) from Iran" masterLastPageNumber="138" masterPageNumber="101" pageNumber="132" updateTime="1725584670209" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="6F4D29A51DC973216E55C2A921BD6650" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A444192B479D1884BABCF45994A7D83D">A taxonomic revision of Cephalaria subgen. Cephalaria sect. Atrocephalae (Caprifoliaceae) from Iran</mods:title>
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<mods:name id="84F3D2B281764BC64D1E0E16A01C8569" type="personal">
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<mods:namePart id="23DDA6AD478C5DAE15EFD83E1958F4B5">Ranjbar, Massoud</mods:namePart>
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<mods:namePart id="6DB5B1CA25A40D4D00092A885E9B50C3">Ranjbar, Zahra</mods:namePart>
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<mods:typeOfResource id="D3A56A01F08A1B1584D837BF3B45E2CD">text</mods:typeOfResource>
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<mods:title id="C3F62E49468C2ECBBA368A27BD03347D">Phytotaxa</mods:title>
</mods:titleInfo>
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<mods:date id="7E89DFBFDFEEC63598F732EA1C888CD5">2017</mods:date>
<mods:detail id="EFC74C3585436D9E57BCBF01B093FE13" type="pubDate">
<mods:number id="D2356B6012B3FC744E020420F8C23528">2017-06-27</mods:number>
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<mods:classification id="D2E05037270581910FD5E4006F66DA40">journal article</mods:classification>
<mods:identifier id="A6DF7E7572D26086C93365FE57352D93" type="DOI">10.11646/phytotaxa.311.2.1</mods:identifier>
<mods:identifier id="25CA48E4003DF454C87A88B59EF34393" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="78F7AD56A04270B9F8F6F8659ECCC916" type="Zenodo-Dep">13700563</mods:identifier>
</mods:mods>
<treatment id="039787EDFF939C4DDEBFD155FBFCA542" LSID="urn:lsid:plazi:treatment:039787EDFF939C4DDEBFD155FBFCA542" httpUri="http://treatment.plazi.org/id/039787EDFF939C4DDEBFD155FBFCA542" lastPageId="33" lastPageNumber="134" pageId="31" pageNumber="132">
<subSubSection id="C3246570FF939C73DEBFD155FCD1A726" box="[136,850,806,833]" pageId="31" pageNumber="132" type="nomenclature">
<paragraph id="8B8136FBFF939C73DEBFD155FCD1A726" blockId="31.[136,1452,806,903]" box="[136,850,806,833]" pageId="31" pageNumber="132">
<heading id="D0C98197FF939C73DEBFD155FCD1A726" box="[136,850,806,833]" fontSize="11" level="2" pageId="31" pageNumber="132" reason="4">
<emphasis id="B94AEAE9FF939C73DEBFD155FF28A726" bold="true" box="[136,171,808,833]" pageId="31" pageNumber="132">16.</emphasis>
<taxonomicName id="4C3E4D78FF939C73DE85D15BFD6DA726" ID-CoL="5XJ6Y" authority="Ranjbar &amp; Z. Ranjbar" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[178,750,806,833]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="31" pageNumber="132" phylum="Tracheophyta" rank="species" species="torbatejamensis" status="sp. nov.">
<emphasis id="B94AEAE9FF939C73DE85D15BFE73A726" bold="true" box="[178,496,806,833]" italics="true" pageId="31" pageNumber="132">Cephalaria torbatejamensis</emphasis>
Ranjbar &amp; Z. Ranjbar
</taxonomicName>
,
<emphasis id="B94AEAE9FF939C73DCCFD155FCD1A726" bold="true" box="[760,850,808,833]" pageId="31" pageNumber="132">
<taxonomicNameLabel id="A2795792FF939C73DCCFD155FCD1A726" box="[760,850,808,833]" pageId="31" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3246570FF939C73DEBFD133FC72AC4A" pageId="31" pageNumber="132" type="materials_examined">
<paragraph id="8B8136FBFF939C73DEBFD133FDEDA7E0" blockId="31.[136,1452,806,903]" pageId="31" pageNumber="132">
Type:—
<collectingCountry id="F329766BFF939C73DEE0D133FE97A704" box="[215,276,846,867]" name="Iran" pageId="31" pageNumber="132">IRAN</collectingCountry>
. Prov.
<collectingRegion id="49FAF819FF939C73DF62D133FD93A704" box="[341,528,846,867]" country="Iran" name="Khorasan-e Razavi" pageId="31" pageNumber="132">Khorasan-e Razavi</collectingRegion>
: Bardu forest, Nasr Abad, near Torbat-e Jam,
<geoCoordinate id="EE0A503CFF939C73DDFED133FBA6A704" box="[969,1061,846,867]" degrees="35" direction="north" minutes="25" orientation="latitude" pageId="31" pageNumber="132" precision="925" value="35.416668">
35º 25
<emphasis id="B94AEAE9FF939C73DA30D133FB8FA704" bold="true" box="[1031,1036,846,867]" italics="true" pageId="31" pageNumber="132">ʹ</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE0A503CFF939C73DA07D133FB09A704" box="[1072,1162,846,867]" degrees="60" direction="east" minutes="18" orientation="longitude" pageId="31" pageNumber="132" precision="925" value="60.3">
60º 18
<emphasis id="B94AEAE9FF939C73DA5AD133FBF1A704" bold="true" box="[1133,1138,846,867]" italics="true" pageId="31" pageNumber="132">ʹ</emphasis>
E
</geoCoordinate>
,
<quantity id="4CC69B1EFF939C73DAA3D133FA98A704" box="[1172,1307,846,867]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="31" pageNumber="132" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500–1600 m</quantity>
,
<date id="FF80103BFF939C73DB12D133FA24A704" box="[1317,1447,846,867]" pageId="31" pageNumber="132" value="2003-05-25">25 May 2003</date>
,
<emphasis id="B94AEAE9FF939C73DE8AD10FFED7A7E0" box="[189,340,882,903]" italics="true" pageId="31" pageNumber="132">Joharchi 34808</emphasis>
(
<collectionCode id="ED2FAE3EFF939C73DF56D10FFE2CA7E0" box="[353,431,882,903]" country="Iran" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="31" pageNumber="132" type="Herbarium">FUMH</collectionCode>
!,
<typeStatus id="54858859FF939C73DF8CD10FFD90A7E0" box="[443,531,882,903]" pageId="31" pageNumber="132" type="holotype">holotype</typeStatus>
).
<figureCitation id="13052A7EFF939C73DC14D10FFDEAA7E0" box="[547,617,882,903]" captionStart="FIGURE 18" captionStartId="32.[136,229,1933,1954]" captionTargetBox="[160,1435,198,1868]" captionTargetId="figure-16@32.[151,1435,190,1908]" captionTargetPageId="32" captionText="FIGURE 18. A. Holotype of Cephalaria torbatejamensis Ranjbar &amp; Z. Ranjbar (Joharchi 34808, FUMH!). B. Capitula. C. Middle leaf. D. Corolla. E. Calyx. F. Involucel. G. Involucral bracts. H. Receptacular bracts." figureDoi="http://doi.org/10.5281/zenodo.13700603" httpUri="https://zenodo.org/record/13700603/files/figure.png" pageId="31" pageNumber="132">Fig. 18</figureCitation>
.
</paragraph>
<paragraph id="8B8136FBFF939C73DEBFD1C5FAE7A36A" blockId="31.[136,1452,952,2093]" pageId="31" pageNumber="132">
Perennial plants, slender, erect. Stems up to
<quantity id="4CC69B1EFF939C73DCB4D1C5FD49A7B6" box="[643,714,952,977]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="132" unit="cm" value="60.0">60 cm</quantity>
tall, many, branched, consisting of 2–3 racemes, rather glabrous or pubescent above. Leaves slightly coriaceous, opposite, blades symmetrical, leaves green when dried, pilose on abaxial surface or slightly hairy only along the midvein and margins. Lower cauline leaves simple or lyrate-pinnatisect, oblanceolate to oblong in outline, 4.5–11 ×
<quantity id="4CC69B1EFF939C73DC50D659FD52A05A" box="[615,721,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="31" pageNumber="132" unit="cm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 cm</quantity>
, with 4 lateral segments, lateral segments linear-oblong, 0.8–2.2 ×
<quantity id="4CC69B1EFF939C73DEBFD635FE85A006" box="[136,262,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="31" pageNumber="132" unit="cm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 cm</quantity>
, entire, acute at apex, terminal segment larger than lateral ones, oblanceolate to spathulate, acute to subacute at apex, serrate, 9–13 teeth (apical and basal sides straight), angle of teeth with reference to the midrib 80º–90º, teeth
<quantity id="4CC69B1EFF939C73DEBFD6EDFF79A0CE" box="[136,250,1168,1193]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="31" pageNumber="132" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
long, midvein gently raised adaxially and sharply keeled abaxially, tertiary veins weakly percurrent, widely spaced, oblique. Middle cauline leaves lyrate, oblanceolate to oblong-lanceolate in outline, 6.5–14 ×
<quantity id="4CC69B1EFF939C73DB36D6C9FAEEA0AA" box="[1281,1389,1204,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="31" pageNumber="132" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
, with 4–6 lateral segments, lateral segments linear-lanceolate, 1–2 ×
<quantity id="4CC69B1EFF939C73DD6ED6A5FC44A096" box="[857,967,1240,1265]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="31" pageNumber="132" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 cm</quantity>
, dentate to serrate, acute at apex, terminal segment larger than lateral ones, lanceolate or oblong to lanceolate, 5–9 ×
<quantity id="4CC69B1EFF939C73DDD4D681FBD2A172" box="[995,1105,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="31" pageNumber="132" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
, acute at apex, dentate, 12–16 teeth (apical side straight, basal side concave), angle of teeth ca. 80º–90º, teeth
<quantity id="4CC69B1EFF939C73DA37D75DFBF0A15E" box="[1024,1139,1312,1337]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="31" pageNumber="132" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, midvein gently raised adaxially and sharply keeled abaxially, tertiary veins reticulate, oblique. Upper cauline leaves simple or lyrate, oblong lanceolate to linear in outline, 4.5–5 ×
<quantity id="4CC69B1EFF939C73DC05D715FD32A1E6" box="[562,689,1384,1409]" metricMagnitude="-3" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="31" pageNumber="132" unit="cm" value="0.6" valueMax="0.8" valueMin="0.4">0.4–0.8 cm</quantity>
, with 2 lateral segments, lateral segments linear, 0.4–1 ×
<quantity id="4CC69B1EFF939C73DB1FD715FA24A1E6" box="[1320,1447,1384,1409]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="31" pageNumber="132" unit="cm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 cm</quantity>
, entire, acute at apex, terminal segment larger than lateral ones, oblanceolate, acute at apex, entire to weakly serrate, 2–6 teeth (apical side straight, basal side concave), angle of teeth with reference to the midrib 30º–45º, teeth
<quantity id="4CC69B1EFF939C73DB0CD7CDFA2FA1AE" box="[1339,1452,1456,1481]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="31" pageNumber="132" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, midvein impressed to flat adaxially and round-keeled abaxially, tertiary veins close together. Peduncle
<quantity id="4CC69B1EFF939C73DB7DD7A9FA28A18A" box="[1354,1451,1492,1517]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="2.3" metricValueMin="0.8" pageId="31" pageNumber="132" unit="cm" value="15.5" valueMax="23.0" valueMin="8.0">8–23 cm</quantity>
long, glabrous. Capitula globose or ovoid,
<quantity id="4CC69B1EFF939C73DC5FD785FD53A276" box="[616,720,1528,1553]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="31" pageNumber="132" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 cm</quantity>
in diameter in flower, subradiant. Involucral bracts in one series, ovate, ca. 5 ×
<quantity id="4CC69B1EFF939C73DF15D461FEE7A252" box="[290,356,1564,1589]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="132" unit="mm" value="3.0">3 mm</quantity>
, green–brown and blackish at apex, tomentose hairs up to
<quantity id="4CC69B1EFF939C73DDDDD461FBA9A252" box="[1002,1066,1564,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="132" unit="mm" value="2.0">2 mm</quantity>
long, obtuse at apex. Receptacular bracts in seven series, rhombic to narrowly ovate to lanceolate, 6–10 ×
<quantity id="4CC69B1EFF939C73DD96D43DFB83A23E" box="[929,1024,1600,1625]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="31" pageNumber="132" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, spine
<quantity id="4CC69B1EFF939C73DA7AD43DFB3DA23E" box="[1101,1214,1600,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="31" pageNumber="132" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, straw-coloured dorsally at base, purple blackish at 2/3 of the bract apex, abruptly acute or acuminate at apex, pilose or sericeous, ciliate at margins, inner bracts recurved at apex. Calyx cupuliform, 1–1.5 ×
<quantity id="4CC69B1EFF939C73DD96D4F5FC78A2C6" box="[929,1019,1672,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="132" unit="mm" value="1.5">1.5 mm</quantity>
, dark green, pubescent, with irregular teeth
<quantity id="4CC69B1EFF939C73DEF1D4D1FEBBA2A2" box="[198,312,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="31" pageNumber="132" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long. Corolla
<quantity id="4CC69B1EFF939C73DFEED4D1FDD2A2A2" box="[473,593,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="31" pageNumber="132" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
long, yellow, densely covered with appressed hairs outside, stamens slightly longer than corolla, anthers deeply yellow to light brown. Involucel 2–2.5 ×
<quantity id="4CC69B1EFF939C73DDDFD4ADFBBCA28E" box="[1000,1087,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="132" unit="mm" value="1.5">1.5 mm</quantity>
in flower, 4-angled, hairy (hairs up to
<quantity id="4CC69B1EFF939C73DEFFD489FE9DA36A" box="[200,286,1780,1805]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="132" unit="mm" value="0.2">0.2 mm</quantity>
long), 4 long and 4 short teeth at apex; longer teeth ca.
<quantity id="4CC69B1EFF939C73DDB9D489FC4CA36A" box="[910,975,1780,1805]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="132" unit="mm" value="2.0">2 mm</quantity>
long, shorter teeth ca.
<quantity id="4CC69B1EFF939C73DAF9D489FAA7A36A" box="[1230,1316,1780,1805]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="132" unit="mm" value="0.5">0.5 mm</quantity>
long.
</paragraph>
<paragraph id="8B8136FBFF939C73DE8AD565FD1EA31E" blockId="31.[136,1452,952,2093]" pageId="31" pageNumber="132">
<materialsCitation id="3B563CA6FF939C73DE8AD565FD37A332" collectionCode="FUMH" collectorName="Ayatollahi &amp; Zangooei" country="Iran" county="Razavi province" elevation="1650" location="Allah Akbar" municipality="Dargaz" pageId="31" pageNumber="132" specimenCount="2" stateProvince="Khorasan-e Shomali">
<emphasis id="B94AEAE9FF939C73DE8AD565FD46A356" bold="true" box="[189,709,1816,1841]" pageId="31" pageNumber="132">
<typeStatus id="54858859FF939C73DE8AD565FDB6A356" box="[189,565,1816,1841]" isOtherTypeStatus="true" pageId="31" pageNumber="132">Additional specimens examined</typeStatus>
(
<typeStatus id="54858859FF939C73DC74D565FD3EA356" box="[579,701,1816,1841]" pageId="31" pageNumber="132" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="F329766BFF939C73DCDFD565FC99A356" box="[744,794,1816,1841]" name="Iran" pageId="31" pageNumber="132">Iran</collectingCountry>
:
<collectingRegion id="49FAF819FF939C73DD14D565FC0DA356" box="[803,910,1816,1841]" country="Iran" name="Khorasan-e Shomali" pageId="31" pageNumber="132">Khorasan</collectingRegion>
<collectingCounty id="62E04E77FF939C73DDA3D565FBCEA356" box="[916,1101,1816,1841]" pageId="31" pageNumber="132">Razavi province</collectingCounty>
,
<collectingMunicipality id="6BE5AC81FF939C73DA60D565FB24A356" box="[1111,1191,1816,1841]" pageId="31" pageNumber="132">Dargaz</collectingMunicipality>
city,
<location id="8EE16020FF939C73DAD6D565FAE9A356" LSID="urn:lsid:plazi:treatment:039787EDFF939C4DDEBFD155FBFCA542:8EE16020FF939C73DAD6D565FAE9A356" box="[1249,1386,1816,1841]" country="Iran" county="Razavi province" municipality="Dargaz" name="Allah Akbar" pageId="31" pageNumber="132" stateProvince="Khorasan-e Shomali">Allah Akbar</location>
neck,
<quantity id="4CC69B1EFF939C73DEBFD541FF63A332" box="[136,224,1852,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="31" pageNumber="132" unit="m" value="1650.0">
<elevation id="0013D1C8FF939C73DEBFD541FF63A332" box="[136,224,1852,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="31" pageNumber="132" unit="m" value="1650.0">1650 m</elevation>
</quantity>
,
<emphasis id="B94AEAE9FF939C73DEDBD541FDBFA332" box="[236,572,1852,1877]" italics="true" pageId="31" pageNumber="132">
<collectorName id="26CB532DFF939C73DEDBD541FEDCA332" box="[236,351,1852,1877]" pageId="31" pageNumber="132">Ayatollahi</collectorName>
&amp;
<collectorName id="26CB532DFF939C73DFB2D541FE6CA332" box="[389,495,1852,1877]" pageId="31" pageNumber="132">Zangooei</collectorName>
15512
</emphasis>
(
<collectionCode id="ED2FAE3EFF939C73DC7BD541FD2BA332" box="[588,680,1852,1877]" country="Iran" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="31" pageNumber="132" type="Herbarium">FUMH</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B563CA6FF939C73DCF7D541FD1AA31E" collectionCode="FUMH" collectorName="S Mashhad &amp; Joharchi" country="Iran" county="Razavi province" elevation="1900" location="Moghan cave" municipality="Mashhad" pageId="31" pageNumber="132" specimenCount="1" stateProvince="Khorasan-e Shomali">
<collectingCountry id="F329766BFF939C73DCF7D541FD72A332" box="[704,753,1852,1877]" name="Iran" pageId="31" pageNumber="132">Iran</collectingCountry>
:
<collectingRegion id="49FAF819FF939C73DCCAD541FCEBA332" box="[765,872,1852,1877]" country="Iran" name="Khorasan-e Shomali" pageId="31" pageNumber="132">Khorasan</collectingRegion>
<collectingCounty id="62E04E77FF939C73DD47D541FBA9A332" box="[880,1066,1852,1877]" pageId="31" pageNumber="132">Razavi province</collectingCounty>
,
<collectorName id="26CB532DFF939C73DA01D541FB34A332" box="[1078,1207,1852,1877]" pageId="31" pageNumber="132">S Mashhad</collectorName>
,
<collectingMunicipality id="6BE5AC81FF939C73DAF5D541FAAAA332" box="[1218,1321,1852,1877]" pageId="31" pageNumber="132">Mashhad</collectingMunicipality>
to
<location id="8EE16020FF939C73DB79D541FF3DA31E" LSID="urn:lsid:plazi:treatment:039787EDFF939C4DDEBFD155FBFCA542:8EE16020FF939C73DB79D541FF3DA31E" country="Iran" county="Razavi province" municipality="Mashhad" name="Moghan cave" pageId="31" pageNumber="132" stateProvince="Khorasan-e Shomali">Moghan cave</location>
,
<quantity id="4CC69B1EFF939C73DEFED51DFEE5A31E" box="[201,358,1888,1913]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="31" pageNumber="132" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<elevation id="0013D1C8FF939C73DEFED51DFEE5A31E" box="[201,358,1888,1913]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="31" pageNumber="132" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</elevation>
</quantity>
,
<emphasis id="B94AEAE9FF939C73DF45D51DFDA1A31E" box="[370,546,1888,1913]" italics="true" pageId="31" pageNumber="132">
<collectorName id="26CB532DFF939C73DF45D51DFE56A31E" box="[370,469,1888,1913]" pageId="31" pageNumber="132">Joharchi</collectorName>
43054
</emphasis>
(
<collectionCode id="ED2FAE3EFF939C73DC06D51DFD0EA31E" box="[561,653,1888,1913]" country="Iran" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="31" pageNumber="132" type="Herbarium">FUMH</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8136FBFF939C73DE8AD5F9FC72AC4A" blockId="31.[136,1452,952,2093]" pageId="31" pageNumber="132">
<emphasis id="B94AEAE9FF939C73DE8AD5F9FE0CA3FA" bold="true" box="[189,399,1924,1949]" pageId="31" pageNumber="132">Taxonomic notes:</emphasis>
—
<taxonomicName id="4C3E4D78FF939C73DF9CD5F9FD67A3FA" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[427,740,1924,1949]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="31" pageNumber="132" phylum="Tracheophyta" rank="species" species="torbatejamensis">
<emphasis id="B94AEAE9FF939C73DF9CD5F9FD67A3FA" box="[427,740,1924,1949]" italics="true" pageId="31" pageNumber="132">Cephalaria torbatejamensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C3E4D78FF939C73DD41D5F9FBACA3FA" box="[886,1071,1924,1949]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="31" pageNumber="132" phylum="Tracheophyta" rank="species" species="microcephala">
<emphasis id="B94AEAE9FF939C73DD41D5F9FBACA3FA" box="[886,1071,1924,1949]" italics="true" pageId="31" pageNumber="132">C. microcephala</emphasis>
</taxonomicName>
in dissection of lower and middle leaves, involucral bracts shape, colour of flower and appendages, and also size and shape of involucel, but differs in some important characters (see
<tableCitation id="C6BC0340FF939C73DC21D5B1FDECA382" box="[534,623,1996,2021]" captionStart="TABLE 4" captionStartId="33.[136,224,334,357]" captionText="TABLE 4. Morphological comparison of Cephalaria axillaris, C. golestanica, C. integerrima, C. microcephala, C. subindivisa, C. sublanata and C. torbatejamensis." pageId="31" pageNumber="132">Table 4</tableCitation>
). Also,
<taxonomicName id="4C3E4D78FF939C73DCFCD5B1FC21A382" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[715,930,1996,2021]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="31" pageNumber="132" phylum="Tracheophyta" rank="species" species="torbatejamensis">
<emphasis id="B94AEAE9FF939C73DCFCD5B1FD5DA382" box="[715,734,1996,2021]" italics="true" pageId="31" pageNumber="132">C</emphasis>
.
<emphasis id="B94AEAE9FF939C73DCD8D5B1FC21A382" box="[751,930,1996,2021]" italics="true" pageId="31" pageNumber="132">torbatejamensis</emphasis>
</taxonomicName>
differs from other related species of
<emphasis id="B94AEAE9FF939C73DB62D5B1FAECA382" box="[1365,1391,1996,2021]" italics="true" pageId="31" pageNumber="132">C.</emphasis>
sect.
<emphasis id="B94AEAE9FF939C73DEBFD58DFE9DAC6E" box="[136,286,2032,2057]" italics="true" pageId="31" pageNumber="132">Atrocephalae</emphasis>
by having oblanceolate to oblong-lanceolate lower and middle cauline leaves, number and length of teeth median leaves, and involucral and receptacular bracts purple blackish at apex.
</paragraph>
</subSubSection>
<caption id="DF416673FFAC9C4CDEBFD5F0FC0AA3A1" ID-DOI="http://doi.org/10.5281/zenodo.13700603" ID-Zenodo-Dep="13700603" httpUri="https://zenodo.org/record/13700603/files/figure.png" pageId="32" pageNumber="133" startId="32.[136,229,1933,1954]" targetBox="[160,1435,198,1868]" targetPageId="32" targetType="figure">
<paragraph id="8B8136FBFFAC9C4CDEBFD5F0FC0AA3A1" blockId="32.[136,1452,1933,1990]" pageId="32" pageNumber="133">
<emphasis id="B94AEAE9FFAC9C4CDEBFD5F0FEA4A3C5" bold="true" box="[136,295,1933,1954]" pageId="32" pageNumber="133">FIGURE 18. A.</emphasis>
Holotype of
<taxonomicName id="4C3E4D78FFAC9C4CDF90D5F0FC0FA3C5" authority="Ranjbar &amp; Z. Ranjbar" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[423,908,1933,1954]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="32" pageNumber="133" phylum="Tracheophyta" rank="species" species="torbatejamensis">
<emphasis id="B94AEAE9FFAC9C4CDF90D5F0FD37A3C5" box="[423,692,1933,1954]" italics="true" pageId="32" pageNumber="133">Cephalaria torbatejamensis</emphasis>
Ranjbar &amp; Z. Ranjbar
</taxonomicName>
(
<emphasis id="B94AEAE9FFAC9C4CDDADD5F0FBB2A3C5" box="[922,1073,1933,1954]" italics="true" pageId="32" pageNumber="133">Joharchi 34808</emphasis>
, FUMH!).
<emphasis id="B94AEAE9FFAC9C4CDAA8D5F0FB36A3C5" bold="true" box="[1183,1205,1933,1954]" pageId="32" pageNumber="133">B.</emphasis>
Capitula.
<emphasis id="B94AEAE9FFAC9C4CDB2FD5F0FAB3A3C5" bold="true" box="[1304,1328,1933,1954]" pageId="32" pageNumber="133">C.</emphasis>
Middle leaf.
<emphasis id="B94AEAE9FFAC9C4CDEBFD5CCFF23A3A1" bold="true" box="[136,160,1969,1990]" pageId="32" pageNumber="133">D.</emphasis>
Corolla.
<emphasis id="B94AEAE9FFAC9C4CDECED5CCFE93A3A1" bold="true" box="[249,272,1969,1990]" pageId="32" pageNumber="133">E.</emphasis>
Calyx.
<emphasis id="B94AEAE9FFAC9C4CDF6CD5CCFEEEA3A1" bold="true" box="[347,365,1969,1990]" pageId="32" pageNumber="133">F.</emphasis>
Involucel.
<emphasis id="B94AEAE9FFAC9C4CDFEDD5CCFE70A3A1" bold="true" box="[474,499,1969,1990]" pageId="32" pageNumber="133">G.</emphasis>
Involucral bracts.
<emphasis id="B94AEAE9FFAC9C4CDC91D5CCFD3CA3A1" bold="true" box="[678,703,1969,1990]" pageId="32" pageNumber="133">H.</emphasis>
Receptacular bracts.
</paragraph>
</caption>
<subSubSection id="C3246570FFAD9C4DDE8AD2DDFBFCA542" pageId="33" pageNumber="134" type="distribution">
<paragraph id="8B8136FBFFAD9C4DDE8AD2DDFBFCA542" blockId="33.[136,1451,160,293]" pageId="33" pageNumber="134">
<emphasis id="B94AEAE9FFAD9C4DDE8AD2DDFE6EA4DE" bold="true" box="[189,493,160,185]" pageId="33" pageNumber="134">Distribution and habitat:</emphasis>
—
<taxonomicName id="4C3E4D78FFAD9C4DDC3ED2DDFCC7A4DE" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[521,836,160,185]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="33" pageNumber="134" phylum="Tracheophyta" rank="species" species="torbatejamensis">
<emphasis id="B94AEAE9FFAD9C4DDC3ED2DDFCC7A4DE" box="[521,836,160,185]" italics="true" pageId="33" pageNumber="134">Cephalaria torbatejamensis</emphasis>
</taxonomicName>
is an Irano-Turanian element and rare endemic to NE
<collectingCountry id="F329766BFFAD9C4DDEBFD2B9FF3AA4BA" box="[136,185,196,221]" name="Iran" pageId="33" pageNumber="134">Iran</collectingCountry>
. It is known only from the mountain meadows, mountain steppe with grasses, thorn-cushions and corn field weed in the mountain zone around Bardu forest, Nasr Abad District, near Torbat-e Jam,
<collectingRegion id="49FAF819FFAD9C4DDA29D295FB71A566" box="[1054,1266,232,257]" country="Iran" name="Khorasan-e Shomali" pageId="33" pageNumber="134">Khorasan province</collectingRegion>
NE
<collectingCountry id="F329766BFFAD9C4DDB12D295FAD5A566" box="[1317,1366,232,257]" name="Iran" pageId="33" pageNumber="134">Iran</collectingCountry>
. This is a special region surrounded by Bardu Mts. with dry climate at elevations of
<quantity id="4CC69B1EFFAD9C4DDDEAD371FBF8A542" box="[989,1147,268,293]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="33" pageNumber="134" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500–1600 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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