1117 lines
134 KiB
XML
1117 lines
134 KiB
XML
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<mods:title id="6467954FBFD4BB3B03D0F0CF6D4B00C2">Synonymic notes in the Southeast Asian genus Lutera Westwood, 1875 (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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<mods:namePart id="7C74A0D33665B0896913502C00E03740">Xu, Hao</mods:namePart>
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<mods:affiliation id="37FB12A7529621A255516B5A483C6D08">Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, 166 Mianxing West Road, Mianyang, Sichuan 621000, P. R. China. & Ecological Security and Protection Key Laboratory of Sichuan Province, Mianyang Normal University, 166 Mianxing West Road, Mianyang, Sichuan 621000, P. R. China.</mods:affiliation>
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<mods:namePart id="29CB7325E7E46D07B3797E6598B69972">Zhao, Ming-Zhi</mods:namePart>
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<mods:affiliation id="2EFFA1ACEEC2D60C5B828E264DD76F8A">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China.</mods:affiliation>
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<taxonomicName id="4C3DFACCC742A30BFF4DF986FDFC3336" ID-CoL="3WG3W" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[151,580,1626,1653]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="nigromaculata">
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<emphasis id="B9495D5DC742A30BFF4DF986FE1A3337" bold="true" box="[151,418,1626,1653]" italics="true" pageId="1" pageNumber="532">Lutera nigromaculata</emphasis>
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<bibRefCitation id="EFACFCBEC742A30BFE73F987FDFC3336" author="Ohaus, F." box="[425,580,1626,1652]" pageId="1" pageNumber="532" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B82814FC742A30BFF4DF9A2FE7033DB" blockId="1.[151,580,1626,1689]" box="[151,456,1663,1689]" pageId="1" pageNumber="532">
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(
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<figureCitation id="13069DCAC742A30BFF45F9A2FEA133DB" box="[159,281,1663,1689]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–16" captionStart-2="FIGURES 17–24" captionStartId-0="2.[151,264,1563,1587]" captionStartId-1="4.[151,264,1903,1927]" captionStartId-2="5.[151,264,1568,1592]" captionTargetBox-0="[222,1365,326,1522]" captionTargetBox-1="[264,1323,329,1857]" captionTargetBox-2="[232,1377,193,1522]" captionTargetId-0="figure-96@2.[213,1375,319,1537]" captionTargetId-1="figure-95@4.[235,1352,309,1880]" captionTargetId-2="figure-18@5.[202,1384,181,1544]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1–6. Habitus of Lutera nigromaculata Ohaus, 1900 from Vietnam. 1, 4, dorsal view; 2, 5, lateral oblique view; 3, 6, ventral view." captionText-1="FIGURES 7–16. Type specimens of Parastasia (Lutera) nigromaculata guttata Frey, 1970. 7–10, lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM); 11–13, paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB); 14–16, paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM)." captionText-2="FIGURES 17–24. Type specimens of Lutera spp. 17–18, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 19–20, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 21–24, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi-0="http://doi.org/10.5281/zenodo.13949860" figureDoi-1="http://doi.org/10.5281/zenodo.13949863" figureDoi-2="http://doi.org/10.5281/zenodo.13949866" httpUri-0="https://zenodo.org/record/13949860/files/figure.png" httpUri-1="https://zenodo.org/record/13949863/files/figure.png" httpUri-2="https://zenodo.org/record/13949866/files/figure.png" pageId="1" pageNumber="532">Figs. 1–18</figureCitation>
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,
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<figureCitation id="13069DCAC742A30BFEFFF9A2FE2F33DB" box="[293,407,1663,1689]" captionStart="FIGURES 31–45" captionStartId="8.[151,264,1924,1948]" captionTargetBox="[189,1398,275,1888]" captionTargetId="figure-77@8.[176,1411,260,1900]" captionTargetPageId="8" captionText="FIGURES 31–45. Male genitalia of Lutera species (31–35, dorsal view; 36–40, lateral view; 41–45, ventral view;). 31, 36, 41, L. nigromaculata Ohaus, 1900 from Thailand; 32–35, 37–40, 42–35, L. nigrita Ohaus, 1930: 32, 37, 42, from Luzon Island; 33, 38, 43, from Mindoro Island; 34, 39, 44, from Negros Island; 35, 40, 45, from Mindanao Island" figureDoi="http://doi.org/10.5281/zenodo.13949870" httpUri="https://zenodo.org/record/13949870/files/figure.png" pageId="1" pageNumber="532">31, 36, 41</figureCitation>
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,
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<figureCitation id="13069DCAC742A30BFE79F9A2FE7833DB" box="[419,448,1663,1689]" captionStart="FIGURE 46" captionStartId="9.[151,250,1970,1994]" captionTargetBox="[164,1423,1173,1947]" captionTargetId="figure-488@9.[164,1423,1173,1947]" captionTargetPageId="9" captionText="FIGURE 46. Known distribution of Lutera nigromaculata Ohaus, 1900 (black dots) and L. nigrita Ohaus, 1930 (black triangles). Type localities are underlined, and the open dot in Nepal is a record from Wada (1998) without precise location." figureDoi="http://doi.org/10.5281/zenodo.13949872" httpUri="https://zenodo.org/record/13949872/files/figure.png" pageId="1" pageNumber="532">46</figureCitation>
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)
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</paragraph>
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<treatmentCitation id="0A9CA75EC742A30BFF4DF919FD8A3399" author="Ohaus, F." box="[151,562,1731,1755]" page="261" pageId="1" pageNumber="532" year="1900">
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<taxonomicName id="4C3DFACCC742A30BFF4DF919FD8A3399" ID-CoL="3WG3W" authority="Ohaus, 1900: 261" authorityName="Ohaus" authorityPageNumber="261" authorityYear="1900" box="[151,562,1731,1755]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="nigromaculata">
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<emphasis id="B9495D5DC742A30BFF4DF919FECE3399" box="[151,374,1731,1755]" italics="true" pageId="1" pageNumber="532">Lutera nigromaculata</emphasis>
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<bibRefCitation id="EFACFCBEC742A30BFEA7F91EFD8A3399" author="Ohaus, F." box="[381,562,1731,1755]" pageId="1" pageNumber="532" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900: 261</bibRefCitation>
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</taxonomicName>
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(type locality: Montagnes de Lakhon [the northern vicinity of Thakhek,
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<collectingRegion id="49F94FADC742A30BFAC3F91EFE9C33B8" country="Laos" name="Khammouan" pageId="1" pageNumber="532">Khammouan Province</collectingRegion>
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,
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<collectingCountry id="F32AC1DFC742A30BFEF7F93FFEDC33B8" box="[301,356,1762,1786]" name="Laos" pageId="1" pageNumber="532">Laos</collectingCountry>
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]);
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<treatmentCitation id="0A9CA75EC742A30BFEA1F93FFDA033B8" author="Ohaus, F." box="[379,536,1762,1786]" page="39" pageId="1" pageNumber="532" year="1918">
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<bibRefCitation id="EFACFCBEC742A30BFEA1F93FFDA033B8" author="Ohaus, F." box="[379,536,1762,1786]" pageId="1" pageNumber="532" pagination="1 - 241" refId="ref7791" refString="Ohaus, F. (1918) Scarabaeidae: Phaenomerinae, Euchirinae, Rutelinae. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum catalogus. Vo. XX. Pars 66. W. Junk, Berlin, pp. 1 - 241." type="book chapter" year="1918">Ohaus 1918: 39</bibRefCitation>
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(catalogued, partim);
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<treatmentCitation id="0A9CA75EC742A30BFD2CF93FFC2733B8" author="Ohaus, F." box="[758,927,1762,1786]" page="112" pageId="1" pageNumber="532" year="1934">
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<bibRefCitation id="EFACFCBEC742A30BFD2CF93FFC2733B8" author="Ohaus, F." box="[758,927,1762,1786]" pageId="1" pageNumber="532" pagination="1 - 172" refId="ref7920" refString="Ohaus, F. (1934) Coleoptera Lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. Erster Teil. In: Wytsman, P. (Ed.), Genera Insectorum. Coleoptera. Fascicule 199 A. Louis Desmet-Verteneuil, Bruxelles, pp. 1 - 172." type="book chapter" year="1934">Ohaus 1934: 112</bibRefCitation>
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</treatmentCitation>
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(catalogued, partim);
|
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<treatmentCitation id="0A9CA75EC742A30BFBA6F93FFA9D33B8" author="Kuijten, P. J." box="[1148,1317,1762,1786]" page="87" pageId="1" pageNumber="532" year="1988">
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<bibRefCitation id="EFACFCBEC742A30BFBA6F93FFA9D33B8" author="Kuijten, P. J." box="[1148,1317,1762,1786]" pageId="1" pageNumber="532" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten 1988: 87</bibRefCitation>
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</treatmentCitation>
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(discussion on taxonomy);
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<treatmentCitation id="0A9CA75EC742A30BFEB8F8DDFDB0325A" author="Wada, K." box="[354,520,1792,1817]" page="369" pageId="1" pageNumber="532" year="1998">
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<bibRefCitation id="EFACFCBEC742A30BFEB8F8DDFDB0325A" author="Wada, K." box="[354,520,1792,1817]" pageId="1" pageNumber="532" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada 1998: 369</bibRefCitation>
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(in key,
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<typeStatus id="54863FEDC742A30BFDB8F8DDFD03325A" box="[610,699,1792,1816]" pageId="1" pageNumber="532" type="holotype">holotype</typeStatus>
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mentioned), figs. 4 (male habitus), 16–17 (male genitalia), 23 (male left fore claws), 26b (distribution map);
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<treatmentCitation id="0A9CA75EC742A30BFDEBF8C2FD623275" author="Krajcik, M." box="[561,730,1823,1847]" page="79" pageId="1" pageNumber="532" year="2007">
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<bibRefCitation id="EFACFCBEC742A30BFDEBF8C2FD623275" author="Krajcik, M." box="[561,730,1823,1847]" pageId="1" pageNumber="532" pagination="1 - 139" refId="ref7338" refString="Krajcik, M. (2007) Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x Supplementu m, 4, 1 - 139." type="journal article" year="2007">Krajčík 2007: 79</bibRefCitation>
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</treatmentCitation>
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(catalogued);
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<treatmentCitation id="0A9CA75EC742A30BFCB1F8C2FB993275" author="Krajcik, M." box="[875,1057,1823,1847]" page="150" pageId="1" pageNumber="532" year="2012">
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<bibRefCitation id="EFACFCBEC742A30BFCB1F8C2FB993275" author="Krajcik, M." box="[875,1057,1823,1847]" pageId="1" pageNumber="532" pagination="1 - 139" refId="ref7376" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. Animma. x Supplementum, 5, 1 - 139." type="journal article" year="2012">Krajčík 2012: 150</bibRefCitation>
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</treatmentCitation>
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(catalogued);
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<treatmentCitation id="0A9CA75EC742A30BFB6BF8C2FAD13275" author="Li, J. K. & Keith, D. & Gao, M. X. & Lin, L." box="[1201,1385,1823,1847]" page="275" pageId="1" pageNumber="532" year="2012">
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<bibRefCitation id="EFACFCBEC742A30BFB6BF8C2FAD13275" author="Li, J. K. & Keith, D. & Gao, M. X. & Lin, L." box="[1201,1385,1823,1847]" pageId="1" pageNumber="532" pagination="273 - 276" refId="ref7447" refString="Li, J. K., Keith, D., Gao, M. X. & Lin, L. (2012) Coleopteres Scarabaeoidea de Pha Ngam (province de Bokeo, Laos). L'Entomologiste, 68 (5), 273 - 276." type="journal article" year="2012">
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Li
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<emphasis id="B9495D5DC742A30BFB17F8FDFB473275" box="[1229,1279,1823,1847]" italics="true" pageId="1" pageNumber="532">et al.</emphasis>
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2012: 275
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</bibRefCitation>
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(Pha Ngam,
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<collectingRegion id="49F94FADC742A30BFEC8F8E3FE0F3214" box="[274,439,1854,1878]" country="Laos" name="Bokeo" pageId="1" pageNumber="532">Bokeo Province</collectingRegion>
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,
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<collectingCountry id="F32AC1DFC742A30BFE1BF8E3FE4E3214" box="[449,502,1854,1878]" name="Laos" pageId="1" pageNumber="532">Laos</collectingCountry>
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).
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||
<treatmentCitationGroup id="AB2DA661C742A30BFF4DF880FBAA3236" box="[151,1042,1884,1909]" pageId="1" pageNumber="532">
|
||
<taxonomicName id="4C3DFACCC742A30BFF4DF880FE433236" ID-CoL="4DCX2" authority="(Ohaus)" baseAuthorityName="Ohaus" box="[151,507,1884,1909]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC742A30BFF4DF880FE273236" box="[151,415,1884,1908]" italics="true" pageId="1" pageNumber="532">Parastasia nigromaculata</emphasis>
|
||
(Ohaus)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A9CA75EC742A30BFDDFF881FD083237" author="Paulian, R." box="[517,688,1884,1909]" page="84" pageId="1" pageNumber="532" year="1959">
|
||
<bibRefCitation id="EFACFCBEC742A30BFDDFF881FD083237" author="Paulian, R." box="[517,688,1884,1909]" pageId="1" pageNumber="532" pagination="73 - 105" refId="ref7972" refString="Paulian, R. (1959) Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). Annales de la Societe entomologique de France, 127, 73 - 105. https: // doi. org / 10.1080 / 21686351.1958.12279112" type="journal article" year="1959">Paulian 1959: 84</bibRefCitation>
|
||
</treatmentCitation>
|
||
(redescription based on
|
||
<typeStatus id="54863FEDC742A30BFC70F881FBBF3236" box="[938,1031,1884,1908]" pageId="1" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B82814FC742A30BFF4DF8A1FC2D32F0" blockId="1.[151,1437,1731,2031]" pageId="1" pageNumber="532">
|
||
<treatmentCitation id="0A9CA75EC742A30BFF4DF8A1FDA832D1" author="Chujo, M." box="[151,528,1915,1939]" page="33" pageId="1" pageNumber="532" year="1966">
|
||
<taxonomicName id="4C3DFACCC742A30BFF4DF8A1FDA832D1" authority="Chujo, 1966: 33" authorityName="Chujo" authorityPageNumber="33" authorityYear="1966" box="[151,528,1915,1939]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="ohausi">
|
||
<emphasis id="B9495D5DC742A30BFF4DF8A1FEED32D1" box="[151,341,1915,1939]" italics="true" pageId="1" pageNumber="532">Parastasia ohausi</emphasis>
|
||
<bibRefCitation id="EFACFCBEC742A30BFEBBF8A6FDA832D1" author="Chujo, M." box="[353,528,1915,1939]" pageId="1" pageNumber="532" pagination="25 - 34" refId="ref6976" refString="Chujo, M. (1966) Coleoptera from Southeast Asia (V). 5. Family Scarabaeidae (2). Memoirs of the Faculty of Education, Kagawa University, Part II (No. 140), 25 - 34." type="book chapter" year="1966">Chûjô, 1966: 33</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
[replacement name for
|
||
<taxonomicName id="4C3DFACCC742A30BFCC3F8A1FB9432D1" ID-CoL="4DCX2" authority="(Ohaus)" baseAuthorityName="Ohaus" box="[793,1068,1915,1939]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC742A30BFCC3F8A1FC7432D1" box="[793,972,1915,1939]" italics="true" pageId="1" pageNumber="532">P. nigromaculata</emphasis>
|
||
(Ohaus)
|
||
</taxonomicName>
|
||
, preoccupied by
|
||
<taxonomicName id="4C3DFACCC742A30BFB33F8A1FE3A32F0" ID-CoL="4DCX2" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC742A30BFB33F8A1FA2432D1" box="[1257,1436,1915,1939]" italics="true" pageId="1" pageNumber="532">P. nigromaculata</emphasis>
|
||
(Blanchard, 1850)
|
||
</taxonomicName>
|
||
; Chomtong,
|
||
<collectingRegion id="49F94FADC742A30BFDDFF847FDC732F0" box="[517,639,1946,1970]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="532">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32AC1DFC742A30BFD53F847FD5132F0" box="[649,745,1946,1970]" name="Thailand" pageId="1" pageNumber="532">Thailand</collectingCountry>
|
||
].
|
||
<emphasis id="B9495D5DC742A30BFD22F847FC2D32F0" bold="true" box="[760,917,1946,1970]" pageId="1" pageNumber="532">New synonym.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B82814FC742A30BFF4DF864FCA032AD" blockId="1.[151,1437,1731,2031]" pageId="1" pageNumber="532">
|
||
<treatmentCitationGroup id="AB2DA661C742A30BFF4DF864FCA032AD" pageId="1" pageNumber="532">
|
||
<treatmentCitation id="0A9CA75EC742A30BFF4DF864FCA83293" author="Frey, G." box="[151,784,1976,2001]" page="174" pageId="1" pageNumber="532" year="1970">
|
||
<taxonomicName id="4C3DFACCC742A30BFF4DF864FCA83293" ID-CoL="7HPST" authority="Frey, 1970: 174" authorityName="Frey" authorityPageNumber="174" authorityYear="1970" box="[151,784,1976,2001]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subGenus="Lutera" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC742A30BFF4DF864FEBD3292" box="[151,261,1977,2000]" italics="true" pageId="1" pageNumber="532">Parastasia</emphasis>
|
||
(
|
||
<emphasis id="B9495D5DC742A30BFEC3F864FEE53292" box="[281,349,1977,2000]" italics="true" pageId="1" pageNumber="532">Lutera</emphasis>
|
||
)
|
||
<emphasis id="B9495D5DC742A30BFEABF865FDE23292" box="[369,602,1976,2000]" italics="true" pageId="1" pageNumber="532">nigromaculata guttata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC742A30BFDBFF864FCA83293" author="Frey, G." box="[613,784,1977,2001]" pageId="1" pageNumber="532" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey, 1970: 174</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="54863FEDC742A30BFCF9F864FCE83293" box="[803,848,1977,2001]" pageId="1" pageNumber="532">type</typeStatus>
|
||
locality:
|
||
<location id="8EE2D794C742A30BFC61F865FBB83292" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C742A30BFC61F865FBB83292" box="[955,1024,1976,2000]" country="Laos" county="New" municipality="Saravan Province" name="Khong" pageId="1" pageNumber="532">Khong</location>
|
||
sedane,
|
||
<location id="8EE2D794C742A30BFBB9F865FAE43292" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C742A30BFBB9F865FAE43292" box="[1123,1372,1976,2000]" country="Laos" county="New" municipality="Saravan Province" name="Wapik-Hamthong Prov." pageId="1" pageNumber="532">Wapik-Hamthong Prov.</location>
|
||
,
|
||
<collectingCountry id="F32AC1DFC742A30BFAB1F864FA243292" box="[1387,1436,1977,2000]" name="Laos" pageId="1" pageNumber="532">Laos</collectingCountry>
|
||
[=
|
||
<location id="8EE2D794C742A30BFF01F80AFEDC32AD" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C742A30BFF01F80AFEDC32AD" box="[219,356,2007,2031]" country="Laos" county="New" municipality="Saravan Province" name="Khongxedon" pageId="1" pageNumber="532">Khongxedon</location>
|
||
,
|
||
<collectingMunicipality id="6BE61B35C742A30BFEB7F80AFD9A32AD" box="[365,546,2007,2031]" pageId="1" pageNumber="532">Saravan Province</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F32AC1DFC742A30BFDF6F80AFDDB32AD" box="[556,611,2007,2031]" name="Laos" pageId="1" pageNumber="532">Laos</collectingCountry>
|
||
]).
|
||
<emphasis id="B9495D5DC742A30BFDA1F80AFCA032AD" bold="true" box="[635,792,2007,2031]" pageId="1" pageNumber="532">
|
||
<collectingCounty id="62E3F9C3C742A30BFDA1F80AFD1332AD" box="[635,683,2007,2031]" pageId="1" pageNumber="532">New</collectingCounty>
|
||
synonym.
|
||
</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B82814FC741A308FF4DFF4AFB30358F" blockId="2.[151,1436,150,297]" pageId="2" pageNumber="533">
|
||
<treatmentCitation id="0A9CA75EC741A308FF4DFF4AFDEB35EC" author="Machatschke, J. W." box="[151,595,150,175]" page="45" pageId="2" pageNumber="533" year="1972">
|
||
<taxonomicName id="4C3DFACCC741A308FF4DFF4AFDEB35EC" authority="Machatschke, 1972: 45" authorityName="Machatschke" authorityPageNumber="45" authorityYear="1972" box="[151,595,150,175]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="pauliana">
|
||
<emphasis id="B9495D5DC741A308FF4DFF4AFEDC35ED" box="[151,356,151,175]" italics="true" pageId="2" pageNumber="533">Parastasia pauliana</emphasis>
|
||
<bibRefCitation id="EFACFCBEC741A308FEB0FF4AFDEB35EC" author="Machatschke, J. W." box="[362,595,150,175]" pageId="2" pageNumber="533" pagination="1 - 361" refId="ref7615" refString="Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus, Supplementa. Pars 66. Fascicle I. Editio Secunda. W. Junk, The Hague, pp. 1 - 361." type="book chapter" year="1972">Machatschke, 1972: 45</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
[replacement name for
|
||
<taxonomicName id="4C3DFACCC741A308FC9BFF4AFBFF35ED" ID-CoL="4DCX2" authority="(Ohaus)" baseAuthorityName="Ohaus" box="[833,1095,151,175]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FC9BFF4AFC5535ED" box="[833,1005,151,175]" italics="true" pageId="2" pageNumber="533">P. nigromaculata</emphasis>
|
||
(Ohaus)
|
||
</taxonomicName>
|
||
, preoccupied by
|
||
<taxonomicName id="4C3DFACCC741A308FB2AFF4AFE3D358F" ID-CoL="4DCX2" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FB2AFF4AFA2435ED" box="[1264,1436,151,175]" italics="true" pageId="2" pageNumber="533">P. nigromaculata</emphasis>
|
||
(Blanchard, 1850)
|
||
</taxonomicName>
|
||
];
|
||
<treatmentCitation id="0A9CA75EC741A308FE4FFF68FD99358F" author="Wada, K." box="[405,545,181,205]" page="5" pageId="2" pageNumber="533" year="1988">
|
||
<bibRefCitation id="EFACFCBEC741A308FE4FFF68FD99358F" author="Wada, K." box="[405,545,181,205]" pageId="2" pageNumber="533" pagination="1 - 19" refId="ref8013" refString="Wada, K. (1988) Notes on the genus Parastasia Westwood, 1841 (Rutelinae, Scarabaeidae, Col.). Lamellicornia, 4, 1 - 19." type="journal article" year="1988">Wada 1988: 5</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<collectingRegion id="49F94FADC741A308FDF4FF68FD12358F" box="[558,682,181,205]" country="Thailand" name="Chiang Mai" pageId="2" pageNumber="533">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32AC1DFC741A308FD6EFF68FCAB358F" box="[692,787,181,205]" name="Thailand" pageId="2" pageNumber="533">Thailand</collectingCountry>
|
||
); Synonymized by
|
||
<treatmentCitation id="0A9CA75EC741A308FC0FFF68FB3C358F" author="Kuijten, P. J." box="[981,1156,181,205]" page="87" pageId="2" pageNumber="533" year="1988">
|
||
<bibRefCitation id="EFACFCBEC741A308FC0FFF68FB3C358F" author="Kuijten, P. J." box="[981,1156,181,205]" pageId="2" pageNumber="533" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten 1988: 87</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC741A308FF4DFF08FE373448" blockId="2.[151,1436,150,297]" pageId="2" pageNumber="533">
|
||
<taxonomicName id="4C3DFACCC741A308FF4DFF08FD8335AE" ID-CoL="7HPST" authority="Frey" authorityName="Frey" authorityYear="1970" box="[151,571,212,236]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC741A308FF4DFF08FE4435AE" box="[151,508,212,236]" italics="true" pageId="2" pageNumber="533">Parastasia nigromaculata guttata</emphasis>
|
||
Frey
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A9CA75EC741A308FD91FF09FCF335AE" author="Machatschke, J. W." box="[587,843,212,236]" page="364" pageId="2" pageNumber="533" year="1974">
|
||
<bibRefCitation id="EFACFCBEC741A308FD91FF09FCF335AE" author="Machatschke, J. W." box="[587,843,212,236]" pageId="2" pageNumber="533" pagination="363 - 429" refId="ref7669" refString="Machatschke, J. W. (1974) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus, Supplementa. Pars 66. Fascicle II. Editio Secunda. W. Junk, The Hague, pp. 363 - 429." type="book chapter" year="1974">Machatschke 1974: 364</bibRefCitation>
|
||
</treatmentCitation>
|
||
[erroneously listed as subspecies of
|
||
<taxonomicName id="4C3DFACCC741A308FB32FF08FE3D3448" ID-CoL="4DCX2" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FB32FF08FA2435AE" box="[1256,1436,212,236]" italics="true" pageId="2" pageNumber="533">P. nigromaculata</emphasis>
|
||
(Blanchard, 1850)
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8B82814FC741A308FF4DFECFFB21346B" blockId="2.[151,1436,150,297]" box="[151,1177,273,297]" pageId="2" pageNumber="533">
|
||
<treatmentCitationGroup id="AB2DA661C741A308FF4DFECFFB21346B" box="[151,1177,273,297]" pageId="2" pageNumber="533">
|
||
<taxonomicName id="4C3DFACCC741A308FF4DFECFFDB4346B" authority="(Frey)" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[151,524,273,297]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC741A308FF4DFECFFE7C346B" box="[151,452,273,297]" italics="true" pageId="2" pageNumber="533">Lutera nigromaculata guttata</emphasis>
|
||
(Frey)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A9CA75EC741A308FDCCFECCFD7A346B" author="Krajcik, M." box="[534,706,273,297]" page="79" pageId="2" pageNumber="533" year="2007">
|
||
<bibRefCitation id="EFACFCBEC741A308FDCCFECCFD7A346B" author="Krajcik, M." box="[534,706,273,297]" pageId="2" pageNumber="533" pagination="1 - 139" refId="ref7338" refString="Krajcik, M. (2007) Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x Supplementu m, 4, 1 - 139." type="journal article" year="2007">Krajčík 2007: 79</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogued);
|
||
<treatmentCitation id="0A9CA75EC741A308FC8EFECCFBB5346B" author="Krajcik, M." box="[852,1037,273,297]" page="150" pageId="2" pageNumber="533" year="2012">
|
||
<bibRefCitation id="EFACFCBEC741A308FC8EFECCFBB5346B" author="Krajcik, M." box="[852,1037,273,297]" pageId="2" pageNumber="533" pagination="1 - 139" refId="ref7376" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. Animma. x Supplementum, 5, 1 - 139." type="journal article" year="2012">Krajčík 2012: 150</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogued).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42D1C7C741A308FF4DF9C6FEA33315" ID-DOI="http://doi.org/10.5281/zenodo.13949860" ID-Zenodo-Dep="13949860" httpUri="https://zenodo.org/record/13949860/files/figure.png" pageId="2" pageNumber="533" startId="2.[151,264,1563,1587]" targetBox="[222,1365,326,1522]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B82814FC741A308FF4DF9C6FEA33315" blockId="2.[151,1437,1563,1623]" pageId="2" pageNumber="533">
|
||
<emphasis id="B9495D5DC741A308FF4DF9C6FE823371" bold="true" box="[151,314,1563,1587]" pageId="2" pageNumber="533">FIGURES 1–6.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C3DFACCC741A308FE6AF9C1FCAD3371" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[432,789,1563,1587]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FE6AF9C1FD363371" box="[432,654,1563,1587]" italics="true" pageId="2" pageNumber="533">Lutera nigromaculata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC741A308FD4EF9C6FCAD3371" author="Ohaus, F." box="[660,789,1563,1587]" pageId="2" pageNumber="533" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
from Vietnam.
|
||
<emphasis id="B9495D5DC741A308FC69F9C6FC673371" bold="true" box="[947,991,1563,1587]" pageId="2" pageNumber="533">1, 4,</emphasis>
|
||
dorsal view;
|
||
<emphasis id="B9495D5DC741A308FBB2F9C6FB2D3371" bold="true" box="[1128,1173,1563,1587]" pageId="2" pageNumber="533">2, 5,</emphasis>
|
||
lateral oblique view;
|
||
<emphasis id="B9495D5DC741A308FAAAF9C6FA253371" bold="true" box="[1392,1437,1563,1587]" pageId="2" pageNumber="533">3, 6,</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327D2C4C741A30EFF4DF974FCBC345F" lastPageId="4" lastPageNumber="535" pageId="2" pageNumber="533" type="materials_examined">
|
||
<paragraph id="8B82814FC741A309FF4DF974FE56313B" blockId="2.[151,1437,1705,2020]" lastBlockId="3.[151,1437,151,2043]" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
|
||
<emphasis id="B9495D5DC741A308FF4DF974FEFE3381" bold="true" box="[151,326,1705,1731]" pageId="2" pageNumber="533">Type material.</emphasis>
|
||
<taxonomicName id="4C3DFACCC741A308FE94F977FD653381" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[334,733,1705,1731]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FE94F977FDFC3381" box="[334,580,1705,1731]" italics="true" pageId="2" pageNumber="533">Lutera nigromaculata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC741A308FD96F974FD653381" author="Ohaus, F." box="[588,733,1705,1731]" pageId="2" pageNumber="533" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<materialsCitation id="3B558B12C741A308FD33F974FA2E3249" collectionCode="MNHN" determinerName="M. L. Jameson" location="Lakhon" pageId="2" pageNumber="533" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
|
||
<emphasis id="B9495D5DC741A308FD33F974FCE83381" bold="true" box="[745,848,1705,1731]" pageId="2" pageNumber="533">
|
||
<typeStatus id="54863FEDC741A308FD33F974FCE83381" box="[745,848,1705,1731]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
, by monotypy,
|
||
<specimenCount id="9D3B4AC6C741A308FBD8F977FBAF3386" box="[1026,1047,1706,1732]" count="1" pageId="2" pageNumber="533" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC741A308FBFCF974FB383381" box="[1062,1152,1705,1731]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="533">MNHN</collectionCode>
|
||
,
|
||
<figureCitation id="13069DCAC741A308FB56F974FAAE3381" box="[1164,1302,1705,1731]" captionStart="FIGURES 17–24" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 17–24. Type specimens of Lutera spp. 17–18, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 19–20, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 21–24, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="2" pageNumber="533">Figs. 17–18</figureCitation>
|
||
): “148 78 // m. de.
|
||
<location id="8EE2D794C741A308FF3CF910FE8633A5" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C741A308FF3CF910FE8633A5" box="[230,318,1741,1767]" name="Lakhon" pageId="2" pageNumber="533">Lakhon</location>
|
||
,
|
||
<location id="8EE2D794C741A308FE91F910FE0B33A5" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C741A308FE91F910FE0B33A5" box="[331,435,1741,1767]" name="Harmand" pageId="2" pageNumber="533">Harmand</location>
|
||
// MUSEUM PARIS, LAKHON, HARMAND 1878 //
|
||
<specimenCount id="9D3B4AC6C741A308FBF0F913FB8733AA" box="[1066,1087,1742,1768]" count="1" pageId="2" pageNumber="533" type="female">♀</specimenCount>
|
||
//
|
||
<taxonomicName id="4C3DFACCC741A308FB85F913FAEE33A5" authorityName="Ohaus" authorityYear="1900" box="[1119,1366,1741,1767]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FB85F913FAEE33A5" box="[1119,1366,1741,1767]" italics="true" pageId="2" pageNumber="533">Lutera nigromaculata</emphasis>
|
||
</taxonomicName>
|
||
, type, Ohaus //
|
||
<taxonomicName id="4C3DFACCC741A308FF26F92CFD733249" authority="OHAUS" authorityName="Ohaus" authorityYear="1900" box="[252,715,1777,1803]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">LUTERA NIGROMACULATA OHAUS</taxonomicName>
|
||
,
|
||
<specimenCount id="9D3B4AC6C741A308FD0CF92FFD53324E" box="[726,747,1778,1804]" count="1" pageId="2" pageNumber="533" type="female">♀</specimenCount>
|
||
, HOLOTYPE, det.
|
||
<determinerName id="7E5E6D8BC741A308FC11F92FFBD2324E" box="[971,1130,1778,1804]" pageId="2" pageNumber="533">M.L. Jameson</determinerName>
|
||
1994 //
|
||
<collectionCode id="ED2C198AC741A308FB1DF92FFAA7324E" box="[1223,1311,1778,1804]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="2" pageNumber="533">MNHN</collectionCode>
|
||
, EC1462”
|
||
</materialsCitation>
|
||
. The number of the type specimens was not given in the original publication of
|
||
<taxonomicName id="4C3DFACCC741A308FBECF8CBFA20326D" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[1078,1432,1813,1839]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC741A308FBECF8CBFB43326D" box="[1078,1275,1813,1839]" italics="true" pageId="2" pageNumber="533">L. nigromaculata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC741A308FADFF8C8FA20326D" author="Ohaus, F." box="[1285,1432,1813,1839]" pageId="2" pageNumber="533" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, but
|
||
<bibRefCitation id="EFACFCBEC741A308FF1FF8E4FED93216" author="Ohaus, F." box="[197,353,1849,1876]" pageId="2" pageNumber="533" pagination="81 - 99" refId="ref7756" refString="Ohaus, F. (1905) Beitrage zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, 1905, 81 - 99. https: // doi. org / 10.1002 / mmnd. 48119050214" type="journal article" year="1905">Ohaus (1905)</bibRefCitation>
|
||
mentioned that he described this species based on a single female. This female
|
||
<typeStatus id="54863FEDC741A308FAD1F8E4FAD53211" box="[1291,1389,1849,1875]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
|
||
(by monotypy) has been examined by
|
||
<bibRefCitation id="EFACFCBEC741A308FDCCF880FD05323A" author="Paulian, R." box="[534,701,1885,1912]" pageId="2" pageNumber="533" pagination="73 - 105" refId="ref7972" refString="Paulian, R. (1959) Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). Annales de la Societe entomologique de France, 127, 73 - 105. https: // doi. org / 10.1080 / 21686351.1958.12279112" type="journal article" year="1959">Paulian (1959)</bibRefCitation>
|
||
, but could not be located when
|
||
<bibRefCitation id="EFACFCBEC741A308FBC4F880FB7B3235" author="Kuijten, P. J." box="[1054,1219,1885,1911]" pageId="2" pageNumber="533" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten (1988)</bibRefCitation>
|
||
tried to examine it, and was later rediscovered in MNHN by M.L. Jameson (see
|
||
<bibRefCitation id="EFACFCBEC741A308FC99F85CFC7E32D9" author="Wada, K." box="[835,966,1921,1947]" pageId="2" pageNumber="533" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada 1998</bibRefCitation>
|
||
). The collecting data given in the original publication (
|
||
<bibRefCitation id="EFACFCBEC741A308FEFDF878FE0A32FD" author="Ohaus, F." box="[295,434,1957,1983]" pageId="2" pageNumber="533" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus 1900</bibRefCitation>
|
||
),
|
||
<emphasis id="B9495D5DC741A308FE1CF87BFE5132FD" box="[454,489,1958,1983]" italics="true" pageId="2" pageNumber="533">i.e.</emphasis>
|
||
, “Lakhon,
|
||
<collectingCountry id="F32AC1DFC741A308FDB2F878FD0F32FD" box="[616,695,1957,1983]" name="Vietnam" pageId="2" pageNumber="533">Tonkin</collectingCountry>
|
||
(Dr. Harmand)”, is somewhat different from the label data of the type specimen. The extra word “
|
||
<collectingCountry id="F32AC1DFC741A308FDDDF814FDE332A1" box="[519,603,1993,2019]" name="Vietnam" pageId="2" pageNumber="533">Tonkin</collectingCountry>
|
||
” implies that Ohaus thought Lakhon belonged to
|
||
<collectingCountry id="F32AC1DFC741A308FB4BF814FB5C32A1" box="[1169,1252,1993,2019]" name="Vietnam" pageId="2" pageNumber="533">Tonkin</collectingCountry>
|
||
. The location of Lakhon is actually not in
|
||
<collectingCountry id="F32AC1DFC740A309FE64FF4AFDB535F3" box="[446,525,151,177]" name="Vietnam" pageId="3" pageNumber="534">Tonkin</collectingCountry>
|
||
(roughly equivalent to northern
|
||
<collectingCountry id="F32AC1DFC740A309FC5EFF4AFC5D35F3" box="[900,997,151,177]" name="Vietnam" pageId="3" pageNumber="534">Vietnam</collectingCountry>
|
||
) or Malay Peninsula (see
|
||
<bibRefCitation id="EFACFCBEC740A309FACBFF4AFA2835F3" author="Yang, W. I." box="[1297,1424,151,177]" pageId="3" pageNumber="534" pagination="147 - 156" refId="ref8194" refString="Yang, W. I. (1936) Descriptions of two new genera and four new species of Pentatomidae in the collection of Paris Museum. Chinese Journal of Entomology, 2, 147 - 156." type="journal article" year="1936">Yang 1936</bibRefCitation>
|
||
), but situated in the valley of the Mekong River. Several attempts have been made by taxonomists (see
|
||
<bibRefCitation id="EFACFCBEC740A309FAC8FF66FF6A35BB" author="Huang, G. - Q. & Liu, B. & Gouverneur, X." pageId="3" pageNumber="534" pagination="553 - 564" refId="ref7216" refString="Huang, G. - Q., Liu, B. & Gouverneur, X. (2015) Note on the genus Neacanista Gressitt, 1940 (Coleoptera: Cerambycidae: Lamiinae: Acanthocinini). Zootaxa, 3981 (4), 553 - 564. https: // doi. org / 10.11646 / zootaxa. 3981.4.6" type="journal article" year="2015">
|
||
Huang
|
||
<emphasis id="B9495D5DC740A309FABEFF66FA2D3596" box="[1380,1429,186,212]" italics="true" pageId="3" pageNumber="534">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
, page 557), but none has been able to conclude the exact site of Lakhon, although it was further located in the vicinity of present-day Thakhek,
|
||
<collectingCountry id="F32AC1DFC740A309FDD3FEDEFDFA345F" box="[521,578,259,285]" name="Laos" pageId="3" pageNumber="534">Laos</collectingCountry>
|
||
, in the boundary of modern
|
||
<collectingCountry id="F32AC1DFC740A309FCA1FEDFFC67345E" box="[891,991,258,284]" name="Thailand" pageId="3" pageNumber="534">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32AC1DFC740A309FBC9FEDEFBF1345F" box="[1043,1097,259,285]" name="Laos" pageId="3" pageNumber="534">Laos</collectingCountry>
|
||
based on the journey and map of François-Jules Harmand (
|
||
<bibRefCitation id="EFACFCBEC740A309FE02FEFBFD733402" author="Geiser, M. & Nagel, P." box="[472,715,294,320]" pageId="3" pageNumber="534" pagination="11 - 46" refId="ref7130" refString="Geiser, M. & Nagel, P. (2013) Coleopterology in Laos - an introduction to the nature of the country and its coleopterological exploration. Entomologica Basiliensia et Collectionis Frey, 34, 11 - 46." type="journal article" year="2013">Geiser & Nagel 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC740A309FD0DFEFAFC1E3403" author="Huang, G. - Q. & Liu, B. & Gouverneur, X." box="[727,934,294,321]" pageId="3" pageNumber="534" pagination="553 - 564" refId="ref7216" refString="Huang, G. - Q., Liu, B. & Gouverneur, X. (2015) Note on the genus Neacanista Gressitt, 1940 (Coleoptera: Cerambycidae: Lamiinae: Acanthocinini). Zootaxa, 3981 (4), 553 - 564. https: // doi. org / 10.11646 / zootaxa. 3981.4.6" type="journal article" year="2015">
|
||
Huang
|
||
<emphasis id="B9495D5DC740A309FCF3FEFAFCE23402" box="[809,858,294,320]" italics="true" pageId="3" pageNumber="534">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
). Fortunately, the most recent archaeological works on the ruins of Muang Kao (near Thakhek) provided a lot of information on the history of Lakhon (Lorrillard 2019;
|
||
<bibRefCitation id="EFACFCBEC740A309FF06FEB3FE2734CA" author="White, J. & Shimizu, N. & Bouasisengpaseuth, B." box="[220,415,366,392]" pageId="3" pageNumber="534" pagination="107 - 111" refId="ref8142" refString="White, J., Shimizu, N. & Bouasisengpaseuth, B. (2019) Preliminary report on the first archaeological investigations in 2018 at the Old City, near Thakhek Khammouane Province, the Lao PDR. Journal of Southeast Asian Archaeology, 39, 107 - 111." type="journal article" year="2019">
|
||
White
|
||
<emphasis id="B9495D5DC740A309FEFCFEB2FEED34CA" box="[294,341,366,392]" italics="true" pageId="3" pageNumber="534">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
). Indeed, the name “Lakhon” is a variant of the Thai or Lao word “Nakhon”, which generally refers to the city, town, or power center, but is also specific to an important city during the period of the Kingdom of Lan Xang (14
|
||
<superScript id="7C482C07C740A309FEF4FE68FE833481" attach="none" box="[302,315,437,451]" fontSize="6" pageId="3" pageNumber="534">th</superScript>
|
||
–18
|
||
<superScript id="7C482C07C740A309FEBFFE68FECA3481" attach="left" box="[357,370,437,451]" fontSize="6" pageId="3" pageNumber="534">th</superScript>
|
||
Century).This ancient city Lakhon (now called Muang Kao locally), was situated five kilometers to the south of Thakhek, but was abandoned at the beginning of the 19
|
||
<superScript id="7C482C07C740A309FC1AFE04FC7534A5" attach="left" box="[960,973,473,487]" fontSize="6" pageId="3" pageNumber="534">th</superScript>
|
||
century. The Dutch merchant Gerrit van Wuysthoff passed this city in the year 1641 and clearly indicated it on the left bank of the Mekong River on his map (Lorrillard 2019), while “Lakhon” was misused for a city on the opposite bank by Harmand and other contemporary French explorers who visited Indo-China around the second half of the 19th century. This later “Lakhon” actually is nowadays
|
||
<collectingRegion id="49F94FADC740A309FED7FDB4FE7E37C1" box="[269,454,617,643]" country="Thailand" name="Nakhon Phanom" pageId="3" pageNumber="534">Nakhon Phanom</collectingRegion>
|
||
in eastern
|
||
<collectingCountry id="F32AC1DFC740A309FDE1FDB4FD2737C1" box="[571,671,617,643]" name="Thailand" pageId="3" pageNumber="534">Thailand</collectingCountry>
|
||
(Lorrillard 2019).According to his journey and map, Harmand sailed upstream along the Mekong River and arrived in Lakhon (
|
||
<emphasis id="B9495D5DC740A309FCF7FD53FCE837E5" box="[813,848,654,679]" italics="true" pageId="3" pageNumber="534">i.e.</emphasis>
|
||
,
|
||
<collectingRegion id="49F94FADC740A309FC84FD50FBA537E5" box="[862,1053,653,679]" country="Thailand" name="Nakhon Phanom" pageId="3" pageNumber="534">Nakhon Phanom</collectingRegion>
|
||
) on
|
||
<date id="FF83A78FC740A309FB95FD50FB4337E5" box="[1103,1275,652,679]" pageId="3" pageNumber="534" value="1877-05-02">
|
||
May 2
|
||
<superScript id="7C482C07C740A309FB43FD51FB1137D8" attach="left" box="[1177,1193,652,666]" fontSize="6" pageId="3" pageNumber="534">nd</superScript>
|
||
, 1877
|
||
</date>
|
||
, and two days later he crossed the Mekong River to collect zoological specimens in “Montagnes de Lakhon”, a karst mountains area (
|
||
<geoCoordinate id="EE09E788C740A309FF0FFD09FECE37AD" box="[213,374,724,751]" degrees="17" direction="north" minutes="34" orientation="latitude" pageId="3" pageNumber="534" precision="1" seconds="56.5" value="17.582361">17°34’56.5”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE09E788C740A309FE5BFD09FD9437AD" box="[385,556,724,751]" degrees="104" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="534" precision="1" seconds="26.7" value="104.857414">104°51’26.7”E</geoCoordinate>
|
||
,
|
||
<figureCitation id="13069DCAC740A309FDEDFD09FD3F37AC" box="[567,647,724,750]" captionStart="FIGURE 46" captionStartId="9.[151,250,1970,1994]" captionTargetBox="[164,1423,1173,1947]" captionTargetId="figure-488@9.[164,1423,1173,1947]" captionTargetPageId="9" captionText="FIGURE 46. Known distribution of Lutera nigromaculata Ohaus, 1900 (black dots) and L. nigrita Ohaus, 1930 (black triangles). Type localities are underlined, and the open dot in Nepal is a record from Wada (1998) without precise location." figureDoi="http://doi.org/10.5281/zenodo.13949872" httpUri="https://zenodo.org/record/13949872/files/figure.png" pageId="3" pageNumber="534">Fig. 46</figureCitation>
|
||
) in the northern vicinity of Thakhek (
|
||
<bibRefCitation id="EFACFCBEC740A309FBF2FD09FB6C37AD" author="Harmand, J." box="[1064,1236,724,751]" pageId="3" pageNumber="534" pagination="241 - 288" refId="ref7171" refString="Harmand, J. (1880) Le Laos et les populations sauvages de l'Indo-Chine, 1877. Deuxieme Partie: De Bassac a Hue. Le Tour du Monde, Nouveau Journal des Voyages, 39 (1), 241 - 288." type="journal article" year="1880">Harmand 1880</bibRefCitation>
|
||
). “M. de Lakhon” on the handwritten label of the type specimen of
|
||
<taxonomicName id="4C3DFACCC740A309FD65FD24FC3A3650" authorityName="Ohaus" authorityYear="1900" box="[703,898,760,786]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC740A309FD65FD24FC3A3650" box="[703,898,760,786]" italics="true" pageId="3" pageNumber="534">L. nigromaculata</emphasis>
|
||
</taxonomicName>
|
||
apparently refers to here. This mountain area is located in the region on the left bank of the Mekong River where the Laotians live, but was also controlled by
|
||
<collectingCountry id="F32AC1DFC740A309FAB9FCC1FA253674" box="[1379,1437,796,822]" name="Thailand" pageId="3" pageNumber="534">Siam</collectingCountry>
|
||
at that time and later evolved into the territory of modern
|
||
<collectingCountry id="F32AC1DFC740A309FCFEFC9CFCE23619" box="[804,858,833,859]" name="Laos" pageId="3" pageNumber="534">Laos</collectingCountry>
|
||
after being colonized by
|
||
<collectingCountry id="F32AC1DFC740A309FBA1FC9CFB733619" box="[1147,1227,833,859]" name="France" pageId="3" pageNumber="534">France</collectingCountry>
|
||
. Lakhon became a well-known type locality because of the large number of zoological specimens collected by Harmand, but the labels of some material were simply written as “Lakhon”, and thus it cannot be determined whether it came from the city or the mountains on the opposite bank of the Mekong. This probably is the reason why the location of Lakhon was generally considered in either
|
||
<collectingCountry id="F32AC1DFC740A309FE35FC12FDEB36AB" box="[495,595,975,1001]" name="Thailand" pageId="3" pageNumber="534">Thailand</collectingCountry>
|
||
or
|
||
<collectingCountry id="F32AC1DFC740A309FDA1FC0DFD0936A8" box="[635,689,976,1002]" name="Laos" pageId="3" pageNumber="534">Laos</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFACFCBEC740A309FD1BFC12FB9D36AB" author="Descarpentries, A. & Villiers, A." box="[705,1061,975,1002]" pageId="3" pageNumber="534" pagination="989 - 1008" refId="ref7053" refString="Descarpentries, A. & Villiers, A. (1967) Catalogue raisonne des Buprestidae d'Indochine. 14. Coraebini (4 e partie). Annales de la Societe entomologique de France, New Series, 3, 989 - 1008." type="journal article" year="1967">Descarpentries & Villiers 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC740A309FBE8FC12FB6636A8" author="Breuning, S." box="[1074,1246,975,1002]" pageId="3" pageNumber="534" pagination="3 - 78" refId="ref6921" refString="Breuning, S. (1978) Revision de la tribu des Acanthocinini de la region Asiato-Australienne (Coleoptera: Cerambycidae), Troisieme partie. Mitteilungen aus dem Zoologischen Museum in Berlin, 54 (1), 3 - 78. https: // doi. org / 10.1002 / mmnz. 19780540102" type="journal article" year="1978">Breuning 1978</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC740A309FB31FC12FA2F36A8" author="Lin, M. & Tavakilian, G. & Montreuil, O. & Yang, X." box="[1259,1431,975,1002]" pageId="3" pageNumber="534" pagination="157 - 176" refId="ref7497" refString="Lin, M., Tavakilian, G., Montreuil, O. & Yang, X. (2009) A study on the indiana & galathea species-group of the genus Glenea, with descriptions of four new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Annales de la Societe entomologique de France, New Series, 45 (2), 157 - 176. https: // doi. org / 10.1080 / 00379271.2009.10697599" type="journal article" year="2009">
|
||
Lin
|
||
<emphasis id="B9495D5DC740A309FAC1FC0DFAEC36AB" box="[1307,1364,975,1001]" italics="true" pageId="3" pageNumber="534">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC740A309FF4DFC2EFDE5314C" author="Alonso-Zarazaga, M. A. & Roca-Cusachs, M." box="[151,605,1011,1038]" pageId="3" pageNumber="534" pagination="457 - 458" refId="ref6827" refString="Alonso-Zarazaga, M. A. & Roca-Cusachs, M. (2017) Svataea nom. nov. for Lakhonia Descarpentries & Villiers, 1967 (Coleoptera: Buprestidae), non Lakhonia Yang, 1936 (Hemiptera: Pentatomidae). Arquivos Entomoloxicos, 17, 457 - 458." type="journal article" year="2017">Alonso-Zarazaga & Roca-Cusachs 2017</bibRefCitation>
|
||
). Harmand returned to Lakhon on May 27
|
||
<superScript id="7C482C07C740A309FBEAFC2EFB853143" attach="none" box="[1072,1085,1011,1025]" fontSize="6" pageId="3" pageNumber="534">th</superScript>
|
||
, and shipped out the specimens before continuing his expedition to the east. These specimens arrived in Phnom Penh (capital of
|
||
<collectingCountry id="F32AC1DFC740A309FB21FBCAFAC83173" box="[1275,1392,1047,1073]" name="Cambodia" pageId="3" pageNumber="534">Cambodia</collectingCountry>
|
||
) 11 months later, and Harmand got back to
|
||
<collectingCountry id="F32AC1DFC740A309FD94FBE1FD223114" box="[590,666,1084,1110]" name="France" pageId="3" pageNumber="534">France</collectingCountry>
|
||
in the year 1878; which matches the number “78” given in the round label of the
|
||
<typeStatus id="54863FEDC740A309FEC6FB82FEC6313B" box="[284,382,1119,1145]" pageId="3" pageNumber="534" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13069DCAC740A309FE57FB82FE59313B" box="[397,481,1119,1145]" captionStart="FIGURES 17–24" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 17–24. Type specimens of Lutera spp. 17–18, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 19–20, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 21–24, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="3" pageNumber="534">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B82814FC740A309FF1DFB59FC8F3036" blockId="3.[151,1437,151,2043]" pageId="3" pageNumber="534">
|
||
<taxonomicName id="4C3DFACCC740A309FF1DFB59FC8031DF" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[199,824,1155,1181]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subGenus="Lutera" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC740A309FF1DFB59FEF831DF" box="[199,320,1156,1181]" italics="true" pageId="3" pageNumber="534">Parastasia</emphasis>
|
||
(
|
||
<emphasis id="B9495D5DC740A309FE8EFB59FE2731DF" box="[340,415,1156,1181]" italics="true" pageId="3" pageNumber="534">Lutera</emphasis>
|
||
)
|
||
<emphasis id="B9495D5DC740A309FE68FB5EFD1731DF" box="[434,687,1155,1181]" italics="true" pageId="3" pageNumber="534">nigromaculata guttata</emphasis>
|
||
<materialsCitation id="3B558B12C740A309FD60FB5EFC8031DF" box="[698,824,1155,1181]" collectingDate="1965-05-16" collectingDateMax="1965-05-30" collectingDateMin="1965-05-16" collectionCode="BPBM, NHMB" country="Laos" county="Wapik-hamthong Prov." determinerName="G. Frey" location="Wapik-hamthong Prov." municipality="Wapik-hamthong Prov." pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-male="2" typeStatus="lectotype">
|
||
<bibRefCitation id="EFACFCBEC740A309FD60FB5EFC8031DF" author="Frey, G." box="[698,824,1155,1181]" pageId="3" pageNumber="534" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9495D5DC740A309FC9CFB5EFC0931DF" bold="true" box="[838,945,1155,1181]" pageId="3" pageNumber="534">
|
||
<typeStatus id="54863FEDC740A309FC9CFB5EFC0931DF" box="[838,945,1155,1181]" pageId="3" pageNumber="534" type="lectotype">lectotype</typeStatus>
|
||
</emphasis>
|
||
, by present designation,
|
||
<specimenCount id="9D3B4AC6C740A309FB0CFB5EFB5531DF" box="[1238,1261,1155,1181]" count="1" pageId="3" pageNumber="534" type="male">♂</specimenCount>
|
||
, (
|
||
<collectionCode id="ED2C198AC740A309FADEFB5EFAEE31DF" box="[1284,1366,1155,1181]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="3" pageNumber="534" type="Museum">BPBM</collectionCode>
|
||
,
|
||
<figureCitation id="13069DCAC740A309FABFFB5EFF6C3183" captionStart="FIGURES 7–16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[264,1323,329,1857]" captionTargetId="figure-95@4.[235,1352,309,1880]" captionTargetPageId="4" captionText="FIGURES 7–16. Type specimens of Parastasia (Lutera) nigromaculata guttata Frey, 1970. 7–10, lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM); 11–13, paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB); 14–16, paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM)." figureDoi="http://doi.org/10.5281/zenodo.13949863" httpUri="https://zenodo.org/record/13949863/files/figure.png" pageId="3" pageNumber="534">Figs. 7–10</figureCitation>
|
||
): “
|
||
<collectingCountry id="F32AC1DFC740A309FF29FB7AFEFA3183" box="[243,322,1191,1217]" name="Laos" pageId="3" pageNumber="534">LAOS</collectingCountry>
|
||
:
|
||
<location id="8EE2D794C740A309FE94FB7AFDEE3183" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FE94FB7AFDEE3183" box="[334,598,1191,1217]" country="Laos" county="Wapik-hamthong Prov." municipality="Wapik-hamthong Prov." name="Wapik-hamthong Prov." pageId="3" pageNumber="534">Wapik-hamthong Prov.</location>
|
||
,
|
||
<location id="8EE2D794C740A309FDB8FB7AFCB33183" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FDB8FB7AFCB33183" box="[610,779,1191,1217]" country="Laos" county="Wapik-hamthong Prov." municipality="Wapik-hamthong Prov." name="Khong Sedone" pageId="3" pageNumber="534">Khong Sedone</location>
|
||
,
|
||
<date id="FF83A78FC740A309FCC2FB7AFC333180" box="[792,907,1191,1218]" pageId="3" pageNumber="534" value="1965-05-30">
|
||
<collectingDate id="EFC75E67C740A309FCC2FB7AFC333180" box="[792,907,1191,1218]" pageId="3" pageNumber="534" value="1965-05-30">30.V.1965</collectingDate>
|
||
</date>
|
||
// Native Collector, BISHOP MUS. // TYPUS //
|
||
<taxonomicName id="4C3DFACCC740A309FF6AFB16FDD931A6" authorityName="Xu & Zhao" authorityYear="2024" box="[176,609,1227,1253]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" status="subsp. nov." subSpecies="guttata">
|
||
<emphasis id="B9495D5DC740A309FF6AFB16FE6E31A7" box="[176,470,1227,1253]" italics="true" pageId="3" pageNumber="534">Parastasia nigromaculata</emphasis>
|
||
ssp.
|
||
<emphasis id="B9495D5DC740A309FDC8FB16FDD931A6" box="[530,609,1227,1252]" italics="true" pageId="3" pageNumber="534">guttata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE026C740A309FDB0FB16FD0D31A7" box="[618,693,1227,1253]" pageId="3" pageNumber="534" rank="subSpecies">n. ssp.</taxonomicNameLabel>
|
||
, det.
|
||
<determinerName id="7E5E6D8BC740A309FD2EFB16FCF431A7" box="[756,844,1227,1253]" pageId="3" pageNumber="534">G. Frey</determinerName>
|
||
, 1968”.
|
||
<emphasis id="B9495D5DC740A309FC74FB16FBE631A7" bold="true" box="[942,1118,1227,1253]" pageId="3" pageNumber="534">
|
||
<typeStatus id="54863FEDC740A309FC74FB16FBE631A7" box="[942,1118,1227,1253]" pageId="3" pageNumber="534" type="paralectotype">Paralectotypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D3B4AC6C740A309FBB4FB16FB2931A7" box="[1134,1169,1227,1253]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FB78FB16FB4231A7" box="[1186,1274,1227,1253]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="534">NHMB</collectionCode>
|
||
,
|
||
<figureCitation id="13069DCAC740A309FADDFB16FA3731A7" box="[1287,1423,1227,1253]" captionStart="FIGURES 7–16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[264,1323,329,1857]" captionTargetId="figure-95@4.[235,1352,309,1880]" captionTargetPageId="4" captionText="FIGURES 7–16. Type specimens of Parastasia (Lutera) nigromaculata guttata Frey, 1970. 7–10, lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM); 11–13, paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB); 14–16, paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM)." figureDoi="http://doi.org/10.5281/zenodo.13949863" httpUri="https://zenodo.org/record/13949863/files/figure.png" pageId="3" pageNumber="534">Figs. 11–13</figureCitation>
|
||
), “
|
||
<collectingCountry id="F32AC1DFC740A309FF78FB32FF49304B" box="[162,241,1263,1289]" name="Laos" pageId="3" pageNumber="534">LAOS</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BE61B35C740A309FED8FB33FDB7304B" box="[258,527,1262,1289]" pageId="3" pageNumber="534">Wapik-hamthong Prov.</collectingMunicipality>
|
||
, Khong Sedone,
|
||
<date id="FF83A78FC740A309FD3BFB33FCEC304B" box="[737,852,1262,1289]" pageId="3" pageNumber="534" value="1965-05-16">
|
||
<collectingDate id="EFC75E67C740A309FD3BFB33FCEC304B" box="[737,852,1262,1289]" pageId="3" pageNumber="534" value="1965-05-16">16.V.1965</collectingDate>
|
||
</date>
|
||
// Native Collector, BISHOP //
|
||
<typeStatus id="54863FEDC740A309FB3BFB32FAC7304B" box="[1249,1407,1263,1289]" pageId="3" pageNumber="534" type="paratype">PARATYPUS</typeStatus>
|
||
//
|
||
<taxonomicName id="4C3DFACCC740A309FF4DFACEFDF6306E" authorityName="Xu & Zhao" authorityYear="2024" box="[151,590,1298,1325]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" status="subsp. nov." subSpecies="guttata">
|
||
<emphasis id="B9495D5DC740A309FF4DFACEFE07306E" box="[151,447,1298,1324]" italics="true" pageId="3" pageNumber="534">Parastasia nigromaculata</emphasis>
|
||
ssp.
|
||
<emphasis id="B9495D5DC740A309FE25FACEFDF6306E" box="[511,590,1299,1324]" italics="true" pageId="3" pageNumber="534">guttata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE026C740A309FD83FACEFD1E306F" box="[601,678,1299,1325]" pageId="3" pageNumber="534" rank="subSpecies">n. ssp.</taxonomicNameLabel>
|
||
, det.
|
||
<determinerName id="7E5E6D8BC740A309FD33FACFFCFB306F" box="[745,835,1298,1325]" pageId="3" pageNumber="534">G. Frey</determinerName>
|
||
, 1968”;
|
||
<specimenCount id="9D3B4AC6C740A309FC73FACFFC74306E" box="[937,972,1298,1324]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FC05FACFFB89306E" box="[991,1073,1298,1324]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="3" pageNumber="534" type="Museum">BPBM</collectionCode>
|
||
,
|
||
<figureCitation id="13069DCAC740A309FB9AFACFFB73306E" box="[1088,1227,1298,1324]" captionStart="FIGURES 7–16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[264,1323,329,1857]" captionTargetId="figure-95@4.[235,1352,309,1880]" captionTargetPageId="4" captionText="FIGURES 7–16. Type specimens of Parastasia (Lutera) nigromaculata guttata Frey, 1970. 7–10, lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM); 11–13, paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB); 14–16, paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM)." figureDoi="http://doi.org/10.5281/zenodo.13949863" httpUri="https://zenodo.org/record/13949863/files/figure.png" pageId="3" pageNumber="534">Figs. 14–16</figureCitation>
|
||
), “
|
||
<collectingCountry id="F32AC1DFC740A309FB37FACFFA84306E" box="[1261,1340,1298,1324]" name="Laos" pageId="3" pageNumber="534">LAOS</collectingCountry>
|
||
:
|
||
<collectingCounty id="62E3F9C3C740A309FA90FACFFEE83013" pageId="3" pageNumber="534">Wapik-hamthong Prov.</collectingCounty>
|
||
, Khong Sedone,
|
||
<date id="FF83A78FC740A309FDC6FAEBFD373013" box="[540,655,1334,1361]" pageId="3" pageNumber="534" value="1965-05-30">
|
||
<collectingDate id="EFC75E67C740A309FDC6FAEBFD373013" box="[540,655,1334,1361]" pageId="3" pageNumber="534" value="1965-05-30">30.V.1965</collectingDate>
|
||
</date>
|
||
// Native Collector, BISHOP MUS. //
|
||
<typeStatus id="54863FEDC740A309FB84FAEBFB443012" box="[1118,1276,1334,1360]" pageId="3" pageNumber="534" type="paratype">PARATYPUS</typeStatus>
|
||
//
|
||
<taxonomicName id="4C3DFACCC740A309FAF9FAEAFE7B3036" authorityName="Xu & Zhao" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" status="subsp. nov." subSpecies="guttata">
|
||
<emphasis id="B9495D5DC740A309FAF9FAEAFE833036" italics="true" pageId="3" pageNumber="534">Parastasia nigromaculata</emphasis>
|
||
ssp.
|
||
<emphasis id="B9495D5DC740A309FEAEFA86FE7B3036" box="[372,451,1371,1396]" italics="true" pageId="3" pageNumber="534">guttata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE026C740A309FE10FA86FDAB3037" box="[458,531,1371,1397]" pageId="3" pageNumber="534" rank="subSpecies">n. ssp.</taxonomicNameLabel>
|
||
, det.
|
||
<determinerName id="7E5E6D8BC740A309FD95FA87FD1D3037" box="[591,677,1370,1397]" pageId="3" pageNumber="534">G. Frey</determinerName>
|
||
, 1968 // +1”.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC740A309FF1DFAA2FE8F332D" blockId="3.[151,1437,151,2043]" pageId="3" pageNumber="534">
|
||
<taxonomicName id="4C3DFACCC740A309FF1DFAA2FCA230DA" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[199,794,1406,1433]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subGenus="Lutera" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC740A309FF1DFAA2FEF830DA" box="[199,320,1407,1432]" italics="true" pageId="3" pageNumber="534">Parastasia</emphasis>
|
||
(
|
||
<emphasis id="B9495D5DC740A309FE95FAA2FE2230DA" box="[335,410,1407,1432]" italics="true" pageId="3" pageNumber="534">Lutera</emphasis>
|
||
)
|
||
<emphasis id="B9495D5DC740A309FE72FAA3FD1830DA" box="[424,672,1406,1432]" italics="true" pageId="3" pageNumber="534">nigromaculata guttata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC740A309FD7FFAA2FCA230DA" author="Frey, G." box="[677,794,1406,1433]" pageId="3" pageNumber="534" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
was described based on three Laotian specimens collected from Khong Sedone (
|
||
<geoCoordinate id="EE09E788C740A309FE48FA7FFE4D30FF" box="[402,501,1442,1469]" degrees="15" direction="north" minutes="34" orientation="latitude" pageId="3" pageNumber="534" precision="925" value="15.566667">15°34’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE09E788C740A309FDDEFA7FFDCB30FF" box="[516,627,1442,1469]" degrees="105" direction="east" minutes="49" orientation="longitude" pageId="3" pageNumber="534" precision="925" value="105.816666">105°49’E</geoCoordinate>
|
||
; as “Khongsedane” in the original publication). This is the only taxon without type designation in the publication of
|
||
<bibRefCitation id="EFACFCBEC740A309FD76FA1BFC8E30A2" author="Frey, G." box="[684,822,1478,1504]" pageId="3" pageNumber="534" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey (1970)</bibRefCitation>
|
||
, although one of the
|
||
<specimenCount id="9D3B4AC6C740A309FBF7FA1BFB2730A2" box="[1069,1183,1478,1504]" count="3" pageId="3" pageNumber="534" type="generic">three type</specimenCount>
|
||
specimens was found bearing a label “TYPUS” and the remaining two had attached a label “
|
||
<typeStatus id="54863FEDC740A309FC12FA37FBD73346" box="[968,1135,1514,1540]" pageId="3" pageNumber="534" type="paratype">PARATYPUS</typeStatus>
|
||
”. According to Art. 72.1.1 (
|
||
<bibRefCitation id="EFACFCBEC740A309FF45F9D3FE9D336A" author="ICZN & International Commission on Zoological Nomenclature" box="[159,293,1550,1576]" pageId="3" pageNumber="534" refId="ref7277" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London, 306 pp. Available from: https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / (accessed 15 March 2023)" type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), all
|
||
<specimenCount id="9D3B4AC6C740A309FE87F9D0FDAA3365" box="[349,530,1549,1575]" count="3" pageId="3" pageNumber="534" type="generic">three specimens</specimenCount>
|
||
should be treated as
|
||
<typeStatus id="54863FEDC740A309FCDAF9D3FCDE336A" box="[768,870,1550,1576]" pageId="3" pageNumber="534" type="syntype">syntypes</typeStatus>
|
||
. Considering the new synonym introduced herein, the male specimen in BPBM with a red label “TYPUS” is here designated as
|
||
<typeStatus id="54863FEDC740A309FBD3F9ECFBC83309" box="[1033,1136,1585,1611]" pageId="3" pageNumber="534" type="lectotype">lectotype</typeStatus>
|
||
to enhance the stability of nomenclature.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC740A30EFF1DF9A4FCBC345F" blockId="3.[151,1437,151,2043]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="535" pageId="3" pageNumber="534">
|
||
<emphasis id="B9495D5DC740A309FF1DF9A4FDF933D1" bold="true" box="[199,577,1657,1683]" pageId="3" pageNumber="534">Additional material examined</emphasis>
|
||
(
|
||
<specimenCount id="9D3B4AC6C740A309FD86F9A4FD1C33D1" box="[604,676,1657,1683]" count="18" pageId="3" pageNumber="534" type="male">18♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3B4AC6C740A309FD60F9A4FD4933D1" box="[698,753,1657,1683]" count="8" pageId="3" pageNumber="534" type="female">8♀♀</specimenCount>
|
||
).
|
||
<emphasis id="B9495D5DC740A309FCCEF9A4FCDE33D1" bold="true" box="[788,870,1657,1683]" pageId="3" pageNumber="534">
|
||
<collectingCountry id="F32AC1DFC740A309FCCEF9A4FCDE33D1" box="[788,870,1657,1683]" name="India" pageId="3" pageNumber="534">INDIA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<materialsCitation id="3B558B12C740A309FC5AF9A4FA2033D1" box="[896,1432,1657,1683]" collectingDate="1990-06-03" collectionCode="ZMPC" collectorName="Darjeeling" country="India" location="Sikkim" pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Sikkim">
|
||
<specimenCount id="9D3B4AC6C740A309FC5AF9A4FC1B33D1" box="[896,931,1657,1683]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FC66F9A4FBB633D1" box="[956,1038,1657,1683]" collectionName="ZMPC" pageId="3" pageNumber="534">ZMPC</collectionCode>
|
||
),
|
||
<date id="FF83A78FC740A309FBF1F9A4FB2733D1" box="[1067,1183,1657,1683]" pageId="3" pageNumber="534" value="1990-06-03">
|
||
<collectingDate id="EFC75E67C740A309FBF1F9A4FB2733D1" box="[1067,1183,1657,1683]" pageId="3" pageNumber="534" value="1990-06-03">1990.VI.3</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FB6FF9A4FA9633D1" box="[1205,1326,1657,1683]" pageId="3" pageNumber="534">Darjeeling</collectorName>
|
||
,
|
||
<collectingRegion id="49F94FADC740A309FA99F9A4FA2033D1" box="[1347,1432,1657,1683]" country="India" name="Sikkim" pageId="3" pageNumber="534">Sikkim</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FF4DF940FC1133FA" box="[151,937,1693,1720]" collectionCode="KWPC" collectorName="Doi Pui" country="Thailand" location="Chiang Mai" pageId="3" pageNumber="534" specimenCount="31" specimenCount-female="25" specimenCount-male="6" stateProvince="Chiang Mai">
|
||
<emphasis id="B9495D5DC740A309FF4DF940FE9633F5" bold="true" box="[151,302,1693,1719]" pageId="3" pageNumber="534">
|
||
<collectingCountry id="F32AC1DFC740A309FF4DF940FE9633F5" box="[151,302,1693,1719]" name="Thailand" pageId="3" pageNumber="534">THAILAND</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D3B4AC6C740A309FE9AF940FEC233F5" box="[320,378,1693,1719]" count="6" pageId="3" pageNumber="534" type="male">6♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3B4AC6C740A309FE53F940FE7633FA" box="[393,462,1693,1720]" count="25" pageId="3" pageNumber="534" type="female">25♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE3AF940FD8C33F5" box="[480,564,1693,1719]" collectionName="KWPC" pageId="3" pageNumber="534">KWPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FD9CF940FD6933F5" box="[582,721,1693,1719]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="534">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectorName id="26C8E499C740A309FD05F940FC8333F5" box="[735,827,1693,1719]" pageId="3" pageNumber="534">Doi Pui</collectorName>
|
||
, 1985.VI
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FC6DF940FEA63399" collectingDate="1989-08-02" collectionCode="KWPC" country="Thailand" location="Maetang" pageId="3" pageNumber="534" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Chiang Mai">
|
||
<specimenCount id="9D3B4AC6C740A309FC6DF940FC6433F5" box="[951,988,1693,1719]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3B4AC6C740A309FC31F940FB9B33F5" box="[1003,1059,1693,1719]" count="7" pageId="3" pageNumber="534" type="female">7♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FBEFF940FB3133F5" box="[1077,1161,1693,1719]" collectionName="KWPC" pageId="3" pageNumber="534">KWPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FB41F940FA9D33F5" box="[1179,1317,1693,1719]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="534">Chiang Mai</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C740A309FAE9F940FA2033F5" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FAE9F940FA2033F5" box="[1331,1432,1693,1719]" country="Thailand" name="Maetang" pageId="3" pageNumber="534" stateProvince="Chiang Mai">Maetang</location>
|
||
,
|
||
<date id="FF83A78FC740A309FF4DF91CFEA63399" box="[151,286,1729,1755]" pageId="3" pageNumber="534" value="1989-08-02">
|
||
<collectingDate id="EFC75E67C740A309FF4DF91CFEA63399" box="[151,286,1729,1755]" pageId="3" pageNumber="534" value="1989-08-02">1989.VIII.2</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FEF3F91CFCBA3399" box="[297,770,1729,1755]" collectionCode="KWPC" collectorName="Fang" country="Thailand" location="Chiang Mai" pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
|
||
<specimenCount id="9D3B4AC6C740A309FEF3F91CFEF43399" box="[297,332,1729,1755]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE83F91CFE153399" box="[345,429,1729,1755]" collectionName="KWPC" pageId="3" pageNumber="534">KWPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FE61F91CFDF93399" box="[443,577,1729,1755]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="534">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectorName id="26C8E499C740A309FD91F91CFD3E3399" box="[587,646,1729,1755]" pageId="3" pageNumber="534">Fang</collectorName>
|
||
, 1993.VIII
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FCD6F91CFC5C33BD" collectingDate="2008-05-26" collectionCode="THNHM" collectorName="T. Jeenthong" country="Thailand" latitude="19.967417" location="Pha Hom Pok National Park" longLatPrecision="1" longitude="99.15381" municipality="Fang Dist." pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Prov.">
|
||
<specimenCount id="9D3B4AC6C740A309FCD6F91CFC973399" box="[780,815,1729,1755]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FCE6F91CFC1E3399" box="[828,934,1729,1755]" pageId="3" pageNumber="534">THNHM</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FC6EF91CFBC23399" box="[948,1146,1729,1755]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="534">Chiang Mai Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BE61B35C740A309FB5EF91CFB423399" box="[1156,1274,1729,1755]" pageId="3" pageNumber="534">Fang Dist.</collectingMunicipality>
|
||
,
|
||
<location id="8EE2D794C740A309FAD9F91CFE8E33BD" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FAD9F91CFE8E33BD" country="Thailand" latitude="19.967417" longLatPrecision="1" longitude="99.15381" municipality="Fang Dist." name="Pha Hom Pok National Park" pageId="3" pageNumber="534" stateProvince="Chiang Mai Prov.">Pha Hom Pok National Park</location>
|
||
,
|
||
<geoCoordinate id="EE09E788C740A309FE9EF939FE5C33BD" box="[324,484,1764,1791]" degrees="19" direction="north" minutes="58" orientation="latitude" pageId="3" pageNumber="534" precision="1" seconds="02.7" value="19.967417">19°58’02.7”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE09E788C740A309FE28F938FD3733BD" box="[498,655,1765,1791]" degrees="99" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="534" precision="1" seconds="13.7" value="99.15381">99°09’13.7”E</geoCoordinate>
|
||
,
|
||
<date id="FF83A78FC740A309FD47F938FCAA33BD" box="[669,786,1765,1791]" pageId="3" pageNumber="534" value="2008-05-26">
|
||
<collectingDate id="EFC75E67C740A309FD47F938FCAA33BD" box="[669,786,1765,1791]" pageId="3" pageNumber="534" value="2008-05-26">2008.V.26</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FCC4F938FC1733BD" box="[798,943,1765,1791]" pageId="3" pageNumber="534">T. Jeenthong</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FC28F938FC5C3260" collectingDate="2019-07-01" collectingDateMax="2019-07-15" collectingDateMin="2019-07-01" collectionCode="ZMPC" collectorName="Ustinov V." country="Thailand" latitude="19.558107" location="Chiangdao Hill Resort" pageId="3" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
|
||
<specimenCount id="9D3B4AC6C740A309FC28F938FBAD33BD" box="[1010,1045,1765,1791]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FBFFF938FBCF33BD" box="[1061,1143,1765,1791]" collectionName="ZMPC" pageId="3" pageNumber="534">ZMPC</collectionCode>
|
||
),
|
||
<collectingRegion id="49F94FADC740A309FB56F938FAAE33BD" box="[1164,1302,1765,1791]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="534">Chiang Mai</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C740A309FAF9F938FEAD3261" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FAF9F938FEAD3261" country="Thailand" latitude="19.558107" name="Chiangdao Hill Resort" pageId="3" pageNumber="534" stateProvince="Chiang Mai">Chiangdao Hill Resort</location>
|
||
, 19°33′29.19″N. 99°04’33.34’E,
|
||
<date id="FF83A78FC740A309FD53F8D5FC883261" box="[649,816,1800,1827]" pageId="3" pageNumber="534" value="2019-07-01" valueMax="2019-07-15" valueMin="2019-07-01">
|
||
<collectingDate id="EFC75E67C740A309FD53F8D5FC883261" box="[649,816,1800,1827]" pageId="3" pageNumber="534" value="2019-07-01" valueMax="2019-07-15" valueMin="2019-07-01">2019.VII.1–15</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FCE1F8D5FC0A3261" box="[827,946,1800,1827]" pageId="3" pageNumber="534">Ustinov V.</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FC35F8D4FD393204" collectingDate="1985-05-18" collectionCode="KWPC" collectorName="H. Akiyama" country="Thailand" location="Wieng Ko Sai National Park" pageId="3" pageNumber="534" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phrae">
|
||
<specimenCount id="9D3B4AC6C740A309FC35F8D4FBAC3261" box="[1007,1044,1801,1827]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3B4AC6C740A309FBFAF8D4FBFB3261" box="[1056,1091,1801,1827]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FB8BF8D4FB1D3261" box="[1105,1189,1801,1827]" collectionName="KWPC" pageId="3" pageNumber="534">KWPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FB69F8D5FB4E3260" box="[1203,1270,1800,1826]" country="Thailand" name="Phrae" pageId="3" pageNumber="534">Phrae</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C740A309FADAF8D5FE8E3204" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FADAF8D5FE8E3204" country="Thailand" name="Wieng Ko Sai National Park" pageId="3" pageNumber="534" stateProvince="Phrae">Wieng Ko Sai National Park</location>
|
||
,
|
||
<date id="FF83A78FC740A309FE98F8F1FE0F3205" box="[322,439,1836,1863]" pageId="3" pageNumber="534" value="1985-05-18">
|
||
<collectingDate id="EFC75E67C740A309FE98F8F1FE0F3205" box="[322,439,1836,1863]" pageId="3" pageNumber="534" value="1985-05-18">1985.V.18</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FE1EF8F0FDF53204" box="[452,589,1836,1863]" pageId="3" pageNumber="534">H. Akiyama</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FD54F8F1FBCC3204" box="[654,1140,1836,1863]" collectionCode="KWPC" collectorName="Wang Chin" country="Thailand" location="Phrae" pageId="3" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Phrae">
|
||
<specimenCount id="9D3B4AC6C740A309FD54F8F1FD093204" box="[654,689,1836,1862]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FD1BF8F1FCAD3204" box="[705,789,1836,1862]" collectionName="KWPC" pageId="3" pageNumber="534">KWPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FCFFF8F1FCD03204" box="[805,872,1836,1862]" country="Thailand" name="Phrae" pageId="3" pageNumber="534">Phrae</collectingRegion>
|
||
,
|
||
<collectorName id="26C8E499C740A309FCA9F8F0FC4D3204" box="[883,1013,1836,1863]" pageId="3" pageNumber="534">Wang Chin</collectorName>
|
||
, 1992.VIII
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FB5BF8F1FE203228" collectingDate="1991-06-20" collectionCode="ZMPC" country="Thailand" location="Wangchin" municipality="Prov" pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Phrae">
|
||
<specimenCount id="9D3B4AC6C740A309FB5BF8F1FB1C3204" box="[1153,1188,1836,1862]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FB6EF8F1FABE3204" box="[1204,1286,1836,1862]" collectionName="ZMPC" pageId="3" pageNumber="534">ZMPC</collectionCode>
|
||
),
|
||
<collectingRegion id="49F94FADC740A309FAC0F8F1FAE23204" box="[1306,1370,1836,1862]" country="Thailand" name="Phrae" pageId="3" pageNumber="534">Phrae</collectingRegion>
|
||
Prov,
|
||
<location id="8EE2D794C740A309FF4DF88DFEB33228" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FF4DF88DFEB33228" box="[151,267,1872,1898]" country="Thailand" municipality="Prov" name="Wangchin" pageId="3" pageNumber="534" stateProvince="Phrae">Wangchin</location>
|
||
,
|
||
<date id="FF83A78FC740A309FECCF88DFE203228" box="[278,408,1872,1898]" pageId="3" pageNumber="534" value="1991-06-20">
|
||
<collectingDate id="EFC75E67C740A309FECCF88DFE203228" box="[278,408,1872,1898]" pageId="3" pageNumber="534" value="1991-06-20">1991.VI.20</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FE7EF88DFF7B32CC" collectingDate="1993-06-17" collectingDateMax="1993-06-21" collectingDateMin="1993-06-17" collectionCode="CZPC" collectorName="J. Schneider" country="Thailand" elevation="1300" location="Thai / Burma border" municipality="Thai" pageId="3" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Mae Hong Son">
|
||
<specimenCount id="9D3B4AC6C740A309FE7EF88DFE7F3228" box="[420,455,1872,1898]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE0FF88DFDA63228" box="[469,542,1872,1898]" collectionName="CZPC" pageId="3" pageNumber="534">CZPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FDF7F88CFD6E3228" box="[557,726,1872,1899]" country="Thailand" name="Mae Hong Son" pageId="3" pageNumber="534">Mae Hong Son</collectingRegion>
|
||
env.,
|
||
<location id="8EE2D794C740A309FCCDF88DFC483228" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FCCDF88DFC483228" box="[791,1008,1872,1898]" country="Thailand" municipality="Thai" name="Thai / Burma border" pageId="3" pageNumber="534" stateProvince="Mae Hong Son">Thai/Burma border</location>
|
||
,
|
||
<quantity id="4CC52CAAC740A309FC20F88DFBE93229" box="[1018,1105,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="534" unit="m" value="1300.0">
|
||
<elevation id="0010667CC740A309FC20F88DFBE93229" box="[1018,1105,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="534" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF83A78FC740A309FB87F88DFAB03228" box="[1117,1288,1872,1898]" pageId="3" pageNumber="534" value="1993-06-17" valueMax="1993-06-21" valueMin="1993-06-17">
|
||
<collectingDate id="EFC75E67C740A309FB87F88DFAB03228" box="[1117,1288,1872,1898]" pageId="3" pageNumber="534" value="1993-06-17" valueMax="1993-06-21" valueMin="1993-06-17">1993.VI.17–21</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FACEF88CFA243228" box="[1300,1436,1872,1899]" pageId="3" pageNumber="534">J. Schneider</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FF15F8A9FE3032F0" collectingDate="2007-06-07" collectingMethod="light trap" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Phu Si Than Wildlife Sanctuary" municipality="Kuchinarai Dist." pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Kalasin Prov.">
|
||
<specimenCount id="9D3B4AC6C740A309FF15F8A9FF4A32CC" box="[207,242,1908,1934]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FEDAF8A9FED232CC" box="[256,362,1908,1934]" pageId="3" pageNumber="534">THNHM</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FEA3F8A9FDAA32CD" box="[377,530,1908,1935]" country="Thailand" name="Kalasin" pageId="3" pageNumber="534">Kalasin Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BE61B35C740A309FDC7F8A9FD6132CC" box="[541,729,1908,1934]" pageId="3" pageNumber="534">Kuchinarai Dist.</collectingMunicipality>
|
||
,
|
||
<location id="8EE2D794C740A309FD39F8A9FBFE32CC" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FD39F8A9FBFE32CC" box="[739,1094,1908,1934]" country="Thailand" municipality="Kuchinarai Dist." name="Phu Si Than Wildlife Sanctuary" pageId="3" pageNumber="534" stateProvince="Kalasin Prov.">Phu Si Than Wildlife Sanctuary</location>
|
||
headquarters,
|
||
<date id="FF83A78FC740A309FB30F8A9FAE632CC" box="[1258,1374,1908,1934]" pageId="3" pageNumber="534" value="2007-06-07">
|
||
<collectingDate id="EFC75E67C740A309FB30F8A9FAE632CC" box="[1258,1374,1908,1934]" pageId="3" pageNumber="534" value="2007-06-07">2007.VI.7</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="527CF958C740A309FAB3F8A9FF7D32F1" pageId="3" pageNumber="534">light trap</collectingMethod>
|
||
,
|
||
<collectorName id="26C8E499C740A309FF08F844FEEB32F0" box="[210,339,1944,1971]" pageId="3" pageNumber="534">W. Jaitrong</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FE4FF845FECD3294" collectingDate="1968-05-25" collectionCode="THNHM" collectorName="S. Sililert" country="Thailand" location="Sakaerat Environmental Research Station" pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Ratchasima Prov.">
|
||
<specimenCount id="9D3B4AC6C740A309FE4FF845FE0032F0" box="[405,440,1944,1970]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE12F845FD8A32F0" box="[456,562,1944,1970]" pageId="3" pageNumber="534">THNHM</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FD98F845FCD432F1" box="[578,876,1944,1971]" country="Thailand" name="Nakhon Ratchasima" pageId="3" pageNumber="534">Nakhon Ratchasima Prov.</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C740A309FCA2F845FAE832F0" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FCA2F845FAE832F0" box="[888,1360,1944,1970]" country="Thailand" name="Sakaerat Environmental Research Station" pageId="3" pageNumber="534" stateProvince="Nakhon Ratchasima Prov.">Sakaerat Environmental Research Station</location>
|
||
,
|
||
<date id="FF83A78FC740A309FA87F845FF753295" pageId="3" pageNumber="534" value="1968-05-25">
|
||
<collectingDate id="EFC75E67C740A309FA87F845FF753295" pageId="3" pageNumber="534" value="1968-05-25">1968. V.25</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FF03F861FEFA3294" box="[217,322,1980,2006]" pageId="3" pageNumber="534">S. Sililert</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C740A309FE5AF861FE0D32B8" collectingDate="2019-07-06" collectionCode="THPC" collectorName="T. Hongsuwong" country="Thailand" location="Sakaerat Environmental Research Station" pageId="3" pageNumber="534" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Ratchasima Prov.">
|
||
<specimenCount id="9D3B4AC6C740A309FE5AF861FE1B3294" box="[384,419,1980,2006]" count="1" pageId="3" pageNumber="534" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE68F861FE453294" box="[434,509,1980,2006]" collectionName="THPC" pageId="3" pageNumber="534">THPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FDD1F861FC893295" box="[523,817,1980,2007]" country="Thailand" name="Nakhon Ratchasima" pageId="3" pageNumber="534">Nakhon Ratchasima Prov.</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C740A309FCE6F861FAB73294" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FCE6F861FAB73294" box="[828,1295,1980,2006]" country="Thailand" name="Sakaerat Environmental Research Station" pageId="3" pageNumber="534" stateProvince="Nakhon Ratchasima Prov.">Sakaerat Environmental Research Station</location>
|
||
,
|
||
<date id="FF83A78FC740A309FAC0F861FA2E3294" box="[1306,1430,1980,2006]" pageId="3" pageNumber="534" value="2019-07-06">
|
||
<collectingDate id="EFC75E67C740A309FAC0F861FA2E3294" box="[1306,1430,1980,2006]" pageId="3" pageNumber="534" value="2019-07-06">2019.VII.6</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C740A309FF4DF83CFEF132B9" box="[151,329,2017,2043]" pageId="3" pageNumber="534">T. Hongsuwong</collectorName>
|
||
et al. leg
|
||
</materialsCitation>
|
||
.;
|
||
<materialsCitation id="3B558B12C740A30EFE1FF83DFCBF35F3" collectingDate="2017-05-14" collectionCode="WPPC" collectorName="W. Pathomwattanurak" country="Thailand" county="Pong Nam Ron Dist." lastPageId="4" lastPageNumber="535" location="Thap Sai Subdist." municipality="Pong Nam Ron Dist." pageId="3" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Chanthaburi Prov.">
|
||
<specimenCount id="9D3B4AC6C740A309FE1FF83DFE5032B8" box="[453,488,2016,2042]" count="1" pageId="3" pageNumber="534" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC740A309FE2DF83DFDFF32B8" box="[503,583,2016,2042]" collectionName="WPPC" pageId="3" pageNumber="534">WPPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC740A309FD8FF83DFC9B32B9" box="[597,803,2016,2043]" country="Thailand" name="Chanthaburi" pageId="3" pageNumber="534">Chanthaburi Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BE61B35C740A309FCF4F83CFBA232B8" box="[814,1050,2016,2043]" pageId="3" pageNumber="534">Pong Nam Ron Dist.</collectingMunicipality>
|
||
,
|
||
<location id="8EE2D794C740A309FBFEF83DFB5632B8" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A309FBFEF83DFB5632B8" box="[1060,1262,2016,2042]" country="Thailand" county="Pong Nam Ron Dist." municipality="Pong Nam Ron Dist." name="Thap Sai Subdist." pageId="3" pageNumber="534" stateProvince="Chanthaburi Prov.">Thap Sai Subdist.</location>
|
||
,
|
||
<location id="8EE2D794C740A30EFB22F83DFEF135F3" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C740A30EFB22F83DFEF135F3" country="Thailand" county="Pong Nam Ron Dist." lastPageId="4" lastPageNumber="535" municipality="Pong Nam Ron Dist." name="Chatrium Golf Resort Soi Dao" pageId="3" pageNumber="534" stateProvince="Chanthaburi Prov.">Chatrium Golf Resort Soi Dao</location>
|
||
,
|
||
<date id="FF83A78FC747A30EFE8CFF4AFE7335F3" box="[342,459,151,177]" pageId="4" pageNumber="535" value="2017-05-14">
|
||
<collectingDate id="EFC75E67C747A30EFE8CFF4AFE7335F3" box="[342,459,151,177]" pageId="4" pageNumber="535" value="2017-05-14">2017.V.14</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C747A30EFE02FF4AFD6A35F3" box="[472,722,151,177]" pageId="4" pageNumber="535">W. Pathomwattanurak</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C747A30EFCCFFF4AFE7A3597" collectingDate="2023-06-04" collectionCode="RBMF" collectorName="K. Jiaranaisakul" country="Thailand" location="Ban Sanephong" pageId="4" pageNumber="535" specimenCount="1" specimenCount-female="1" stateProvince="Kanchanaburi Prov.">
|
||
<specimenCount id="9D3B4AC6C747A30EFCCFFF4AFC8035F3" box="[789,824,151,177]" count="1" pageId="4" pageNumber="535" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC747A30EFC93FF4AFC2235F3" box="[841,922,151,177]" collectionName="RBMF" pageId="4" pageNumber="535">RBMF</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC747A30EFC71FF4AFB3635F3" box="[939,1166,151,177]" country="Thailand" name="Kanchanaburi" pageId="4" pageNumber="535">Kanchanaburi Prov.</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C747A30EFB41FF4AFAE835F3" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C747A30EFB41FF4AFAE835F3" box="[1179,1360,151,177]" country="Thailand" name="Ban Sanephong" pageId="4" pageNumber="535" stateProvince="Kanchanaburi Prov.">Ban Sanephong</location>
|
||
,
|
||
<date id="FF83A78FC747A30EFA87FF4AFF743597" pageId="4" pageNumber="535" value="2023-06-04">
|
||
<collectingDate id="EFC75E67C747A30EFA87FF4AFF743597" pageId="4" pageNumber="535" value="2023-06-04">2023. VI.4</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C8E499C747A30EFF03FF66FE363597" box="[217,398,187,213]" pageId="4" pageNumber="535">K. Jiaranaisakul</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B558B12C747A30EFE15FF67FF0735BB" collectionCode="MYNU" collectorName="Dong Tien" country="Vietnam" location="Binh Thuan Prov." pageId="4" pageNumber="535" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Binh Thuan Prov.">
|
||
<emphasis id="B9495D5DC747A30EFE15FF67FDEE3596" bold="true" box="[463,598,186,212]" pageId="4" pageNumber="535">
|
||
<collectingCountry id="F32AC1DFC747A30EFE15FF67FDEE3596" box="[463,598,186,212]" name="Vietnam" pageId="4" pageNumber="535">VIETNAM</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D3B4AC6C747A30EFDBCFF66FD323597" box="[614,650,187,213]" count="1" pageId="4" pageNumber="535" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3B4AC6C747A30EFD4DFF66FD023597" box="[663,698,187,213]" count="1" pageId="4" pageNumber="535" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC747A30EFD10FF66FC9B3597" box="[714,803,187,213]" collectionName="MYNU" pageId="4" pageNumber="535">MYNU</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC747A30EFCE8FF66FC453597" box="[818,1021,187,213]" country="Vietnam" name="Binh Thuan" pageId="4" pageNumber="535">Binh Thuan Prov.</collectingRegion>
|
||
,
|
||
<collectorName id="26C8E499C747A30EFBD3FF66FB3B3597" box="[1033,1155,187,213]" pageId="4" pageNumber="535">Dong Tien</collectorName>
|
||
, 2020.V, native collector leg
|
||
</materialsCitation>
|
||
.;
|
||
<materialsCitation id="3B558B12C747A30EFF09FF02FB8F35BB" box="[211,1079,223,249]" collectionCode="MYNU" country="Vietnam" location="Dong Tien" pageId="4" pageNumber="535" specimenCount="2" specimenCount-male="2" stateProvince="Binh Thuan Prov.">
|
||
<specimenCount id="9D3B4AC6C747A30EFF09FF02FEB335BB" box="[211,267,223,249]" count="2" pageId="4" pageNumber="535" type="male">2♂♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC747A30EFEC7FF02FECE35BB" box="[285,374,223,249]" collectionName="MYNU" pageId="4" pageNumber="535">MYNU</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC747A30EFE52FF02FDE035BB" box="[392,600,223,249]" country="Vietnam" name="Binh Thuan" pageId="4" pageNumber="535">Binh Thuan Prov.</collectingRegion>
|
||
,
|
||
<location id="8EE2D794C747A30EFDBCFF02FD5B35BB" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C747A30EFDBCFF02FD5B35BB" box="[614,739,223,249]" country="Vietnam" name="Dong Tien" pageId="4" pageNumber="535" stateProvince="Binh Thuan Prov.">Dong Tien</location>
|
||
, 2022.V, native collector leg
|
||
</materialsCitation>
|
||
.;
|
||
<materialsCitation id="3B558B12C747A30EFB90FF03FCBC345F" collectingDate="2024-05-17" collectionCode="ZMPC" country="Laos" location="Hoi Khua" municipality="Pak Long" pageId="4" pageNumber="535" specimenCount="3" specimenCount-male="3" stateProvince="Luang Prabang">
|
||
<emphasis id="B9495D5DC747A30EFB90FF03FB2E35BA" bold="true" box="[1098,1174,222,248]" pageId="4" pageNumber="535">
|
||
<collectingCountry id="F32AC1DFC747A30EFB90FF03FB2E35BA" box="[1098,1174,222,248]" name="Laos" pageId="4" pageNumber="535">LAOS</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D3B4AC6C747A30EFB73FF02FB5935BB" box="[1193,1249,223,249]" count="3" pageId="4" pageNumber="535" type="male">3♂♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED2C198AC747A30EFB29FF02FAFB35BB" box="[1267,1347,223,249]" collectionName="ZMPC" pageId="4" pageNumber="535">ZMPC</collectionCode>
|
||
)
|
||
<collectingRegion id="49F94FADC747A30EFA8FFF02FF4E345F" country="Laos" name="Louangphabang" pageId="4" pageNumber="535">Luang Prabang</collectingRegion>
|
||
, near
|
||
<collectingMunicipality id="6BE61B35C747A30EFEE2FEDEFE1F345F" box="[312,423,259,285]" pageId="4" pageNumber="535">Pak Long</collectingMunicipality>
|
||
,
|
||
<location id="8EE2D794C747A30EFE68FEDEFD9A345F" LSID="urn:lsid:plazi:treatment:03943059C742A30CFF4DF986FC3E329F:8EE2D794C747A30EFE68FEDEFD9A345F" box="[434,546,259,285]" country="Laos" municipality="Pak Long" name="Hoi Khua" pageId="4" pageNumber="535" stateProvince="Luang Prabang">Hoi Khua</location>
|
||
, at light,
|
||
<date id="FF83A78FC747A30EFD50FEDEFCBC345F" box="[650,772,259,285]" pageId="4" pageNumber="535" value="2024-05-17">
|
||
<collectingDate id="EFC75E67C747A30EFD50FEDEFCBC345F" box="[650,772,259,285]" pageId="4" pageNumber="535" value="2024-05-17">2024.V.17.</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42D1C7C747A30EFF4DF8B2FE0732B6" ID-DOI="http://doi.org/10.5281/zenodo.13949863" ID-Zenodo-Dep="13949863" httpUri="https://zenodo.org/record/13949863/files/figure.png" pageId="4" pageNumber="535" startId="4.[151,264,1903,1927]" targetBox="[264,1323,329,1857]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B82814FC747A30EFF4DF8B2FE0732B6" blockId="4.[151,1437,1903,2036]" pageId="4" pageNumber="535">
|
||
<emphasis id="B9495D5DC747A30EFF4DF8B2FEFF32C5" bold="true" box="[151,327,1903,1927]" pageId="4" pageNumber="535">FIGURES 7–16.</emphasis>
|
||
Type specimens of
|
||
<taxonomicName id="4C3DFACCC747A30EFDCBF8ADFB8F32CA" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[529,1079,1904,1928]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="535" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subGenus="Lutera" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC747A30EFDCBF8ADFDC732C5" box="[529,639,1904,1927]" italics="true" pageId="4" pageNumber="535">Parastasia</emphasis>
|
||
(
|
||
<emphasis id="B9495D5DC747A30EFD57F8ADFD6932C5" box="[653,721,1904,1927]" italics="true" pageId="4" pageNumber="535">Lutera</emphasis>
|
||
)
|
||
<emphasis id="B9495D5DC747A30EFD05F8ADFC7A32C5" box="[735,962,1904,1928]" italics="true" pageId="4" pageNumber="535">nigromaculata guttata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC747A30EFC12F8ADFB8F32CA" author="Frey, G." box="[968,1079,1904,1928]" pageId="4" pageNumber="535" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9495D5DC747A30EFB9AF8B2FBC132C5" bold="true" box="[1088,1145,1903,1927]" pageId="4" pageNumber="535">7–10,</emphasis>
|
||
lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM);
|
||
<emphasis id="B9495D5DC747A30EFCA1F84EFC0732E9" bold="true" box="[891,959,1939,1963]" pageId="4" pageNumber="535">11–13,</emphasis>
|
||
paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB);
|
||
<emphasis id="B9495D5DC747A30EFD66F86AFCBA328D" bold="true" box="[700,770,1975,1999]" pageId="4" pageNumber="535">14–16,</emphasis>
|
||
paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF42D1C7C746A30FFF4DF9FDFB1633C2" ID-DOI="http://doi.org/10.5281/zenodo.13949866" ID-Zenodo-Dep="13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="5" pageNumber="536" startId="5.[151,264,1568,1592]" targetBox="[232,1377,193,1522]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B82814FC746A30FFF4DF9FDFB1633C2" blockId="5.[151,1437,1568,1665]" pageId="5" pageNumber="536">
|
||
<emphasis id="B9495D5DC746A30FFF4DF9FDFEE1337A" bold="true" box="[151,345,1568,1592]" pageId="5" pageNumber="536">FIGURES 17–24.</emphasis>
|
||
Type specimens of
|
||
<taxonomicName id="4C3DFACCC746A30FFDEDF9FCFD09337A" box="[567,689,1569,1592]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B9495D5DC746A30FFDEDF9FCFDC3337A" box="[567,635,1569,1592]" italics="true" pageId="5" pageNumber="536">Lutera</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
<emphasis id="B9495D5DC746A30FFD61F9FDFCB9337A" bold="true" box="[699,769,1568,1592]" pageId="5" pageNumber="536">17–18,</emphasis>
|
||
female holotype of
|
||
<taxonomicName id="4C3DFACCC746A30FFC3BF9FCFA9E337A" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[993,1318,1568,1592]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC746A30FFC3BF9FCFB2D337A" box="[993,1173,1568,1592]" italics="true" pageId="5" pageNumber="536">L. nigromaculata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC746A30FFB7BF9FDFA9E337A" author="Ohaus, F." box="[1185,1318,1568,1592]" pageId="5" pageNumber="536" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
and labels (©MNHN);
|
||
<emphasis id="B9495D5DC746A30FFECCF999FEE4331E" bold="true" box="[278,348,1604,1628]" pageId="5" pageNumber="536">19–20,</emphasis>
|
||
inauthentic type of
|
||
<taxonomicName id="4C3DFACCC746A30FFDFDF998FD6D331E" authorityName="Ohaus" authorityYear="1900" box="[551,725,1604,1628]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC746A30FFDFDF998FD6D331E" box="[551,725,1604,1628]" italics="true" pageId="5" pageNumber="536">L. nigromaculata</emphasis>
|
||
</taxonomicName>
|
||
in MFNB (photograph by Bernd Jäger);
|
||
<emphasis id="B9495D5DC746A30FFBACF999FB04331E" bold="true" box="[1142,1212,1604,1628]" pageId="5" pageNumber="536">21–24,</emphasis>
|
||
female holotype of
|
||
<taxonomicName id="4C3DFACCC746A30FFA52F998FEDB33C2" authority="Ohaus, 1930" authorityName="Ohaus" authorityYear="1930" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="nigrita">
|
||
<emphasis id="B9495D5DC746A30FFA52F998FF6433C2" italics="true" pageId="5" pageNumber="536">L. nigrita</emphasis>
|
||
<bibRefCitation id="EFACFCBEC746A30FFF38F9B5FEDB33C2" author="Ohaus, F." box="[226,355,1640,1664]" pageId="5" pageNumber="536" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
(in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327D2C4C746A30CFF1DF973FEA83187" lastPageId="6" lastPageNumber="537" pageId="5" pageNumber="536" type="description">
|
||
<paragraph id="8B82814FC746A30FFF1DF973FA2532AB" blockId="5.[151,1437,1710,2025]" pageId="5" pageNumber="536">
|
||
<emphasis id="B9495D5DC746A30FFF1DF973FE5B338A" bold="true" box="[199,483,1710,1736]" pageId="5" pageNumber="536">Redescription. General.</emphasis>
|
||
Body broadly ovoid and strongly convex. All external setae yellowish brown, dorsal surface almost glabrous.
|
||
<emphasis id="B9495D5DC746A30FFE68F90FFE4133AE" bold="true" box="[434,505,1746,1772]" pageId="5" pageNumber="536">Head.</emphasis>
|
||
Dorsal surface yellow with two black spots on frons, marginal portions darkened. Clypeus nearly semicircular, uneven, apex weakly emarginated, anterior margin moderately raised; clypeus and two black spots on frons densely striolate, other portions of frons bearing sparse and elongated punctures. Frontoclypeal suture broadly interrupted medially. Eye canthus simple, not extending beyond outermost point of eye. Antenna reddish brown, length of antennal club equal to the combined length of antennomeres 2–7. Labrum not exposed dorsally. Mandible with blackish brown and upright apex, anterior margin mesially with a small and blunt protrusion. Galea and maxillary palp reddish brown, both mandible and maxilla with dense and long setae at outer margin.
|
||
<collectingRegion id="49F94FADC746A30FFF4DF812FF4E32AB" box="[151,246,1999,2025]" country="Myanmar" name="Chin" pageId="5" pageNumber="536">Mentum</collectingRegion>
|
||
yellowish brown, margins darkened, anterior margin weakly trilobed, surface with sparse short setae.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF4DFF4AFC61368B" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF4DFF4AFEAF35F3" bold="true" box="[151,279,151,177]" pageId="6" pageNumber="537">Pronotum.</emphasis>
|
||
Yellow to brown, darkened at anterior and posterior margins. Strongly convex, about 1.58 times as wide as long, widest near middle. Anterior and posterior margins broadly and strongly protruding respectively; anterior marginal membrane absent. Sides weakly convergent posteriad in posterior half, broadly curved at middle, then strongly convergent anteriad in anterior half. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Anterior angle weakly protruding, posterior angle rounded. Surface with 11 evenly distributed large black spots: one row including four spots at anterior third and another including five spots at basal third, near each lateral margin a single spot at middle; punctures on surface sparse and minute, but dense and rather coarse on the three lateral spots, all marginal lines with elongated punctures.
|
||
<emphasis id="B9495D5DC745A30CFC27FE4FFBC434EE" bold="true" box="[1021,1148,402,428]" pageId="6" pageNumber="537">Scutellum.</emphasis>
|
||
Yellow to brown, margin black. Semicircular in shape, anterior margin slightly sinuate, lateral margin somewhat rounded. Surface with sparse and minute punctures.
|
||
<emphasis id="B9495D5DC745A30CFE4FFE06FE5034B7" bold="true" box="[405,488,475,501]" pageId="6" pageNumber="537">Elytra.</emphasis>
|
||
Yellow to brown. Convex, almost as wide as long, widest at middle. Surface uneven, humeral umbones weakly prominent. Strial punctures large, punctures on primary striae as large as punctures on secondary striae, primary stria I well defined by a longitudinal row of regular punctures, other striae unrecognizable; the whole surface with sparse small punctures, denser on lateral portions. Epipleura with a row of dense, short to moderately long setae.
|
||
<emphasis id="B9495D5DC745A30CFE40FDB6FD7937C6" bold="true" box="[410,705,618,645]" pageId="6" pageNumber="537">Ventral thoracic surface.</emphasis>
|
||
Yellow, anterior part of prosternum and three markings on center of metasternum brown, narrow margins dark brown; sparsely punctate, with moderately long setae, mesosternum with transverse striolation and dense, short setae; mesosternal process not produced beyond mesocoxae, apex wide and truncate, weakly curved.
|
||
<emphasis id="B9495D5DC745A30CFDF6FD0BFD1037B2" bold="true" box="[556,680,726,752]" pageId="6" pageNumber="537">Abdomen.</emphasis>
|
||
Abdominal ventrites not abbreviated at middle, with shallow and transverse striolation, each ventrite with a transverse row of moderately long setae near posterior margin; pygidium sparsely punctate, shallowly striolate basally, with some long setae at apex.
|
||
<emphasis id="B9495D5DC745A30CFC2CFCC2FB8D367B" bold="true" box="[1014,1077,799,825]" pageId="6" pageNumber="537">Legs.</emphasis>
|
||
Tibiae with brown longitudinal patches; protibia strongly tridentate; protarsus thickened, small internomedial protuberance of tarsomere 5 absent in protarsus but present in meso- and metatarsi; all larger tarsal claws split at apex, lower branch of inner protarsal claws conspicuously longer and wider than upper branch, with a rounded apex.
|
||
<emphasis id="B9495D5DC745A30CFBC1FC56FB7536E6" bold="true" box="[1051,1229,906,933]" pageId="6" pageNumber="537">Male genitalia.</emphasis>
|
||
As in
|
||
<figureCitation id="13069DCAC745A30CFACDFC56FF0E368B" captionStart="FIGURES 31–45" captionStartId="8.[151,264,1924,1948]" captionTargetBox="[189,1398,275,1888]" captionTargetId="figure-77@8.[176,1411,260,1900]" captionTargetPageId="8" captionText="FIGURES 31–45. Male genitalia of Lutera species (31–35, dorsal view; 36–40, lateral view; 41–45, ventral view;). 31, 36, 41, L. nigromaculata Ohaus, 1900 from Thailand; 32–35, 37–40, 42–35, L. nigrita Ohaus, 1930: 32, 37, 42, from Luzon Island; 33, 38, 43, from Mindoro Island; 34, 39, 44, from Negros Island; 35, 40, 45, from Mindanao Island" figureDoi="http://doi.org/10.5281/zenodo.13949870" httpUri="https://zenodo.org/record/13949870/files/figure.png" pageId="6" pageNumber="537">Figs 31, 36, 41</figureCitation>
|
||
. Parameres symmetric, concavity of ventral margin varies significantly.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF1DFC0FFE2F311B" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFC0FFE0136AE" bold="true" box="[199,441,978,1004]" pageId="6" pageNumber="537">Sexual dimorphism.</emphasis>
|
||
Clypeus more triangular in female (
|
||
<figureCitation id="13069DCAC745A30CFC83FC0EFC7F36AF" box="[857,967,978,1005]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1563,1587]" captionTargetBox="[222,1365,326,1522]" captionTargetId="figure-96@2.[213,1375,319,1537]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Lutera nigromaculata Ohaus, 1900 from Vietnam. 1, 4, dorsal view; 2, 5, lateral oblique view; 3, 6, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13949860" httpUri="https://zenodo.org/record/13949860/files/figure.png" pageId="6" pageNumber="537">Figs. 4–5</figureCitation>
|
||
), basal third weakly convergent anteriad, apical third rather strongly convergent to the strongly bilobed apex, apical angle each rounded and upright; Lateral portions of frons also strongly striolate. Protarsi not thickened, lower branch of protarsal claw equal to the upper one and slightly wider.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF1DFBBEFC10313F" blockId="6.[151,1437,150,2013]" box="[199,936,1123,1149]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFBBEFEC6313F" bold="true" box="[199,382,1123,1149]" pageId="6" pageNumber="537">Measurements.</emphasis>
|
||
Body length 14.0–
|
||
<quantity id="4CC52CAAC745A30CFD8FFBBEFD02313F" box="[597,698,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="6" pageNumber="537" unit="mm" value="19.8">19.8 mm</quantity>
|
||
, width
|
||
<quantity id="4CC52CAAC745A30CFCD6FBBEFC1B313F" box="[780,931,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.19" metricValueMin="0.83" pageId="6" pageNumber="537" unit="mm" value="10.1" valueMax="11.9" valueMin="8.3">8.3–11.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF1DFB5BFEA83187" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFB5BFEF531E2" bold="true" box="[199,333,1158,1184]" pageId="6" pageNumber="537">Variability.</emphasis>
|
||
The size of the black spots on the dorsal surface is variable, and sometimes the adjacent two are connected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327D2C4C745A30CFF1DFB13FB77304F" pageId="6" pageNumber="537" type="diagnosis">
|
||
<paragraph id="8B82814FC745A30CFF1DFB13FB77304F" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFB13FE7631AA" bold="true" box="[199,462,1230,1256]" pageId="6" pageNumber="537">Differential diagnosis.</emphasis>
|
||
This species is easily distinguished from the congeners by the unique pattern of black spots on the dorsal surface and the presence of a black groove along the sutural costa on elytron.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327D2C4C745A30CFF1DFACBFBE23073" box="[199,1114,1302,1329]" pageId="6" pageNumber="537" type="distribution">
|
||
<paragraph id="8B82814FC745A30CFF1DFACBFBE23073" blockId="6.[151,1437,150,2013]" box="[199,1114,1302,1329]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFACBFED93072" bold="true" box="[199,353,1302,1328]" pageId="6" pageNumber="537">Distribution.</emphasis>
|
||
<collectingCountry id="F32AC1DFC745A30CFEBDFACAFE153073" box="[359,429,1303,1329]" name="Nepal" pageId="6" pageNumber="537">Nepal</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32AC1DFC745A30CFE62FACAFDA63073" box="[440,542,1303,1329]" name="Thailand" pageId="6" pageNumber="537">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32AC1DFC745A30CFDF3FACAFDDA3073" box="[553,610,1303,1329]" name="Laos" pageId="6" pageNumber="537">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32AC1DFC745A30CFDB7FACAFD743073" box="[621,716,1303,1329]" name="Vietnam" pageId="6" pageNumber="537">Vietnam</collectingCountry>
|
||
(
|
||
<emphasis id="B9495D5DC745A30CFD06FACAFCDA3073" bold="true" box="[732,866,1303,1329]" pageId="6" pageNumber="537">new record</emphasis>
|
||
),
|
||
<collectingCountry id="F32AC1DFC745A30CFCA0FACAFC0C3073" box="[890,948,1303,1329]" name="India" pageId="6" pageNumber="537">India</collectingCountry>
|
||
(
|
||
<emphasis id="B9495D5DC745A30CFC19FACAFBF23073" bold="true" box="[963,1098,1303,1329]" pageId="6" pageNumber="537">new record</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327D2C4C745A30CFF1DFAE6FC3E329F" pageId="6" pageNumber="537" type="discussion">
|
||
<paragraph id="8B82814FC745A30CFF1DFAE6FDD033FF" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<emphasis id="B9495D5DC745A30CFF1DFAE6FE833017" bold="true" box="[199,315,1339,1365]" pageId="6" pageNumber="537">Remarks.</emphasis>
|
||
<bibRefCitation id="EFACFCBEC745A30CFE9BFAE6FE623017" author="Arrow, G. J." box="[321,474,1339,1365]" pageId="6" pageNumber="537" refId="ref6879" refString="Arrow, G. J. (1917) The Fauna of British India, Including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor & Francis, London, 387 pp." type="book" year="1917">Arrow (1917)</bibRefCitation>
|
||
merged
|
||
<taxonomicName id="4C3DFACCC745A30CFDE2FAE6FD3B3016" authorityName="Westwood" authorityYear="1875" box="[568,643,1339,1364]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9495D5DC745A30CFDE2FAE6FD3B3016" box="[568,643,1339,1364]" italics="true" pageId="6" pageNumber="537">Lutera</emphasis>
|
||
</taxonomicName>
|
||
into the genus
|
||
<taxonomicName id="4C3DFACCC745A30CFCF2FAE6FC193016" baseAuthorityName="sensu Arrow" baseAuthorityYear="1917" box="[808,929,1339,1364]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9495D5DC745A30CFCF2FAE6FC193016" box="[808,929,1339,1364]" italics="true" pageId="6" pageNumber="537">Parastasia</emphasis>
|
||
</taxonomicName>
|
||
, resulting in a secondary homonymy between
|
||
<taxonomicName id="4C3DFACCC745A30CFF4DFA82FE57303B" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[151,495,1375,1401]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC745A30CFF4DFA82FEE2303B" box="[151,346,1375,1401]" italics="true" pageId="6" pageNumber="537">L. nigromaculata</emphasis>
|
||
<bibRefCitation id="EFACFCBEC745A30CFEBBFA82FE57303B" author="Ohaus, F." box="[353,495,1375,1401]" pageId="6" pageNumber="537" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3DFACCC745A30CFDFDFA82FC07303B" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[551,959,1374,1401]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC745A30CFDFDFA82FD50303B" box="[551,744,1375,1401]" italics="true" pageId="6" pageNumber="537">P. nigromaculata</emphasis>
|
||
(Blanchard, 1850)
|
||
</taxonomicName>
|
||
. However, he did not review the names of congeners distributed outside
|
||
<collectingCountry id="F32AC1DFC745A30CFE30FA5EFDC030DF" box="[490,632,1411,1437]" name="India" pageId="6" pageNumber="537">British India</collectingCountry>
|
||
to propose replacement names for potential homonyms. Ohaus did not agree with this synonymization and did not place these two species in the same genus in the two world catalogues he compiled (
|
||
<bibRefCitation id="EFACFCBEC745A30CFEEDFA16FE7B30A7" author="Ohaus, F." box="[311,451,1483,1509]" pageId="6" pageNumber="537" pagination="1 - 241" refId="ref7791" refString="Ohaus, F. (1918) Scarabaeidae: Phaenomerinae, Euchirinae, Rutelinae. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum catalogus. Vo. XX. Pars 66. W. Junk, Berlin, pp. 1 - 241." type="book chapter" year="1918">Ohaus 1918</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFACFCBEC745A30CFE0BFA16FDB530A7" author="Ohaus, F." box="[465,525,1483,1509]" pageId="6" pageNumber="537" pagination="1 - 172" refId="ref7920" refString="Ohaus, F. (1934) Coleoptera Lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. Erster Teil. In: Wytsman, P. (Ed.), Genera Insectorum. Coleoptera. Fascicule 199 A. Louis Desmet-Verteneuil, Bruxelles, pp. 1 - 172." type="book chapter" year="1934">1934</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFACFCBEC745A30CFDF9FA16FD0430A7" author="Chujo, M." box="[547,700,1483,1509]" pageId="6" pageNumber="537" pagination="25 - 34" refId="ref6976" refString="Chujo, M. (1966) Coleoptera from Southeast Asia (V). 5. Family Scarabaeidae (2). Memoirs of the Faculty of Education, Kagawa University, Part II (No. 140), 25 - 34." type="book chapter" year="1966">Chûjô (1966)</bibRefCitation>
|
||
noticed this homonymy earlier than
|
||
<bibRefCitation id="EFACFCBEC745A30CFBB3FA16FADC30A7" author="Machatschke, J. W." box="[1129,1380,1483,1509]" pageId="6" pageNumber="537" pagination="1 - 361" refId="ref7615" refString="Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus, Supplementa. Pars 66. Fascicle I. Editio Secunda. W. Junk, The Hague, pp. 1 - 361." type="book chapter" year="1972">Machatschke’s (1972)</bibRefCitation>
|
||
new catalogue, but the new name he proposed,
|
||
<taxonomicName id="4C3DFACCC745A30CFDB4FA32FD6C334B" authorityName="Chujo" authorityYear="1966" box="[622,724,1519,1545]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="ohausi">
|
||
<emphasis id="B9495D5DC745A30CFDB4FA32FD6C334B" box="[622,724,1519,1545]" italics="true" pageId="6" pageNumber="537">P. ohausi</emphasis>
|
||
</taxonomicName>
|
||
, was published in a Japanese journal that was not indexed in the Zoological Record. Several European coleopterists, including Vladimír Balthasar (1897–1978), collaborated with Chûjô on several reports in the same issue of this journal. Nevertheless, for unknown reasons, this paper, which includes not only this new name but also a new species of the genus
|
||
<taxonomicName id="4C3DFACCC745A30CFC1AF986FB713337" authority="Dejean, 1833" authorityName="Dejean" authorityYear="1833" box="[960,1225,1627,1653]" class="Insecta" family="Scarabaeidae" genus="Adoretus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9495D5DC745A30CFC1AF986FB9C3337" box="[960,1060,1627,1653]" italics="true" pageId="6" pageNumber="537">Adoretus</emphasis>
|
||
Dejean, 1833
|
||
</taxonomicName>
|
||
, has hitherto been overlooked. Presently, the validity of the genus
|
||
<taxonomicName id="4C3DFACCC745A30CFD71F9A2FD4E33DA" authorityName="Westwood" authorityYear="1875" box="[683,758,1663,1688]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9495D5DC745A30CFD71F9A2FD4E33DA" box="[683,758,1663,1688]" italics="true" pageId="6" pageNumber="537">Lutera</emphasis>
|
||
</taxonomicName>
|
||
has been widely accepted, and thus the replacement name
|
||
<taxonomicName id="4C3DFACCC745A30CFA52F9A2FECB33FF" authority="Chujo, 1966" authorityName="Chujo" authorityYear="1966" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="ohausi">
|
||
<emphasis id="B9495D5DC745A30CFA52F9A2FF5A33FF" italics="true" pageId="6" pageNumber="537">P. ohausi</emphasis>
|
||
<bibRefCitation id="EFACFCBEC745A30CFF33F97EFECB33FF" author="Chujo, M." box="[233,371,1699,1725]" pageId="6" pageNumber="537" pagination="25 - 34" refId="ref6976" refString="Chujo, M. (1966) Coleoptera from Southeast Asia (V). 5. Family Scarabaeidae (2). Memoirs of the Faculty of Education, Kagawa University, Part II (No. 140), 25 - 34." type="book chapter" year="1966">Chûjô, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
is synonymized here.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF0FF91AFA8B320F" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
<bibRefCitation id="EFACFCBEC745A30CFF0FF91AFED933A3" author="Frey, G." box="[213,353,1735,1761]" pageId="6" pageNumber="537" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey (1970)</bibRefCitation>
|
||
named a subspecies
|
||
<taxonomicName id="4C3DFACCC745A30CFDBDF91AFC3533A2" authorityName="Frey" authorityYear="1970" box="[615,909,1735,1761]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="guttata">
|
||
<emphasis id="B9495D5DC745A30CFDBDF91AFC3533A2" box="[615,909,1735,1761]" italics="true" pageId="6" pageNumber="537">L. nigromaculata guttata</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F32AC1DFC745A30CFB97F91AFB3F33A3" box="[1101,1159,1735,1761]" name="Laos" pageId="6" pageNumber="537">Laos</collectingCountry>
|
||
. Based on the original description and the examination of the
|
||
<typeStatus id="54863FEDC745A30CFD92F936FDC03247" box="[584,632,1771,1797]" pageId="6" pageNumber="537">type</typeStatus>
|
||
specimens (
|
||
<figureCitation id="13069DCAC745A30CFCD8F936FCC43247" box="[770,892,1771,1797]" captionStart="FIGURES 7–16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[264,1323,329,1857]" captionTargetId="figure-95@4.[235,1352,309,1880]" captionTargetPageId="4" captionText="FIGURES 7–16. Type specimens of Parastasia (Lutera) nigromaculata guttata Frey, 1970. 7–10, lectotype (in dorsal, ventral, and frontal view) and labels (photograph by James H. Boone, BPBM); 11–13, paralectotype (in dorsal and lateral view) and labels (photograph by Christopher German, NHMB); 14–16, paralectotype (in dorsal and lateral view) and labels (photograph by James H. Boone, BPBM)." figureDoi="http://doi.org/10.5281/zenodo.13949863" httpUri="https://zenodo.org/record/13949863/files/figure.png" pageId="6" pageNumber="537">Figs. 7–16</figureCitation>
|
||
), this subspecies is undoubtedly a synonym of
|
||
<taxonomicName id="4C3DFACCC745A30CFA5CF936FE35326B" authority="Ohaus" authorityName="Ohaus" authorityYear="1900" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC745A30CFA5CF936FE83326B" italics="true" pageId="6" pageNumber="537">L. nigromaculata</emphasis>
|
||
Ohaus
|
||
</taxonomicName>
|
||
, as the putative distinctive features stated by Frey,
|
||
<emphasis id="B9495D5DC745A30CFC0BF8D2FC4C326A" box="[977,1012,1807,1832]" italics="true" pageId="6" pageNumber="537">i.e.</emphasis>
|
||
, larger maculae on the dorsal surface and an additional black line along the elytral suture, are absolutely identical to those of
|
||
<taxonomicName id="4C3DFACCC745A30CFBB1F8EEFA94320F" authorityName="Ohaus" authorityYear="1900" box="[1131,1324,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="nigromaculata">
|
||
<emphasis id="B9495D5DC745A30CFBB1F8EEFA94320F" box="[1131,1324,1843,1869]" italics="true" pageId="6" pageNumber="537">L. nigromaculata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B82814FC745A30CFF1DF88AFC3E329F" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="537">
|
||
This beetle was previously known to occur only in
|
||
<collectingCountry id="F32AC1DFC745A30CFCD9F88AFCDF3233" box="[771,871,1879,1905]" name="Thailand" pageId="6" pageNumber="537">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32AC1DFC745A30CFC47F88AFC6B3233" box="[925,979,1879,1905]" name="Laos" pageId="6" pageNumber="537">Laos</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFACFCBEC745A30CFC38F88AFBD23233" author="Chujo, M." box="[994,1130,1879,1905]" pageId="6" pageNumber="537" pagination="25 - 34" refId="ref6976" refString="Chujo, M. (1966) Coleoptera from Southeast Asia (V). 5. Family Scarabaeidae (2). Memoirs of the Faculty of Education, Kagawa University, Part II (No. 140), 25 - 34." type="book chapter" year="1966">Chûjô 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC745A30CFBACF88AFB533233" author="Frey, G." box="[1142,1259,1879,1905]" pageId="6" pageNumber="537" pagination="170 - 183" refId="ref7097" refString="Frey, G. (1970) Neue Ruteliden aus Indochina (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 170 - 183." type="journal article" year="1970">Frey 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC745A30CFB2DF88AFAC03233" author="Wada, K." box="[1271,1400,1879,1905]" pageId="6" pageNumber="537" pagination="1 - 19" refId="ref8013" refString="Wada, K. (1988) Notes on the genus Parastasia Westwood, 1841 (Rutelinae, Scarabaeidae, Col.). Lamellicornia, 4, 1 - 19." type="journal article" year="1988">Wada 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFACFCBEC745A30CFA59F88AFEAA32D7" author="Li, J. K. & Keith, D. & Gao, M. X. & Lin, L." pageId="6" pageNumber="537" pagination="273 - 276" refId="ref7447" refString="Li, J. K., Keith, D., Gao, M. X. & Lin, L. (2012) Coleopteres Scarabaeoidea de Pha Ngam (province de Bokeo, Laos). L'Entomologiste, 68 (5), 273 - 276." type="journal article" year="2012">
|
||
Li
|
||
<emphasis id="B9495D5DC745A30CFF4DF8A6FF7732D7" box="[151,207,1915,1941]" italics="true" pageId="6" pageNumber="537">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). According to the distribution map drawn by
|
||
<bibRefCitation id="EFACFCBEC745A30CFCD5F8A6FC1932D7" author="Wada, K." box="[783,929,1915,1941]" pageId="6" pageNumber="537" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada (1998)</bibRefCitation>
|
||
, it is widely distributed in Indo-China as well as
|
||
<collectingCountry id="F32AC1DFC745A30CFF6EF842FF4F32FB" box="[180,247,1951,1977]" name="Nepal" pageId="6" pageNumber="537">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="13069DCAC745A30CFEDCF842FEE132FB" box="[262,345,1951,1977]" captionStart="FIGURE 46" captionStartId="9.[151,250,1970,1994]" captionTargetBox="[164,1423,1173,1947]" captionTargetId="figure-488@9.[164,1423,1173,1947]" captionTargetPageId="9" captionText="FIGURE 46. Known distribution of Lutera nigromaculata Ohaus, 1900 (black dots) and L. nigrita Ohaus, 1930 (black triangles). Type localities are underlined, and the open dot in Nepal is a record from Wada (1998) without precise location." figureDoi="http://doi.org/10.5281/zenodo.13949872" httpUri="https://zenodo.org/record/13949872/files/figure.png" pageId="6" pageNumber="537">Fig. 46</figureCitation>
|
||
). However, no precise localities have been provided by Wada. This species is newly recorded from South
|
||
<collectingCountry id="F32AC1DFC745A30CFF05F81EFE86329F" box="[223,318,1987,2013]" name="Vietnam" pageId="6" pageNumber="537">Vietnam</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32AC1DFC745A30CFEAEF81EFE16329F" box="[372,430,1987,2013]" name="India" pageId="6" pageNumber="537">India</collectingCountry>
|
||
here with detailed collecting information.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |