266 lines
54 KiB
XML
266 lines
54 KiB
XML
<document id="70252B1072182B591DFB653B8E9EDA77" ID-CLB-Dataset="264211" ID-DOI="10.5852/ejt.2023.889.2245" ID-GBIF-Dataset="db71145d-18b1-4ad1-a2a4-72d7b8928a7d" ID-ISSN="2118-9773" ID-Zenodo-Dep="8269451" ID-ZooBank="52E11625-6E26-4E4F-8B15-05195087BAB4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692626252704" checkinUser="felipe" docAuthor="Domahovski, Alexandre C. & Cavichioli, Rodney R." docDate="2023" docId="66347308575FFF8DFDF695B0FC2CFE47" docLanguage="en" docName="ejt-889_1-53.pdf" docOrigin="European Journal of Taxonomy 889 (1)" docSource="http://zoobank.org/52e11625-6e26-4e4f-8b15-05195087bab4" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costanana obtusa Domahovski & Cavichioli 2023, sp. nov." docType="treatment" docUuid="6C9FE23A-E57D-440D-8C41-93B9E2934678" docUuidSource="ZooBank" docVersion="8" lastPageNumber="18" masterDocId="9A0D0B70574FFF9EFF8A935BFF99FFB7" masterDocTitle="Eight new species of Costanana DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov." masterLastPageNumber="53" masterPageNumber="1" pageNumber="17" updateTime="1699595446126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ADB9D727969EBE5E1E9E418CFD13831F">Eight new species of Costanana DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov.</mods:title>
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<mods:name id="EF9604209FE577CE0E6A79349BD6EEAA" type="personal">
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<mods:roleTerm id="00309E812FF3901A0B49A7A13A1349F7">Author</mods:roleTerm>
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<mods:namePart id="8B4789E571C2DA23BD56D76C74EA748A">Domahovski, Alexandre C.</mods:namePart>
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<mods:affiliation id="06EAAFE4F831392D0994C9DEF765A24E">Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="DF5E088E4013AB66DFFCB9ADCFB9228C" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="7BF8C2B1DB46180A586192B7A8904A12">Cavichioli, Rodney R.</mods:namePart>
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<mods:affiliation id="058859E3668B171995A3522FEB61AE44">Departamento de Zoologia, Universidade Federal do Paraná. Caixa Postal 19020, Curitiba, 81531 - 980, PR, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="2163BCB3D8273B43F8B24F17A4FDAE3E" type="email">cavich@ufpr.br</mods:nameIdentifier>
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<mods:title id="8ADD48DCE6B55CB96C688822A3811AB7">European Journal of Taxonomy</mods:title>
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<mods:part id="F32C07A19D9A695565C641D6BD4446AE">
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<mods:date id="A8E171EB7FED5426F369B74DE734BFCE">2023</mods:date>
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<mods:number id="357C01E8E4F3D8E96B78FAEFF98DEFF1">2023-08-21</mods:number>
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<treatment id="66347308575FFF8DFDF695B0FC2CFE47" ID-DOI="http://doi.org/10.5281/zenodo.8271507" ID-GBIF-Taxon="212449071" ID-Zenodo-Dep="8271507" LSID="urn:lsid:zoobank.org:act:6C9FE23A-E57D-440D-8C41-93B9E2934678" httpUri="http://treatment.plazi.org/id/66347308575FFF8DFDF695B0FC2CFE47" lastPageId="19" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="A6879195575FFF8EFDF695B0FC21F8B1" box="[636,952,1771,1798]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="EE22C21E575FFF8EFDF695B0FC21F8B1" blockId="16.[370,1217,1771,1865]" box="[636,952,1771,1798]" pageId="16" pageNumber="17">
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<heading id="B56A7572575FFF8EFDF695B0FC21F8B1" box="[636,952,1771,1798]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
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<taxonomicName id="299DB99D575FFF8EFDF695B0FCCFF8B1" ID-CoL="130be42e-788c-46f9-83fb-cc59b22d50a2" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[636,854,1771,1798]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="obtusa" status="sp. nov.">
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<emphasis id="DCE91E0C575FFF8EFDF695B0FCCFF8B1" bold="true" box="[636,854,1771,1798]" italics="true" pageId="16" pageNumber="17">Costanana obtusa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C7DAA377575FFF8EFCD795B7FC21F8B1" box="[861,952,1772,1798]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="A6879195575FFF8EFEF89456FCE9F8FE" pageId="16" pageNumber="17" type="description">
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<uri id="9A0CCE1C575FFF8EFEF89456FB58F89F" box="[370,1217,1805,1832]" pageId="16" pageNumber="17">
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urn:lsid:zoobank.org:act:
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<uuid id="9A3BF8CB575FFF8EFD1B9456FB58F89F" box="[657,1217,1805,1832]" pageId="16" pageNumber="17">6C9FE23A-E57D-440D-8C41-93B9E2934678</uuid>
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</uri>
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</paragraph>
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<paragraph id="EE22C21E575FFF8EFD4E9474FCE9F8FE" blockId="16.[370,1217,1771,1865]" box="[708,880,1839,1865]" pageId="16" pageNumber="17">
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<figureCitation id="76A6DE9B575FFF8EFD4E9474FC96F8FE" box="[708,783,1839,1865]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="16" pageNumber="17">Figs 7</figureCitation>
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,
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<figureCitation id="76A6DE9B575FFF8EFC919474FCE9F8FE" box="[795,880,1839,1865]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="16" pageNumber="17">12A–B</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A6879195575FFF8FFF379435FC71FED0" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="EE22C21E575FFF8EFF379435FEAFF83F" blockId="16.[189,310,1902,1928]" box="[189,310,1902,1928]" pageId="16" pageNumber="17">
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<heading id="B56A7572575FFF8EFF379435FEAFF83F" bold="true" box="[189,310,1902,1928]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
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<emphasis id="DCE91E0C575FFF8EFF379435FEAFF83F" bold="true" box="[189,310,1902,1928]" pageId="16" pageNumber="17">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575FFF8FFF3794C7FC71FED0" blockId="16.[189,1400,1948,2009]" lastBlockId="17.[189,1398,265,360]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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Coloration (
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<figureCitation id="76A6DE9B575FFF8EFEC694C7FE20F801" box="[332,441,1948,1974]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="16" pageNumber="17">Fig. 12A</figureCitation>
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) of head and pronotum brown with black maculae, forewing costal margin with broad light yellow macula on anterior third and large depigmented (white) macula on apical third. Style (
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<figureCitation id="76A6DE9B575EFF8FFF4C9252FEBCFE94" box="[198,293,265,291]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7K</figureCitation>
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) with ventral dentiform process near base of blade; apex truncated. Aedeagus (
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<figureCitation id="76A6DE9B575EFF8FFB639252FAF7FE94" box="[1257,1390,265,291]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7L–M</figureCitation>
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) without processes near base; shaft slender with pair of long filiform subapical processes directed basally, with one or two spurs near base; apex of shaft spatulate and rounded.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6879195575EFF8FFF3792D5FB11FE62" box="[189,1160,398,470]" pageId="17" pageNumber="18" type="etymology">
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<paragraph id="EE22C21E575EFF8FFF3792D5FEDDFE1F" blockId="17.[189,324,398,424]" box="[189,324,398,424]" pageId="17" pageNumber="18">
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<heading id="B56A7572575EFF8FFF3792D5FEDDFE1F" bold="true" box="[189,324,398,424]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="DCE91E0C575EFF8FFF3792D5FEDDFE1F" bold="true" box="[189,324,398,424]" pageId="17" pageNumber="18">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575EFF8FFF3792E0FB11FE62" blockId="17.[189,1160,443,470]" box="[189,1160,443,470]" pageId="17" pageNumber="18">
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The species epithet ‘
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<taxonomicName id="299DB99D575EFF8FFE2592E0FE64FE62" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[431,509,443,469]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="DCE91E0C575EFF8FFE2592E0FE64FE62" box="[431,509,443,469]" italics="true" pageId="17" pageNumber="18">obtusa</emphasis>
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</taxonomicName>
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’ refers to the unusual apex of the style being truncated.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6879195575EFF8FFF3792A0FB55FDD1" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="EE22C21E575EFF8FFF3792A0FE33FDA2" blockId="17.[189,426,507,533]" box="[189,426,507,533]" pageId="17" pageNumber="18">
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<heading id="B56A7572575EFF8FFF3792A0FE33FDA2" bold="true" box="[189,426,507,533]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="DCE91E0C575EFF8FFF3792A0FE33FDA2" bold="true" box="[189,426,507,533]" pageId="17" pageNumber="18">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575EFF8FFF609172FB55FDD1" blockId="17.[189,1228,553,614]" pageId="17" pageNumber="18">
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<materialsCitation id="5EF5C843575EFF8FFF609172FB55FDD1" ID-GBIF-Occurrence="4399003346" collectingDate="1980-12" collectionCode="DZUP" collectorName="M. Alvarenga" country="Brazil" location="Encruzilhada" pageId="17" pageNumber="18" specimenCode="DZUP 215492" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="holotype">
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<emphasis id="DCE91E0C575EFF8FFF609172FEC5FDF4" bold="true" box="[234,348,553,579]" pageId="17" pageNumber="18">
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<typeStatus id="31267CBC575EFF8FFF609172FEC5FDF4" box="[234,348,553,579]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="968A828E575EFF8FFF379110FEBFFDD2" box="[189,294,587,613]" name="Brazil" pageId="17" pageNumber="18">BRAZIL</collectingCountry>
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•
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<specimenCount id="F89B0997575EFF8FFEB59117FEC1FDD1" box="[319,344,588,614]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
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; “Brasil,
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<collectingRegion id="2C590CFC575EFF8FFE4C9110FD94FDD2" box="[454,525,587,613]" country="Brazil" name="Bahia" pageId="17" pageNumber="18">Bahia</collectingRegion>
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, \
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<location id="EB4294C5575EFF8FFDA39110FD5DFDD2" LSID="urn:lsid:plazi:treatment:66347308575FFF8DFDF695B0FC2CFE47:EB4294C5575EFF8FFDA39110FD5DFDD2" box="[553,708,587,613]" country="Brazil" name="Encruzilhada" pageId="17" pageNumber="18" stateProvince="Bahia">Encruzilhada</location>
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\
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<date id="9A23E4DE575EFF8FFD509110FCDCFDD2" box="[730,837,587,613]" pageId="17" pageNumber="18" value="1980-12">
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<collectingDate id="8A671D36575EFF8FFD509110FCDCFDD2" box="[730,837,587,613]" pageId="17" pageNumber="18" value="1980-12">XII.1980</collectingDate>
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</date>
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\
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<collectorName id="4368A7C8575EFF8FFCD69117FB9AFDD2" box="[860,1027,587,614]" pageId="17" pageNumber="18">M. Alvarenga</collectorName>
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”;
|
||
<specimenCode id="BE3B6A65575EFF8FFB939117FB51FDD1" box="[1049,1224,587,614]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="17" pageNumber="18" type="University or college">DZUP 215492</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6879195575EFF8FFF3791D0FEA0F8FB" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="EE22C21E575EFF8FFF3791D0FED7FD12" blockId="17.[189,334,651,677]" box="[189,334,651,677]" pageId="17" pageNumber="18">
|
||
<heading id="B56A7572575EFF8FFF3791D0FED7FD12" bold="true" box="[189,334,651,677]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="DCE91E0C575EFF8FFF3791D0FED7FD12" bold="true" box="[189,334,651,677]" pageId="17" pageNumber="18">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF6091E2FEB3FD64" blockId="17.[189,808,697,758]" box="[234,298,697,723]" pageId="17" pageNumber="18">
|
||
<emphasis id="DCE91E0C575EFF8FFF6091E2FEB3FD64" bold="true" box="[234,298,697,723]" pageId="17" pageNumber="18">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF379187FCB1FD41" blockId="17.[189,808,697,758]" box="[189,808,731,758]" pageId="17" pageNumber="18">
|
||
<smallCapsWord id="E8C454C2575EFF8FFF379187FEF7FD43" baselines="751,751" box="[189,366,732,758]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="17" pageNumber="18">MEASUREMENTS</smallCapsWord>
|
||
.
|
||
<typeStatus id="31267CBC575EFF8FFEF79180FE73FD42" box="[381,490,731,757]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
|
||
male: total length
|
||
<quantity id="29656FFB575EFF8FFD4C9180FCBAFD41" box="[710,803,731,758]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="17" pageNumber="18" unit="mm" value="6.8">6.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF379040FAEEFBB5" blockId="17.[188,1399,795,1027]" pageId="17" pageNumber="18">
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||
<smallCapsWord id="E8C454C2575EFF8FFF379040FED1FC83" baselines="814,815" box="[189,328,795,821]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="17" pageNumber="18">COLORATION</smallCapsWord>
|
||
. Dorsal portion of head and thorax brown (
|
||
<figureCitation id="76A6DE9B575EFF8FFCF99040FB9AFC82" box="[883,1027,795,822]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Figs 7A, C</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B575EFF8FFB9F9040FBD2FC82" box="[1045,1099,795,821]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="17" pageNumber="18">12A</figureCitation>
|
||
); ventral portion yellow (
|
||
<figureCitation id="76A6DE9B575EFF8FFF4F9066FED1FCEF" box="[197,328,829,856]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Figs 7B–C</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B575EFF8FFEDF9065FE10FCEF" box="[341,393,830,856]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="17" pageNumber="18">12B</figureCitation>
|
||
). Crown (
|
||
<figureCitation id="76A6DE9B575EFF8FFE759066FDC5FCEF" box="[511,604,829,856]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7A</figureCitation>
|
||
) almost completely black, except median line and posterior margin brown; pair of rounded black spots behind ocelli, near posterior margin; eyes and ocelli red. Pronotum (
|
||
<figureCitation id="76A6DE9B575EFF8FFF4C90D9FEBBFC2B" box="[198,290,898,924]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7A</figureCitation>
|
||
) with black irregular maculae near anterior margin; lateral margins yellow. Face (
|
||
<figureCitation id="76A6DE9B575EFF8FFB5090D9FAAAFC2B" box="[1242,1331,898,924]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7B</figureCitation>
|
||
) with frons brownish. Forewing (
|
||
<figureCitation id="76A6DE9B575EFF8FFE7490FFFDC3FC09" box="[510,602,932,958]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7D</figureCitation>
|
||
) with large triangular light yellow macula on anterior half of costal margin and a large rounded depigmented (white) macula at level of outer anteapical cell, remaining portion of costal cell, apical cells, apex of clavus and appendix dark brown; apical margin yellow-brown.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF379772FB7FFB1E" blockId="17.[189,1398,1064,1193]" pageId="17" pageNumber="18">
|
||
<smallCapsWord id="E8C454C2575EFF8FFF379772FEA3FBF6" baselines="1084,1084" box="[189,314,1065,1091]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Structure" pageId="17" pageNumber="18">STRUCTURE</smallCapsWord>
|
||
. Head in dorsal view (
|
||
<figureCitation id="76A6DE9B575EFF8FFDEB9773FD5CFBF4" box="[609,709,1064,1091]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7A</figureCitation>
|
||
): median length of crown as long as interocular width; transocular width of head 8.2 tenths humeral width of pronotum; in lateral view (
|
||
<figureCitation id="76A6DE9B575EFF8FFBE59710FB52FBD2" box="[1135,1227,1099,1125]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7C</figureCitation>
|
||
), with anterior margin thick, with 7–8 well developed carinae. Forewing (
|
||
<figureCitation id="76A6DE9B575EFF8FFC109736FC65FB30" box="[922,1020,1133,1159]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7D</figureCitation>
|
||
) inner discal cell open, m-cu 2 crossvein absent; apex broadly rounded. Other characteristics as in the generic description.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF37978BFDD0F95E" blockId="17.[189,1401,1231,1770]" pageId="17" pageNumber="18">
|
||
<smallCapsWord id="E8C454C2575EFF8FFF37978BFF66FB5F" baselines="1251,1251" box="[189,255,1232,1258]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="17" pageNumber="18">MALE</smallCapsWord>
|
||
<smallCapsWord id="E8C454C2575EFF8FFE8F978EFE1FFB5F" baselines="1251" box="[261,390,1237,1256]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="17" pageNumber="18">TERMINALIA</smallCapsWord>
|
||
. Sternite VIII (
|
||
<figureCitation id="76A6DE9B575EFF8FFDBE9794FD13FB5E" box="[564,650,1231,1257]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7E</figureCitation>
|
||
) 1.1× as long as wide; lateral margins excavated at base, forming a triangular projection; apex rounded. Valve (
|
||
<figureCitation id="76A6DE9B575EFF8FFD6A97AAFCA3FABB" box="[736,826,1265,1292]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7F</figureCitation>
|
||
) 2.2× as wide as long; posterior margin almost straight and weakly emarginated medially. Pygofer (
|
||
<figureCitation id="76A6DE9B575EFF8FFCAB9648FCE6FA99" box="[801,895,1299,1326]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7G</figureCitation>
|
||
), in lateral view, 1.8× as long as maximum height; basodorsal process present but weakly developed; ventral margin with rounded protrusion near midlength; posterodorsal and ventral margins rounded; external surface near ventral margin with short filiform setae; macrosetae distributed on posterior half; apex broadly rounded. Subgenital plate (
|
||
<figureCitation id="76A6DE9B575EFF8FFF4C96C7FEBEFA01" box="[198,295,1436,1462]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7G</figureCitation>
|
||
), in lateral view, slightly surpassing half length of pygofer; in ventral view (
|
||
<figureCitation id="76A6DE9B575EFF8FFB4296C7FAB0FA01" box="[1224,1321,1436,1462]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7H</figureCitation>
|
||
), 3.5× as long as wide; inner margin straight; external margin slightly rounded on basal half; ventral surface and external margin with long filiform setae; apex tapered and weakly sclerotized. Connective (
|
||
<figureCitation id="76A6DE9B575EFF8FFA9596BBFAF4FA4C" box="[1311,1389,1504,1531]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7I</figureCitation>
|
||
) approximately linear; arms not produced laterally; stalk longer than maximum width of arms, widened apically; median keel weakly developed. Style (
|
||
<figureCitation id="76A6DE9B575EFF8FFD74957EFCCDF988" box="[766,852,1573,1599]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7J</figureCitation>
|
||
), in dorsal view, with outer lobe reduced and subtruncated; in lateral view (
|
||
<figureCitation id="76A6DE9B575EFF8FFDA0951CFD12F9D6" box="[554,651,1607,1633]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7K</figureCitation>
|
||
), blade broadened medially; ventral margin not serrated, with dentiform process near base of blade; apex truncated. Aedeagus (
|
||
<figureCitation id="76A6DE9B575EFF8FFC369532FBA6F934" box="[956,1087,1641,1667]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Fig. 7L–M</figureCitation>
|
||
) preatrium not developed; dorsal apodeme strongly elongated laterally, forming divergent and truncated apices; shaft slender, curved dorsally; pair of long filiform subapical processes directed basally, with one or two spurs near base; apex spatulate and rounded.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF60944BFEDCF89D" blockId="17.[189,325,1808,1868]" box="[234,325,1808,1834]" pageId="17" pageNumber="18">
|
||
<emphasis id="DCE91E0C575EFF8FFF60944BFEDCF89D" bold="true" box="[234,325,1808,1834]" pageId="17" pageNumber="18">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8FFF379469FEA0F8FB" blockId="17.[189,325,1808,1868]" box="[189,313,1842,1868]" pageId="17" pageNumber="18">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6879195575EFF8DFF379429FC2CFE47" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="EE22C21E575EFF8FFF379429FEB6F83B" blockId="17.[189,303,1906,1932]" box="[189,303,1906,1932]" pageId="17" pageNumber="18">
|
||
<heading id="B56A7572575EFF8FFF379429FEB6F83B" bold="true" box="[189,303,1906,1932]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="DCE91E0C575EFF8FFF379429FEB6F83B" bold="true" box="[189,303,1906,1932]" pageId="17" pageNumber="18">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575EFF8DFF3794FBFC2CFE47" blockId="17.[189,1399,1952,2012]" lastBlockId="19.[188,1399,265,496]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
|
||
<taxonomicName id="299DB99D575EFF8FFF3794FBFE0BF80D" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[189,402,1952,1978]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obtusa" status="sp. nov.">
|
||
<emphasis id="DCE91E0C575EFF8FFF3794FBFE0BF80D" box="[189,402,1952,1978]" italics="true" pageId="17" pageNumber="18">Costanana obtusa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7DAA377575EFF8FFE1094FBFE6CF80D" box="[410,501,1952,1978]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="299DB99D575EFF8FFD4594FBFC0DF80E" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[719,916,1952,1978]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="piraquarensis" status="sp. nov.">
|
||
<emphasis id="DCE91E0C575EFF8FFD4594FBFC0DF80E" box="[719,916,1952,1978]" italics="true" pageId="17" pageNumber="18">C. piraquarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7DAA377575EFF8FFC1194FBFC6FF80D" box="[923,1014,1952,1978]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having a similar color pattern (
|
||
<figureCitation id="76A6DE9B575EFF8FFF4C9499FED6F86B" box="[198,335,1986,2012]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="17" pageNumber="18">Fig.12A–D</figureCitation>
|
||
), especially the pair of large maculae on the costal margin of the forewing (
|
||
<figureCitation id="76A6DE9B575EFF8FFB529499FAA2F86B" box="[1240,1339,1986,2012]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="17" pageNumber="18">Figs 7D</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B575EFF8FFAC39499FAF7F86B" box="[1353,1390,1986,2012]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-19@20.[189,1398,268,1810]" captionTargetPageId="20" captionText="Fig. 8. Costanana piraquarensis sp. nov., holotype, ♂ (DZUP 215485). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269475" httpUri="https://zenodo.org/record/8269475/files/figure.png" pageId="17" pageNumber="18">8D</figureCitation>
|
||
) and the general shape of the pygofer, style, connective and aedeagus (
|
||
<figureCitation id="76A6DE9B575CFF8DFC739252FB1CFE94" box="[1017,1157,265,291]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Figs 7G–M</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B575CFF8DFB1B9252FB7AFE94" box="[1169,1251,265,291]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-19@20.[189,1398,268,1810]" captionTargetPageId="20" captionText="Fig. 8. Costanana piraquarensis sp. nov., holotype, ♂ (DZUP 215485). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269475" httpUri="https://zenodo.org/record/8269475/files/figure.png" pageId="19" pageNumber="20">8G–M</figureCitation>
|
||
).
|
||
<taxonomicName id="299DB99D575CFF8DFB729252FE92FEF2" authorityName="Domahovski & Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obtusa">
|
||
<emphasis id="DCE91E0C575CFF8DFB729252FE92FEF2" italics="true" pageId="19" pageNumber="20">Costanana obtusa</emphasis>
|
||
</taxonomicName>
|
||
is separated from
|
||
<taxonomicName id="299DB99D575CFF8DFE7E9277FD25FEF2" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[500,700,300,326]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="piraquarensis">
|
||
<emphasis id="DCE91E0C575CFF8DFE7E9277FD25FEF2" box="[500,700,300,326]" italics="true" pageId="19" pageNumber="20">C. piraquarensis</emphasis>
|
||
</taxonomicName>
|
||
in having the following differences: (1) apex of pygofer rounded (
|
||
<figureCitation id="76A6DE9B575CFF8DFEA79216FE12FED0" box="[301,395,333,359]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Fig. 7G</figureCitation>
|
||
); (2) connective longer (
|
||
<figureCitation id="76A6DE9B575CFF8DFD219216FD64FED0" box="[683,765,333,359]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Fig. 7I</figureCitation>
|
||
); (3) style with apex truncated (
|
||
<figureCitation id="76A6DE9B575CFF8DFBF99216FB48FED0" box="[1139,1233,333,359]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Fig. 7K</figureCitation>
|
||
); (4) aedeagal shaft slenderer (
|
||
<figureCitation id="76A6DE9B575CFF8DFEF09234FD99FE3D" box="[378,512,367,394]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Fig. 7L–M</figureCitation>
|
||
); and (5) subapical pair of processes having one or two spurs near the base (
|
||
<figureCitation id="76A6DE9B575CFF8DFF4C92CAFEB0FE1B" box="[198,297,401,428]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1397,268,1766]" captionTargetId="figure-19@18.[189,1398,266,1771]" captionTargetPageId="18" captionText="Fig. 7. Costanana obtusa sp. nov., holotype, ♂ (DZUP 215492). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269473" httpUri="https://zenodo.org/record/8269473/files/figure.png" pageId="19" pageNumber="20">Fig. 7M</figureCitation>
|
||
). Unfortunately the legs of the
|
||
<typeStatus id="31267CBC575CFF8DFD1A92CAFD6EFE1C" box="[656,759,401,427]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
are missing but, due to the distinctly different genitalia and the lack of importance for species delimitation of the morphology of the legs, we suppose this does not prejudge the description or future recognition of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |