treatments-xml/data/25/2C/87/252C87918B332C09FF20FE1ABA213DDE.xml

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<mods:title id="162583112D193CA83E7F45B4C673B373">Descriptions of two new shell-dwelling species of Metriaclima (Cichlidae) from Lake Malaŵi, Africa</mods:title>
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<mods:namePart id="A2BC248388E3D34BF64D7FFF60A036C5">Miller, Torin A.</mods:namePart>
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<mods:namePart id="5EA4D80AD8C9340BD573ACFE83A709B1">Konings, Adrianus F.</mods:namePart>
<mods:affiliation id="5FDC1DE52EF27DFEF0AA2AB880B25D18">Cichlid Press, PO Box 13608, El Paso, TX 79913.</mods:affiliation>
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<mods:namePart id="D4924CE89560B7299BDBC496C6D8D734">Stauffer, Jay R.</mods:namePart>
<mods:affiliation id="1AD2A5DB48C552D0C37C2C9DB57D60FE">Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802. &amp; Honorary Research Associate, South African Institute for Aquatic Biodiversity, Makhanda, RSA.</mods:affiliation>
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<heading id="F67281EB8B332C0CFF20FE1AB9033E83" bold="true" box="[151,550,402,429]" fontSize="11" level="1" pageId="3" pageNumber="422" reason="1">
<emphasis id="9FF1EA958B332C0CFF20FE1AB9033E83" bold="true" box="[151,550,402,429]" pageId="3" pageNumber="422">
<taxonomicName id="6A854D048B332C0CFF20FE1ABAAD3E82" authority="Miller &amp; Konings &amp; Stauffer, 2021" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[151,392,402,429]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae" status="sp. nov.">
<emphasis id="9FF1EA958B332C0CFF20FE1ABAAD3E82" bold="true" box="[151,392,402,429]" italics="true" pageId="3" pageNumber="422">Metriaclima ngarae</emphasis>
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, 
<taxonomicNameLabel id="84C257EE8B332C0CFE21FE1BB9033E83" box="[406,550,402,429]" pageId="3" pageNumber="422" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="35BE2A028B332C0CFF20FE3FBA273EFE" box="[151,258,439,465]" captionStart="FIGURE 2" captionStartId="3.[151,250,1958,1982]" captionTargetBox="[151,1436,1295,1933]" captionTargetId="figure-403@3.[151,1436,1295,1933]" captionTargetPageId="3" captionText="FIGURE 2. Metriaclima ngarae. Holotype, PSU 13365, adult male, 59.1 mm SL, Ngara, Lake Malaŵi." figureDoi="http://doi.org/10.5281/zenodo.5572222" httpUri="https://zenodo.org/record/5572222/files/figure.png" pageId="3" pageNumber="422">Figures 2</figureCitation>
, 
<figureCitation id="35BE2A028B332C0CFEB9FE3FBA383EFE" box="[270,285,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1190,1214]" captionTargetBox="[167,1420,181,1166]" captionTargetId="figure-30@4.[167,1420,181,1166]" captionTargetPageId="4" captionText="FIGURE 3. CT scan of the cranium of Metriaclima ngarae Holotype, PSU 13365, adult male, 59.1 mm SL, Ngara, Lake Malaŵi; angle of ethmovomerine block is 46.4° with parasphenoid." figureDoi="http://doi.org/10.5281/zenodo.5572224" httpUri="https://zenodo.org/record/5572224/files/figure.png" pageId="3" pageNumber="422">3</figureCitation>
, 
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; 
<tableCitation id="E007033C8B332C0CFEF1FE3FBABC3EFE" box="[326,409,439,465]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1351,228,2023]" captionTargetPageId="5" captionText="TABLE 1. Morphological and meristic data for Metriaclima ngarae from Ngara, Lake Malaŵi, PSU 13365, 13366, n=14. Ranges include holotype." httpUri="http://table.plazi.org/id/F9FA660F8B352C0AFF20FF1FBA833FFC" pageId="3" pageNumber="422" tableUuid="F9FA660F8B352C0AFF20FF1FBA833FFC">Table 1</tableCitation>
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<taxonomicName id="6A854D048B332C0CFF20FE73BA8E3D3C" authorityName="Burgess" authorityYear="1976" box="[151,427,507,531]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="lanisticola">
<emphasis id="9FF1EA958B332C0CFF20FE73BA8E3D3C" box="[151,427,507,531]" italics="true" pageId="3" pageNumber="422">Pseudotropheus lanisticola</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B332C0CFDA7FE73B9BA3D3C" author="Konings, A. F." box="[528,671,507,531]" pageId="3" pageNumber="422" refId="ref6746" refString="Konings, A. F. (1989) Malawi cichlids in their natural habitat. Verduijn Cichlids &amp; Lake Fish Movies, Zevenhuizen, 304 pp." type="book" year="1989">Konings 1989</bibRefCitation>
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<taxonomicName id="6A854D048B332C0CFF20FD92BA973D1D" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[151,434,538,562]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="livingstonii">
<emphasis id="9FF1EA958B332C0CFF20FD92BA973D1D" box="[151,434,538,562]" italics="true" pageId="3" pageNumber="422">Pseudotropheus livingstonii</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B332C0CFDA0FD92B9833D1D" author="Konings, A. F." box="[535,678,538,562]" pageId="3" pageNumber="422" refId="ref6774" refString="Konings, A. F. (1995) Malaw ˆ i cichlids in their natural habitat. 2 nd Edition. Cichlid Press, El Paso, Texas, 303 pp." type="book" year="1995">Konings 1995</bibRefCitation>
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<taxonomicName id="6A854D048B332C0CFF20FDB0BAA83D7F" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[151,397,568,592]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="livingstonii">
<emphasis id="9FF1EA958B332C0CFF20FDB0BAA83D7F" box="[151,397,568,592]" italics="true" pageId="3" pageNumber="422">Metriaclima livingstonii</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B332C0CFE45FDB0B9A43D7E" author="Konings, A." box="[498,641,568,593]" pageId="3" pageNumber="422" refId="ref6807" refString="Konings, A. (2001) Malawi cichlids in their natural habitat. 3 rd Edition. Cichlid Press, El Paso, Texas, 352 pp." type="book" year="2001">Konings 2001</bibRefCitation>
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<taxonomicName id="6A854D048B332C0CFF20FDDFBA1D3D40" box="[151,312,599,623]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="9FF1EA958B332C0CFF20FDDFBA313D40" box="[151,276,599,623]" italics="true" pageId="3" pageNumber="422">Metriaclima</emphasis>
sp.
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‘lanisticola north’ 
<bibRefCitation id="C9144B768B332C0CFE40FDDFB9A33D40" author="Konings, A." box="[503,646,599,623]" pageId="3" pageNumber="422" refId="ref6836" refString="Konings, A. (2007) Malawi cichlids in their natural habitat. 4 th Edition. Cichlid Press, El Paso, Texas, 424 pp." type="book" year="2007">Konings 2007</bibRefCitation>
</paragraph>
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<typeStatus id="723E88258B332C0CFF20FD15BA233D98" box="[151,262,669,695]" pageId="3" pageNumber="422" type="holotype">Holotype</typeStatus>
.
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13365, adult male, 
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SL, (
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, 
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, 
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<collectingDate id="C97FE9AF8B332C0CFF20FD4ABA233DF4" box="[151,262,705,732]" pageId="3" pageNumber="422" value="2000-01">Jan. 2000</collectingDate>
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, 
<collectorName id="007053518B332C0CFEA6FD49BAA53DF4" box="[273,384,705,731]" pageId="3" pageNumber="422">S. Grant’s</collectorName>
crew.
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</paragraph>
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<typeStatus id="723E88258B332C0CFF70FD6DBA643DD0" box="[199,321,741,767]" pageId="3" pageNumber="422" type="paratype">Paratypes</typeStatus>
.
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<collectionCode id="CB94AE428B332C0CFEFAFD6DBAA53DD0" box="[333,384,741,767]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="3" pageNumber="422" type="Museum">PSU</collectionCode>
13366, 
<specimenCode id="FD239EFC8B332C0CFE6CFD6DBADC3DD0" box="[475,505,741,767]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="3" pageNumber="422" type="Museum">14</specimenCode>
, (
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– 
<quantity id="6A7D9B628B332C0CFD36FD6DB9C13C2F" box="[641,740,741,768]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" pageId="3" pageNumber="422" unit="mm" value="62.4">62.4 mm</quantity>
SL), data as for holotype
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.
</paragraph>
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<subSubSection id="E59F650C8B332C0CFF70FC81B87D3BD8" pageId="3" pageNumber="422" type="diagnosis">
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<emphasis id="9FF1EA958B332C0CFF70FC81BA673C0C" bold="true" box="[199,322,777,803]" pageId="3" pageNumber="422">Diagnosis.</emphasis>
The moderately sloped vomer (45.4º in 
<typeStatus id="723E88258B332C0CFCA5FC81B8533C0B" box="[786,886,777,804]" pageId="3" pageNumber="422" type="holotype">holotype</typeStatus>
) with a swollen rostral tip (
<figureCitation id="35BE2A028B332C0CFB01FC81BFDF3C0C" box="[1206,1274,777,803]" captionStart="FIGURE 3" captionStartId="4.[151,250,1190,1214]" captionTargetBox="[167,1420,181,1166]" captionTargetId="figure-30@4.[167,1420,181,1166]" captionTargetPageId="4" captionText="FIGURE 3. CT scan of the cranium of Metriaclima ngarae Holotype, PSU 13365, adult male, 59.1 mm SL, Ngara, Lake Malaŵi; angle of ethmovomerine block is 46.4° with parasphenoid." figureDoi="http://doi.org/10.5281/zenodo.5572224" httpUri="https://zenodo.org/record/5572224/files/figure.png" pageId="3" pageNumber="422">Fig. 3</figureCitation>
) and bicuspid teeth in the anterior portion of the outer row of both upper and lower jaws place this species in 
<taxonomicName id="6A854D048B332C0CFB6FFCA5BE473C68" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[1240,1378,813,839]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="genus">
<emphasis id="9FF1EA958B332C0CFB6FFCA5BE473C68" box="[1240,1378,813,839]" italics="true" pageId="3" pageNumber="422">Metriaclima</emphasis>
</taxonomicName>
. The presence of a black submarginal band (
<figureCitation id="35BE2A028B332C0CFDE3FCD9B9BD3C44" box="[596,664,849,875]" captionStart="FIGURE 4" captionStartId="6.[151,250,1763,1787]" captionTargetBox="[151,1436,777,1740]" captionTargetId="figure-282@6.[151,1436,777,1740]" captionTargetPageId="6" captionText="FIGURE 4. A male Metriaclima ngarae at a submerged reef near Ngara (10.233397 S, 34.106592 E)." figureDoi="http://doi.org/10.5281/zenodo.5572226" httpUri="https://zenodo.org/record/5572226/files/figure.png" pageId="3" pageNumber="422">Fig. 4</figureCitation>
) in the dorsal and anal fin distinguishes 
<taxonomicName id="6A854D048B332C0CFBD0FCDABFF83C44" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1127,1245,850,875]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFBD0FCDABFF83C44" box="[1127,1245,850,875]" italics="true" pageId="3" pageNumber="422">M. ngarae</emphasis>
</taxonomicName>
from most other species in 
<taxonomicName id="6A854D048B332C0CFEBEFCFDBAB63CA0" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[265,403,885,911]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="genus">
<emphasis id="9FF1EA958B332C0CFEBEFCFDBAB63CA0" box="[265,403,885,911]" italics="true" pageId="3" pageNumber="422">Metriaclima</emphasis>
</taxonomicName>
, except for 
<taxonomicName id="6A854D048B332C0CFDA3FCFEBF6D3CBF" authority="(Stauffer, Bowers, Kellogg, &amp; McKaye)" baseAuthorityName="Stauffer, Bowers, Kellogg, &amp; McKaye" box="[532,1096,885,912]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="phaeos">
<emphasis id="9FF1EA958B332C0CFDA3FCFEB9A23CA0" box="[532,647,885,911]" italics="true" pageId="3" pageNumber="422">M. phaeos</emphasis>
(Stauffer, Bowers, Kellogg, &amp; McKaye)
</taxonomicName>
, 
<taxonomicName id="6A854D048B332C0CFBE3FCFEBA373C9C" authority="Stauffer, Black, &amp; Konings" authorityName="Stauffer, Black, &amp; Konings" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="lundoense">
<emphasis id="9FF1EA958B332C0CFBE3FCFEBFCF3CA0" box="[1108,1258,885,911]" italics="true" pageId="3" pageNumber="422">M. lundoense</emphasis>
Stauffer, Black, &amp; Konings
</taxonomicName>
, 
<taxonomicName id="6A854D048B332C0CFE97FC12B8343C9C" authority="Stauffer, Black, &amp; Konings" authorityName="Stauffer, Black, &amp; Konings" box="[288,785,921,947]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="nigrodorsalis">
<emphasis id="9FF1EA958B332C0CFE97FC12BAFE3C9C" box="[288,475,921,947]" italics="true" pageId="3" pageNumber="422">M. nigrodorsalis</emphasis>
Stauffer, Black, &amp; Konings
</taxonomicName>
, 
<taxonomicName id="6A854D048B332C0CFCA8FC12BF263C9C" authority="Stauffer" authorityName="Stauffer" authorityYear="2018" box="[799,1027,921,947]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9FF1EA958B332C0CFCA8FC12B8863C9C" box="[799,931,921,947]" italics="true" pageId="3" pageNumber="422">M. koningsi</emphasis>
Stauffer
</taxonomicName>
, 
<taxonomicName id="6A854D048B332C0CFBA7FC12BF833C9C" authority="Miller &amp; Konings &amp; Stauffer, 2021" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1040,1190,921,947]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae" status="sp. nov.">
<emphasis id="9FF1EA958B332C0CFBA7FC12BF833C9C" box="[1040,1190,921,947]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84C257EE8B332C0CFB1BFC12BFCD3C9B" box="[1196,1256,922,948]" pageId="3" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
, and from some populations of 
<taxonomicName id="6A854D048B332C0CFE88FC36B9153CF7" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[319,560,957,984]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="zebra">
<emphasis id="9FF1EA958B332C0CFE88FC36BA873CF8" box="[319,418,957,983]" italics="true" pageId="3" pageNumber="422">M. zebra</emphasis>
(Boulenger)
</taxonomicName>
, 
<taxonomicName id="6A854D048B332C0CFD89FC36B82E3CF8" authority="Tawil" authorityName="Tawil" box="[574,779,957,983]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="pulpican">
<emphasis id="9FF1EA958B332C0CFD89FC36B9E03CF8" box="[574,709,957,983]" italics="true" pageId="3" pageNumber="422">M. pulpican</emphasis>
Tawil
</taxonomicName>
, and 
<taxonomicName id="6A854D048B332C0CFCFFFC36BFC83CF7" authority="Li, Konings, &amp; Stauffer. It" authorityName="Li, Konings, &amp; Stauffer. It" authorityYear="2016" box="[840,1261,957,984]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="usisyae">
<emphasis id="9FF1EA958B332C0CFCFFFC36B89A3CF8" box="[840,959,958,983]" italics="true" pageId="3" pageNumber="422">M. usisyae</emphasis>
Li, Konings, &amp; Stauffer. It
</taxonomicName>
differs from all of these except 
<taxonomicName id="6A854D048B332C0CFEFDFC6ABAC73CD4" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[330,482,993,1019]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B332C0CFEFDFC6ABAC73CD4" box="[330,482,993,1019]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
by the number of vertical bars below the dorsal fin which is 4-5 for 
<taxonomicName id="6A854D048B332C0CFB42FC6ABE4E3CD4" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1269,1387,994,1019]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFB42FC6ABE4E3CD4" box="[1269,1387,994,1019]" italics="true" pageId="3" pageNumber="422">M. ngarae</emphasis>
</taxonomicName>
and 6-10 for the other species or none at all for 
<taxonomicName id="6A854D048B332C0CFDCCFB8EB9DA3B30" authorityName="Stauffer" authorityYear="2018" box="[635,767,1029,1055]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9FF1EA958B332C0CFDCCFB8EB9DA3B30" box="[635,767,1029,1055]" italics="true" pageId="3" pageNumber="422">M. koningsi</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="6A854D048B332C0CFC05FB8EBF623B30" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[946,1095,1029,1055]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B332C0CFC05FB8EBF623B30" box="[946,1095,1029,1055]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
by a greater interorbital width (30.7–37.4% vs. 23.6–29.7% HL) and by a greater ratio of the snout length in the distance between snout tip and pelvic fin origin (28.7–32.8% vs. 20.6–27.8% SNP2). On average, the ratio of premaxillary pedicel length in snout length is smaller in 
<taxonomicName id="6A854D048B332C0CFEC0FBFABAC93BA4" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[375,492,1138,1163]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFEC0FBFABAC93BA4" box="[375,492,1138,1163]" italics="true" pageId="3" pageNumber="422">M. ngarae</emphasis>
</taxonomicName>
at 35.1% vs. 44.6 %SNL in 
<taxonomicName id="6A854D048B332C0CFC87FBFAB8E23BA4" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[816,967,1137,1163]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B332C0CFC87FBFAB8E23BA4" box="[816,967,1137,1163]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
(ranges 20.3–45.2 vs. 33.1–62.7 %SNL). Males of 
<taxonomicName id="6A854D048B332C0CFEB7FB1EBA563B80" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[256,371,1174,1199]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFEB7FB1EBA563B80" box="[256,371,1174,1199]" italics="true" pageId="3" pageNumber="422">M. ngarae</emphasis>
</taxonomicName>
have a color pattern similar to that of females but males of 
<taxonomicName id="6A854D048B332C0CFBB1FB1EBFB93B80" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1030,1180,1173,1199]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B332C0CFBB1FB1EBFB93B80" box="[1030,1180,1173,1199]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
are dark brown to blue without visible bar pattern. Female 
<taxonomicName id="6A854D048B332C0CFD91FB32B9BF3BFC" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[550,666,1210,1235]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFD91FB32B9BF3BFC" box="[550,666,1210,1235]" italics="true" pageId="3" pageNumber="422">M. ngarae</emphasis>
</taxonomicName>
have a similar color pattern to those of 
<taxonomicName id="6A854D048B332C0CFBEEFB32BFD53BFC" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1113,1264,1209,1235]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B332C0CFBEEFB32BFD53BFC" box="[1113,1264,1209,1235]" italics="true" pageId="3" pageNumber="422">M. gallireyae</emphasis>
</taxonomicName>
but the bars on the flank are less prominent and do not extend to the abdomen.
</paragraph>
</subSubSection>
<caption id="F9FA660F8B332C0CFF20F82EBF923891" ID-DOI="http://doi.org/10.5281/zenodo.5572222" ID-Zenodo-Dep="5572222" box="[151,1207,1958,1982]" httpUri="https://zenodo.org/record/5572222/files/figure.png" pageId="3" pageNumber="422" startId="3.[151,250,1958,1982]" targetBox="[151,1436,1295,1933]" targetPageId="3">
<paragraph id="AD3A36878B332C0CFF20F82EBF923891" blockId="3.[151,1207,1958,1982]" box="[151,1207,1958,1982]" pageId="3" pageNumber="422">
<emphasis id="9FF1EA958B332C0CFF20F82EBA363891" bold="true" box="[151,275,1958,1982]" pageId="3" pageNumber="422">FIGURE 2.</emphasis>
<taxonomicName id="6A854D048B332C0CFEAEF82EBAC03891" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[281,485,1958,1982]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B332C0CFEAEF82EBAC03891" box="[281,485,1958,1982]" italics="true" pageId="3" pageNumber="422">Metriaclima ngarae</emphasis>
</taxonomicName>
. Holotype, PSU 13365, adult male, 59.1 mm SL, Ngara, Lake Malaŵi.
</paragraph>
</caption>
<caption id="F9FA660F8B342C0BFF20FB2EB8613BCD" ID-DOI="http://doi.org/10.5281/zenodo.5572224" ID-Zenodo-Dep="5572224" httpUri="https://zenodo.org/record/5572224/files/figure.png" pageId="4" pageNumber="423" startId="4.[151,250,1190,1214]" targetBox="[167,1420,181,1166]" targetPageId="4">
<paragraph id="AD3A36878B342C0BFF20FB2EB8613BCD" blockId="4.[151,1437,1190,1250]" pageId="4" pageNumber="423">
<emphasis id="9FF1EA958B342C0BFF20FB2EBA333B91" bold="true" box="[151,278,1190,1214]" pageId="4" pageNumber="423">FIGURE 3.</emphasis>
CT scan of the cranium of 
<taxonomicName id="6A854D048B342C0BFDF7FB2EB82B3B91" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[576,782,1190,1214]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B342C0BFDF7FB2EB82B3B91" box="[576,782,1190,1214]" italics="true" pageId="4" pageNumber="423">Metriaclima ngarae</emphasis>
</taxonomicName>
Holotype, PSU 13365, adult male, 59.1 mm SL, Ngara, Lake Malaŵi; angle of ethmovomerine block is 46.4° with parasphenoid.
</paragraph>
</caption>
<subSubSection id="E59F650C8B342C09FF70FA98BAC03FD6" lastPageId="6" lastPageNumber="425" pageId="4" pageNumber="423" type="description">
<paragraph id="AD3A36878B342C0BFF70FA98B95B392D" blockId="4.[151,1437,1296,2043]" pageId="4" pageNumber="423">
<emphasis id="9FF1EA958B342C0BFF70FA98BA7C3A05" bold="true" box="[199,345,1296,1322]" pageId="4" pageNumber="423">Description.</emphasis>
Morphometric and meristic data in 
<tableCitation id="E007033C8B342C0BFD4DFA98B8753A05" box="[762,848,1296,1322]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1351,228,2023]" captionTargetPageId="5" captionText="TABLE 1. Morphological and meristic data for Metriaclima ngarae from Ngara, Lake Malaŵi, PSU 13365, 13366, n=14. Ranges include holotype." httpUri="http://table.plazi.org/id/F9FA660F8B352C0AFF20FF1FBA833FFC" pageId="4" pageNumber="423" tableUuid="F9FA660F8B352C0AFF20FF1FBA833FFC">Table 1</tableCitation>
. Small-sized mbuna, ovoid body (mean BD 38.7% SL) with greatest body depth at about 3 
<superScript id="5AF09BCF8B342C0BFDF8FABCB9783A6D" attach="left" box="[591,605,1332,1346]" fontSize="6" pageId="4" pageNumber="423">rd</superScript>
and 4 
<superScript id="5AF09BCF8B342C0BFD29FABCB98E3A6D" attach="left" box="[670,683,1332,1346]" fontSize="6" pageId="4" pageNumber="423">th</superScript>
dorsal-fin spine. Dorsal body profile with gradual curve downward posteriorly with highest point at 6 
<superScript id="5AF09BCF8B342C0BFDA6FAD0B93B3A49" attach="left" box="[529,542,1368,1382]" fontSize="6" pageId="4" pageNumber="423">th</superScript>
or 7 
<superScript id="5AF09BCF8B342C0BFDE7FAD0B9783A49" attach="left" box="[592,605,1368,1382]" fontSize="6" pageId="4" pageNumber="423">th</superScript>
dorsal-fin spine, more pronounced towards posterior end of dorsal fin and beginning caudal peduncle; ventral body profile nearly straight between pelvic fins and base of anal fin with slight upward curve to caudal fin. Dorsal head profile round with continuous curve between interorbital and dorsal-fin origin; short snout with isognathous jaws; teeth in upper and lower jaws in 2–3 rows (mode 3); teeth in outer row bicuspid, inner rows unicuspid or tricuspid.
</paragraph>
<paragraph id="AD3A36878B342C0BFF70F984BAC83869" blockId="4.[151,1437,1296,2043]" pageId="4" pageNumber="423">
Dorsal fin with XVI or XVII (mode XVII) spines and 7–8 (mode 8) rays. Anal fin with III spines and 8 rays. First 5–6 dorsal-fin spines gradually longer posteriorly with 6 
<superScript id="5AF09BCF8B342C0BFCF2F9B8B8773911" attach="left" box="[837,850,1584,1598]" fontSize="6" pageId="4" pageNumber="423">th</superScript>
spine about twice as long as first; last 11 spines only slightly greater in length posteriorly with last spine longest; soft dorsal fin with rounded to subacuminate tip, second or third ray longest, to approximately ¼ length caudal fin. Anal-fin spines progressively longer posteriorly; 5th or 6th anal-fin ray longest, tip to base of caudal fin in females and to ½ length of caudal fin in males, equal to or slightly further posterior than tip dorsal fin. Caudal fin subtruncate to emarginate. Pectoral fin rounded, paddle-shaped, short, extending to vertical through base of 8th or 9th dorsal-fin spine. Flank scales ctenoid with abrupt change to small scales on breast and belly; 30–32 (mode 31) lateral-line scales, cheek with 4 rows of small scales. Tiny scales over proximal ¼ of caudal fin.
</paragraph>
<paragraph id="AD3A36878B342C0BFF70F8D8BFE938D5" blockId="4.[151,1437,1296,2043]" pageId="4" pageNumber="423">Head of males gray with two light gray interorbital bars; black opercular spot; throat yellow. Flank gray with light blue spots; 4–6 light bars below lateral line; ventrally white. Dorsal fin clear with orange markings, black submarginal band and white lappets. Caudal-fin rays orange with blue membranes. Anal fin with black marginal band encompassing all spines and through rays over 2/3 of distal portion fin; proximal portion of rays light blue; one yellow ocellus. Pelvic fin with spine and first two rays black; remaining clear. Pectoral fin clear.</paragraph>
<caption id="F9FA660F8B352C0AFF20FF1FBA833FFC" ID-Table-UUID="F9FA660F8B352C0AFF20FF1FBA833FFC" httpUri="http://table.plazi.org/id/F9FA660F8B352C0AFF20FF1FBA833FFC" pageId="5" pageNumber="424" startId="5.[151,239,151,175]" targetBox="[166,1351,228,2023]" targetIsTable="true" targetPageId="5">
<paragraph id="AD3A36878B352C0AFF20FF1FBA833FFC" blockId="5.[151,1437,151,212]" pageId="5" pageNumber="424">
<emphasis id="9FF1EA958B352C0AFF20FF1FBA2C3F80" bold="true" box="[151,265,151,175]" pageId="5" pageNumber="424">TABLE 1.</emphasis>
Morphological and meristic data for 
<taxonomicName id="6A854D048B352C0AFD26FF1FB8433F80" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[657,870,151,175]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B352C0AFD26FF1FB8433F80" box="[657,870,151,175]" italics="true" pageId="5" pageNumber="424">Metriaclima ngarae</emphasis>
</taxonomicName>
from Ngara, Lake Malaŵi, PSU 13365, 13366, n=14. Ranges include holotype.
</paragraph>
</caption>
<paragraph id="AD3A36878B352C0AFF11FF6CBFDE38C8" pageId="5" pageNumber="424">
<table id="DF85C4278B35D3F0FF11FF6CBE6238C8" box="[166,1351,228,2023]" colsContinueIn="9.[166,1350,228,2023]" gridcols="5" gridrows="45" pageId="5" pageNumber="424">
<tr id="13B534C58B35D3F0FF11FF6CBE623FD3" box="[166,1351,228,252]" gridrow="0" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FF6CB92E3FD3" box="[166,523,228,252]" gridcol="0" gridrow="0" pageId="5" pageNumber="424">Variable</th>
<th id="50645DB98B35D3F0FD00FF6CB8303FD3" box="[695,789,228,252]" gridcol="1" gridrow="0" pageId="5" pageNumber="424">Holotype</th>
<th id="50645DB98B35D3F0FCE8FF6CB8BF3FD3" box="[863,922,228,252]" gridcol="2" gridrow="0" pageId="5" pageNumber="424">Mean</th>
<th id="50645DB98B35D3F0FC57FF6CBF523FD3" box="[992,1143,228,252]" gridcol="3" gridrow="0" pageId="5" pageNumber="424">Std Dev</th>
<th id="50645DB98B35D3F0FB62FF6CBE623FD3" box="[1237,1351,228,252]" gridcol="4" gridrow="0" pageId="5" pageNumber="424">Range</th>
</tr>
<tr id="13B534C58B35D3F0FF11FE84BE623E0B" box="[166,1351,268,292]" gridrow="1" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FE84B92E3E0B" box="[166,523,268,292]" gridcol="0" gridrow="1" pageId="5" pageNumber="424">Standard length (mm)</th>
<td id="50645DB98B35D3F0FD00FE84B8303E0B" box="[695,789,268,292]" gridcol="1" gridrow="1" pageId="5" pageNumber="424">59.1</td>
<td id="50645DB98B35D3F0FCE8FE84B8BF3E0B" box="[863,922,268,292]" gridcol="2" gridrow="1" pageId="5" pageNumber="424">56.6</td>
<td id="50645DB98B35D3F0FC57FE84BF523E0B" box="[992,1143,268,292]" gridcol="3" gridrow="1" pageId="5" pageNumber="424">3.7</td>
<td id="50645DB98B35D3F0FB62FE84BE623E0B" box="[1237,1351,268,292]" gridcol="4" gridrow="1" pageId="5" pageNumber="424">49.6–62.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FEBCBE623E63" box="[166,1351,308,332]" gridrow="2" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FEBCB92E3E63" box="[166,523,308,332]" gridcol="0" gridrow="2" pageId="5" pageNumber="424">Head length (mm)</th>
<td id="50645DB98B35D3F0FD00FEBCB8303E63" box="[695,789,308,332]" gridcol="1" gridrow="2" pageId="5" pageNumber="424">19.1</td>
<td id="50645DB98B35D3F0FCE8FEBCB8BF3E63" box="[863,922,308,332]" gridcol="2" gridrow="2" pageId="5" pageNumber="424">18.7</td>
<td id="50645DB98B35D3F0FC57FEBCBF523E63" box="[992,1143,308,332]" gridcol="3" gridrow="2" pageId="5" pageNumber="424">1.2</td>
<td id="50645DB98B35D3F0FB62FEBCBE623E63" box="[1237,1351,308,332]" gridcol="4" gridrow="2" pageId="5" pageNumber="424">17.0–21.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FED4BE623E5A" box="[166,1351,348,373]" gridrow="3" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FED4BE623E5A" box="[166,1351,348,373]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="5" pageNumber="424">Percent head length (%)</th>
</tr>
<tr id="13B534C58B35D3F0FF11FE0DBE623EB2" box="[166,1351,389,413]" gridrow="4" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FE0DB92E3EB2" box="[166,523,389,413]" gridcol="0" gridrow="4" pageId="5" pageNumber="424">Snout length</th>
<td id="50645DB98B35D3F0FD00FE0DB8303EB2" box="[695,789,389,413]" gridcol="1" gridrow="4" pageId="5" pageNumber="424">34.6</td>
<td id="50645DB98B35D3F0FCE8FE0DB8BF3EB2" box="[863,922,389,413]" gridcol="2" gridrow="4" pageId="5" pageNumber="424">34.9</td>
<td id="50645DB98B35D3F0FC57FE0DBF523EB2" box="[992,1143,389,413]" gridcol="3" gridrow="4" pageId="5" pageNumber="424">1.2</td>
<td id="50645DB98B35D3F0FB62FE0DBE623EB2" box="[1237,1351,389,413]" gridcol="4" gridrow="4" pageId="5" pageNumber="424">30.8–36.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FE25BE623EEA" box="[166,1351,429,453]" gridrow="5" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FE25B92E3EEA" box="[166,523,429,453]" gridcol="0" gridrow="5" pageId="5" pageNumber="424">Postorbital head length</th>
<td id="50645DB98B35D3F0FD00FE25B8303EEA" box="[695,789,429,453]" gridcol="1" gridrow="5" pageId="5" pageNumber="424">39.5</td>
<td id="50645DB98B35D3F0FCE8FE25B8BF3EEA" box="[863,922,429,453]" gridcol="2" gridrow="5" pageId="5" pageNumber="424">40.8</td>
<td id="50645DB98B35D3F0FC57FE25BF523EEA" box="[992,1143,429,453]" gridcol="3" gridrow="5" pageId="5" pageNumber="424">1.7</td>
<td id="50645DB98B35D3F0FB62FE25BE623EEA" box="[1237,1351,429,453]" gridcol="4" gridrow="5" pageId="5" pageNumber="424">36.8–42.7</td>
</tr>
<tr id="13B534C58B35D3F0FF11FE5DBE623EC1" box="[166,1351,469,494]" gridrow="6" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FE5DB92E3EC1" box="[166,523,469,494]" gridcol="0" gridrow="6" pageId="5" pageNumber="424">Horizontal eye diameter</th>
<td id="50645DB98B35D3F0FD00FE5DB8303EC1" box="[695,789,469,494]" gridcol="1" gridrow="6" pageId="5" pageNumber="424">30.6</td>
<td id="50645DB98B35D3F0FCE8FE5DB8BF3EC1" box="[863,922,469,494]" gridcol="2" gridrow="6" pageId="5" pageNumber="424">30.1</td>
<td id="50645DB98B35D3F0FC57FE5DBF523EC1" box="[992,1143,469,494]" gridcol="3" gridrow="6" pageId="5" pageNumber="424">1.6</td>
<td id="50645DB98B35D3F0FB62FE5DBE623EC1" box="[1237,1351,469,494]" gridcol="4" gridrow="6" pageId="5" pageNumber="424">26.7–32.3</td>
</tr>
<tr id="13B534C58B35D3F0FF11FE75BE623D39" box="[166,1351,509,534]" gridrow="7" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FE75B92E3D39" box="[166,523,509,534]" gridcol="0" gridrow="7" pageId="5" pageNumber="424">Vertical eye diameter</th>
<td id="50645DB98B35D3F0FD00FE75B8303D39" box="[695,789,509,534]" gridcol="1" gridrow="7" pageId="5" pageNumber="424">28.9</td>
<td id="50645DB98B35D3F0FCE8FE75B8BF3D39" box="[863,922,509,534]" gridcol="2" gridrow="7" pageId="5" pageNumber="424">28.0</td>
<td id="50645DB98B35D3F0FC57FE75BF523D39" box="[992,1143,509,534]" gridcol="3" gridrow="7" pageId="5" pageNumber="424">1.5</td>
<td id="50645DB98B35D3F0FB62FE75BE623D39" box="[1237,1351,509,534]" gridcol="4" gridrow="7" pageId="5" pageNumber="424">24.4–29.9</td>
</tr>
<tr id="13B534C58B35D3F0FF11FDAEBE623D11" box="[166,1351,550,574]" gridrow="8" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FDAEB92E3D11" box="[166,523,550,574]" gridcol="0" gridrow="8" pageId="5" pageNumber="424">Preorbital depth</th>
<td id="50645DB98B35D3F0FD00FDAEB8303D11" box="[695,789,550,574]" gridcol="1" gridrow="8" pageId="5" pageNumber="424">18.7</td>
<td id="50645DB98B35D3F0FCE8FDAEB8BF3D11" box="[863,922,550,574]" gridcol="2" gridrow="8" pageId="5" pageNumber="424">19.3</td>
<td id="50645DB98B35D3F0FC57FDAEBF523D11" box="[992,1143,550,574]" gridcol="3" gridrow="8" pageId="5" pageNumber="424">1.1</td>
<td id="50645DB98B35D3F0FB62FDAEBE623D11" box="[1237,1351,550,574]" gridcol="4" gridrow="8" pageId="5" pageNumber="424">17.9–21.2</td>
</tr>
<tr id="13B534C58B35D3F0FF11FDC6BE623D49" box="[166,1351,590,614]" gridrow="9" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FDC6B92E3D49" box="[166,523,590,614]" gridcol="0" gridrow="9" pageId="5" pageNumber="424">Cheek depth</th>
<td id="50645DB98B35D3F0FD00FDC6B8303D49" box="[695,789,590,614]" gridcol="1" gridrow="9" pageId="5" pageNumber="424">35.3</td>
<td id="50645DB98B35D3F0FCE8FDC6B8BF3D49" box="[863,922,590,614]" gridcol="2" gridrow="9" pageId="5" pageNumber="424">34.6</td>
<td id="50645DB98B35D3F0FC57FDC6BF523D49" box="[992,1143,590,614]" gridcol="3" gridrow="9" pageId="5" pageNumber="424">2.5</td>
<td id="50645DB98B35D3F0FB62FDC6BE623D49" box="[1237,1351,590,614]" gridcol="4" gridrow="9" pageId="5" pageNumber="424">30.3–38.6</td>
</tr>
<tr id="13B534C58B35D3F0FF11FDFEBE623DA1" box="[166,1351,630,654]" gridrow="10" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FDFEB92E3DA1" box="[166,523,630,654]" gridcol="0" gridrow="10" pageId="5" pageNumber="424">Lower jaw length</th>
<td id="50645DB98B35D3F0FD00FDFEB8303DA1" box="[695,789,630,654]" gridcol="1" gridrow="10" pageId="5" pageNumber="424">25.0</td>
<td id="50645DB98B35D3F0FCE8FDFEB8BF3DA1" box="[863,922,630,654]" gridcol="2" gridrow="10" pageId="5" pageNumber="424">26.0</td>
<td id="50645DB98B35D3F0FC57FDFEBF523DA1" box="[992,1143,630,654]" gridcol="3" gridrow="10" pageId="5" pageNumber="424">2.8</td>
<td id="50645DB98B35D3F0FB62FDFEBE623DA1" box="[1237,1351,630,654]" gridcol="4" gridrow="10" pageId="5" pageNumber="424">20.7–30.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FD16BE623D98" box="[166,1351,670,695]" gridrow="11" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FD16B92E3D98" box="[166,523,670,695]" gridcol="0" gridrow="11" pageId="5" pageNumber="424">Head depth</th>
<td id="50645DB98B35D3F0FD00FD16B8303D98" box="[695,789,670,695]" gridcol="1" gridrow="11" pageId="5" pageNumber="424">109.9</td>
<td id="50645DB98B35D3F0FCE8FD16B8BF3D98" box="[863,922,670,695]" gridcol="2" gridrow="11" pageId="5" pageNumber="424">104.1</td>
<td id="50645DB98B35D3F0FC57FD16BF523D98" box="[992,1143,670,695]" gridcol="3" gridrow="11" pageId="5" pageNumber="424">4.6</td>
<td id="50645DB98B35D3F0FB62FD16BE623D98" box="[1237,1351,670,695]" gridcol="4" gridrow="11" pageId="5" pageNumber="424">98.5–113.0</td>
</tr>
<tr id="13B534C58B35D3F0FF11FD4FBE623DF0" box="[166,1351,711,735]" gridrow="12" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FD4FB92E3DF0" box="[166,523,711,735]" gridcol="0" gridrow="12" pageId="5" pageNumber="424">Inter-orbital width</th>
<td id="50645DB98B35D3F0FD00FD4FB8303DF0" box="[695,789,711,735]" gridcol="1" gridrow="12" pageId="5" pageNumber="424">34.4</td>
<td id="50645DB98B35D3F0FCE8FD4FB8BF3DF0" box="[863,922,711,735]" gridcol="2" gridrow="12" pageId="5" pageNumber="424">33.4</td>
<td id="50645DB98B35D3F0FC57FD4FBF523DF0" box="[992,1143,711,735]" gridcol="3" gridrow="12" pageId="5" pageNumber="424">2.1</td>
<td id="50645DB98B35D3F0FB62FD4FBE623DF0" box="[1237,1351,711,735]" gridcol="4" gridrow="12" pageId="5" pageNumber="424">30.7–37.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FD67BE623C28" box="[166,1351,751,775]" gridrow="13" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FD67B92E3C28" box="[166,523,751,775]" gridcol="0" gridrow="13" pageId="5" pageNumber="424">Premaxillary pedicel length</th>
<td id="50645DB98B35D3F0FD00FD67B8303C28" box="[695,789,751,775]" gridcol="1" gridrow="13" pageId="5" pageNumber="424">15.0</td>
<td id="50645DB98B35D3F0FCE8FD67B8BF3C28" box="[863,922,751,775]" gridcol="2" gridrow="13" pageId="5" pageNumber="424">12.2</td>
<td id="50645DB98B35D3F0FC57FD67BF523C28" box="[992,1143,751,775]" gridcol="3" gridrow="13" pageId="5" pageNumber="424">2.9</td>
<td id="50645DB98B35D3F0FB62FD67BE623C28" box="[1237,1351,751,775]" gridcol="4" gridrow="13" pageId="5" pageNumber="424">7.0–15.8</td>
</tr>
<tr id="13B534C58B35D3F0FF11FC9FBE623C00" box="[166,1351,791,815]" gridrow="14" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FC9FBE623C00" box="[166,1351,791,815]" colspan="5" colspanRight="4" gridcol="0" gridrow="14" pageId="5" pageNumber="424">Percent standard length (%)</th>
</tr>
<tr id="13B534C58B35D3F0FF11FCB7BE623C77" box="[166,1351,831,856]" gridrow="15" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FCB7B92E3C77" box="[166,523,831,856]" gridcol="0" gridrow="15" pageId="5" pageNumber="424">Head length</th>
<td id="50645DB98B35D3F0FD00FCB7B8303C77" box="[695,789,831,856]" gridcol="1" gridrow="15" pageId="5" pageNumber="424">32.3</td>
<td id="50645DB98B35D3F0FCE8FCB7B8BF3C77" box="[863,922,831,856]" gridcol="2" gridrow="15" pageId="5" pageNumber="424">33.1</td>
<td id="50645DB98B35D3F0FC57FCB7BF523C77" box="[992,1143,831,856]" gridcol="3" gridrow="15" pageId="5" pageNumber="424">0.9</td>
<td id="50645DB98B35D3F0FB62FCB7BE623C77" box="[1237,1351,831,856]" gridcol="4" gridrow="15" pageId="5" pageNumber="424">32.0–34.5</td>
</tr>
<tr id="13B534C58B35D3F0FF11FCE0BE623CAF" box="[166,1351,872,896]" gridrow="16" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FCE0B92E3CAF" box="[166,523,872,896]" gridcol="0" gridrow="16" pageId="5" pageNumber="424">Body depth</th>
<td id="50645DB98B35D3F0FD00FCE0B8303CAF" box="[695,789,872,896]" gridcol="1" gridrow="16" pageId="5" pageNumber="424">39.1</td>
<td id="50645DB98B35D3F0FCE8FCE0B8BF3CAF" box="[863,922,872,896]" gridcol="2" gridrow="16" pageId="5" pageNumber="424">38.7</td>
<td id="50645DB98B35D3F0FC57FCE0BF523CAF" box="[992,1143,872,896]" gridcol="3" gridrow="16" pageId="5" pageNumber="424">0.8</td>
<td id="50645DB98B35D3F0FB62FCE0BE623CAF" box="[1237,1351,872,896]" gridcol="4" gridrow="16" pageId="5" pageNumber="424">37.1–40.1</td>
</tr>
<tr id="13B534C58B35D3F0FF11FC18BE623C87" box="[166,1351,912,936]" gridrow="17" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FC18B92E3C87" box="[166,523,912,936]" gridcol="0" gridrow="17" pageId="5" pageNumber="424">Snout to dorsal fin origin</th>
<td id="50645DB98B35D3F0FD00FC18B8303C87" box="[695,789,912,936]" gridcol="1" gridrow="17" pageId="5" pageNumber="424">34.9</td>
<td id="50645DB98B35D3F0FCE8FC18B8BF3C87" box="[863,922,912,936]" gridcol="2" gridrow="17" pageId="5" pageNumber="424">35.7</td>
<td id="50645DB98B35D3F0FC57FC18BF523C87" box="[992,1143,912,936]" gridcol="3" gridrow="17" pageId="5" pageNumber="424">1.3</td>
<td id="50645DB98B35D3F0FB62FC18BE623C87" box="[1237,1351,912,936]" gridcol="4" gridrow="17" pageId="5" pageNumber="424">33.9–37.7</td>
</tr>
<tr id="13B534C58B35D3F0FF11FC30BE623CFF" box="[166,1351,952,976]" gridrow="18" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FC30B92E3CFF" box="[166,523,952,976]" gridcol="0" gridrow="18" pageId="5" pageNumber="424">Snout to pelvic fin origin</th>
<td id="50645DB98B35D3F0FD00FC30B8303CFF" box="[695,789,952,976]" gridcol="1" gridrow="18" pageId="5" pageNumber="424">38.4</td>
<td id="50645DB98B35D3F0FCE8FC30B8BF3CFF" box="[863,922,952,976]" gridcol="2" gridrow="18" pageId="5" pageNumber="424">37.5</td>
<td id="50645DB98B35D3F0FC57FC30BF523CFF" box="[992,1143,952,976]" gridcol="3" gridrow="18" pageId="5" pageNumber="424">1.1</td>
<td id="50645DB98B35D3F0FB62FC30BE623CFF" box="[1237,1351,952,976]" gridcol="4" gridrow="18" pageId="5" pageNumber="424">36.3–39.9</td>
</tr>
<tr id="13B534C58B35D3F0FF11FC68BE623CD7" box="[166,1351,992,1016]" gridrow="19" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FC68B92E3CD7" box="[166,523,992,1016]" gridcol="0" gridrow="19" pageId="5" pageNumber="424">Dorsal fin base length</th>
<td id="50645DB98B35D3F0FD00FC68B8303CD7" box="[695,789,992,1016]" gridcol="1" gridrow="19" pageId="5" pageNumber="424">59.8</td>
<td id="50645DB98B35D3F0FCE8FC68B8BF3CD7" box="[863,922,992,1016]" gridcol="2" gridrow="19" pageId="5" pageNumber="424">58.3</td>
<td id="50645DB98B35D3F0FC57FC68BF523CD7" box="[992,1143,992,1016]" gridcol="3" gridrow="19" pageId="5" pageNumber="424">1.4</td>
<td id="50645DB98B35D3F0FB62FC68BE623CD7" box="[1237,1351,992,1016]" gridcol="4" gridrow="19" pageId="5" pageNumber="424">56.0–60.2</td>
</tr>
<tr id="13B534C58B35D3F0FF11FB80BE623B0E" box="[166,1351,1032,1057]" gridrow="20" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FB80B92E3B0E" box="[166,523,1032,1057]" gridcol="0" gridrow="20" pageId="5" pageNumber="424">Anterior dorsal to anterior anal</th>
<td id="50645DB98B35D3F0FD00FB80B8303B0E" box="[695,789,1032,1057]" gridcol="1" gridrow="20" pageId="5" pageNumber="424">50.1</td>
<td id="50645DB98B35D3F0FCE8FB80B8BF3B0E" box="[863,922,1032,1057]" gridcol="2" gridrow="20" pageId="5" pageNumber="424">50.1</td>
<td id="50645DB98B35D3F0FC57FB80BF523B0E" box="[992,1143,1032,1057]" gridcol="3" gridrow="20" pageId="5" pageNumber="424">0.8</td>
<td id="50645DB98B35D3F0FB62FB80BE623B0E" box="[1237,1351,1032,1057]" gridcol="4" gridrow="20" pageId="5" pageNumber="424">48.8–51.5</td>
</tr>
<tr id="13B534C58B35D3F0FF11FBB9BE623B66" box="[166,1351,1073,1097]" gridrow="21" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FBB9B92E3B66" box="[166,523,1073,1097]" gridcol="0" gridrow="21" pageId="5" pageNumber="424">Anterior dorsal to posterior anal</th>
<td id="50645DB98B35D3F0FD00FBB9B8303B66" box="[695,789,1073,1097]" gridcol="1" gridrow="21" pageId="5" pageNumber="424">63.5</td>
<td id="50645DB98B35D3F0FCE8FBB9B8BF3B66" box="[863,922,1073,1097]" gridcol="2" gridrow="21" pageId="5" pageNumber="424">62.9</td>
<td id="50645DB98B35D3F0FC57FBB9BF523B66" box="[992,1143,1073,1097]" gridcol="3" gridrow="21" pageId="5" pageNumber="424">1.3</td>
<td id="50645DB98B35D3F0FB62FBB9BE623B66" box="[1237,1351,1073,1097]" gridcol="4" gridrow="21" pageId="5" pageNumber="424">60.8–64.6</td>
</tr>
<tr id="13B534C58B35D3F0FF11FBD1BE623B5E" box="[166,1351,1113,1137]" gridrow="22" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FBD1B92E3B5E" box="[166,523,1113,1137]" gridcol="0" gridrow="22" pageId="5" pageNumber="424">Posterior dorsal to anterior anal</th>
<td id="50645DB98B35D3F0FD00FBD1B8303B5E" box="[695,789,1113,1137]" gridcol="1" gridrow="22" pageId="5" pageNumber="424">36.6</td>
<td id="50645DB98B35D3F0FCE8FBD1B8BF3B5E" box="[863,922,1113,1137]" gridcol="2" gridrow="22" pageId="5" pageNumber="424">36.0</td>
<td id="50645DB98B35D3F0FC57FBD1BF523B5E" box="[992,1143,1113,1137]" gridcol="3" gridrow="22" pageId="5" pageNumber="424">1.2</td>
<td id="50645DB98B35D3F0FB62FBD1BE623B5E" box="[1237,1351,1113,1137]" gridcol="4" gridrow="22" pageId="5" pageNumber="424">34.2–37.7</td>
</tr>
<tr id="13B534C58B35D3F0FF11FB09BE623BB6" box="[166,1351,1153,1177]" gridrow="23" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FB09B92E3BB6" box="[166,523,1153,1177]" gridcol="0" gridrow="23" pageId="5" pageNumber="424">Posterior dorsal to posterior anal</th>
<td id="50645DB98B35D3F0FD00FB09B8303BB6" box="[695,789,1153,1177]" gridcol="1" gridrow="23" pageId="5" pageNumber="424">19.3</td>
<td id="50645DB98B35D3F0FCE8FB09B8BF3BB6" box="[863,922,1153,1177]" gridcol="2" gridrow="23" pageId="5" pageNumber="424">19.4</td>
<td id="50645DB98B35D3F0FC57FB09BF523BB6" box="[992,1143,1153,1177]" gridcol="3" gridrow="23" pageId="5" pageNumber="424">0.9</td>
<td id="50645DB98B35D3F0FB62FB09BE623BB6" box="[1237,1351,1153,1177]" gridcol="4" gridrow="23" pageId="5" pageNumber="424">18.1–20.3</td>
</tr>
<tr id="13B534C58B35D3F0FF11FB21BE623BED" box="[166,1351,1193,1218]" gridrow="24" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FB21B92E3BED" box="[166,523,1193,1218]" gridcol="0" gridrow="24" pageId="5" pageNumber="424">Posterior dorsal to ventral caudal</th>
<td id="50645DB98B35D3F0FD00FB21B8303BED" box="[695,789,1193,1218]" gridcol="1" gridrow="24" pageId="5" pageNumber="424">21.1</td>
<td id="50645DB98B35D3F0FCE8FB21B8BF3BED" box="[863,922,1193,1218]" gridcol="2" gridrow="24" pageId="5" pageNumber="424">20.6</td>
<td id="50645DB98B35D3F0FC57FB21BF523BED" box="[992,1143,1193,1218]" gridcol="3" gridrow="24" pageId="5" pageNumber="424">0.9</td>
<td id="50645DB98B35D3F0FB62FB21BE623BED" box="[1237,1351,1193,1218]" gridcol="4" gridrow="24" pageId="5" pageNumber="424">19.3–22.3</td>
</tr>
<tr id="13B534C58B35D3F0FF11FB5ABE623BC5" box="[166,1351,1234,1258]" gridrow="25" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FB5AB92E3BC5" box="[166,523,1234,1258]" gridcol="0" gridrow="25" pageId="5" pageNumber="424">Posterior anal to dorsal caudal</th>
<td id="50645DB98B35D3F0FD00FB5AB8303BC5" box="[695,789,1234,1258]" gridcol="1" gridrow="25" pageId="5" pageNumber="424">25.1</td>
<td id="50645DB98B35D3F0FCE8FB5AB8BF3BC5" box="[863,922,1234,1258]" gridcol="2" gridrow="25" pageId="5" pageNumber="424">23.2</td>
<td id="50645DB98B35D3F0FC57FB5ABF523BC5" box="[992,1143,1234,1258]" gridcol="3" gridrow="25" pageId="5" pageNumber="424">1.1</td>
<td id="50645DB98B35D3F0FB62FB5ABE623BC5" box="[1237,1351,1234,1258]" gridcol="4" gridrow="25" pageId="5" pageNumber="424">20.6–25.1</td>
</tr>
<tr id="13B534C58B35D3F0FF11FB72BE623A3D" box="[166,1351,1274,1298]" gridrow="26" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FB72B92E3A3D" box="[166,523,1274,1298]" gridcol="0" gridrow="26" pageId="5" pageNumber="424">Anterior dorsal to pelvic-fin origin</th>
<td id="50645DB98B35D3F0FD00FB72B8303A3D" box="[695,789,1274,1298]" gridcol="1" gridrow="26" pageId="5" pageNumber="424">39.4</td>
<td id="50645DB98B35D3F0FCE8FB72B8BF3A3D" box="[863,922,1274,1298]" gridcol="2" gridrow="26" pageId="5" pageNumber="424">38.5</td>
<td id="50645DB98B35D3F0FC57FB72BF523A3D" box="[992,1143,1274,1298]" gridcol="3" gridrow="26" pageId="5" pageNumber="424">0.9</td>
<td id="50645DB98B35D3F0FB62FB72BE623A3D" box="[1237,1351,1274,1298]" gridcol="4" gridrow="26" pageId="5" pageNumber="424">37.5–40.2</td>
</tr>
<tr id="13B534C58B35D3F0FF11FAAABE623A15" box="[166,1351,1314,1338]" gridrow="27" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FAAAB92E3A15" box="[166,523,1314,1338]" gridcol="0" gridrow="27" pageId="5" pageNumber="424">Posterior dorsal to pelvic-fin origin</th>
<td id="50645DB98B35D3F0FD00FAAAB8303A15" box="[695,789,1314,1338]" gridcol="1" gridrow="27" pageId="5" pageNumber="424">65.2</td>
<td id="50645DB98B35D3F0FCE8FAAAB8BF3A15" box="[863,922,1314,1338]" gridcol="2" gridrow="27" pageId="5" pageNumber="424">61.5</td>
<td id="50645DB98B35D3F0FC57FAAABF523A15" box="[992,1143,1314,1338]" gridcol="3" gridrow="27" pageId="5" pageNumber="424">1.4</td>
<td id="50645DB98B35D3F0FB62FAAABE623A15" box="[1237,1351,1314,1338]" gridcol="4" gridrow="27" pageId="5" pageNumber="424">58.7–65.2</td>
</tr>
<tr id="13B534C58B35D3F0FF11FAC2BE623A4C" box="[166,1351,1354,1379]" gridrow="28" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FAC2B92E3A4C" box="[166,523,1354,1379]" gridcol="0" gridrow="28" pageId="5" pageNumber="424">Caudal peduncle length</th>
<td id="50645DB98B35D3F0FD00FAC2B8303A4C" box="[695,789,1354,1379]" gridcol="1" gridrow="28" pageId="5" pageNumber="424">14.9</td>
<td id="50645DB98B35D3F0FCE8FAC2B8BF3A4C" box="[863,922,1354,1379]" gridcol="2" gridrow="28" pageId="5" pageNumber="424">14.0</td>
<td id="50645DB98B35D3F0FC57FAC2BF523A4C" box="[992,1143,1354,1379]" gridcol="3" gridrow="28" pageId="5" pageNumber="424">1.0</td>
<td id="50645DB98B35D3F0FB62FAC2BE623A4C" box="[1237,1351,1354,1379]" gridcol="4" gridrow="28" pageId="5" pageNumber="424">11.8–15.4</td>
</tr>
<tr id="13B534C58B35D3F0FF11FAFBBE623AA4" box="[166,1351,1395,1419]" gridrow="29" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FAFBB92E3AA4" box="[166,523,1395,1419]" gridcol="0" gridrow="29" pageId="5" pageNumber="424">Least caudal peduncle depth</th>
<td id="50645DB98B35D3F0FD00FAFBB8303AA4" box="[695,789,1395,1419]" gridcol="1" gridrow="29" pageId="5" pageNumber="424">16.3</td>
<td id="50645DB98B35D3F0FCE8FAFBB8BF3AA4" box="[863,922,1395,1419]" gridcol="2" gridrow="29" pageId="5" pageNumber="424">16.2</td>
<td id="50645DB98B35D3F0FC57FAFBBF523AA4" box="[992,1143,1395,1419]" gridcol="3" gridrow="29" pageId="5" pageNumber="424">0.6</td>
<td id="50645DB98B35D3F0FB62FAFBBE623AA4" box="[1237,1351,1395,1419]" gridcol="4" gridrow="29" pageId="5" pageNumber="424">15.5–17.3</td>
</tr>
<tr id="13B534C58B35D3F0FF11FA13BE623A9B" box="[166,1351,1435,1460]" gridrow="30" pageId="5" pageNumber="424" rowspan-1="1">
<th id="50645DB98B35D3F0FF11FA13B92E3A9B" box="[166,523,1435,1460]" gridcol="0" gridrow="30" pageId="5" pageNumber="424">Meristics</th>
<td id="50645DB98B35D3F0FCE8FA13B8BF3A9B" box="[863,922,1435,1460]" gridcol="2" gridrow="30" pageId="5" pageNumber="424">Mode</td>
<td id="50645DB98B35D3F0FC57FA13BF523A9B" box="[992,1143,1435,1460]" gridcol="3" gridrow="30" pageId="5" pageNumber="424">Frequency (%)</td>
<td id="50645DB98B35D3F0FB62FA13BE623A9B" box="[1237,1351,1435,1460]" gridcol="4" gridrow="30" pageId="5" pageNumber="424">Range</td>
</tr>
<tr id="13B534C58B35D3F0FF11FA4BBE623AF4" box="[166,1351,1475,1499]" gridrow="31" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FA4BB92E3AF4" box="[166,523,1475,1499]" gridcol="0" gridrow="31" pageId="5" pageNumber="424">Dorsal-fin spines</th>
<td id="50645DB98B35D3F0FD00FA4BB8303AF4" box="[695,789,1475,1499]" gridcol="1" gridrow="31" pageId="5" pageNumber="424">17</td>
<td id="50645DB98B35D3F0FCE8FA4BB8BF3AF4" box="[863,922,1475,1499]" gridcol="2" gridrow="31" pageId="5" pageNumber="424">17</td>
<td id="50645DB98B35D3F0FC57FA4BBF523AF4" box="[992,1143,1475,1499]" gridcol="3" gridrow="31" pageId="5" pageNumber="424">71.4</td>
<td id="50645DB98B35D3F0FB62FA4BBE623AF4" box="[1237,1351,1475,1499]" gridcol="4" gridrow="31" pageId="5" pageNumber="424">16–17</td>
</tr>
<tr id="13B534C58B35D3F0FF11FA63BE62392B" box="[166,1351,1515,1540]" gridrow="32" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11FA63B92E392B" box="[166,523,1515,1540]" gridcol="0" gridrow="32" pageId="5" pageNumber="424">Dorsal-fin rays</th>
<td id="50645DB98B35D3F0FD00FA63B830392B" box="[695,789,1515,1540]" gridcol="1" gridrow="32" pageId="5" pageNumber="424">8</td>
<td id="50645DB98B35D3F0FCE8FA63B8BF392B" box="[863,922,1515,1540]" gridcol="2" gridrow="32" pageId="5" pageNumber="424">8</td>
<td id="50645DB98B35D3F0FC57FA63BF52392B" box="[992,1143,1515,1540]" gridcol="3" gridrow="32" pageId="5" pageNumber="424">71.4</td>
<td id="50645DB98B35D3F0FB62FA63BE62392B" box="[1237,1351,1515,1540]" gridcol="4" gridrow="32" pageId="5" pageNumber="424">7–8</td>
</tr>
<tr id="13B534C58B35D3F0FF11F99CBE623903" box="[166,1351,1556,1580]" gridrow="33" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F99CB92E3903" box="[166,523,1556,1580]" gridcol="0" gridrow="33" pageId="5" pageNumber="424">Anal-fin spines</th>
<td id="50645DB98B35D3F0FD00F99CB8303903" box="[695,789,1556,1580]" gridcol="1" gridrow="33" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FCE8F99CB8BF3903" box="[863,922,1556,1580]" gridcol="2" gridrow="33" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FC57F99CBF523903" box="[992,1143,1556,1580]" gridcol="3" gridrow="33" pageId="5" pageNumber="424">100.0</td>
<td id="50645DB98B35D3F0FB62F99CBE623903" box="[1237,1351,1556,1580]" gridcol="4" gridrow="33" pageId="5" pageNumber="424">3–3</td>
</tr>
<tr id="13B534C58B35D3F0FF11F9B4BE62397A" box="[166,1351,1596,1621]" gridrow="34" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F9B4B92E397A" box="[166,523,1596,1621]" gridcol="0" gridrow="34" pageId="5" pageNumber="424">Anal-fin rays</th>
<td id="50645DB98B35D3F0FD00F9B4B830397A" box="[695,789,1596,1621]" gridcol="1" gridrow="34" pageId="5" pageNumber="424">8</td>
<td id="50645DB98B35D3F0FCE8F9B4B8BF397A" box="[863,922,1596,1621]" gridcol="2" gridrow="34" pageId="5" pageNumber="424">8</td>
<td id="50645DB98B35D3F0FC57F9B4BF52397A" box="[992,1143,1596,1621]" gridcol="3" gridrow="34" pageId="5" pageNumber="424">100.0</td>
<td id="50645DB98B35D3F0FB62F9B4BE62397A" box="[1237,1351,1596,1621]" gridcol="4" gridrow="34" pageId="5" pageNumber="424">8–8</td>
</tr>
<tr id="13B534C58B35D3F0FF11F9ECBE623952" box="[166,1351,1636,1661]" gridrow="35" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F9ECB92E3952" box="[166,523,1636,1661]" gridcol="0" gridrow="35" pageId="5" pageNumber="424">Pectoral-fin rays</th>
<td id="50645DB98B35D3F0FD00F9ECB8303952" box="[695,789,1636,1661]" gridcol="1" gridrow="35" pageId="5" pageNumber="424">12</td>
<td id="50645DB98B35D3F0FCE8F9ECB8BF3952" box="[863,922,1636,1661]" gridcol="2" gridrow="35" pageId="5" pageNumber="424">14</td>
<td id="50645DB98B35D3F0FC57F9ECBF523952" box="[992,1143,1636,1661]" gridcol="3" gridrow="35" pageId="5" pageNumber="424">85.7</td>
<td id="50645DB98B35D3F0FB62F9ECBE623952" box="[1237,1351,1636,1661]" gridcol="4" gridrow="35" pageId="5" pageNumber="424">12–15</td>
</tr>
<tr id="13B534C58B35D3F0FF11F904BE62398A" box="[166,1351,1676,1701]" gridrow="36" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F904B92E398A" box="[166,523,1676,1701]" gridcol="0" gridrow="36" pageId="5" pageNumber="424">Pelvic-fin rays</th>
<td id="50645DB98B35D3F0FD00F904B830398A" box="[695,789,1676,1701]" gridcol="1" gridrow="36" pageId="5" pageNumber="424">7</td>
<td id="50645DB98B35D3F0FCE8F904B8BF398A" box="[863,922,1676,1701]" gridcol="2" gridrow="36" pageId="5" pageNumber="424">7</td>
<td id="50645DB98B35D3F0FC57F904BF52398A" box="[992,1143,1676,1701]" gridcol="3" gridrow="36" pageId="5" pageNumber="424">85.7</td>
<td id="50645DB98B35D3F0FB62F904BE62398A" box="[1237,1351,1676,1701]" gridcol="4" gridrow="36" pageId="5" pageNumber="424">6–7</td>
</tr>
<tr id="13B534C58B35D3F0FF11F93DBE6239E2" box="[166,1351,1717,1741]" gridrow="37" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F93DB92E39E2" box="[166,523,1717,1741]" gridcol="0" gridrow="37" pageId="5" pageNumber="424">Lateral line scales</th>
<td id="50645DB98B35D3F0FD00F93DB83039E2" box="[695,789,1717,1741]" gridcol="1" gridrow="37" pageId="5" pageNumber="424">32</td>
<td id="50645DB98B35D3F0FCE8F93DB8BF39E2" box="[863,922,1717,1741]" gridcol="2" gridrow="37" pageId="5" pageNumber="424">31</td>
<td id="50645DB98B35D3F0FC57F93DBF5239E2" box="[992,1143,1717,1741]" gridcol="3" gridrow="37" pageId="5" pageNumber="424">57.1</td>
<td id="50645DB98B35D3F0FB62F93DBE6239E2" box="[1237,1351,1717,1741]" gridcol="4" gridrow="37" pageId="5" pageNumber="424">30–32</td>
</tr>
<tr id="13B534C58B35D3F0FF11F955BE6239DA" box="[166,1351,1757,1781]" gridrow="38" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F955B92E39DA" box="[166,523,1757,1781]" gridcol="0" gridrow="38" pageId="5" pageNumber="424">Pored scales caudal</th>
<td id="50645DB98B35D3F0FD00F955B83039DA" box="[695,789,1757,1781]" gridcol="1" gridrow="38" pageId="5" pageNumber="424">2</td>
<td id="50645DB98B35D3F0FCE8F955B8BF39DA" box="[863,922,1757,1781]" gridcol="2" gridrow="38" pageId="5" pageNumber="424">2</td>
<td id="50645DB98B35D3F0FC57F955BF5239DA" box="[992,1143,1757,1781]" gridcol="3" gridrow="38" pageId="5" pageNumber="424">50.0</td>
<td id="50645DB98B35D3F0FB62F955BE6239DA" box="[1237,1351,1757,1781]" gridcol="4" gridrow="38" pageId="5" pageNumber="424">0–2</td>
</tr>
<tr id="13B534C58B35D3F0FF11F88DBE623832" box="[166,1351,1797,1821]" gridrow="39" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F88DB92E3832" box="[166,523,1797,1821]" gridcol="0" gridrow="39" pageId="5" pageNumber="424">Cheek scale rows</th>
<td id="50645DB98B35D3F0FD00F88DB8303832" box="[695,789,1797,1821]" gridcol="1" gridrow="39" pageId="5" pageNumber="424">4</td>
<td id="50645DB98B35D3F0FCE8F88DB8BF3832" box="[863,922,1797,1821]" gridcol="2" gridrow="39" pageId="5" pageNumber="424">4</td>
<td id="50645DB98B35D3F0FC57F88DBF523832" box="[992,1143,1797,1821]" gridcol="3" gridrow="39" pageId="5" pageNumber="424">100.0</td>
<td id="50645DB98B35D3F0FB62F88DBE623832" box="[1237,1351,1797,1821]" gridcol="4" gridrow="39" pageId="5" pageNumber="424">4–4</td>
</tr>
<tr id="13B534C58B35D3F0FF11F8A5BE623869" box="[166,1351,1837,1862]" gridrow="40" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F8A5B92E3869" box="[166,523,1837,1862]" gridcol="0" gridrow="40" pageId="5" pageNumber="424">Gill rakers 1st ceratobranchial</th>
<td id="50645DB98B35D3F0FD00F8A5B8303869" box="[695,789,1837,1862]" gridcol="1" gridrow="40" pageId="5" pageNumber="424">10</td>
<td id="50645DB98B35D3F0FCE8F8A5B8BF3869" box="[863,922,1837,1862]" gridcol="2" gridrow="40" pageId="5" pageNumber="424">11</td>
<td id="50645DB98B35D3F0FC57F8A5BF523869" box="[992,1143,1837,1862]" gridcol="3" gridrow="40" pageId="5" pageNumber="424">57.1</td>
<td id="50645DB98B35D3F0FB62F8A5BE623869" box="[1237,1351,1837,1862]" gridcol="4" gridrow="40" pageId="5" pageNumber="424">9–12</td>
</tr>
<tr id="13B534C58B35D3F0FF11F8DEBE623841" box="[166,1351,1878,1902]" gridrow="41" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F8DEB92E3841" box="[166,523,1878,1902]" gridcol="0" gridrow="41" pageId="5" pageNumber="424">Gill rakers 1st epibranchial</th>
<td id="50645DB98B35D3F0FD00F8DEB8303841" box="[695,789,1878,1902]" gridcol="1" gridrow="41" pageId="5" pageNumber="424">5</td>
<td id="50645DB98B35D3F0FCE8F8DEB8BF3841" box="[863,922,1878,1902]" gridcol="2" gridrow="41" pageId="5" pageNumber="424">5</td>
<td id="50645DB98B35D3F0FC57F8DEBF523841" box="[992,1143,1878,1902]" gridcol="3" gridrow="41" pageId="5" pageNumber="424">50.0</td>
<td id="50645DB98B35D3F0FB62F8DEBE623841" box="[1237,1351,1878,1902]" gridcol="4" gridrow="41" pageId="5" pageNumber="424">4–5</td>
</tr>
<tr id="13B534C58B35D3F0FF11F8F6BE6238B9" box="[166,1351,1918,1942]" gridrow="42" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F8F6B92E38B9" box="[166,523,1918,1942]" gridcol="0" gridrow="42" pageId="5" pageNumber="424">Teeth outer left lower jaw</th>
<td id="50645DB98B35D3F0FD00F8F6B83038B9" box="[695,789,1918,1942]" gridcol="1" gridrow="42" pageId="5" pageNumber="424">8</td>
<td id="50645DB98B35D3F0FCE8F8F6B8BF38B9" box="[863,922,1918,1942]" gridcol="2" gridrow="42" pageId="5" pageNumber="424">7</td>
<td id="50645DB98B35D3F0FC57F8F6BF5238B9" box="[992,1143,1918,1942]" gridcol="3" gridrow="42" pageId="5" pageNumber="424">35.7</td>
<td id="50645DB98B35D3F0FB62F8F6BE6238B9" box="[1237,1351,1918,1942]" gridcol="4" gridrow="42" pageId="5" pageNumber="424">4–10</td>
</tr>
<tr id="13B534C58B35D3F0FF11F82EBE623891" box="[166,1351,1958,1982]" gridrow="43" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F82EB92E3891" box="[166,523,1958,1982]" gridcol="0" gridrow="43" pageId="5" pageNumber="424">Tooth rows upper jaw</th>
<td id="50645DB98B35D3F0FD00F82EB8303891" box="[695,789,1958,1982]" gridcol="1" gridrow="43" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FCE8F82EB8BF3891" box="[863,922,1958,1982]" gridcol="2" gridrow="43" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FC57F82EBF523891" box="[992,1143,1958,1982]" gridcol="3" gridrow="43" pageId="5" pageNumber="424">92.9</td>
<td id="50645DB98B35D3F0FB62F82EBE623891" box="[1237,1351,1958,1982]" gridcol="4" gridrow="43" pageId="5" pageNumber="424">2–3</td>
</tr>
<tr id="13B534C58B35D3F0FF11F846BE6238C8" box="[166,1351,1998,2023]" gridrow="44" pageId="5" pageNumber="424">
<th id="50645DB98B35D3F0FF11F846B92E38C8" box="[166,523,1998,2023]" gridcol="0" gridrow="44" pageId="5" pageNumber="424">Tooth rows lower jaw</th>
<td id="50645DB98B35D3F0FD00F846B83038C8" box="[695,789,1998,2023]" gridcol="1" gridrow="44" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FCE8F846B8BF38C8" box="[863,922,1998,2023]" gridcol="2" gridrow="44" pageId="5" pageNumber="424">3</td>
<td id="50645DB98B35D3F0FC57F846BF5238C8" box="[992,1143,1998,2023]" gridcol="3" gridrow="44" pageId="5" pageNumber="424">64.3</td>
<td id="50645DB98B35D3F0FB62F846BE6238C8" box="[1237,1351,1998,2023]" gridcol="4" gridrow="44" pageId="5" pageNumber="424">2–3</td>
</tr>
</table>
</paragraph>
<paragraph id="AD3A36878B362C09FF70FF1FBAC03FD6" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="425">Head of females dark gray with two faint green interorbital bars; ventral half pale yellow; black opercular spot; pale yellow throat. Flank yellowish, white below lateral line with light gray bars above lateral line. Fin coloration same as males, not as intense.</paragraph>
</subSubSection>
<subSubSection id="E59F650C8B362C09FF70FE8AB9003D86" pageId="6" pageNumber="425" type="distribution">
<paragraph id="AD3A36878B362C09FF70FE8AB9FC3E4A" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="425">
<emphasis id="9FF1EA958B362C09FF70FE8ABA443E33" bold="true" box="[199,353,258,284]" pageId="6" pageNumber="425">Distribution.</emphasis>
<taxonomicName id="6A854D048B362C09FEDEFE8BB9693E33" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[361,588,259,285]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B362C09FEDEFE8BB9693E33" box="[361,588,259,285]" italics="true" pageId="6" pageNumber="425">Metriaclima ngarae</emphasis>
</taxonomicName>
is found on submerged reefs near Ngara (
<geoCoordinate id="C8B150408B362C09FB87FE8BBFE73E32" box="[1072,1218,259,285]" degrees="10.233397" direction="south" orientation="latitude" pageId="6" pageNumber="425" precision="1" value="-10.233397">10.233397 S</geoCoordinate>
, 
<geoCoordinate id="C8B150408B362C09FB66FE8ABE473E32" box="[1233,1378,258,285]" degrees="34.106592" direction="east" orientation="longitude" pageId="6" pageNumber="425" precision="1" value="34.10659">34.106592 E</geoCoordinate>
) and Mdoka (
<geoCoordinate id="C8B150408B362C09FF40FEAFBAAE3E6E" box="[247,395,295,321]" degrees="10.280608" direction="south" orientation="latitude" pageId="6" pageNumber="425" precision="1" value="-10.280608">10.280608 S</geoCoordinate>
, 
<geoCoordinate id="C8B150408B362C09FE2DFEAEB90B3E6E" box="[410,558,294,321]" degrees="34.179156" direction="east" orientation="longitude" pageId="6" pageNumber="425" precision="1" value="34.179157">34.179156 E</geoCoordinate>
), and has also been reported from Chesese (
<geoCoordinate id="C8B150408B362C09FB9AFEAFBFE43E6E" box="[1069,1217,295,321]" degrees="10.280608" direction="south" orientation="latitude" pageId="6" pageNumber="425" precision="1" value="-10.280608">10.280608 S</geoCoordinate>
, 
<geoCoordinate id="C8B150408B362C09FB67FEAEBE473E6E" box="[1232,1378,294,321]" degrees="34.179756" direction="east" orientation="longitude" pageId="6" pageNumber="425" precision="1" value="34.179756">34.179756 E</geoCoordinate>
) and Kaporo (
<geoCoordinate id="C8B150408B362C09FF4DFEC3BA593E4A" box="[250,380,331,357]" degrees="9.731972" direction="south" orientation="latitude" pageId="6" pageNumber="425" precision="1" value="-9.731972">9.731972 S</geoCoordinate>
, 
<geoCoordinate id="C8B150408B362C09FE3EFEC2B9283E4A" box="[393,525,330,357]" degrees="33.94415" direction="east" orientation="longitude" pageId="6" pageNumber="425" precision="1" value="33.94415">33.94415 E</geoCoordinate>
), 
<collectingCountry id="D59276178B362C09FD96FEC3B9523E4A" box="[545,631,331,357]" name="Malawi" pageId="6" pageNumber="425">Malaŵi</collectingCountry>
(
<figureCitation id="35BE2A028B362C09FD31FEC3B9ED3E4A" box="[646,712,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1776,1800]" captionTargetBox="[161,1425,463,1748]" captionTargetId="figure-160@2.[151,1436,453,1753]" captionTargetPageId="2" captionText="FIGURE 1. Map of Lake Malaŵi with the localities marked that are mentioned in the text." figureDoi="http://doi.org/10.5281/zenodo.5572220" httpUri="https://zenodo.org/record/5572220/files/figure.png" pageId="6" pageNumber="425">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="AD3A36878B362C09FF70FEE6B9003D86" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="425">
<emphasis id="9FF1EA958B362C09FF70FEE6BA8D3EA7" bold="true" box="[199,424,366,392]" pageId="6" pageNumber="425">Field observations.</emphasis>
<taxonomicName id="6A854D048B362C09FE18FEE7B9AB3EA7" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[431,654,367,393]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B362C09FE18FEE7B9AB3EA7" box="[431,654,367,393]" italics="true" pageId="6" pageNumber="425">Metriaclima ngarae</emphasis>
</taxonomicName>
is most often encountered near empty shells of the gastropod 
<taxonomicName id="6A854D048B362C09FA88FEE7BA7B3E82" authority="Dohrn" authorityName="Dohrn" authorityYear="1865" class="Gastropoda" family="Ampullariidae" genus="Lanistes" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="425" phylum="Mollusca" rank="species" species="nyassanus">
<emphasis id="9FF1EA958B362C09FA88FEE7BA2F3E83" italics="true" pageId="6" pageNumber="425">Lanistes nyassanus</emphasis>
Dohrn
</taxonomicName>
that lie scattered on the sandy bottom at a depth ranging between 
<quantity id="6A7D9B628B362C09FB34FE1ABFCE3E82" box="[1155,1259,402,429]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="425" unit="m" value="17.5" valueMax="25.0" valueMin="10.0">10–25 m</quantity>
(
<figureCitation id="35BE2A028B362C09FB48FE1BBE6C3E82" box="[1279,1353,403,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1763,1787]" captionTargetBox="[151,1436,777,1740]" captionTargetId="figure-282@6.[151,1436,777,1740]" captionTargetPageId="6" captionText="FIGURE 4. A male Metriaclima ngarae at a submerged reef near Ngara (10.233397 S, 34.106592 E)." figureDoi="http://doi.org/10.5281/zenodo.5572226" httpUri="https://zenodo.org/record/5572226/files/figure.png" pageId="6" pageNumber="425">Fig. 4</figureCitation>
). Most observations occurred near rocky habitats, but it was also found at a depth of about 
<quantity id="6A7D9B628B362C09FBE2FE3FBFAA3EFE" box="[1109,1167,439,465]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="425" unit="m" value="20.0">20 m</quantity>
near the outflow of the Songwe River that forms the border between 
<collectingCountry id="D59276178B362C09FD2CFE53B9D43EDA" box="[667,753,475,501]" name="Malawi" pageId="6" pageNumber="425">Malawi</collectingCountry>
and 
<collectingCountry id="D59276178B362C09FC9FFE53B8B53EDA" box="[808,912,475,501]" name="Tanzania" pageId="6" pageNumber="425">Tanzania</collectingCountry>
, with no known rocky reefs within 
<quantity id="6A7D9B628B362C09FA94FE53BE4B3EDA" box="[1315,1390,475,501]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="425" unit="km" value="30.0">30 km</quantity>
. No stomach analyses were performed but individuals were observed picking at targeted sites on the sand as well as combing the algal matrix covering the empty shells, small rocks, and other objects that lie on the open sand. Most individuals encountered were solitary and always in association with an empty shell of 
<taxonomicName id="6A854D048B362C09FB3AFDCFBE423D4F" authorityName="Dohrn" authorityYear="1865" box="[1165,1383,583,608]" class="Gastropoda" family="Ampullariidae" genus="Lanistes" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="425" phylum="Mollusca" rank="species" species="nyassanus">
<emphasis id="9FF1EA958B362C09FB3AFDCFBE423D4F" box="[1165,1383,583,608]" italics="true" pageId="6" pageNumber="425">Lanistes nyassanus</emphasis>
</taxonomicName>
into which it would retreat when threatened. Mouth-brooding females stay near their shell and probably release their offspring inside such shells as well.
</paragraph>
</subSubSection>
<subSubSection id="E59F650C8B362C09FF70FD3BBA213DDE" pageId="6" pageNumber="425" type="etymology">
<paragraph id="AD3A36878B362C09FF70FD3BBA213DDE" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="425">
<emphasis id="9FF1EA958B362C09FF70FD3BBA6B3DE2" bold="true" box="[199,334,691,717]" pageId="6" pageNumber="425">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="6A854D048B362C09FDE0FD3BB9833DE3" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[599,678,691,716]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B362C09FDE0FD3BB9833DE3" box="[599,678,691,716]" italics="true" pageId="6" pageNumber="425">ngarae</emphasis>
</taxonomicName>
in the genitive of Ngara, 
<collectingCountry id="D59276178B362C09FC44FD3BBF6E3DE2" box="[1011,1099,691,717]" name="Malawi" pageId="6" pageNumber="425">Malawi</collectingCountry>
, where the specimens were collected.
</paragraph>
</subSubSection>
</treatment>
</document>