204 lines
41 KiB
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204 lines
41 KiB
XML
<document id="5DDF3E49E07EF0D4016B335A6189174E" ID-CLB-Dataset="7777" ID-DOI="10.5281/zenodo.4503863" ID-GBIF-Dataset="5de89f6c-aa0e-4f40-bd33-883dce6bb1ba" ID-ISSN="0374-1036" ID-Zenodo-Dep="4503863" ID-ZooBank="D69294C8-0182-4F35-9565-3C1823B9FD17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612462066712" checkinUser="felipe" docAuthor="Gnezdilov, Vladimir M." docDate="2015" docId="03E12E3507346576818EFCA328153879" docLanguage="en" docName="ActaEntMusNatPra.55.1.9-18.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Bruneastrum Gnezdilov 2015, gen. nov." docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FFD8564D073565748054FFB22A723E32" masterDocTitle="Description of a new genus and species of Hemisphaeriini from Brunei with an identification key to the Bornean species of the tribe (Hemiptera: Fulgoroidea: Issidae)" masterLastPageNumber="18" masterPageNumber="9" pageNumber="10" updateTime="1698927216983" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="741737BD8D8E2EA5CDB1000719AE2569">
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<mods:title id="D48E15D2269159DC4C1977A780A4CAEB">Description of a new genus and species of Hemisphaeriini from Brunei with an identification key to the Bornean species of the tribe (Hemiptera: Fulgoroidea: Issidae)</mods:title>
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<mods:name id="43647F96134FDFD606F3E8D9D105EE08" type="personal">
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<mods:roleTerm id="07154D1CBBCCD65DA274427D175D3810">Author</mods:roleTerm>
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<mods:namePart id="01475127E3CEA11B88712888AC8A8D3D">Gnezdilov, Vladimir M.</mods:namePart>
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<mods:typeOfResource id="32085B0FFAE4B8759C1133CD3E509ED2">text</mods:typeOfResource>
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<mods:title id="794AC2637200C13D9C27292BBAD4B557">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="FF5393303EA9614C08FBCD59F66EFDD0">
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<mods:date id="C6274DA3180F5226CF141A334EB27C20">2015</mods:date>
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<mods:number id="3CCB197C3761DC15C1C9D1588D7ABCCB">2015-06-01</mods:number>
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<mods:number id="163B7CB1796808B8DCDA1DECE3DB5D23">55</mods:number>
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<mods:classification id="2CEFE6C560546464D5837615371D8C23">journal article</mods:classification>
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<mods:identifier id="544BE399A501826DD981B4AD988C661B" type="DOI">10.5281/zenodo.4503863</mods:identifier>
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<treatment id="03E12E3507346576818EFCA328153879" ID-DOI="http://doi.org/10.5281/zenodo.4601034" ID-GBIF-Taxon="180328955" ID-Zenodo-Dep="4601034" LSID="urn:lsid:plazi:treatment:03E12E3507346576818EFCA328153879" httpUri="http://treatment.plazi.org/id/03E12E3507346576818EFCA328153879" lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10">
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<subSubSection id="C352CCA807346575818EFCA328AA3D1B" box="[474,728,785,809]" pageId="1" pageNumber="10" type="nomenclature">
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<paragraph id="8BF79F2307346575818EFCA328AA3D1B" blockId="1.[474,728,785,809]" box="[474,728,785,809]" pageId="1" pageNumber="10">
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<heading id="D0BF284F07346575818EFCA328AA3D1B" bold="true" box="[474,728,785,809]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="10" reason="3">
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<emphasis id="B93C433107346575818EFCA328AA3D1B" bold="true" box="[474,728,785,809]" pageId="1" pageNumber="10">
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<taxonomicName id="4C48E4A007346575818EFCA3281F3D1A" ID-CoL="3DG8" authority="Gnezdilov, 2015" authorityName="Gnezdilov" authorityYear="2015" box="[474,621,785,808]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B93C433107346575818EFCA3281F3D1A" bold="true" box="[474,621,785,808]" italics="true" pageId="1" pageNumber="10">Bruneastrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20FFE4A073465758220FCA028AA3D1B" box="[628,728,786,809]" pageId="1" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352CCA8073465758025FCF329653D6A" box="[113,791,833,856]" pageId="1" pageNumber="10" type="type_taxon">
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<paragraph id="8BF79F23073465758025FCF329653D6A" blockId="1.[113,791,833,856]" box="[113,791,833,856]" pageId="1" pageNumber="10">
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<emphasis id="B93C4331073465758025FCF32B773D6A" bold="true" box="[113,261,833,856]" pageId="1" pageNumber="10">
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<typeStatus id="54F32181073465758025FCF32ADB3D6A" box="[113,169,833,856]" pageId="1" pageNumber="10">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C48E4A0073465758158FCF328713D6A" ID-CoL="NFHT" authority="Gnezdilov, 2015" authorityName="Gnezdilov" authorityYear="2015" box="[268,515,833,856]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="10" phylum="Arthropoda" rank="species" species="cardinale" status="sp. nov.">
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<emphasis id="B93C4331073465758158FCF328713D6A" box="[268,515,833,856]" italics="true" pageId="1" pageNumber="10">Bruneastrum cardinale</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20FFE4A07346575825DFCF3282C3D6A" box="[521,606,833,856]" pageId="1" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C352CCA8073465758025FCC229E33A75" pageId="1" pageNumber="10" type="diagnosis">
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<paragraph id="8BF79F23073465758025FCC229E33A75" blockId="1.[113,1089,880,1607]" pageId="1" pageNumber="10">
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<emphasis id="B93C4331073465758025FCC22A943DB5" bold="true" box="[113,230,880,903]" pageId="1" pageNumber="10">Diagnosis.</emphasis>
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Metope smooth, with no intermediate carinae visible (
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<figureCitation id="137383A607346575836EFCC229F33DB5" box="[826,897,880,903]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 3</figureCitation>
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,
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<figureCitation id="137383A60734657583DAFCC229D93DB5" box="[910,939,880,903]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">13</figureCitation>
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). Metope and coryphe joining at obtuse angle in lateral view (
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<figureCitation id="137383A6073465758209FC2228D83D95" box="[605,682,912,935]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">Fig. 12</figureCitation>
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). Coryphe transverse (
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<figureCitation id="137383A60734657583C0FC2229A53D95" box="[916,983,912,935]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 4</figureCitation>
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,
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<figureCitation id="137383A60734657583B5FC22298C3D95" box="[993,1022,912,935]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">14</figureCitation>
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). Fore wings without hypocostal plate, with basal angle of costal margin distinctly convex (semicircular) (
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<figureCitation id="137383A60734657580EAFC622B733DD5" box="[190,257,976,999]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 1</figureCitation>
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,
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<figureCitation id="137383A607346575815BFC622B5F3DD5" box="[271,301,976,999]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">12</figureCitation>
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). Hind wings unilobed (
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<figureCitation id="137383A607346575827BFC6228023DD5" box="[559,624,976,999]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Fig. 2</figureCitation>
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), reaching abdominal apex. Hind tibia with 2 lateral spines. First metatarsomere with 5 intermediate spines in full row. Phallobase with a pair of long, curved latero-apical processes, narrowing apically (
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<figureCitation id="137383A607346575836CFBA2290B3A15" box="[824,889,1040,1063]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="1" pageNumber="10">Fig. 6</figureCitation>
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, laphp) and with a pair of ventro-lateral lobes (
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<figureCitation id="137383A60734657581C8FB822BAE3A75" box="[412,476,1072,1095]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="1" pageNumber="10">Fig. 6</figureCitation>
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, vlphl). Aedeagus lacking ventral hooks.
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</paragraph>
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</subSubSection>
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<subSubSection id="C352CCA8073465768025FBE2297A3839" lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10" type="description">
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<paragraph id="8BF79F23073465768025FBE2297A3839" blockId="1.[113,1089,880,1607]" lastBlockId="2.[151,1127,1044,1611]" lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10">
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<emphasis id="B93C4331073465758025FBE22A8E3A55" bold="true" box="[113,252,1104,1127]" pageId="1" pageNumber="10">Description.</emphasis>
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Metope smooth, with rudimentary median carina only in its apical part and without sublateral carinae, elongate, enlarged above the clypeus (
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<figureCitation id="137383A6073465758366FBC2290A3AB5" box="[818,888,1136,1159]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 3</figureCitation>
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,
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<figureCitation id="137383A60734657583D1FBC229D03AB5" box="[901,930,1136,1159]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">13</figureCitation>
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). Lateral keels (margins) of metope visor shading the antennae in dorsal view (
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<figureCitation id="137383A6073465758340FB2229FC3A95" box="[788,910,1168,1191]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">Figs 11, 14</figureCitation>
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). Metopoclypeal suture distinct, straight. Postclypeus large, smooth, with thick median carina running through the anteclypeus as well. Metope and coryphe joining at obtuse angle (in lateral view); metope visible from above (
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<figureCitation id="137383A607346575811EFB422BB53B35" box="[330,455,1264,1287]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">Figs 11, 12</figureCitation>
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). Coryphe transverse, twice as wide as long medially, with anterior margin convex and posterior margin weakly concave (
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<figureCitation id="137383A6073465758376FAA229183B15" box="[802,874,1296,1319]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 4</figureCitation>
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,
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<figureCitation id="137383A607346575832CFAA229E43B15" box="[888,918,1296,1319]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">14</figureCitation>
|
||
). Ocelli absent. Pedicel 1.5 times as long as wide, cylindrical. Pronotum very short, nearly 5 times shorter than mesonotum, without carinae; anterior and posterior margins keel-shaped. Paradiscal fields of pronotum very narrow, not visible behind the eyes. Paranotal lobes large, without carinae. Tegulae large. Mesonotum smooth, without carinae. Fore wings smooth, wide, narrowing apically, with distinctly convex (semicircular) basal angle of costal margin (
|
||
<figureCitation id="137383A60734657583A2FA022E483BF5" box="[1014,1082,1456,1479]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="137383A6073465758025FA622AFC3BD5" box="[113,142,1488,1511]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="1" pageNumber="10">12</figureCitation>
|
||
); fore wings without hypocostal plate, with setae on costal margin. Claval suture of fore wing distinct only basally, disappearing after ¼ of wing length (
|
||
<figureCitation id="137383A607346575835FFA42293A3835" box="[779,840,1520,1543]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Fig. 1</figureCitation>
|
||
). Hind wings unilobed, well-developed, reaching abdominal apex, with long and dense setae on costal margin (
|
||
<figureCitation id="137383A6073465758441F9A22AF33875" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="1" pageNumber="10">Fig. 2</figureCitation>
|
||
). Hind tibiae with 2 lateral spines distally. First metatarsomere with 2 lateral and 5 intermediate spines in full row apically. Second metatarsomere with only 2 latero-apical spines.
|
||
<emphasis id="B93C43310737657680E1FB812B213A79" bold="true" box="[181,339,1075,1099]" italics="true" pageId="2" pageNumber="11">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="137383A6073765768134FB862BBC3A79" box="[352,462,1076,1099]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Figs 5–10</figureCitation>
|
||
). Pygofer wide, with convex hind margin (in lateral view) (
|
||
<figureCitation id="137383A607376576846FFB862AD53A59" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 6</figureCitation>
|
||
). Anal tube wide (in dorsal view), with deep apical concavity (
|
||
<figureCitation id="137383A607376576831EFBE629FE3A59" box="[842,908,1108,1131]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 9</figureCitation>
|
||
). Anal column long, 0.3 times as long as whole anal tube (
|
||
<figureCitation id="137383A607376576827EFBC628083AB9" box="[554,634,1140,1163]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 10</figureCitation>
|
||
). Phallobase curved at obtuse angle (in lateral view), wide (in ventral view) (
|
||
<figureCitation id="137383A60737657681B5FB2628513A99" box="[481,547,1172,1195]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 5</figureCitation>
|
||
), with a pair of long latero-apical processes curved at right angle and narrowing apically (
|
||
<figureCitation id="137383A607376576825BFB06283D3AF9" box="[527,591,1204,1227]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 6</figureCitation>
|
||
, laphp) and with a pair of ventro-lateral lobes (
|
||
<figureCitation id="137383A607376576846FFB062AD43AD9" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 6</figureCitation>
|
||
, vlphl). Ventral phallobase lobe wide, reaching phallobase apex. Aedeagus lacking ventral hooks. Style massive, with hind margin concave (
|
||
<figureCitation id="137383A60737657682C6FB4628A23B39" box="[658,720,1268,1291]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 7</figureCitation>
|
||
, hms). Capitulum of style on short neck (
|
||
<figureCitation id="137383A60737657680CBFAA62A923B19" box="[159,224,1300,1323]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 7</figureCitation>
|
||
, n), wide (in dorsal view) (
|
||
<figureCitation id="137383A6073765768254FAA628323B19" box="[512,576,1300,1323]" captionStart="Figs 5–10" captionStartId="3.[113,150,1129,1148]" captionTargetBox="[115,1087,224,1098]" captionTargetId="figure-11@3.[113,1088,218,1104]" captionTargetPageId="3" captionText="Figs 5–10. Bruneastrum cardinale gen. & sp. nov., holotype, male genitalia. 5 – penis, ventral view; 6 – genital block, lateral view; 7 – style, lateral view; 8 – style, dorsal view; 9 – anal tube, dorsal view; 10 – same, lateral view. Abbreviations: ac – anal column; cps – capitulum of style; hms – hind margin of style; laphp – latero-apical processes of phallobase; n – neck of capitulum; vlphl – ventro-lateral phallobase lobe; vphl – ventral phallobase lobe." figureDoi="http://doi.org/10.5281/zenodo.4503867" httpUri="https://zenodo.org/record/4503867/files/figure.png" pageId="2" pageNumber="11">Fig. 8</figureCitation>
|
||
), with distinct lateral tooth, apical tooth very weak.
|
||
<emphasis id="B93C43310737657680C3FA862BE03B79" bold="true" box="[151,402,1332,1355]" pageId="2" pageNumber="11">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C48E4A00737657681CDFA8628563B79" authority="Gnezdilov, 2015" authorityName="Gnezdilov" authorityYear="2015" box="[409,548,1332,1355]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93C43310737657681CDFA8628563B79" box="[409,548,1332,1355]" italics="true" pageId="2" pageNumber="11">Bruneastrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20FFE4A073765768278FA8628FC3B79" box="[556,654,1332,1355]" pageId="2" pageNumber="11" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is similar to the genus
|
||
<taxonomicName id="4C48E4A00737657683DDFA862ABE3B59" authority="Distant, 1909" authorityName="Distant" authorityYear="1909" class="Insecta" family="Issidae" genus="Mongoliana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C43310737657683DDFA862E793B79" box="[905,1035,1332,1355]" italics="true" pageId="2" pageNumber="11">Mongoliana</emphasis>
|
||
<bibRefCitation id="EFD9E2D2073765768447FA862ABE3B59" author="DISTANT W. L." pageId="2" pageNumber="11" pagination="73 - 87" refId="ref3279" refString="DISTANT W. L. 1909: Rhynchotal notes. XLVIII. Annales and Magazine of Natural History, Series 4 8: 73 - 87." type="journal article" year="1909">Distant, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
with which it shares a distinctly convex (semicircular) basal angle of the fore wing costal margin (
|
||
<figureCitation id="137383A60737657680BBFAC62B433BB9" box="[239,305,1396,1419]" captionStart="Figs 1–4" captionStartId="2.[151,188,932,951]" captionTargetBox="[230,1054,230,897]" captionTargetId="figure-10@2.[208,1069,198,911]" captionTargetPageId="2" captionText="Figs 1–4. Bruneastrum cardinale gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus." figureDoi="http://doi.org/10.5281/zenodo.4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="2" pageNumber="11">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="137383A6073765768169FAC62BF03BB9" box="[317,386,1396,1419]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="2" pageNumber="11">11–12</figureCitation>
|
||
), but differs from the latter by the absence of aedeagal ventral hooks in males.All species of
|
||
<taxonomicName id="4C48E4A0073765768127FA262B873B99" authorityName="Distant" authorityYear="1909" box="[371,501,1428,1451]" class="Insecta" family="Issidae" genus="Mongoliana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C4331073765768127FA262B873B99" box="[371,501,1428,1451]" italics="true" pageId="2" pageNumber="11">Mongoliana</emphasis>
|
||
</taxonomicName>
|
||
(known from mainland
|
||
<collectingCountry id="F35FDFB30737657682A4FA2629433B99" box="[752,817,1428,1451]" name="China" pageId="2" pageNumber="11">China</collectingCountry>
|
||
) have well-developed ventral hooks of aedeagus (
|
||
<bibRefCitation id="EFD9E2D2073765768138FA062B8F3BF9" author="FENNAH R. G." box="[364,509,1460,1483]" pageId="2" pageNumber="11" pagination="441 - 527" refId="ref3306" refString="FENNAH R. G. 1956: Fulgoroidea from Southern China. Proceedings of the California Academy of Science, Series 4 28: 441 - 527." type="journal article" year="1956">FENNAH 1956</bibRefCitation>
|
||
, fig. 18B;
|
||
<bibRefCitation id="EFD9E2D2073765768239FA0629663BF9" author="CHE Y. - L. & WANG Y. - L. & CHOU I." box="[621,788,1460,1483]" pageId="2" pageNumber="11" pagination="35 - 44" refId="ref3200" refString="CHE Y. - L., WANG Y. - L. & CHOU I. 2003: Taxonomic study of the genus Mongoliana Distant (Homoptera: Fulgoroidea: Issidae). Entomotaxonomia 25: 35 - 44." type="journal article" year="2003">CHE et al. 2003</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C48E4A0073765768372FA0629C33BF9" authority="Gnezdilov, 2015" authorityName="Gnezdilov" authorityYear="2015" box="[806,945,1460,1483]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93C4331073765768372FA0629C33BF9" box="[806,945,1460,1483]" italics="true" pageId="2" pageNumber="11">Bruneastrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20FFE4A0737657683EDFA062E693BF9" box="[953,1051,1460,1483]" pageId="2" pageNumber="11" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
differs from the genera
|
||
<taxonomicName id="4C48E4A007376576811CFA6628F83BD9" authority="Schaum, 1850" authorityName="Schaum" authorityYear="1850" box="[328,650,1492,1515]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C433107376576811CFA662B9B3BD9" box="[328,489,1492,1515]" italics="true" pageId="2" pageNumber="11">Hemisphaerius</emphasis>
|
||
Schaum, 1850
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C48E4A00737657682EBFA6629EE3BD9" authority="Stal, 1870" authorityName="Stal" authorityYear="1870" box="[703,924,1492,1515]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C43310737657682EBFA66295B3BD9" box="[703,809,1492,1515]" italics="true" pageId="2" pageNumber="11">Gergithus</emphasis>
|
||
Stål, 1870
|
||
</taxonomicName>
|
||
(both known from Borneo, see below) by more elongate fore wings (
|
||
<figureCitation id="137383A60737657682F8FA46288E3839" box="[684,764,1524,1547]" captionStart="Figs 11–14" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 11–14. Bruneastrum cardinale gen. & sp. nov., holotype (BMNH), body length 4.1 mm. 11 – dorsal view; 12 – lateral view; 13 – frontal view; 14 – head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="2" pageNumber="11">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF37CFAB0737657680C3FC162E2F3DE0" ID-DOI="http://doi.org/10.5281/zenodo.4503865" ID-Zenodo-Dep="4503865" httpUri="https://zenodo.org/record/4503865/files/figure.png" pageId="2" pageNumber="11" startId="2.[151,188,932,951]" targetBox="[230,1054,230,897]" targetPageId="2">
|
||
<paragraph id="8BF79F230737657680C3FC162E2F3DE0" blockId="2.[151,1132,932,978]" pageId="2" pageNumber="11">
|
||
Figs 1–4.
|
||
<taxonomicName id="4C48E4A00737657680B8FC162BC03D85" authorityName="Gnezdilov" authorityYear="2015" box="[236,434,932,951]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="species" species="cardinale">
|
||
<emphasis id="B93C43310737657680B8FC162BC03D85" box="[236,434,932,951]" italics="true" pageId="2" pageNumber="11">Bruneastrum cardinale</emphasis>
|
||
</taxonomicName>
|
||
gen. & sp. nov., holotype. 1 – fore wing; 2 – hind wing; 3 – metope and clypeus; 4 – coryphe.Abbreviations: cs – claval suture; mcm – median carina of metope; mcc – median carina of clypeus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C352CCA80737657680C3F9A628153879" pageId="2" pageNumber="11" type="etymology">
|
||
<paragraph id="8BF79F230737657680C3F9A628153879" blockId="2.[151,1127,1044,1611]" pageId="2" pageNumber="11">
|
||
<emphasis id="B93C43310737657680C3F9A62B6A3819" bold="true" box="[151,280,1556,1579]" pageId="2" pageNumber="11">Etymology.</emphasis>
|
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Derived from the combination of the Greek noun άστρο (star) and
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<collectingCountry id="F35FDFB3073765768383F9A62E523819" box="[983,1056,1556,1579]" name="Brunei" pageId="2" pageNumber="11">Brunei</collectingCountry>
|
||
which means a “star of
|
||
<collectingCountry id="F35FDFB307376576811EF9862BE73879" box="[330,405,1588,1611]" name="Brunei" pageId="2" pageNumber="11">Brunei</collectingCountry>
|
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” (gender neutrum).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |