271 lines
36 KiB
XML
271 lines
36 KiB
XML
<document id="BEEF7BE1276DDA64C79F66AD0FB1A246" ID-CLB-Dataset="29085" ID-DOI="10.11646/zootaxa.4455.3.7" ID-GBIF-Dataset="e805d477-ca64-4ea3-8990-285677f540b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457495" ID-ZooBank="6B5ED1A9-FC34-4142-85AD-B35C73A427DE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539263023203" checkinUser="plazi" docAuthor="Fanti, Fabrizio & Pankowski, Mary K." docDate="2018" docId="03DC87CAFF8AFFD9FF2F13C8FC84FC85" docLanguage="en" docName="zootaxa.4455.3.7.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malthodes (Malthodes) josephi Fanti & Pankowski 2018" docType="treatment" docVersion="7" lastPageNumber="521" masterDocId="FFE5FFB2FF8CFFD1FFB8106EFFD6FFBD" masterDocTitle="Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae)" masterLastPageNumber="524" masterPageNumber="513" pageNumber="519" updateTime="1698576187060" updateUser="ExternalLinkService">
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<mods:mods id="A5FB7EA403308D096900722A3F571649" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="23492D034537459594CDE541075BA526">
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<mods:title id="2257D794F57A0E77BFF5C1B95DCEB1BB">Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C3538A4252FB630F79012D113A230DF1" type="personal">
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<mods:roleTerm id="C1DDFAA8C171168C2FF6BE37D8D9832A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7411268EE962238CA30416AE422E8F81">Fanti, Fabrizio</mods:namePart>
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</mods:name>
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<mods:name id="E1E8C885FBF81DE6E49A50FC563B111E" type="personal">
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<mods:role id="042AEF1CFA9FF5C3EEB8AEFE43BEC1D4">
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<mods:roleTerm id="75D5070E08DD611E69591E3F796C8512">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1E8E1EB27623EEFB94F40005DBA9B3C0">Pankowski, Mary K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A6F8595EF9289C6F5C1C3B6F252A3EBF">text</mods:typeOfResource>
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<mods:titleInfo id="1AAFBBDA3509ED618AFF3B36B0EB9110">
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<mods:title id="289C173377AD9F865F1EE06330EE8F2C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="39DFB0FAF1D00BE02EB983C1F0C10B1E">
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<mods:date id="0ABBD1D8F0452D14956C978C98EA460C">2018</mods:date>
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<mods:number id="2B40719184A5FC8DDA7A4D2C7B396211">2018-08-03</mods:number>
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<mods:detail id="763407E253ED27CEEAEC6EA36B4BF7AC" type="volume">
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<mods:number id="4D413A3D0088FAEFF1B516273CD2DE46">4455</mods:number>
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</mods:detail>
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<mods:detail id="C0CC6386957F7715AB5B8A799800AE11" type="issue">
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<mods:number id="94A75333845BDD7C372D8045E1EB78EE">3</mods:number>
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<mods:start id="C2A277AC932FAC48F381C43AF37EB319">513</mods:start>
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<mods:classification id="A807D811C9C74241D10FD2D90B2319FD">journal article</mods:classification>
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<mods:identifier id="00B14BBDB67C12895C41BA9EB2EB36D9" type="CLB-Dataset">29085</mods:identifier>
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<mods:identifier id="32A62D30ABB32BC727474E674237D30B" type="DOI">10.11646/zootaxa.4455.3.7</mods:identifier>
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<mods:identifier id="55A0BE6EDB0C5701C0FB075AA0AA770B" type="GBIF-Dataset">e805d477-ca64-4ea3-8990-285677f540b1</mods:identifier>
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<mods:identifier id="BA913E5133828CBC113AC116015DA3D1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="83AFE774229136A630F0BCE233A8C68A" type="Zenodo-Dep">1457495</mods:identifier>
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</mods:mods>
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<treatment id="03DC87CAFF8AFFD9FF2F13C8FC84FC85" ID-DOI="http://doi.org/10.5281/zenodo.5961870" ID-GBIF-Taxon="148691446" ID-Zenodo-Dep="5961870" LSID="urn:lsid:plazi:treatment:03DC87CAFF8AFFD9FF2F13C8FC84FC85" httpUri="http://treatment.plazi.org/id/03DC87CAFF8AFFD9FF2F13C8FC84FC85" lastPageId="8" lastPageNumber="521" pageId="6" pageNumber="519">
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<subSubSection id="C36F6557FF8AFFD7FF2F13C8FEC4FC5F" pageId="6" pageNumber="519" type="nomenclature">
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<paragraph id="8BCA36DCFF8AFFD7FF2F13C8FBD7FC7D" blockId="6.[151,1025,934,994]" box="[151,1025,934,960]" pageId="6" pageNumber="519">
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<heading id="D08281B0FF8AFFD7FF2F13C8FBD7FC7D" bold="true" box="[151,1025,934,960]" fontSize="11" level="1" pageId="6" pageNumber="519" reason="1">
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||
<emphasis id="B901EACEFF8AFFD7FF2F13C8FBD7FC7D" bold="true" box="[151,1025,934,960]" pageId="6" pageNumber="519">
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||
<taxonomicName id="4C754D5FFF8AFFD7FF2F13C8FDDBFC7D" authorityName="Fanti & Pankowski" authorityYear="2018" box="[151,525,934,960]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="519" phylum="Arthropoda" rank="species" species="josephi" subGenus="Malthodes">
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<emphasis id="B901EACEFF8AFFD7FF2F13C8FEC0FC7D" bold="true" box="[151,278,934,960]" italics="true" pageId="6" pageNumber="519">Malthodes</emphasis>
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(
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<emphasis id="B901EACEFF8AFFD7FE9F13C8FE70FC7D" bold="true" box="[295,422,934,960]" italics="true" pageId="6" pageNumber="519">Malthodes</emphasis>
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)
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<emphasis id="B901EACEFF8AFFD7FE0F13C8FDDBFC7D" bold="true" box="[439,525,934,960]" italics="true" pageId="6" pageNumber="519">josephi</emphasis>
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</taxonomicName>
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FANTI & M. K. PANKOWSKI sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA36DCFF8AFFD7FF2F13A7FEC4FC5F" blockId="6.[151,1025,934,994]" box="[151,274,969,994]" pageId="6" pageNumber="519">
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||
(
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<figureCitation id="134E2A59FF8AFFD7FF2713A7FEDFFC5F" box="[159,265,969,994]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="7.[151,250,1110,1132]" captionStartId-1="7.[151,250,1908,1930]" captionStartId-2="8.[151,250,1805,1827]" captionTargetBox-0="[216,1371,193,1089]" captionTargetBox-1="[348,1246,1329,1842]" captionTargetBox-2="[340,1247,877,1784]" captionTargetId-0="figure@7.[216,1371,193,1089]" captionTargetId-1="figure@7.[281,1305,1246,1886]" captionTargetId-2="figure@8.[340,1247,877,1784]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 4. Malthodes josephi sp. nov. in Baltic amber. A: Holotype, lateral view, scale bar = 0.5 mm. B: Holotype, lateral view, scale bar = 0.1 mm. C: Holotype, detail of terminalia, scale bar = 0.1 mm. D: Holotype, detail of terminalia, scale bar = 0.2 mm." captionText-1="FIGURE 5. Malthodes josephi sp. nov. in Baltic amber. Line drawing of the last abdominal segments, scale bar = 0.1 mm." captionText-2="FIGURE 6. Malthodes josephi sp. nov. in Baltic amber. Arrows shows cuticular vesicles extruded from the abdominal segments." httpUri-0="https://zenodo.org/record/1457503/files/figure.png" httpUri-1="https://zenodo.org/record/1457505/files/figure.png" httpUri-2="https://zenodo.org/record/1457507/files/figure.png" pageId="6" pageNumber="519">Figs. 4–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F6557FF8AFFD7FF2F147FFBF5F8FF" pageId="6" pageNumber="519" type="description">
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<paragraph id="8BCA36DCFF8AFFD7FF2F147FFBF5F8FF" blockId="6.[151,1437,1041,2038]" pageId="6" pageNumber="519">
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<emphasis id="B901EACEFF8AFFD7FF2F147FFEFFFB97" bold="true" box="[151,297,1041,1066]" pageId="6" pageNumber="519">Description.</emphasis>
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Adult, winged. Male, based on the last abdominal segments modified and the long antennae. Body length:
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<quantity id="4C8D9B39FF8AFFD7FF481458FE9AFBF0" box="[240,332,1078,1102]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="519" unit="mm" value="2.8">2.8 mm</quantity>
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; elytra:
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<quantity id="4C8D9B39FF8AFFD7FE161458FDDDFBF0" box="[430,523,1078,1102]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="519" unit="mm" value="1.2">1.2 mm</quantity>
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; antennae:
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<quantity id="4C8D9B39FF8AFFD7FD291458FD38FBF0" box="[657,750,1078,1102]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="519" unit="mm" value="2.2">2.2 mm</quantity>
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. Entirely dark brown including legs and antennae, without yellow spots on elytra. Head completely exposed, strongly rounded, larger than pronotum, equipped with shallow punctation. Eyes rounded, very large and prominent, inserted laterally to the head. Mandibles elongated, falciform, large at base and thin apically, concave on the internal side. Maxillary palps 4-segmented, unequal in length, with the terminal palpomere globular and distally pointed. Labial palps 3-segmented with the last segment globular and pointed. Antennae filiform, 11-segmented, long, surpassing the elytra, not reaching the last abdominal segments; antennomere I (scape) elongated, not club-shaped; antennomeres II–III filiform and approximately 1.4 times shorter than scape; antennomeres IV–X elongated, filiform, subequal in length, slightly longer than antennomeres II–III; antennomere XI thinner and longer than previous one, apically rounded; antennomeres with small setae. Pronotum large, transverse, surface undulating and with shallow punctation, posterior margin straight, anterior margin strongly bordered, sides sinuous, corners rounded. Scutellum triangular-shaped. Elytra wider than pronotum, short, slender, parallel-sided, reaching until the fifth abdominal segment, wide at base and narrower posteriorly to the humeral zone, apex rounded, surface pubescent. Posterior wings wide and long, dark, surpassing the elytra and the abdominal segments. Anterior and median legs very short, posterior legs long; coxae robust; trochanters elongated; femora enlarged; pro- and mesotibiae as long as pro- and mesofemora, metatibiae longer than metafemora, all the tibiae cylindrical and very thin. Tarsi 5-segmented; tarsomere I long and more of two times longer than second; tarsomere III slightly shorter than second; tarsomere IV bilobed; tarsomere V slender; claws simple. Penultimate tergite (tergite 9) elongated and slightly bent at sides; last tergite (tergite 10) short and wide with the sides bent, apically the margin is straight; last sternite (sternite 9) long and flat, large at base and curved after few millimeters, from which start two elongated and separated lobes with rounded apex; two wide and irregular with rounded apex urophysis, each inserted by the penultimate sternite (sternite 8) and located on both sides of the sternite 9. Sternal surface of abdomen punctate and with pubescence.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F6557FF8AFFD7FF7F1723FC1EF837" pageId="6" pageNumber="519" type="etymology">
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<paragraph id="8BCA36DCFF8AFFD7FF7F1723FC1EF837" blockId="6.[151,1437,1041,2038]" pageId="6" pageNumber="519">
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<emphasis id="B901EACEFF8AFFD7FF7F1723FE98F8DB" bold="true" box="[199,334,1869,1894]" pageId="6" pageNumber="519">Etymology.</emphasis>
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Species named after Joseph M. Pankowski, paternal grandfather of the second author, who always puts his family first, is generous beyond words and who is a friend to all.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F6557FF8AFFD6FF7F17FBFD02F84E" lastPageId="7" lastPageNumber="520" pageId="6" pageNumber="519" type="materials_examined">
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<paragraph id="8BCA36DCFF8AFFD7FF7F17FBFA51F813" blockId="6.[151,1437,1041,2038]" box="[199,1415,1941,1966]" pageId="6" pageNumber="519">
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<emphasis id="B901EACEFF8AFFD7FF7F17FBFEEDF813" bold="true" box="[199,315,1941,1966]" pageId="6" pageNumber="519">
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<typeStatus id="54CE887EFF8AFFD7FF7F17FBFEE0F813" box="[199,310,1941,1966]" pageId="6" pageNumber="519" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, adult specimen included in Baltic amber, accession No.
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<collectionCode id="ED64AE19FF8AFFD7FC4517F8FB98F813" box="[1021,1102,1942,1966]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="519">USNM</collectionCode>
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<collectionCode id="ED64AE19FF8AFFD7FBEC17F8FB50F813" box="[1108,1158,1942,1966]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15292" name="Herbarium Mediterraneum Panormitanum" pageId="6" pageNumber="519">PAL</collectionCode>
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<quantity id="4C8D9B39FF8AFFD7FB3617FBFB29F813" box="[1166,1279,1941,1966]" metricMagnitude="4" metricUnit="m" metricValue="1.80984144" pageId="6" pageNumber="519" unit="in" value="712536.0">712536 in</quantity>
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the
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<collectionCode id="ED64AE19FF8AFFD7FA9717F8FA52F813" box="[1327,1412,1942,1966]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="519">USNM</collectionCode>
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.
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</paragraph>
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<paragraph id="8BCA36DCFF8AFFD7FF7F17D7FE9EF84B" blockId="6.[151,1437,1041,2038]" pageId="6" pageNumber="519">
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<materialsCitation id="3B1D3C81FF8AFFD7FF7F17D7FE9EF84B" ID-GBIF-Occurrence="1917075940" country="Russia" county="Peninsula" location="Yantarny" municipality="Samland" pageId="6" pageNumber="519" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="B901EACEFF8AFFD7FF7F17D7FEB9F86F" bold="true" box="[199,367,1977,2002]" pageId="6" pageNumber="519">
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<typeStatus id="54CE887EFF8AFFD7FF7F17D7FED4F86F" box="[199,258,1977,2002]" pageId="6" pageNumber="519">Type</typeStatus>
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locality.
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</emphasis>
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<location id="8EAA6007FF8AFFD7FEC717D4FE32F86F" LSID="urn:lsid:plazi:treatment:03DC87CAFF8AFFD9FF2F13C8FC84FC85:8EAA6007FF8AFFD7FEC717D4FE32F86F" box="[383,484,1978,2002]" country="Russia" county="Peninsula" municipality="Samland" name="Yantarny" pageId="6" pageNumber="519" stateProvince="Kaliningrad">Yantarny</location>
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settlement (formerly
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<location id="8EAA6007FF8AFFD7FD4D17D7FCA8F86F" LSID="urn:lsid:plazi:treatment:03DC87CAFF8AFFD9FF2F13C8FC84FC85:8EAA6007FF8AFFD7FD4D17D7FCA8F86F" box="[757,894,1977,2002]" country="Russia" county="Peninsula" municipality="Samland" name="Palmnicken" pageId="6" pageNumber="519" stateProvince="Kaliningrad">Palmnicken</location>
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),
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<location id="8EAA6007FF8AFFD7FC2217D7FC2AF86F" LSID="urn:lsid:plazi:treatment:03DC87CAFF8AFFD9FF2F13C8FC84FC85:8EAA6007FF8AFFD7FC2217D7FC2AF86F" box="[922,1020,1977,2002]" country="Russia" county="Peninsula" municipality="Samland" name="Sambian" pageId="6" pageNumber="519" stateProvince="Kaliningrad">Sambian</location>
|
||
(
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<collectingMunicipality id="6BAEACA6FF8AFFD7FBAC17D7FBAFF86F" box="[1044,1145,1977,2002]" pageId="6" pageNumber="519">Samland</collectingMunicipality>
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||
)
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<collectingCounty id="62AB4E50FF8AFFD7FB2A17D7FAD4F86F" box="[1170,1282,1977,2002]" pageId="6" pageNumber="519">Peninsula</collectingCounty>
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||
,
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<collectingRegion id="49B1F83EFF8AFFD7FAAE17D7FA4AF86F" box="[1302,1436,1977,2002]" country="Russia" name="Kaliningrad" pageId="6" pageNumber="519">Kaliningrad</collectingRegion>
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||
Region,
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<collectingCountry id="F362764CFF8AFFD7FF4E17B3FE92F84B" box="[246,324,2013,2038]" name="Russia" pageId="6" pageNumber="519">Russia</collectingCountry>
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||
.
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</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF0A6654FF8BFFD6FF2F1438FF3DFB17" httpUri="https://zenodo.org/record/1457503/files/figure.png" pageId="7" pageNumber="520" startId="7.[151,250,1110,1132]" targetBox="[216,1371,193,1089]" targetPageId="7">
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<paragraph id="8BCA36DCFF8BFFD6FF2F1438FF3DFB17" blockId="7.[151,1436,1110,1194]" pageId="7" pageNumber="520">
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<emphasis id="B901EACEFF8BFFD6FF2F1438FEC2FBD1" bold="true" box="[151,276,1110,1132]" pageId="7" pageNumber="520">FIGURE 4.</emphasis>
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<taxonomicName id="4C754D5FFF8BFFD6FEA41438FE0EFBD1" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[284,472,1110,1132]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="520" phylum="Arthropoda" rank="species" species="josephi" status="sp. nov.">
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<emphasis id="B901EACEFF8BFFD6FEA41438FE0EFBD1" box="[284,472,1110,1132]" italics="true" pageId="7" pageNumber="520">Malthodes josephi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257B5FF8BFFD6FE581439FDE4FBD1" box="[480,562,1111,1132]" pageId="7" pageNumber="520" rank="species">
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||
<emphasis id="B901EACEFF8BFFD6FE581439FDE4FBD1" bold="true" box="[480,562,1111,1132]" pageId="7" pageNumber="520">sp. nov.</emphasis>
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</taxonomicNameLabel>
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in Baltic amber. A: Holotype, lateral view, scale bar = 0.5 mm. B: Holotype, lateral view, scale bar = 0.1 mm. C: Holotype, detail of
|
||
<emphasis id="B901EACEFF8BFFD6FD31141BFD25FB36" box="[649,755,1141,1163]" italics="true" pageId="7" pageNumber="520">terminalia</emphasis>
|
||
, scale bar = 0.1 mm. D: Holotype, detail of
|
||
<emphasis id="B901EACEFF8BFFD6FB0E141BFAF6FB36" box="[1206,1312,1141,1163]" italics="true" pageId="7" pageNumber="520">terminalia</emphasis>
|
||
, scale bar = 0.2 mm.
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</paragraph>
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</caption>
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<caption id="DF0A6654FF8BFFD6FF2F171AFAA3F837" box="[151,1397,1908,1930]" httpUri="https://zenodo.org/record/1457505/files/figure.png" pageId="7" pageNumber="520" startId="7.[151,250,1908,1930]" targetBox="[348,1246,1329,1842]" targetPageId="7">
|
||
<paragraph id="8BCA36DCFF8BFFD6FF2F171AFAA3F837" blockId="7.[151,1397,1908,1930]" box="[151,1397,1908,1930]" pageId="7" pageNumber="520">
|
||
<emphasis id="B901EACEFF8BFFD6FF2F171AFEC4F834" bold="true" box="[151,274,1908,1930]" pageId="7" pageNumber="520">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C754D5FFF8BFFD6FEA1171AFE05F837" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[281,467,1908,1930]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="520" phylum="Arthropoda" rank="species" species="josephi" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8BFFD6FEA1171AFE05F837" box="[281,467,1908,1930]" italics="true" pageId="7" pageNumber="520">Malthodes josephi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8BFFD6FE61171AFDFFF834" box="[473,553,1908,1929]" pageId="7" pageNumber="520" rank="species">
|
||
<emphasis id="B901EACEFF8BFFD6FE61171AFDFFF834" bold="true" box="[473,553,1908,1929]" pageId="7" pageNumber="520">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in Baltic amber. Line drawing of the last abdominal segments, scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA36DCFF8BFFD6FF7F17D8FD02F84E" blockId="7.[199,1409,1974,2035]" pageId="7" pageNumber="520">
|
||
<emphasis id="B901EACEFF8BFFD6FF7F17D8FE83F872" bold="true" box="[199,341,1974,1999]" pageId="7" pageNumber="520">
|
||
<typeStatus id="54CE887EFF8BFFD6FF7F17D8FED4F872" box="[199,258,1974,1999]" pageId="7" pageNumber="520">Type</typeStatus>
|
||
strata.
|
||
</emphasis>
|
||
Baltic amber, Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.8–33.9 Mya.
|
||
<emphasis id="B901EACEFF8BFFD6FF7F17B4FEA7F84E" bold="true" box="[199,369,2010,2035]" pageId="7" pageNumber="520">Syninclusions.</emphasis>
|
||
Wood remains and air bubbles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF84FFD9FF7F10F9FEF0FD4D" pageId="8" pageNumber="521" type="diagnosis">
|
||
<paragraph id="8BCA36DCFF84FFD9FF7F10F9FEF0FD4D" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="521">
|
||
<emphasis id="B901EACEFF84FFD9FF7F10F9FE06FF0D" bold="true" box="[199,464,151,176]" pageId="8" pageNumber="521">Differential diagnosis.</emphasis>
|
||
Characters such as the globular and pointed last palpomere, the short elytra and the strongly modified last abdominal segments (both tergites and sternites) suggest that the new species belongs to the genus
|
||
<taxonomicName id="4C754D5FFF84FFD9FF6710B1FE83FF45" authorityName="Kiesenwetter" authorityYear="1852" box="[223,341,223,248]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF84FFD9FF6710B1FE83FF45" box="[223,341,223,248]" italics="true" pageId="8" pageNumber="521">Malthodes</emphasis>
|
||
</taxonomicName>
|
||
. No other fossil of
|
||
<taxonomicName id="4C754D5FFF84FFD9FD9610B1FD72FF45" authorityName="Kiesenwetter" authorityYear="1852" box="[558,676,223,248]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF84FFD9FD9610B1FD72FF45" box="[558,676,223,248]" italics="true" pageId="8" pageNumber="521">Malthodes</emphasis>
|
||
</taxonomicName>
|
||
shows the same features of the
|
||
<emphasis id="B901EACEFF84FFD9FBB110B1FBA8FF45" box="[1033,1150,223,248]" italics="true" pageId="8" pageNumber="521">terminalia</emphasis>
|
||
of the new species (
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FAD910B1FEE1FEA1" pageId="8" pageNumber="521" refId="ref5639" refString="Fanti, F. & Vitali, F. (2017) Key to fossil Malthininae, with description of two new species in Baltic amber (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 17 (1), 19 - 27." type="journal article">Fanti & Vitali 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FEFA116AFD80FEA1" box="[322,598,260,285]" pageId="8" pageNumber="521" refId="ref5544" refString="Fanti, F. & Michalski, A. R. (2018) An unusual fossil Malthodes with long elytra (Insecta Coleoptera Cantharidae). Giornale italiano di Entomologia, 15 (63), 127 - 132." type="journal article">Fanti & Michalski 2018</bibRefCitation>
|
||
), and no extant species of Central-North Europe and Alps is similar. Only the group of
|
||
<taxonomicName id="4C754D5FFF84FFD9FE9D1147FD53FEFD" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" box="[293,645,295,320]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="alpicola">
|
||
<emphasis id="B901EACEFF84FFD9FE9D1147FE70FEFD" box="[293,422,295,320]" italics="true" pageId="8" pageNumber="521">M. alpicola</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FE151149FD53FEFD" box="[429,645,295,320]" pageId="8" pageNumber="521" refId="ref6256" refString="Kiesenwetter, E. A. H. von (1852) Beitrage zu einer Monographie der Malthinen. Linnaea entomologica, 7, 239 - 324, tabs. 1 - 2." type="journal article">Kiesenwetter, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C754D5FFF84FFD9FD2D1147FCC0FEFD" authorityName="Kiesenwetter" authorityYear="1852" box="[661,790,295,320]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="alpicola">
|
||
<emphasis id="B901EACEFF84FFD9FD2D1147FCC0FEFD" box="[661,790,295,320]" italics="true" pageId="8" pageNumber="521">M. alpicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C754D5FFF84FFD9FC9B1147FBAAFEFD" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" box="[803,1148,295,320]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="guttifer">
|
||
<emphasis id="B901EACEFF84FFD9FC9B1147FC4DFEFD" box="[803,923,296,320]" italics="true" pageId="8" pageNumber="521">M. guttifer</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FC191149FBAAFEFD" box="[929,1148,295,320]" pageId="8" pageNumber="521" refId="ref6256" refString="Kiesenwetter, E. A. H. von (1852) Beitrage zu einer Monographie der Malthinen. Linnaea entomologica, 7, 239 - 324, tabs. 1 - 2." type="journal article">Kiesenwetter, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C754D5FFF84FFD9FB3F1147FF04FED8" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="spretus">
|
||
<emphasis id="B901EACEFF84FFD9FB3F1147FB2AFEFD" box="[1159,1276,297,320]" italics="true" pageId="8" pageNumber="521">M. spretus</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FABA1149FF04FED8" pageId="8" pageNumber="521" refId="ref6256" refString="Kiesenwetter, E. A. H. von (1852) Beitrage zu einer Monographie der Malthinen. Linnaea entomologica, 7, 239 - 324, tabs. 1 - 2." type="journal article">Kiesenwetter, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C754D5FFF84FFD9FF671123FE2FFED8" authority="Fiori, 1905" authorityName="Fiori" authorityYear="1905" box="[223,505,332,357]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="bertolinii">
|
||
<emphasis id="B901EACEFF84FFD9FF671123FEB9FED8" box="[223,367,332,357]" italics="true" pageId="8" pageNumber="521">M. bertolinii</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FEC11122FE2FFED8" box="[377,505,332,357]" pageId="8" pageNumber="521" refId="ref5726" refString="Fiori, A. (1905) Revisione delle specie italiane a me note del genere Malthodes Kiesw. Rivista Coleotterologica Italiana, 3 (12), 221 - 252, 2 tabs." type="journal article">Fiori, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
) might be seen as similar, although not from a strictly anatomical point of view, because of the last sternite (sternite 9) that is shaped as a wide, apically forked membrane (
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FB051101FA99FE35" box="[1213,1359,367,392]" pageId="8" pageNumber="521" refId="ref6551" refString="Liberti, G. (2011) Le specie di Malthodes Kiesenwetter, 1852 delle Alpi Marittime e Liguri (Coleoptera, Cantharidae). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 103, 147 - 246." type="journal article">Liberti 2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FAE51101FA4EFE35" box="[1373,1432,367,392]" pageId="8" pageNumber="521" refId="ref6597" refString="Liberti, G. (2015) Le specie di Malthodes Kiesenwetter, 1852 d'Italia nord-occidentale (Coleoptera, Cantharidae) (Malthodes di Lombardia: 2 ° contributo alla conoscenza dei Malthodes italiani). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 107, 1 - 151." type="journal article">2015</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FF2F11FAFF07FE10" box="[151,209,404,429]" pageId="8" pageNumber="521" refId="ref6654" refString="Liberti, G. (2016) Le specie di Malthodes Kiesenwetter, 1852 delle Alpi (Coleoptera, Cantharidae) (Malthodes del Nord-Est: 3 ° contributo alla conoscenza del genere Malthodes in Italia). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 108, 89 - 198." type="journal article">2016</bibRefCitation>
|
||
) and possessing a similar last tergite (tergite 10). However, these species are much bigger in size, and tendentially have a yellow spot (more or less evident) on each elytral apex. The group of
|
||
<taxonomicName id="4C754D5FFF84FFD9FB2F11D7FF0EFE48" authority="(Linnaeus, 1758)" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B901EACEFF84FFD9FB2F11D7FACAFE6D" box="[1175,1308,441,464]" italics="true" pageId="8" pageNumber="521">M. minimus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FA9511D9FF06FE48" pageId="8" pageNumber="521" refId="ref6771" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp." type="book">Linnaeus, 1758</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is only vaguely similar, with the last tergite lobe-shaped and little developed, and the last sternite without median membrane. The important diagnostic characters of
|
||
<taxonomicName id="4C754D5FFF84FFD9FCF61191FC12FDA5" authorityName="Kiesenwetter" authorityYear="1852" box="[846,964,511,536]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF84FFD9FCF61191FC12FDA5" box="[846,964,511,536]" italics="true" pageId="8" pageNumber="521">Malthodes</emphasis>
|
||
</taxonomicName>
|
||
are based on the male’s terminalia and habitus, as the morphology of the fossils is nearly identical to the living species. It is, however, interesting to note that, out of all the fossil species known, only
|
||
<taxonomicName id="4C754D5FFF84FFD9FD191227FC6BFDE2" authority="Kazantsev" authorityName="Kazantsev" box="[673,957,583,608]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="perkovskyi">
|
||
<emphasis id="B901EACEFF84FFD9FD191227FC96FDDD" box="[673,832,583,608]" italics="true" pageId="8" pageNumber="521">M. perkovskyi</emphasis>
|
||
Kazantsev
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49B1F83EFF84FFD9FBBC1226FB99FDE2" box="[1028,1103,584,607]" country="Ukraine" name="Rivne" pageId="8" pageNumber="521">Rovno</collectingRegion>
|
||
amber (
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FB151226FAB1FDDD" box="[1197,1383,584,608]" pageId="8" pageNumber="521" refId="ref6052" refString="Kazantsev, S. V. (2010) New Malthodes (Insecta: Cantharidae: Coleoptera) from the Rovno Amber (Upper Eocene of Ukraine). Russian Entomological Journal, 19 (2), 105 - 107." type="journal article">Kazantsev 2010</bibRefCitation>
|
||
) has the elytra with two yellow spots apically (
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FD3B1202FCCCFD39" box="[643,794,620,644]" pageId="8" pageNumber="521" refId="ref5803" refString="Forster, B. (1891) Die Insekten des " Plattigen Steinmergels " von Brunstatt. Abhandlungen zur Geologischen Specialkarte von Elsass-Lothringen, 3 (5), 335 - 593 + [1], 6 pls. (+ 6), + [1]." type="book chapter">Förster 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FC911202FB58FD38" box="[809,1166,620,645]" pageId="8" pageNumber="521" refId="ref6518" refString="Kuska, A. & Kupryjanowicz, J. (2005) Soldier beetles (Coleoptera: Cantharidae) from Baltic amber. Polskie Pismo Entomologiczne, 74, 309 - 316." type="journal article">Kuśka & Kupryjanowicz 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FB251202FA41FD39" box="[1181,1431,620,645]" pageId="8" pageNumber="521" refId="ref6484" refString="Kuska, A. & Kania, I. (2010) New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa, 2400, 49 - 56." type="journal article">Kuśka & Kania 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FF2F12FEFE20FD15" box="[151,502,655,680]" pageId="8" pageNumber="521" refId="ref6207" refString="Kazantsev, S. V. & Perkovsky, E. E. (2014) A new Malthodes and some other interesting soldier beetles (Coleoptera: Cantharidae) from Late Eocene Rovno amber. Russian Entomological Journal, 23 (2), 113 - 116." type="journal article">Kazantsev & Perkovsky 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FDBE12E1FD42FD15" box="[518,660,655,680]" pageId="8" pageNumber="521" refId="ref5322" refString="Fanti, F. (2017 b) Malthodes michalskii: a new species of Cantharidae from Baltic amber (Coleoptera). Giornale italiano di Entomologia, 14 (62), 685 - 690." type="journal article">Fanti 2017b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FD1C12E1FC46FD15" box="[676,912,655,680]" pageId="8" pageNumber="521" refId="ref5639" refString="Fanti, F. & Vitali, F. (2017) Key to fossil Malthininae, with description of two new species in Baltic amber (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 17 (1), 19 - 27." type="journal article">Fanti & Vitali 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FC1812E1FB69FD15" box="[928,1215,655,680]" pageId="8" pageNumber="521" refId="ref5544" refString="Fanti, F. & Michalski, A. R. (2018) An unusual fossil Malthodes with long elytra (Insecta Coleoptera Cantharidae). Giornale italiano di Entomologia, 15 (63), 127 - 132." type="journal article">Fanti & Michalski 2018</bibRefCitation>
|
||
). These are indeed present, even if sometimes difficult to see, in many living species (
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FCC712DAFBC6FD70" box="[895,1040,692,717]" pageId="8" pageNumber="521" refId="ref5684" refString="Fender, K. M. (1951) The Malthini of North America (Coleoptera-Cantharidae). The American Midland Naturalist, 46 (3), 513 - 629. https: // doi. org / 10.2307 / 2421804" type="journal article">Fender 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FBA412DAFB6CFD71" box="[1052,1210,692,717]" pageId="8" pageNumber="521" refId="ref7201" refString="Wittmer, W. (1970) Zur Kenntnis der Gattung Malthodes Kies. (Col., Cantharidae) (48. Beitrag zur Kenntnis der palaearktischen Cantharidae). Entomologische Arbeiten aus dem Museum G. Frey, 21, 13 - 107." type="journal article">Wittmer 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FB7D12DAFA84FD71" box="[1221,1362,692,717]" pageId="8" pageNumber="521" refId="ref6551" refString="Liberti, G. (2011) Le specie di Malthodes Kiesenwetter, 1852 delle Alpi Marittime e Liguri (Coleoptera, Cantharidae). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 103, 147 - 246." type="journal article">Liberti 2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FAE512DAFA4EFD70" box="[1373,1432,692,717]" pageId="8" pageNumber="521" refId="ref6597" refString="Liberti, G. (2015) Le specie di Malthodes Kiesenwetter, 1852 d'Italia nord-occidentale (Coleoptera, Cantharidae) (Malthodes di Lombardia: 2 ° contributo alla conoscenza dei Malthodes italiani). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 107, 1 - 151." type="journal article">2015</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FF2F12B9FF04FD4D" box="[151,210,727,752]" pageId="8" pageNumber="521" refId="ref6654" refString="Liberti, G. (2016) Le specie di Malthodes Kiesenwetter, 1852 delle Alpi (Coleoptera, Cantharidae) (Malthodes del Nord-Est: 3 ° contributo alla conoscenza del genere Malthodes in Italia). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 108, 89 - 198." type="journal article">2016</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44B2DFF84FFD9FF6512B6FECCFD4D" box="[221,282,728,752]" pageId="8" pageNumber="521" refId="ref6713" refString="Liberti, G. (2017) Le specie appenniniche di Malthodes Kiesenwetter, 1852 (Coleoptera, Cantharidae) (Malthodes d'Italia peninsulare: 4 ° contributo alla conoscenza del genere Malthodes in Italia). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 110, 33 - 164." type="journal article">2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF84FFD9FF7F1292FC84FC85" pageId="8" pageNumber="521" type="discussion">
|
||
<paragraph id="8BCA36DCFF84FFD9FF7F1292FC84FC85" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="521">
|
||
<emphasis id="B901EACEFF84FFD9FF7F1292FEEDFCA8" bold="true" box="[199,315,764,789]" pageId="8" pageNumber="521">Remarks.</emphasis>
|
||
The amber piece measures
|
||
<date id="FFCB101CFF84FFD9FD321292FD07FCA9" box="[650,721,764,788]" pageId="8" pageNumber="521" value="1922-10-28">28x22</date>
|
||
x
|
||
<quantity id="4C8D9B39FF84FFD9FD671292FCE2FCA9" box="[735,820,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="521" unit="mm" value="10.0">10 mm</quantity>
|
||
, is rectangularly shaped and weighs
|
||
<quantity id="4C8D9B39FF84FFD9FB531292FAB7FCA8" box="[1259,1377,764,789]" metricMagnitude="-3" metricUnit="kg" metricValue="2.8" pageId="8" pageNumber="521" unit="g" value="2.8">2.8 grams</quantity>
|
||
. The amber is opaque yellow and there are scratches on the surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |