180 lines
22 KiB
XML
180 lines
22 KiB
XML
<document id="5FB02AAFD2A07FA546C446FE8D48F728" ID-DOI="10.5852/ejt.2024.925.2465" ID-ISSN="2118-9773" ID-Zenodo-Dep="10824968" ID-ZooBank="5848AF9A-F01D-45F1-8626-E489AC15EE7F" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1710590167793" checkinUser="plazi" docAuthor="Génier, François & Darling, James D. G." docDate="2024" docId="03DC87B7EF26FFBDFDA9FD73C8D7E5A0" docLanguage="en" docName="EJT.2024.925.220-267.pdf" docOrigin="European Journal of Taxonomy 925" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2465/10955" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andinocopris Génier & Darling 2024, gen. nov." docType="treatment" docUuid="DB47A9F1-784F-4431-A7AA-2044987EF94D" docUuidSource="ZooBank" docVersion="4" lastPageNumber="223" masterDocId="FFE5FFCFEF25FFBEFFCFFF9FC93BE277" masterDocTitle="Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe)" masterLastPageNumber="267" masterPageNumber="220" pageNumber="223" updateTime="1710872936379" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="13EDC110E485A6195B61B4CD0F193A82">Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe)</mods:title>
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<mods:namePart id="CFE6AA1EC1C40077343FC50547B16303">Génier, François</mods:namePart>
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<mods:affiliation id="8C40715BC7C22FD31746E1A4CB89C807">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:nameIdentifier id="EC825B592A87549525E9343C9FB9C3E1" type="email">FGenier@nature.ca</mods:nameIdentifier>
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<mods:namePart id="A60EC1555072EDE9B7F9F581968DF32B">Darling, James D. G.</mods:namePart>
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<mods:affiliation id="C55527BBDBC6F2CB9EF14FBBCDFFAD5A">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:nameIdentifier id="F8025AF0EFCD8F405943C140880F5857" type="email">james21darling@gmail.com</mods:nameIdentifier>
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<mods:title id="A0385D12E14095DB20E97A9456E058C4">European Journal of Taxonomy</mods:title>
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<mods:part id="C93A3C631E850E0775C134AF5EA58F0F">
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<mods:date id="B2AEB1C680512A9695FA7C060EB02AB6">2024</mods:date>
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<mods:number id="5912C133583911A6F5230B7230D739FB">2024-03-15</mods:number>
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<mods:number id="C4552E12E1E22B3684CA6018571FAA2E">925</mods:number>
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<mods:identifier id="EE92CEF3276DFC021273B065EE03826C" type="DOI">10.5852/ejt.2024.925.2465</mods:identifier>
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<treatment id="03DC87B7EF26FFBDFDA9FD73C8D7E5A0" ID-DOI="http://doi.org/10.5281/zenodo.10824976" ID-Zenodo-Dep="10824976" LSID="urn:lsid:zoobank.org:act:DB47A9F1-784F-4431-A7AA-2044987EF94D" httpUri="http://treatment.plazi.org/id/03DC87B7EF26FFBDFDA9FD73C8D7E5A0" lastPageNumber="223" pageId="3" pageNumber="223">
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<subSubSection id="C36F652AEF26FFBDFDA9FD73CAF5E170" box="[614,974,747,775]" pageId="3" pageNumber="223" type="nomenclature">
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<paragraph id="8BCA36A1EF26FFBDFDA9FD73CAF5E170" blockId="3.[367,1221,747,807]" box="[614,974,747,775]" pageId="3" pageNumber="223">
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<heading id="D08281CDEF26FFBDFDA9FD73CAF5E170" box="[614,974,747,775]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="223" reason="2">
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Genus
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<taxonomicName id="4C754D22EF26FFBDFD76FD74CA67E171" authority="Génier & Darling, 2024" authorityName="Génier & Darling" authorityYear="2024" box="[697,860,747,774]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B901EAB3EF26FFBDFD76FD74CA67E171" bold="true" box="[697,860,747,774]" italics="true" pageId="3" pageNumber="223">Andinocopris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257C8EF26FFBDFCACFD72CAF5E170" box="[867,974,749,775]" pageId="3" pageNumber="223" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C36F652AEF26FFBDFEA0FC92CDFEE150" box="[367,1221,781,807]" pageId="3" pageNumber="223" type="description">
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<paragraph id="8BCA36A1EF26FFBDFEA0FC92CDFEE150" blockId="3.[367,1221,747,807]" box="[367,1221,781,807]" pageId="3" pageNumber="223">
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<uri id="FFE43AA3EF26FFBDFEA0FC92CDFEE150" box="[367,1221,781,807]" pageId="3" pageNumber="223">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD30C74EF26FFBDFD42FC92CDFEE150" box="[653,1221,781,807]" pageId="3" pageNumber="223">DB47A9F1-784F-4431-A7AA-2044987EF94D</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF26FFBDFF72FCD3C863E111" box="[189,344,844,870]" pageId="3" pageNumber="223" type="type_taxon">
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<paragraph id="8BCA36A1EF26FFBDFF72FCD3C863E111" blockId="3.[189,344,844,870]" box="[189,344,844,870]" pageId="3" pageNumber="223">
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<heading id="D08281CDEF26FFBDFF72FCD3C863E111" bold="true" box="[189,344,844,870]" fontSize="11" level="3" pageId="3" pageNumber="223" reason="3">
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<emphasis id="B901EAB3EF26FFBDFF72FCD3C863E111" bold="true" box="[189,344,844,870]" pageId="3" pageNumber="223">
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<typeStatus id="54CE8803EF26FFBDFF72FCD3C9C1E111" box="[189,250,844,870]" pageId="3" pageNumber="223">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF26FFBDFF72FCE5CA19E1E4" box="[189,802,889,915]" pageId="3" pageNumber="223" type="reference_group">
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<paragraph id="8BCA36A1EF26FFBDFF72FCE5CA19E1E4" blockId="3.[189,802,889,915]" box="[189,802,889,915]" pageId="3" pageNumber="223">
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<taxonomicName id="4C754D22EF26FFBDFF72FCE5CB12E1E4" ID-CoL="4HZSV" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[189,553,889,915]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="species" species="achamas">
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<emphasis id="B901EAB3EF26FFBDFF72FCE5C8BDE1E4" box="[189,390,889,915]" italics="true" pageId="3" pageNumber="223">Pinotus achamas</emphasis>
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<bibRefCitation id="EFE44B50EF26FFBDFE41FCE6CB12E1E4" author="Harold E. von" box="[398,553,889,915]" pageId="3" pageNumber="223" pagination="95 - 104" refId="ref26084" refString="Harold E. von 1867. Diagnosen neuer Coprophagen. Coleopterologische Hefte 2: 95 - 104. Available from https: // www. biodiversitylibrary. org / page / 16069770 [accessed 6 Feb. 2024]." type="journal article" year="1867">Harold, 1867</bibRefCitation>
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</taxonomicName>
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, present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF26FFBDFF72FC27CAE7E6D3" pageId="3" pageNumber="223" type="diagnosis">
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<paragraph id="8BCA36A1EF26FFBDFF72FC27C8F2E1A5" blockId="3.[189,457,952,978]" box="[189,457,952,978]" pageId="3" pageNumber="223">
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<heading id="D08281CDEF26FFBDFF72FC27C8F2E1A5" bold="true" box="[189,457,952,978]" fontSize="11" level="3" pageId="3" pageNumber="223" reason="3">
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<emphasis id="B901EAB3EF26FFBDFF72FC27C8F2E1A5" bold="true" box="[189,457,952,978]" pageId="3" pageNumber="223">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA36A1EF26FFBDFF72FC7BCAE7E6D3" blockId="3.[189,1400,996,1188]" pageId="3" pageNumber="223">
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Dorsal ocular width greater than one quarter of interocular distance. Lateral pronotal carina and pronotal edge joined anteriorly and posteriorly, forming closed ellipse. Pronotal armament in the form of an overhanging ridge or bifurcating projection in large individuals of both sexes. Elytral stria 8 usually absent, remnant present on apical elytral hump and connected to stria five in some individuals of
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<taxonomicName id="4C754D22EF26FFBDFF2FFBF6C857E6F4" authorityName="Harold" authorityYear="1867" box="[224,364,1129,1155]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="species" species="achamas">
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<emphasis id="B901EAB3EF26FFBDFF2FFBF6C857E6F4" box="[224,364,1129,1155]" italics="true" pageId="3" pageNumber="223">A. achamas</emphasis>
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</taxonomicName>
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(
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<figureCitation id="134E2A24EF26FFBDFE4FFBF6C8E0E6F4" box="[384,475,1129,1155]" captionStart="Figs 58–64. 58" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[234,1353,267,1780]" captionTargetId="figure-17@31.[232,1356,265,1815]" captionTargetPageId="31" captionText="Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing." figureDoi="http://doi.org/10.5281/zenodo.10824994" httpUri="https://zenodo.org/record/10824994/files/figure.png" pageId="3" pageNumber="223">Fig. 62</figureCitation>
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). Meso-metasternal suture posteriorly arcuate (
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<figureCitation id="134E2A24EF26FFBDFBDAFBF6CD4BE6F4" box="[1045,1136,1129,1155]" captionStart="Figs 58–64. 58" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[234,1353,267,1780]" captionTargetId="figure-17@31.[232,1356,265,1815]" captionTargetPageId="31" captionText="Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing." figureDoi="http://doi.org/10.5281/zenodo.10824994" httpUri="https://zenodo.org/record/10824994/files/figure.png" pageId="3" pageNumber="223">Fig. 60</figureCitation>
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).
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<specimenCount id="9D73FD28EF26FFBDFB46FBF6CDA4E6F4" box="[1161,1183,1129,1155]" count="1" pageId="3" pageNumber="223" type="male">♂</specimenCount>
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FLP endophallite sigmoidal, abruptly narrowing and inflecting medially (
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<figureCitation id="134E2A24EF26FFBDFC89FB15CAF6E6D3" box="[838,973,1162,1188]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="3" pageNumber="223">Figs 31–32</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF26FFBDFF72FB56CA0EE725" pageId="3" pageNumber="223" type="etymology">
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<paragraph id="8BCA36A1EF26FFBDFF72FB56C87FE694" blockId="3.[189,324,1225,1251]" box="[189,324,1225,1251]" pageId="3" pageNumber="223">
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<heading id="D08281CDEF26FFBDFF72FB56C87FE694" bold="true" box="[189,324,1225,1251]" fontSize="11" level="3" pageId="3" pageNumber="223" reason="3">
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<emphasis id="B901EAB3EF26FFBDFF72FB56C87FE694" bold="true" box="[189,324,1225,1251]" pageId="3" pageNumber="223">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA36A1EF26FFBDFF72FB6ACA0EE725" blockId="3.[189,1399,1269,1362]" pageId="3" pageNumber="223">
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||
<taxonomicName id="4C754D22EF26FFBDFF72FB6AC860E778" authorityName="Génier & Darling" authorityYear="2024" box="[189,347,1269,1295]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B901EAB3EF26FFBDFF72FB6AC860E778" box="[189,347,1269,1295]" italics="true" pageId="3" pageNumber="223">Andinocopris</emphasis>
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||
</taxonomicName>
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||
is a combination of the New Latin adjective ‘
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<emphasis id="B901EAB3EF26FFBDFCADFB6ACA84E778" box="[866,959,1269,1295]" italics="true" pageId="3" pageNumber="223">andinus</emphasis>
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’ and the Ancient Greek noun ‘κόπρος’ (‘
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<emphasis id="B901EAB3EF26FFBDFF1EFA88C81BE746" box="[209,288,1303,1329]" italics="true" pageId="3" pageNumber="223">kópros</emphasis>
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’). The former refers to the Andes, where the genus occurs at high elevations from
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<collectingCountry id="F3627631EF26FFBDFB2AFA88CC62E746" box="[1253,1369,1303,1329]" name="Colombia" pageId="3" pageNumber="223">Colombia</collectingCountry>
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to
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<collectingCountry id="F3627631EF26FFBDFF72FAA7C9CDE725" box="[189,246,1336,1362]" name="Peru" pageId="3" pageNumber="223">Peru</collectingCountry>
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, and the latter refers to its association with dung.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF26FFBDFF72FAE8CA9EE51C" pageId="3" pageNumber="223" type="description">
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||
<paragraph id="8BCA36A1EF26FFBDFF72FAE8C875E7E6" blockId="3.[189,334,1399,1425]" box="[189,334,1399,1425]" pageId="3" pageNumber="223">
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||
<heading id="D08281CDEF26FFBDFF72FAE8C875E7E6" bold="true" box="[189,334,1399,1425]" fontSize="11" level="3" pageId="3" pageNumber="223" reason="3">
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||
<emphasis id="B901EAB3EF26FFBDFF72FAE8C875E7E6" bold="true" box="[189,334,1399,1425]" pageId="3" pageNumber="223">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA36A1EF26FFBDFF72FA3CCA9EE51C" blockId="3.[187,1399,1443,1900]" pageId="3" pageNumber="223">
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Body length
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<quantity id="4C8D9B44EF26FFBDFEAAFA3CC8D4E7C9" box="[357,495,1443,1470]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.4" metricValueMin="2.0" pageId="3" pageNumber="223" unit="mm" value="27.0" valueMax="34.0" valueMin="20.0">20–34 mm</quantity>
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. Anteroventral clypeal tooth, rather small, transverse and varying from truncated to acute in frontal view. Dorsal ocular width greater than one-quarter interocular distance. Anterior pronotal edge straight (
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<taxonomicName id="4C754D22EF26FFBDFD9BFA79CBD8E788" authorityName="Harold" authorityYear="1867" box="[596,739,1509,1536]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="species" species="achamas">
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<emphasis id="B901EAB3EF26FFBDFD9BFA79CBD8E788" box="[596,739,1509,1536]" italics="true" pageId="3" pageNumber="223">A. achamas</emphasis>
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</taxonomicName>
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) or slightly sinuous (
|
||
<taxonomicName id="4C754D22EF26FFBDFC35FA79CDB8E788" authorityName="Génier & Darling" authorityYear="2024" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[1018,1155,1509,1536]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF26FFBDFC35FA79CDB8E788" box="[1018,1155,1509,1536]" italics="true" pageId="3" pageNumber="223">A. buckleyi</emphasis>
|
||
</taxonomicName>
|
||
) medially. Anterior pronotal membrane very wide laterally, transition between membrane and anterior edge of pronotum ill-defined medially. Anteromedial pronotal surface smooth. Lateral pronotal carina and pronotal edge joined anteriorly and posteriorly, forming closed carinated ellipse. Pronotal armament in large individuals of both sexes forming overhanging ridge or bifurcating projection; ventral surface of overhanging armament uneven. Anterior apex of prosternum broadly rounded. Underside of elytron with long semi-erected setae. Elytral stria 8 usually absent, remanent present on apical elytral hump and connected to stria five in some individuals of
|
||
<taxonomicName id="4C754D22EF26FFBDFDE1F952CB8EE490" authorityName="Harold" authorityYear="1867" box="[558,693,1741,1767]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="223" phylum="Arthropoda" rank="species" species="achamas">
|
||
<emphasis id="B901EAB3EF26FFBDFDE1F952CB8EE490" box="[558,693,1741,1767]" italics="true" pageId="3" pageNumber="223">A. achamas</emphasis>
|
||
</taxonomicName>
|
||
. Hindwing with AP reduced (
|
||
<figureCitation id="134E2A24EF26FFBDFBC5F952CDA8E490" box="[1034,1171,1741,1767]" captionStart="Figs 37–41. 37–38" captionStartId="23.[189,242,1892,1918]" captionTargetBox="[257,1314,294,1816]" captionTargetId="figure-17@23.[221,1366,265,1843]" captionTargetPageId="23" captionText="Figs 37–41. 37–38. Genital segment, dorsal view. 37. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 38. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 39–41. Hindwing, dorsal view. 39. Andinocopris achamas (Harold, 1867) gen. et comb. nov. 40. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov. 41. Homocopris torulosus." figureDoi="http://doi.org/10.5281/zenodo.10824988" httpUri="https://zenodo.org/record/10824988/files/figure.png" pageId="3" pageNumber="223">Figs 39–40</figureCitation>
|
||
). Meso-metasternal suture posteriorly arcuate (
|
||
<figureCitation id="134E2A24EF26FFBDFE38F971CB74E57F" box="[503,591,1774,1800]" captionStart="Figs 58–64. 58" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[234,1353,267,1780]" captionTargetId="figure-17@31.[232,1356,265,1815]" captionTargetPageId="31" captionText="Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing." figureDoi="http://doi.org/10.5281/zenodo.10824994" httpUri="https://zenodo.org/record/10824994/files/figure.png" pageId="3" pageNumber="223">Fig. 60</figureCitation>
|
||
). Surface of median metasternal lobe convex, some individuals with shallow and narrow longitudinal sulcus; surface setigerous anterolaterally, impunctate or finely punctate throughout. Pygidial surface finely punctate.
|
||
<specimenCount id="9D73FD28EF26FFBDFD18F8AECBD6E53C" box="[727,749,1841,1867]" count="1" pageId="3" pageNumber="223" type="male">♂</specimenCount>
|
||
paramere tapering slightly in lateral view; apex blunt. FLP endophallite sigmoidal, narrowing and inflecting centrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF26FFBDFF72F80FC8D7E5A0" box="[189,492,1936,2007]" pageId="3" pageNumber="223" type="distribution">
|
||
<paragraph id="8BCA36A1EF26FFBDFF72F80FC86DE5DD" blockId="3.[189,342,1936,1962]" box="[189,342,1936,1962]" pageId="3" pageNumber="223">
|
||
<heading id="D08281CDEF26FFBDFF72F80FC86DE5DD" bold="true" box="[189,342,1936,1962]" fontSize="11" level="3" pageId="3" pageNumber="223" reason="3">
|
||
<emphasis id="B901EAB3EF26FFBDFF72F80FC86DE5DD" bold="true" box="[189,342,1936,1962]" pageId="3" pageNumber="223">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF26FFBDFF72F822C8D7E5A0" blockId="3.[189,492,1981,2007]" box="[189,492,1981,2007]" pageId="3" pageNumber="223">
|
||
<collectingCountry id="F3627631EF26FFBDFF72F822C80FE5A0" box="[189,308,1981,2007]" name="Colombia" pageId="3" pageNumber="223">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3627631EF26FFBDFE8FF822C89FE5A0" box="[320,420,1981,2007]" name="Ecuador" pageId="3" pageNumber="223">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3627631EF26FFBDFE60F822C8D2E5A0" box="[431,489,1981,2007]" name="Peru" pageId="3" pageNumber="223">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |