treatments-xml/data/03/DC/87/03DC8795FFC2FFCEFEAAFB67FB06FE3D.xml

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<mods:title>Clarias microspilus, a new walking catfish (Teleostei: Clariidae) from northern Sumatra, Indonesia</mods:title>
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<mods:namePart>Ng, Heok Hee</mods:namePart>
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<mods:namePart>Hadiaty, Renny K.</mods:namePart>
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<mods:date>2011</mods:date>
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<emphasis bold="true" box="[170,337,1280,1307]" pageId="1" pageNumber="1578">Type material</emphasis>
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: 
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SL; 
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: Nangroe 
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, Kabupaten 
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Selatan, Sungai Lembang at Pasilembang (
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’N &amp; 
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’E), coll. 
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et al., 
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<collectingDate box="[600,717,1430,1456]" pageId="1" pageNumber="1578" value="1999-02-26">26.ii.1999</collectingDate>
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.
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<typeStatus box="[208,321,1468,1494]" pageId="1" pageNumber="1578">Paratypes</typeStatus>
: 
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(3), 
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SL
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; 
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(1), 
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SL; 
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: 
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<collectingMunicipality box="[237,479,1542,1568]" pageId="1" pageNumber="1578">Darussalam province</collectingMunicipality>
. 
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<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFD3FF9F1FCF8F987" box="[685,770,1542,1568]" county="Nangroe" municipality="Darussalam province" name="Selatan" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Selatan</location>
, 
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(
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’N &amp; 
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’E), coll. 
<collectorName box="[621,771,1579,1606]" pageId="1" pageNumber="1578">H.B. Munaf</collectorName>
, 
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. 
<materialsCitation collectionCode="MZB" collectorName="Selatan &amp; Sungai Lembang &amp; Pucuk Lembang" county="Nangroe" location="Kabupaten" municipality="Darussalam Province" pageId="1" pageNumber="1578" specimenCode="MZB 8705" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
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(1), 
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SL; 
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: 
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<collectingRegion box="[288,347,1654,1680]" country="Indonesia" name="Aceh" pageId="1" pageNumber="1578">Aceh</collectingRegion>
<collectingMunicipality box="[369,626,1654,1680]" pageId="1" pageNumber="1578">Darussalam Province</collectingMunicipality>
, 
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, 
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at 
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(
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’N &amp; 
<superScript attach="none" box="[283,351,1728,1755]" fontSize="6" pageId="1" pageNumber="1578">97028</superScript>
’E), coll
</materialsCitation>
. 
<materialsCitation collectingDate="1997-09-02" collectionCode="R" collectorName="K. Hadiaty &amp; Mun'im" pageId="1" pageNumber="1578" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<collectionCode box="[455,477,1729,1755]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="1578">R</collectionCode>
. 
<collectorName box="[481,605,1728,1755]" pageId="1" pageNumber="1578">K. Hadiaty</collectorName>
&amp; A. 
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, 
<date box="[170,293,1766,1792]" pageId="1" pageNumber="1578" value="1997-09-02">
<collectingDate box="[170,293,1766,1792]" pageId="1" pageNumber="1578" value="1997-09-02">02.ix.1997</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation collectingDate="1999-02-21" collectionCode="MZB" collectorName="Suaq Belimbing &amp; S. Wirjoatmodjo" county="Nangroe" location="Kabupaten" municipality="Darussalam Province" pageId="1" pageNumber="1578" specimenCode="MZB 8713" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<specimenCode box="[309,438,1766,1792]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="1578" type="Museum">MZB 8713</specimenCode>
(1), 
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SL; 
<collectingRegion box="[674,772,1766,1792]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="1578">Sumatra</collectingRegion>
: 
<collectingCounty box="[170,266,1804,1830]" pageId="1" pageNumber="1578">Nangroe</collectingCounty>
<collectingRegion box="[272,331,1803,1829]" country="Indonesia" name="Aceh" pageId="1" pageNumber="1578">Aceh</collectingRegion>
<collectingMunicipality box="[337,578,1803,1829]" pageId="1" pageNumber="1578">Darussalam Province</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFDDFF8FCFD3CF882" box="[589,710,1803,1829]" county="Nangroe" municipality="Darussalam Province" name="Kabupaten" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Kabupaten</location>
<collectingRegion box="[716,775,1803,1829]" country="Indonesia" name="Aceh" pageId="1" pageNumber="1578">Aceh</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFF38F8C7FF05F8ED" box="[170,255,1840,1866]" county="Nangroe" municipality="Darussalam Province" name="Selatan" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Selatan</location>
, swamp at 
<collectorName box="[375,555,1840,1867]" pageId="1" pageNumber="1578">Suaq Belimbing</collectorName>
(
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’N &amp; 
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’E), coll. 
<collectorName box="[233,429,1878,1904]" pageId="1" pageNumber="1578">S. Wirjoatmodjo</collectorName>
, 
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<collectingDate box="[447,564,1878,1904]" pageId="1" pageNumber="1578" value="1999-02-21">21.ii.1999</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation accessionNumber="ZRC 51917" collectionCode="ZRC" collectorName="Tapaktuan &amp; Subulussalam" county="Nangroe" location="Kabupaten" municipality="Darussalam Province" pageId="1" pageNumber="1578" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
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51917
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(4), 
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SL; 
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: 
<collectingCounty box="[596,692,1916,1942]" pageId="1" pageNumber="1578">Nangroe</collectingCounty>
<collectingRegion box="[716,775,1915,1941]" country="Indonesia" name="Aceh" pageId="1" pageNumber="1578">Aceh</collectingRegion>
<collectingMunicipality box="[170,413,1952,1978]" pageId="1" pageNumber="1578">Darussalam Province</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFE3BF857FDD8F81D" box="[425,546,1952,1978]" county="Nangroe" municipality="Darussalam Province" name="Kabupaten" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Kabupaten</location>
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<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFDFFF857FD38F81D" box="[621,706,1952,1978]" county="Nangroe" municipality="Darussalam Province" name="Selatan" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Selatan</location>
, 
<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFD5DF856FEB5F847" county="Nangroe" municipality="Darussalam Province" name="Desa Ujung Padang" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Desa Ujung Padang</location>
, blackwater swamp along 
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– 
<collectorName box="[813,969,235,261]" pageId="1" pageNumber="1578">Subulussalam</collectorName>
road (
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’55.2”N &amp; 
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’17.6”E), coll
</materialsCitation>
. 
<materialsCitation box="[868,1139,272,299]" collectingDate="2009-04-18" collectionCode="T" location="Sim" pageId="1" pageNumber="1578" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<collectionCode box="[868,886,273,299]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="1578">T</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:03DC8795FFC2FFCEFEAAFB67FB06FE3D:8EAA6058FFC2FFCBFC13FEE7FC54FE8D" box="[897,942,272,298]" name="Sim" pageId="1" pageNumber="1578" stateProvince="Sumatera Utara">Sim</location>
et al., 
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<collectingDate box="[1018,1139,272,298]" pageId="1" pageNumber="1578" value="2009-04-18">18.iv.2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="1578" type="diagnosis">
<paragraph blockId="1.[813,1425,347,1531]" box="[813,928,347,373]" pageId="1" pageNumber="1578">
<emphasis bold="true" box="[813,928,347,373]" pageId="1" pageNumber="1578">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="1.[813,1425,347,1531]" pageId="1" pageNumber="1578">
<taxonomicName authorityName="Ng &amp; Hadiaty" authorityYear="2011" box="[850,1074,384,410]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="microspilus">
<emphasis box="[850,1074,384,410]" italics="true" pageId="1" pageNumber="1578">Clarias microspilus</emphasis>
</taxonomicName>
can be distinguished from all Southeast Asian congeners, except for 
<taxonomicName box="[1246,1410,422,448]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="intermedius">
<emphasis box="[1246,1410,422,448]" italics="true" pageId="1" pageNumber="1578">C. intermedius</emphasis>
</taxonomicName>
, 
<taxonomicName box="[813,951,459,485]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="insolitus">
<emphasis box="[813,951,459,485]" italics="true" pageId="1" pageNumber="1578">C. insolitus</emphasis>
</taxonomicName>
, 
<taxonomicName box="[975,1144,459,485]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="meladerma">
<emphasis box="[975,1144,459,485]" italics="true" pageId="1" pageNumber="1578">C. meladerma</emphasis>
</taxonomicName>
, 
<taxonomicName box="[1168,1317,459,485]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="olivaceus">
<emphasis box="[1168,1317,459,485]" italics="true" pageId="1" pageNumber="1578">C. olivaceus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="planiceps">
<emphasis italics="true" pageId="1" pageNumber="1578">C. planiceps</emphasis>
</taxonomicName>
, in having a serrated (vs. smooth or rugose, with irregular bumps) anterior margin of the pectoral spine. It differs from 
<taxonomicName box="[1045,1208,571,597]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="intermedius">
<emphasis box="[1045,1208,571,597]" italics="true" pageId="1" pageNumber="1578">C. intermedius</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1260,1417,571,597]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="meladerma">
<emphasis box="[1260,1417,571,597]" italics="true" pageId="1" pageNumber="1578">C. meladerma</emphasis>
</taxonomicName>
by a longer distance between the tip of the occipital process and the base of the first dorsal-fin ray (6.5–9.2 % SL vs. 2.8–5.6), from 
<taxonomicName box="[1090,1217,683,709]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="insolitus">
<emphasis box="[1090,1217,683,709]" italics="true" pageId="1" pageNumber="1578">C. insolitus</emphasis>
</taxonomicName>
, 
<taxonomicName box="[1231,1370,683,709]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="olivaceus">
<emphasis box="[1231,1370,683,709]" italics="true" pageId="1" pageNumber="1578">C. olivaceus</emphasis>
</taxonomicName>
and 
<taxonomicName box="[813,950,720,747]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="planiceps">
<emphasis box="[813,950,720,747]" italics="true" pageId="1" pageNumber="1578">C. planiceps</emphasis>
</taxonomicName>
by a deeper body (14.9–18.9 % SL vs. 9.7– 15.2) and wider head (18.6–21.7 % SL vs. 14.0–18.7). 
<taxonomicName authorityName="Ng &amp; Hadiaty" authorityYear="2011" box="[813,1032,795,821]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="microspilus">
<emphasis box="[813,1032,795,821]" italics="true" pageId="1" pageNumber="1578">Clarias microspilus</emphasis>
</taxonomicName>
further differs from 
<taxonomicName box="[1254,1417,795,821]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="intermedius">
<emphasis box="[1254,1417,795,821]" italics="true" pageId="1" pageNumber="1578">C. intermedius</emphasis>
</taxonomicName>
in having fewer rays in the median fins (64–68 vs. 70– 72 dorsal-fin rays and 51–56 vs. 61–62 anal-fin rays), from 
<taxonomicName box="[874,1002,907,933]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="insolitus">
<emphasis box="[874,1002,907,933]" italics="true" pageId="1" pageNumber="1578">C. insolitus</emphasis>
</taxonomicName>
in having a shorter distance between the tip of the occipital process and the base of the first dorsal-fin ray (6.5–9.2 % SL vs. 10.3–12.4), and from 
<taxonomicName box="[813,975,1019,1045]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="meladerma">
<emphasis box="[813,975,1019,1045]" italics="true" pageId="1" pageNumber="1578">C. meladerma</emphasis>
</taxonomicName>
in having more serrations (22–34 vs. 14–22) on the anterior edge of the pectoral spine. It is further distinguished from 
<taxonomicName box="[1143,1283,1094,1120]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="olivaceus">
<emphasis box="[1143,1283,1094,1120]" italics="true" pageId="1" pageNumber="1578">C. olivaceus</emphasis>
</taxonomicName>
in having a more posteriorly situated frontal fontanel (the anterior tip of the frontal fontanel reaching to a line through the middle of the orbit vs. to anterior orbital margin), a smooth (vs. weakly serrated) posterodorsal margin of the pectoral spine, a shorter distance between the tip of the occipital process and the base of the first dorsal-fin ray (6.5–9.2 % SL vs. 9.3–11.1) and a wider occipital process (31.7–40.8 %HL vs. 25.5–31.5), and from 
<taxonomicName class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="1578" phylum="Chordata" rank="species" species="planiceps">
<emphasis italics="true" pageId="1" pageNumber="1578">C. planiceps</emphasis>
</taxonomicName>
in having a deeper head (12.9–16.0 % SL vs. 9.5–11.5) and smaller interorbital distance (40.5–44.5 % HL vs. 46.4–49.9).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="1581" pageId="1" pageNumber="1578" type="description">
<paragraph blockId="1.[813,1418,1579,2016]" box="[813,951,1579,1605]" pageId="1" pageNumber="1578">
<emphasis bold="true" box="[813,951,1579,1605]" pageId="1" pageNumber="1578">Description</emphasis>
</paragraph>
<paragraph blockId="1.[813,1418,1579,2016]" lastBlockId="2.[170,775,235,672]" lastPageId="2" lastPageNumber="1579" pageId="1" pageNumber="1578">
Biometric data in 
<tableCitation box="[1052,1136,1616,1643]" captionStart="Table 1" captionStartId="2.[170,224,743,763]" captionText="Table 1. Biometric data for Clarias microspilus (n = 11)." pageId="1" pageNumber="1578">Table 1</tableCitation>
. Head depressed; dorsal profile slightly convex and ventral profile almost straight. Bony elements of dorsal surface of head covered with thick skin; bones not readily visible, but sutures sometimes evident. Anterior pair of nostrils tubular and medial to maxillary barbel base. Posterior pair of nostrils bordered by nasal barbels anteriorly and fleshy membrane posteriorly; posteromedial to maxillary barbel base. Eye ovoid, horizontal axis longest, subcutaneous; located dorsolaterally on head. Anterior fontanel short and squat (“shoe-shaped” of 
<bibRefCitation author="Teugels, G. G." box="[170,334,235,262]" pageId="2" pageNumber="1579" pagination="1 - 199" refId="ref6393" refString="Teugels, G. G. (1986). A systematic revision of the African species of the genus Clarias (Pisces; Clariidae). Annales du Musee Royal de l'Afrique Centrale (Zoologie) 247: 1 - 199." type="journal article" year="1986">Teugels, 1986</bibRefCitation>
); anterior tip reaching to line through middle of orbits. Occipital process rounded. Gill openings narrow, extending from dorsal-most point of pectoral-fin base to isthmus. Gill membranes free from isthmus but united to each other with 8 (n=5) branchiostegal rays. First branchial arch with 5+13 (n=2) or 5+14* (n=3) gill rakers.
</paragraph>
<paragraph blockId="2.[170,775,235,672]" lastBlockId="3.[170,775,1580,2017]" lastPageId="3" lastPageNumber="1580" pageId="2" pageNumber="1579">Mouth subterminal, with fleshy, plicate lips. Oral teeth small and in irregular rows on all tooth-bearing surfaces. Premaxillary tooth band rectangular, with median notch on posterior edge. Dentary tooth band much narrower than premaxillary tooth band at symphysis, tapering laterally. Vomerine tooth band unpaired, continuous across midline, crescentic and smoothly arched along anterior and posterior margins. Premaxillary and dentary teeth viliform; vomerine teeth subgranular. Barbels in four pairs; long and slender with thick fleshy bases. Maxillary barbel extending to base of fifth or sixth dorsal-fin ray. Nasal barbel, extending nearly to level of base of last pectoral-fin ray. Inner mandibular-barbel origin close to midline; barbel thicker and longer than nasal barbel and extending beyond base of last pectoral-fin ray. Outer mandibular barbel originating posterolateral to inner mandibular barbel, extending nearly to base of first pelvic-fin ray.</paragraph>
<caption box="[170,725,743,763]" pageId="2" pageNumber="1579" startId="2.[170,224,743,763]">
<paragraph blockId="2.[170,725,743,763]" box="[170,725,743,763]" pageId="2" pageNumber="1579">
Table 1. Biometric data for 
<taxonomicName authorityName="Ng &amp; Hadiaty" authorityYear="2011" box="[442,639,743,763]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="1579" phylum="Chordata" rank="species" species="microspilus">
<emphasis bold="true" box="[442,639,743,763]" italics="true" pageId="2" pageNumber="1579">Clarias microspilus</emphasis>
</taxonomicName>
(n = 11).
</paragraph>
</caption>
<paragraph pageId="2" pageNumber="1579">
<table box="[186,976,788,2002]" gridcols="4" gridrows="32" pageId="2" pageNumber="1579">
<tr box="[186,976,788,808]" gridrow="0" pageId="2" pageNumber="1579" rowspan-0="1">
<td box="[543,634,788,808]" gridcol="1" gridrow="0" pageId="2" pageNumber="1579">
<emphasis bold="true" box="[543,634,788,808]" pageId="2" pageNumber="1579">Holotype</emphasis>
</td>
<td box="[686,804,788,808]" gridcol="2" gridrow="0" pageId="2" pageNumber="1579">
<emphasis bold="true" box="[713,778,788,808]" pageId="2" pageNumber="1579">Range</emphasis>
</td>
<td box="[847,976,788,808]" gridcol="3" gridrow="0" pageId="2" pageNumber="1579">
<emphasis bold="true" box="[857,965,788,808]" pageId="2" pageNumber="1579">Mean ± SD</emphasis>
</td>
</tr>
<tr box="[186,976,826,846]" gridrow="1" pageId="2" pageNumber="1579">
<td box="[186,976,826,846]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="2" pageNumber="1579">
<emphasis bold="true" box="[186,232,826,846]" pageId="2" pageNumber="1579">%SL</emphasis>
</td>
</tr>
<tr box="[186,976,864,884]" gridrow="2" pageId="2" pageNumber="1579">
<td box="[186,480,864,884]" gridcol="0" gridrow="2" pageId="2" pageNumber="1579">Predorsal length</td>
<td box="[543,634,864,884]" gridcol="1" gridrow="2" pageId="2" pageNumber="1579">36.4</td>
<td box="[686,804,864,884]" gridcol="2" gridrow="2" pageId="2" pageNumber="1579">31.4–36.4</td>
<td box="[847,976,864,884]" gridcol="3" gridrow="2" pageId="2" pageNumber="1579">33.8 ± 1.91</td>
</tr>
<tr box="[186,976,902,922]" gridrow="3" pageId="2" pageNumber="1579">
<td box="[186,480,902,922]" gridcol="0" gridrow="3" pageId="2" pageNumber="1579">Preanal length</td>
<td box="[543,634,902,922]" gridcol="1" gridrow="3" pageId="2" pageNumber="1579">49.0</td>
<td box="[686,804,902,922]" gridcol="2" gridrow="3" pageId="2" pageNumber="1579">49.0–55.0</td>
<td box="[847,976,902,922]" gridcol="3" gridrow="3" pageId="2" pageNumber="1579">51.7 ± 2.13</td>
</tr>
<tr box="[186,976,940,960]" gridrow="4" pageId="2" pageNumber="1579">
<td box="[186,480,940,960]" gridcol="0" gridrow="4" pageId="2" pageNumber="1579">Prepelvic length</td>
<td box="[543,634,940,960]" gridcol="1" gridrow="4" pageId="2" pageNumber="1579">42.4</td>
<td box="[686,804,940,960]" gridcol="2" gridrow="4" pageId="2" pageNumber="1579">42.4–46.7</td>
<td box="[847,976,940,960]" gridcol="3" gridrow="4" pageId="2" pageNumber="1579">44.5 ± 1.71</td>
</tr>
<tr box="[186,976,977,997]" gridrow="5" pageId="2" pageNumber="1579">
<td box="[186,480,977,997]" gridcol="0" gridrow="5" pageId="2" pageNumber="1579">Prepectoral length</td>
<td box="[543,634,977,997]" gridcol="1" gridrow="5" pageId="2" pageNumber="1579">22.0</td>
<td box="[686,804,977,997]" gridcol="2" gridrow="5" pageId="2" pageNumber="1579">20.3–24.3</td>
<td box="[847,976,977,997]" gridcol="3" gridrow="5" pageId="2" pageNumber="1579">22.3 ± 1.54</td>
</tr>
<tr box="[186,976,1015,1035]" gridrow="6" pageId="2" pageNumber="1579">
<td box="[186,480,1015,1035]" gridcol="0" gridrow="6" pageId="2" pageNumber="1579">Length of dorsal-fin base</td>
<td box="[543,634,1015,1035]" gridcol="1" gridrow="6" pageId="2" pageNumber="1579">68.3</td>
<td box="[686,804,1015,1035]" gridcol="2" gridrow="6" pageId="2" pageNumber="1579">64.7–69.0</td>
<td box="[847,976,1015,1035]" gridcol="3" gridrow="6" pageId="2" pageNumber="1579">67.3 ± 1.71</td>
</tr>
<tr box="[186,976,1053,1073]" gridrow="7" pageId="2" pageNumber="1579">
<td box="[186,480,1053,1073]" gridcol="0" gridrow="7" pageId="2" pageNumber="1579">Anal-fin length</td>
<td box="[543,634,1053,1073]" gridcol="1" gridrow="7" pageId="2" pageNumber="1579">48.6</td>
<td box="[686,804,1053,1073]" gridcol="2" gridrow="7" pageId="2" pageNumber="1579">47.4–50.5</td>
<td box="[847,976,1053,1073]" gridcol="3" gridrow="7" pageId="2" pageNumber="1579">49.2 ± 1.21</td>
</tr>
<tr box="[186,976,1091,1111]" gridrow="8" pageId="2" pageNumber="1579">
<td box="[186,480,1091,1111]" gridcol="0" gridrow="8" pageId="2" pageNumber="1579">Pelvic-fin length</td>
<td box="[543,634,1091,1111]" gridcol="1" gridrow="8" pageId="2" pageNumber="1579">10.9</td>
<td box="[686,804,1091,1111]" gridcol="2" gridrow="8" pageId="2" pageNumber="1579">9.2–10.9</td>
<td box="[847,976,1091,1111]" gridcol="3" gridrow="8" pageId="2" pageNumber="1579">9.8 ± 0.66</td>
</tr>
<tr box="[186,976,1129,1149]" gridrow="9" pageId="2" pageNumber="1579">
<td box="[186,480,1129,1149]" gridcol="0" gridrow="9" pageId="2" pageNumber="1579">Pectoral-fin length</td>
<td box="[543,634,1129,1149]" gridcol="1" gridrow="9" pageId="2" pageNumber="1579">17.0</td>
<td box="[686,804,1129,1149]" gridcol="2" gridrow="9" pageId="2" pageNumber="1579">12.7–17.2</td>
<td box="[847,976,1129,1149]" gridcol="3" gridrow="9" pageId="2" pageNumber="1579">14.9 ± 1.94</td>
</tr>
<tr box="[186,976,1166,1186]" gridrow="10" pageId="2" pageNumber="1579">
<td box="[186,480,1166,1186]" gridcol="0" gridrow="10" pageId="2" pageNumber="1579">Pectoral-spine length</td>
<td box="[543,634,1166,1186]" gridcol="1" gridrow="10" pageId="2" pageNumber="1579">12.5</td>
<td box="[686,804,1166,1186]" gridcol="2" gridrow="10" pageId="2" pageNumber="1579">9.2–12.5</td>
<td box="[847,976,1166,1186]" gridcol="3" gridrow="10" pageId="2" pageNumber="1579">10.5 ± 1.16</td>
</tr>
<tr box="[186,976,1204,1224]" gridrow="11" pageId="2" pageNumber="1579">
<td box="[186,480,1204,1224]" gridcol="0" gridrow="11" pageId="2" pageNumber="1579">Caudal-fin length</td>
<td box="[543,634,1204,1224]" gridcol="1" gridrow="11" pageId="2" pageNumber="1579">18.7</td>
<td box="[686,804,1204,1224]" gridcol="2" gridrow="11" pageId="2" pageNumber="1579">14.3–19.2</td>
<td box="[847,976,1204,1224]" gridcol="3" gridrow="11" pageId="2" pageNumber="1579">17.2 ± 1.75</td>
</tr>
<tr box="[186,976,1240,1286]" gridrow="12" pageId="2" pageNumber="1579">
<td box="[186,480,1240,1286]" gridcol="0" gridrow="12" pageId="2" pageNumber="1579">Distance between occipital process and dorsal fin</td>
<td box="[543,634,1240,1286]" gridcol="1" gridrow="12" pageId="2" pageNumber="1579">8.9</td>
<td box="[686,804,1240,1286]" gridcol="2" gridrow="12" pageId="2" pageNumber="1579">6.5–9.2</td>
<td box="[847,976,1240,1286]" gridcol="3" gridrow="12" pageId="2" pageNumber="1579">7.9 ± 1.07</td>
</tr>
<tr box="[186,976,1302,1322]" gridrow="13" pageId="2" pageNumber="1579">
<td box="[186,480,1302,1322]" gridcol="0" gridrow="13" pageId="2" pageNumber="1579">Body depth at anus</td>
<td box="[543,634,1302,1322]" gridcol="1" gridrow="13" pageId="2" pageNumber="1579">18.9</td>
<td box="[686,804,1302,1322]" gridcol="2" gridrow="13" pageId="2" pageNumber="1579">14.9–18.9</td>
<td box="[847,976,1302,1322]" gridcol="3" gridrow="13" pageId="2" pageNumber="1579">17.2 ± 1.48</td>
</tr>
<tr box="[186,976,1340,1360]" gridrow="14" pageId="2" pageNumber="1579">
<td box="[186,480,1340,1360]" gridcol="0" gridrow="14" pageId="2" pageNumber="1579">Caudal peduncle depth</td>
<td box="[543,634,1340,1360]" gridcol="1" gridrow="14" pageId="2" pageNumber="1579">8.1</td>
<td box="[686,804,1340,1360]" gridcol="2" gridrow="14" pageId="2" pageNumber="1579">7.7–9.4</td>
<td box="[847,976,1340,1360]" gridcol="3" gridrow="14" pageId="2" pageNumber="1579">8.4 ± 0.62</td>
</tr>
<tr box="[186,976,1378,1398]" gridrow="15" pageId="2" pageNumber="1579">
<td box="[186,480,1378,1398]" gridcol="0" gridrow="15" pageId="2" pageNumber="1579">Head length</td>
<td box="[543,634,1378,1398]" gridcol="1" gridrow="15" pageId="2" pageNumber="1579">28.1</td>
<td box="[686,804,1378,1398]" gridcol="2" gridrow="15" pageId="2" pageNumber="1579">25.3–28.1</td>
<td box="[847,976,1378,1398]" gridcol="3" gridrow="15" pageId="2" pageNumber="1579">26.9 ± 0.94</td>
</tr>
<tr box="[186,976,1416,1436]" gridrow="16" pageId="2" pageNumber="1579">
<td box="[186,480,1416,1436]" gridcol="0" gridrow="16" pageId="2" pageNumber="1579">Head width</td>
<td box="[543,634,1416,1436]" gridcol="1" gridrow="16" pageId="2" pageNumber="1579">21.7</td>
<td box="[686,804,1416,1436]" gridcol="2" gridrow="16" pageId="2" pageNumber="1579">18.6–21.7</td>
<td box="[847,976,1416,1436]" gridcol="3" gridrow="16" pageId="2" pageNumber="1579">19.8 ± 1.04</td>
</tr>
<tr box="[186,976,1453,1473]" gridrow="17" pageId="2" pageNumber="1579">
<td box="[186,480,1453,1473]" gridcol="0" gridrow="17" pageId="2" pageNumber="1579">Head depth</td>
<td box="[543,634,1453,1473]" gridcol="1" gridrow="17" pageId="2" pageNumber="1579">15.5</td>
<td box="[686,804,1453,1473]" gridcol="2" gridrow="17" pageId="2" pageNumber="1579">12.9–16.0</td>
<td box="[847,976,1453,1473]" gridcol="3" gridrow="17" pageId="2" pageNumber="1579">14.6 ± 1.18</td>
</tr>
<tr box="[186,976,1491,1511]" gridrow="18" pageId="2" pageNumber="1579">
<td box="[186,976,1491,1511]" colspan="4" colspanRight="3" gridcol="0" gridrow="18" pageId="2" pageNumber="1579">
<emphasis bold="true" box="[186,233,1491,1511]" pageId="2" pageNumber="1579">%HL</emphasis>
</td>
</tr>
<tr box="[186,976,1529,1549]" gridrow="19" pageId="2" pageNumber="1579">
<td box="[186,480,1529,1549]" gridcol="0" gridrow="19" pageId="2" pageNumber="1579">Snout length</td>
<td box="[543,634,1529,1549]" gridcol="1" gridrow="19" pageId="2" pageNumber="1579">31.2</td>
<td box="[686,804,1529,1549]" gridcol="2" gridrow="19" pageId="2" pageNumber="1579">26.6–32.0</td>
<td box="[847,976,1529,1549]" gridcol="3" gridrow="19" pageId="2" pageNumber="1579">29.4 ± 2.09</td>
</tr>
<tr box="[186,976,1567,1587]" gridrow="20" pageId="2" pageNumber="1579">
<td box="[186,480,1567,1587]" gridcol="0" gridrow="20" pageId="2" pageNumber="1579">Interorbital distance</td>
<td box="[543,634,1567,1587]" gridcol="1" gridrow="20" pageId="2" pageNumber="1579">44.0</td>
<td box="[686,804,1567,1587]" gridcol="2" gridrow="20" pageId="2" pageNumber="1579">40.5–44.5</td>
<td box="[847,976,1567,1587]" gridcol="3" gridrow="20" pageId="2" pageNumber="1579">43.0 ± 1.42</td>
</tr>
<tr box="[186,976,1605,1625]" gridrow="21" pageId="2" pageNumber="1579">
<td box="[186,480,1605,1625]" gridcol="0" gridrow="21" pageId="2" pageNumber="1579">Eye diameter</td>
<td box="[543,634,1605,1625]" gridcol="1" gridrow="21" pageId="2" pageNumber="1579">6.3</td>
<td box="[686,804,1605,1625]" gridcol="2" gridrow="21" pageId="2" pageNumber="1579">6.0–7.2</td>
<td box="[847,976,1605,1625]" gridcol="3" gridrow="21" pageId="2" pageNumber="1579">6.3 ± 0.43</td>
</tr>
<tr box="[186,976,1642,1662]" gridrow="22" pageId="2" pageNumber="1579">
<td box="[186,480,1642,1662]" gridcol="0" gridrow="22" pageId="2" pageNumber="1579">Frontal fontanel length</td>
<td box="[543,634,1642,1662]" gridcol="1" gridrow="22" pageId="2" pageNumber="1579">17.3</td>
<td box="[686,804,1642,1662]" gridcol="2" gridrow="22" pageId="2" pageNumber="1579">11.5–17.3</td>
<td box="[847,976,1642,1662]" gridcol="3" gridrow="22" pageId="2" pageNumber="1579">14.4 ± 2.16</td>
</tr>
<tr box="[186,976,1680,1700]" gridrow="23" pageId="2" pageNumber="1579">
<td box="[186,480,1680,1700]" gridcol="0" gridrow="23" pageId="2" pageNumber="1579">Frontal fontanel width</td>
<td box="[543,634,1680,1700]" gridcol="1" gridrow="23" pageId="2" pageNumber="1579">3.4</td>
<td box="[686,804,1680,1700]" gridcol="2" gridrow="23" pageId="2" pageNumber="1579">3.4–6.7</td>
<td box="[847,976,1680,1700]" gridcol="3" gridrow="23" pageId="2" pageNumber="1579">5.3 ± 1.15</td>
</tr>
<tr box="[186,976,1718,1738]" gridrow="24" pageId="2" pageNumber="1579">
<td box="[186,480,1718,1738]" gridcol="0" gridrow="24" pageId="2" pageNumber="1579">Occipital fontanel length</td>
<td box="[543,634,1718,1738]" gridcol="1" gridrow="24" pageId="2" pageNumber="1579">8.4</td>
<td box="[686,804,1718,1738]" gridcol="2" gridrow="24" pageId="2" pageNumber="1579">7.1–12.8</td>
<td box="[847,976,1718,1738]" gridcol="3" gridrow="24" pageId="2" pageNumber="1579">9.6 ± 1.80</td>
</tr>
<tr box="[186,976,1755,1776]" gridrow="25" pageId="2" pageNumber="1579">
<td box="[186,480,1755,1776]" gridcol="0" gridrow="25" pageId="2" pageNumber="1579">Occipital fontanel width</td>
<td box="[543,634,1755,1776]" gridcol="1" gridrow="25" pageId="2" pageNumber="1579">4.2</td>
<td box="[686,804,1755,1776]" gridcol="2" gridrow="25" pageId="2" pageNumber="1579">3.9–5.7</td>
<td box="[847,976,1755,1776]" gridcol="3" gridrow="25" pageId="2" pageNumber="1579">4.8 ± 0.61</td>
</tr>
<tr box="[186,976,1793,1813]" gridrow="26" pageId="2" pageNumber="1579">
<td box="[186,480,1793,1813]" gridcol="0" gridrow="26" pageId="2" pageNumber="1579">Occipital process length</td>
<td box="[543,634,1793,1813]" gridcol="1" gridrow="26" pageId="2" pageNumber="1579">9.4</td>
<td box="[686,804,1793,1813]" gridcol="2" gridrow="26" pageId="2" pageNumber="1579">7.7–14.8</td>
<td box="[847,976,1793,1813]" gridcol="3" gridrow="26" pageId="2" pageNumber="1579">11.0 ± 2.39</td>
</tr>
<tr box="[186,976,1831,1851]" gridrow="27" pageId="2" pageNumber="1579">
<td box="[186,480,1831,1851]" gridcol="0" gridrow="27" pageId="2" pageNumber="1579">Occipital process width</td>
<td box="[543,634,1831,1851]" gridcol="1" gridrow="27" pageId="2" pageNumber="1579">40.8</td>
<td box="[686,804,1831,1851]" gridcol="2" gridrow="27" pageId="2" pageNumber="1579">31.7–40.8</td>
<td box="[847,976,1831,1851]" gridcol="3" gridrow="27" pageId="2" pageNumber="1579">36.8 ± 3.47</td>
</tr>
<tr box="[186,976,1869,1889]" gridrow="28" pageId="2" pageNumber="1579">
<td box="[186,480,1869,1889]" gridcol="0" gridrow="28" pageId="2" pageNumber="1579">Nasal barbel length</td>
<td box="[543,634,1869,1889]" gridcol="1" gridrow="28" pageId="2" pageNumber="1579">105.0</td>
<td box="[686,804,1869,1889]" gridcol="2" gridrow="28" pageId="2" pageNumber="1579">84.0–105.0</td>
<td box="[847,976,1869,1889]" gridcol="3" gridrow="28" pageId="2" pageNumber="1579">94.8 ± 7.17</td>
</tr>
<tr box="[186,976,1907,1927]" gridrow="29" pageId="2" pageNumber="1579">
<td box="[186,480,1907,1927]" gridcol="0" gridrow="29" pageId="2" pageNumber="1579">Maxillary barbel length</td>
<td box="[543,634,1907,1927]" gridcol="1" gridrow="29" pageId="2" pageNumber="1579">143.7</td>
<td box="[686,804,1907,1927]" gridcol="2" gridrow="29" pageId="2" pageNumber="1579">128.0–154.9</td>
<td box="[847,976,1907,1927]" gridcol="3" gridrow="29" pageId="2" pageNumber="1579">139.8 ± 11.30</td>
</tr>
<tr box="[186,976,1945,1965]" gridrow="30" pageId="2" pageNumber="1579">
<td box="[186,480,1945,1965]" gridcol="0" gridrow="30" pageId="2" pageNumber="1579">Inner mandibular barbel length</td>
<td box="[543,634,1945,1965]" gridcol="1" gridrow="30" pageId="2" pageNumber="1579">91.1</td>
<td box="[686,804,1945,1965]" gridcol="2" gridrow="30" pageId="2" pageNumber="1579">83.3–97.1</td>
<td box="[847,976,1945,1965]" gridcol="3" gridrow="30" pageId="2" pageNumber="1579">90.7 ± 5.80</td>
</tr>
<tr box="[186,976,1982,2002]" gridrow="31" pageId="2" pageNumber="1579">
<td box="[186,480,1982,2002]" gridcol="0" gridrow="31" pageId="2" pageNumber="1579">Outer mandibular barbel length</td>
<td box="[543,634,1982,2002]" gridcol="1" gridrow="31" pageId="2" pageNumber="1579">123.6</td>
<td box="[686,804,1982,2002]" gridcol="2" gridrow="31" pageId="2" pageNumber="1579">110.3–130.6</td>
<td box="[847,976,1982,2002]" gridcol="3" gridrow="31" pageId="2" pageNumber="1579">121.6 ± 7.52</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4986819" ID-Zenodo-Dep="4986819" box="[170,1245,1478,1498]" httpUri="https://zenodo.org/record/4986819/files/figure.png" pageId="3" pageNumber="1580" startId="3.[170,232,1478,1498]" targetBox="[170,1151,236,1466]" targetPageId="3">
<paragraph blockId="3.[170,1245,1478,1498]" box="[170,1245,1478,1498]" pageId="3" pageNumber="1580">
Image 1. 
<taxonomicName authority="Ng &amp; Hadiaty, 2011" authorityName="Ng &amp; Hadiaty" authorityYear="2011" box="[261,458,1478,1498]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="1580" phylum="Chordata" rank="species" species="microspilus" status="sp. nov.">
<emphasis bold="true" box="[261,458,1478,1498]" italics="true" pageId="3" pageNumber="1580">Clarias microspilus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[464,546,1478,1498]" pageId="3" pageNumber="1580" rank="species">sp. nov.</taxonomicNameLabel>
, MZB 8706, holotype, 136.1mm SL; Sumatra: Aceh, Sungai Lembang.
</paragraph>
</caption>
<paragraph blockId="3.[170,775,1580,2017]" pageId="3" pageNumber="1580">Body cylindrical, becoming compressed towards caudal peduncle. Dorsal profile rising very gently from tip of snout to origin of dorsal fin and thereafter almost horizontal to end of caudal peduncle. Ventral profile slightly convex to middle of head and thereafter almost horizontal to end of caudal peduncle. Skin smooth. Lateral line complete and midlateral in position. Vertebrae 20+40=60* (n=1), 19+42=61 (n=3), or 20+42=62 (n=1).</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4986821" ID-Zenodo-Dep="4986821" httpUri="https://zenodo.org/record/4986821/files/figure.png" pageId="3" pageNumber="1580" startId="3.[813,875,1965,1985]" targetBox="[813,1418,1729,1946]" targetPageId="3">
<paragraph blockId="3.[813,1391,1964,2010]" pageId="3" pageNumber="1580">
Image 2. Left pectoral spine of 
<taxonomicName authority=", MZB" authorityName="MZB" box="[1126,1330,1964,1985]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="1580" phylum="Chordata" rank="species" species="microspilus">
<emphasis bold="true" box="[1126,1273,1964,1985]" italics="true" pageId="3" pageNumber="1580">C. microspilus</emphasis>
, MZB
</taxonomicName>
8705, paratype, 175.7mm SL.
</paragraph>
</caption>
<paragraph blockId="3.[170,775,1580,2017]" lastBlockId="4.[170,775,235,822]" lastPageId="4" lastPageNumber="1581" pageId="3" pageNumber="1580">
Dorsal fin with long base, spanning posterior three- quarters of body; with 64* (n=1), 66 (n=3), or 68 (n=1) rays covered by thick layer of skin and without spine. Dorsal-fin margin straight, parallel to dorsal edge of body. Pectoral fin with small spine, sharply pointed at tip, and 8,i (n=5) rays. Almost entire length of anterior spine margin with 22–34 prominent serrations; posterior spine margin smooth or with uneven asperities (
<figureCitation box="[299,404,422,448]" captionStart="Image 2" captionStartId="3.[813,875,1965,1985]" captionTargetBox="[813,1418,1729,1946]" captionTargetId="figure-131@3.[801,1425,1593,2062]" captionTargetPageId="3" captionText="Image 2. Left pectoral spine of C. microspilus, MZB 8705, paratype, 175.7mm SL." figureDoi="http://doi.org/10.5281/zenodo.4986821" httpUri="https://zenodo.org/record/4986821/files/figure.png" pageId="4" pageNumber="1581">Image 2</figureCitation>
). Pectoral-fin margin straight anteriorly, convex posteriorly. Pelvic-fin origin at anterior third of body, with i,5 (n=5) rays and convex margin; tip of adpressed fin reaching base of first two or three anal-fin rays. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Anal fin with long base and 51* (n=1), 54 (n=2), or 56 (n=2) rays covered by thick layer of skin; margin straight and parallel to ventral edge of body. Caudal peduncle very short. Caudal fin rounded, with i,7,7,i (n=5) principal rays.
</paragraph>
<paragraph blockId="4.[170,776,869,1158]" box="[170,299,869,895]" pageId="4" pageNumber="1581">
<emphasis bold="true" box="[170,299,869,895]" pageId="4" pageNumber="1581">Coloration</emphasis>
</paragraph>
<paragraph blockId="4.[170,776,869,1158]" pageId="4" pageNumber="1581">In 70% ethanol: Dorsal and lateral surfaces of head and body violet-gray, fading to pale gray on ventral surfaces. Dorsal, anal and caudal fins violet-gray with very thin hyaline distal margin. Pectoral-fin rays violet-gray, with hyaline interradial membranes. Pelvic fins hyaline. Barbels and pectoral spines violetgray dorsally and light grey ventrally.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1581" type="distribution">
<paragraph blockId="4.[170,775,1205,1307]" box="[170,316,1205,1231]" pageId="4" pageNumber="1581">
<emphasis bold="true" box="[170,316,1205,1231]" pageId="4" pageNumber="1581">Distribution</emphasis>
</paragraph>
<paragraph blockId="4.[170,775,1205,1307]" pageId="4" pageNumber="1581">
Known from the short coastal rivers that drain the western face of the Leuser Mountain Range (
<figureCitation box="[675,741,1280,1307]" captionStart="Figure 1" captionStartId="4.[170,235,1990,2010]" captionTargetBox="[170,1417,1435,1972]" captionTargetId="figure-448@4.[170,1417,1435,1974]" captionTargetPageId="4" captionText="Figure 1. Collection localities of Clarias microspilus." figureDoi="http://doi.org/10.5281/zenodo.4986823" httpUri="https://zenodo.org/record/4986823/files/figure.png" pageId="4" pageNumber="1581">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1581" type="etymology">
<paragraph blockId="4.[813,1418,235,410]" box="[813,942,235,261]" pageId="4" pageNumber="1581">
<emphasis bold="true" box="[813,942,235,261]" pageId="4" pageNumber="1581">Etymology</emphasis>
</paragraph>
<paragraph blockId="4.[813,1418,235,410]" pageId="4" pageNumber="1581">From the Greek, mikros, meaning small, and, spilos, meaning spot. The name is used in reference to the very small white spots arranged in a longitudinal and several transverse series on the body.</paragraph>
</subSubSection>
</treatment>
</document>