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<document id="A307C718D303A3F212BC6A99498139A9" ID-DOI="10.11646/zootaxa.3869.4.6" ID-GBIF-Dataset="5781fab6-f58a-42d7-b9a2-891a99f1d51e" ID-ISSN="1175-5326" ID-Zenodo-Dep="230830" ID-ZooBank="39555505-B707-4172-BED0-06242CB096EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460924755262" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Vidthayanon, Chavalit" docDate="2014" docId="03DC878AFFC4FFFAFF067A15FA9AF5B0" docLanguage="en" docName="zt03869p434.pdf" docOrigin="Zootaxa 3869 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exostoma peregrinator Ng &amp; Vidthayanon, 2014, new species" docType="treatment" docVersion="8" lastPageNumber="430" masterDocId="FFE5FFF2FFCCFFF0FF917B3DFFC3F258" masterDocTitle="A review of the glyptosternine catfish genus Exostoma Blyth 1860 from Thailand, with descriptions of two new species (Teleostei: Siluriformes)" masterLastPageNumber="434" masterPageNumber="420" pageNumber="428" updateTime="1698362302441" updateUser="plazi">
<mods:mods id="92946F06ECECE53CF9A97E7E8925FA8E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="31AEDB0EF86D7F99F4590C18F4B74BAB">A review of the glyptosternine catfish genus Exostoma Blyth 1860 from Thailand, with descriptions of two new species (Teleostei: Siluriformes)</mods:title>
</mods:titleInfo>
<mods:name id="9B6025BC16DA26430E167115A15EE7EB" type="personal">
<mods:role id="1E13676763FDC7D8FA2A822023CAC58E">
<mods:roleTerm id="B6FAFA140738D0A4E3C220982D130161">Author</mods:roleTerm>
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<mods:namePart id="9DEB1B0D60E0621BF92979676EAE01D9">Ng, Heok Hee</mods:namePart>
</mods:name>
<mods:name id="4221FE3FF2A8E39E18081BE1E1399448" type="personal">
<mods:role id="630F090A1AA3E25B2BD5CAAB8B9820AD">
<mods:roleTerm id="7FF051ACFB092DCE2AE384EA3F34F7BA">Author</mods:roleTerm>
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<mods:namePart id="B61834BD28628CB00FA2FE4FB19DD100">Vidthayanon, Chavalit</mods:namePart>
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<mods:typeOfResource id="6B37CE609ED85C3D84E43CD976EDA952">text</mods:typeOfResource>
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<mods:titleInfo id="4D4C8126C7605FF3174D7FFB47933A3C">
<mods:title id="A19ACC88675EA38C400FEEA38805FB51">Zootaxa</mods:title>
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<mods:part id="ECF2DA4DFF1DA9BB818ABE88D7D4D09A">
<mods:date id="BEE0382983A2521AC8802383B2D9E6C6">2014</mods:date>
<mods:detail id="824FBFE3454E0CFEB8F06B2D4A871E5A" type="volume">
<mods:number id="F6DD9F4C9E5A38CCD885C66EC642FC8A">3869</mods:number>
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<mods:detail id="DCFFEA7933C35AC66FE1B84F14CBEE69" type="issue">
<mods:number id="E95F0C4BF2B91591C4E38B1ACF6262C4">4</mods:number>
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<mods:classification id="9B212E60E8746D632EE58FD506168D2A">journal article</mods:classification>
<mods:identifier id="B62E59BACC934990D38B7F4E13BEBD0B" type="DOI">10.11646/zootaxa.3869.4.6</mods:identifier>
<mods:identifier id="9BE80A91B4AE157DB8F3539FD28CE02B" type="GBIF-Dataset">5781fab6-f58a-42d7-b9a2-891a99f1d51e</mods:identifier>
<mods:identifier id="038AAF5FFB814A163C0F113DF8D97FC3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="663B66B94EC0ED2C6442F395BF9E9B15" type="Zenodo-Dep">230830</mods:identifier>
<mods:identifier id="0F6376F61D149052A319B484F3665E06" type="ZooBank">39555505-B707-4172-BED0-06242CB096EC</mods:identifier>
</mods:mods>
<treatment id="03DC878AFFC4FFFAFF067A15FA9AF5B0" ID-DOI="http://doi.org/10.5281/zenodo.5615980" ID-GBIF-Taxon="119589431" ID-Zenodo-Dep="5615980" LSID="urn:lsid:plazi:treatment:03DC878AFFC4FFFAFF067A15FA9AF5B0" httpUri="http://treatment.plazi.org/id/03DC878AFFC4FFFAFF067A15FA9AF5B0" lastPageId="10" lastPageNumber="430" pageId="8" pageNumber="428">
<subSubSection id="C36F6517FFC4FFF8FF067A15FC2CF3F2" pageId="8" pageNumber="428" type="nomenclature">
<paragraph id="8BCA369CFFC4FFF8FF067A15FDF4F318" blockId="8.[151,567,295,357]" box="[151,567,295,320]" pageId="8" pageNumber="428">
<heading id="D08281F0FFC4FFF8FF067A15FDF4F318" bold="true" box="[151,567,295,320]" fontSize="10" level="2" pageId="8" pageNumber="428" reason="2">
<emphasis id="B901EA8EFFC4FFF8FF067A15FDF4F318" bold="true" box="[151,567,295,320]" pageId="8" pageNumber="428">
<taxonomicName id="4C754D1FFFC4FFF8FF067A15FE5CF318" ID-CoL="3DQD6" box="[151,415,296,320]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="428" phylum="Chordata" rank="species" species="peregrinator" status="sp. nov.">
<emphasis id="B901EA8EFFC4FFF8FF067A15FE5CF318" bold="true" box="[151,415,296,320]" italics="true" pageId="8" pageNumber="428">Exostoma peregrinator</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23257F5FFC4FFF8FE3C7A1AFDF4F318" box="[429,567,295,320]" pageId="8" pageNumber="428" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA369CFFC4FFF8FF067A71FED1F33C" blockId="8.[151,567,295,357]" box="[151,274,332,357]" pageId="8" pageNumber="428">
(
<figureCitation id="134E2A19FFC4FFF8FF0E7A71FF2DF33D" box="[159,238,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[151,1436,655,1949]" captionTargetId="figure@8.[151,1436,655,1949]" captionTargetPageId="8" captionText="FIGURE 6. Exostoma peregrinator, NIFI 3831, holotype, 70.2 mm SL; Thailand: Ban Tun." httpUri="https://zenodo.org/record/230836/files/figure.png" pageId="8" pageNumber="428">Figs. 6</figureCitation>
,
<figureCitation id="134E2A19FFC4FFF8FF6B7A71FECAF33C" box="[250,265,332,356]" captionStart="FIGURE 7" captionStartId="10.[151,250,683,705]" captionTargetBox="[185,1396,221,590]" captionTargetId="figure@10.[150,1436,171,661]" captionTargetPageId="10" captionText="FIGURE 7. Exostoma peregrinator, NIFI 3831, holotype, 70.2 mm SL; photograph taken immediately after preservation, showing color in life." httpUri="https://zenodo.org/record/230837/files/figure.png" pageId="8" pageNumber="428">7</figureCitation>
)
</paragraph>
<paragraph id="8BCA369CFFC4FFF8FF067AA8FC2CF3F2" blockId="8.[151,1007,404,426]" box="[151,1007,404,426]" pageId="8" pageNumber="428">
<treatmentCitationGroup id="AB6511B2FFC4FFF8FF067AA8FC2CF3F2" box="[151,1007,404,426]" pageId="8" pageNumber="428">
<taxonomicName id="4C754D1FFFC4FFF8FF067AA8FEAAF3F2" ID-CoL="3DQCP" box="[151,361,404,426]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="428" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B901EA8EFFC4FFF8FF067AA8FEAAF3F2" box="[151,361,404,426]" italics="true" pageId="8" pageNumber="428">Exostoma berdmorei</emphasis>
</taxonomicName>
(non Blyth)—
<treatmentCitation id="0AD4108DFFC4FFF8FE6A7AA9FCE9F3F2" author="Vidthayanon" box="[507,810,404,426]" page="106" pageId="8" pageNumber="428" year="2005">
<bibRefCitation id="EFE44B6DFFC4FFF8FE6A7AA9FD39F3F2" author="Vidthayanon" box="[507,762,404,426]" pageId="8" pageNumber="428" refString="Vidthayanon, C., Termvidchakorn, A. &amp; Pe, M. (2005) Inland fishes of Myanmar. SEAFDEC, Bangkok, 160 pp." type="book" year="2005">
Vidthayanon
<emphasis id="B901EA8EFFC4FFF8FD147AA8FD7BF3F2" box="[645,696,404,426]" italics="true" pageId="8" pageNumber="428">et al.</emphasis>
, 2005
</bibRefCitation>
: 106
</treatmentCitation>
(photograph only).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36F6517FFC4FFF8FF067AEBFAC4F003" pageId="8" pageNumber="428" type="materials_examined">
<paragraph id="8BCA369CFFC4FFF8FF067AEBFC83F04B" blockId="8.[151,1436,470,603]" pageId="8" pageNumber="428">
<emphasis id="B901EA8EFFC4FFF8FF067AEBFE86F3B7" bold="true" box="[151,325,470,495]" pageId="8" pageNumber="428">
<typeStatus id="54CE883EFFC4FFF8FF067AEBFF11F3B7" box="[151,210,470,495]" pageId="8" pageNumber="428">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54CE883EFFC4FFF8FEDC7AEBFE79F3B7" box="[333,442,470,495]" pageId="8" pageNumber="428" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED64AE59FFC4FFF8FE557AEAFE39F3B6" box="[452,506,471,494]" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="8" pageNumber="428">NIFI</collectionCode>
3831,
<quantity id="4C8D9B79FFC4FFF8FDD87AEAFD6DF3B6" box="[585,686,471,495]" metricMagnitude="-2" metricUnit="m" metricValue="7.02" pageId="8" pageNumber="428" unit="mm" value="70.2">70.2 mm</quantity>
<collectionCode id="ED64AE59FFC4FFF8FD247AEAFD1AF3B7" box="[693,729,471,495]" pageId="8" pageNumber="428">SL</collectionCode>
;
<collectingCountry id="F362760CFFC4FFF8FD747AEBFC8EF3B7" box="[741,845,470,495]" name="Thailand" pageId="8" pageNumber="428">Thailand</collectingCountry>
: Mae Hong Son Province, Mae La Noi District, Ban Tun,
<geoCoordinate id="EE41505BFFC4FFF8FF407AC6FE96F04B" box="[209,341,507,531]" direction="north" orientation="latitude" pageId="8" pageNumber="428" precision="15" value="18.30861">18°18'31&quot;N</geoCoordinate>
<geoCoordinate id="EE41505BFFC4FFF8FECD7AC6FE11F04B" box="[348,466,507,531]" direction="east" orientation="longitude" pageId="8" pageNumber="428" precision="15" value="98.14361">98°8'37&quot;E</geoCoordinate>
; C. Vidthayanon,
<date id="FFCB105CFFC4FFF8FD0A7AC6FCF8F04B" box="[667,827,506,531]" pageId="8" pageNumber="428" value="1998-04-28">28 April 1998</date>
.
</paragraph>
<paragraph id="8BCA369CFFC4FFF8FF567922FAC4F003" blockId="8.[151,1436,470,603]" pageId="8" pageNumber="428">
<typeStatus id="54CE883EFFC4FFF8FF567922FEFAF06F" box="[199,313,543,567]" pageId="8" pageNumber="428" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED64AE59FFC4FFF8FED57922FEB8F06E" box="[324,379,543,566]" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="8" pageNumber="428">NIFI</collectionCode>
3762 (4),
<quantity id="4C8D9B79FFC4FFF8FE657923FD5BF06E" box="[500,664,542,567]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" metricValueMax="6.56" metricValueMin="4.56" pageId="8" pageNumber="428" unit="mm" value="55.599999999999994" valueMax="65.6" valueMin="45.6">45.665.6 mm</quantity>
<collectionCode id="ED64AE59FFC4FFF8FD317922FD07F06F" box="[672,708,543,567]" pageId="8" pageNumber="428">SL</collectionCode>
; locality as for
<typeStatus id="54CE883EFFC4FFF8FCE67923FC1EF06F" box="[887,989,542,567]" pageId="8" pageNumber="428" type="holotype">holotype</typeStatus>
; Mae Hong Son Fisheries Station staff, 2005.
<collectionCode id="ED64AE59FFC4FFF8FF4C797EFED0F002" box="[221,275,579,602]" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="8" pageNumber="428">NIFI</collectionCode>
3764 (8),
<quantity id="4C8D9B79FFC4FFF8FE19797FFDE8F002" box="[392,555,578,603]" metricMagnitude="-2" metricUnit="m" metricValue="4.535" metricValueMax="6.74" metricValueMin="2.33" pageId="8" pageNumber="428" unit="mm" value="45.35" valueMax="67.4" valueMin="23.3">23.367.4 mm</quantity>
<collectionCode id="ED64AE59FFC4FFF8FDA0797EFD96F003" box="[561,597,579,603]" pageId="8" pageNumber="428">SL</collectionCode>
;
<collectionCode id="ED64AE59FFC4FFF8FDF0797EFD7BF002" box="[609,696,579,602]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="428">UMMZ</collectionCode>
250051 (6),
<quantity id="4C8D9B79FFC4FFF8FCD8797FFC2FF002" box="[841,1004,578,603]" metricMagnitude="-2" metricUnit="m" metricValue="4.5649999999999995" metricValueMax="6.51" metricValueMin="2.62" pageId="8" pageNumber="428" unit="mm" value="45.65" valueMax="65.1" valueMin="26.2">26.265.1 mm</quantity>
<collectionCode id="ED64AE59FFC4FFF8FC63797EFBD5F003" box="[1010,1046,579,603]" pageId="8" pageNumber="428">SL</collectionCode>
; data as for
<typeStatus id="54CE883EFFC4FFF8FB0C797FFAC0F003" box="[1181,1283,578,603]" pageId="8" pageNumber="428" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF0A6614FFC4FFF8FF067C8EFBFFF591" box="[151,1084,1971,1993]" httpUri="https://zenodo.org/record/230836/files/figure.png" pageId="8" pageNumber="428" targetBox="[151,1436,655,1949]" targetPageId="8">
<paragraph id="8BCA369CFFC4FFF8FF067C8EFBFFF591" blockId="8.[151,1084,1971,1993]" box="[151,1084,1971,1993]" pageId="8" pageNumber="428">
<emphasis id="B901EA8EFFC4FFF8FF067C8EFED1F590" bold="true" box="[151,274,1971,1993]" pageId="8" pageNumber="428">FIGURE 6.</emphasis>
<taxonomicName id="4C754D1FFFC4FFF8FE887C89FDC1F591" box="[281,514,1972,1993]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="428" phylum="Chordata" rank="species" species="peregrinator">
<emphasis id="B901EA8EFFC4FFF8FE887C89FDC1F591" box="[281,514,1972,1993]" italics="true" pageId="8" pageNumber="428">Exostoma peregrinator</emphasis>
</taxonomicName>
, NIFI 3831, holotype, 70.2 mm SL; Thailand: Ban Tun.
</paragraph>
</caption>
<subSubSection id="C36F6517FFC5FFF9FF567BAAFC09F345" pageId="9" pageNumber="429" type="diagnosis">
<paragraph id="8BCA369CFFC5FFF9FF567BAAFC09F345" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="429">
<emphasis id="B901EA8EFFC5FFF9FF567BAAFE82F2E8" bold="true" box="[199,321,151,176]" pageId="9" pageNumber="429">Diagnosis.</emphasis>
<taxonomicName id="4C754D1FFFC5FFF9FED87BA4FD8DF2F7" box="[329,590,152,176]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="429" phylum="Chordata" rank="species" species="peregrinator">
<emphasis id="B901EA8EFFC5FFF9FED87BA4FD8DF2F7" box="[329,590,152,176]" italics="true" pageId="9" pageNumber="429">Exostoma peregrinator</emphasis>
</taxonomicName>
is distinguished from congeners in having a combination of the following characters: eye diameter 9.312.8% HL; dorsal-to-adipose distance 8.213.1% SL; length of adipose-fin base 31.734.6% SL; adipose fin adnate to upper procurrent caudal-fin rays; body depth at anus 9.913.1% SL; caudal peduncle depth 6.28.5% SL; lunate caudal fin; 2224 preanal vertebrae.
</paragraph>
</subSubSection>
<subSubSection id="C36F6517FFC5FFFAFF567A1AFCDDF557" lastPageId="10" lastPageNumber="430" pageId="9" pageNumber="429" type="description">
<paragraph id="8BCA369CFFC5FFF9FF567A1AFD65F038" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="429">
<emphasis id="B901EA8EFFC5FFF9FF567A1AFE9AF318" bold="true" box="[199,345,295,320]" pageId="9" pageNumber="429">Description.</emphasis>
Morphometric data as in
<tableCitation id="C6F70327FFC5FFF9FDE67A1AFD0FF318" box="[631,716,295,320]" captionStart="TABLE 3" captionStartId="9.[151,239,656,679]" captionTargetBox="[159,1290,706,1916]" captionTargetPageId="9" captionText="TABLE 3. Morphometric data for Exostoma peregrinator (NIFI 3831, NIFI 3762, NIFI 3764, UMMZ 250051; n = 19)" httpUri="http://table.plazi.org/id/DF0A6614FFC5FFF9FF0679ADFABCF0FF" pageId="9" pageNumber="429" tableUuid="DF0A6614FFC5FFF9FF0679ADFABCF0FF">Table 3</tableCitation>
. Head and body broad, very strongly depressed, with paired fins greatly enlarged to form elongate ovoid adhesive disc with body. Rostral margin rounded in dorsal view, moderately curved in lateral view. Dorsal profile rising gently and evenly from anterior orbital margin to origin of dorsal fin, then sloping gently ventrally to end of caudal peduncle. Ventral profile flat to anal-fin base, then sloping gently dorsally to end of caudal peduncle. Anus and urogenital openings located at vertical through posterior margin of adpressed pelvic fin. Mouth ventrally directed with broad, thin, papillate lips. Most of premaxillary tooth patches exposed when mouth is closed. Lower lip with prominent labial fold bearing entire posterior margin, notched at insertions of inner mandibular barbels. Postlabial groove on lower jaw present and uninterrupted. Dorsal surface of head covered with conical tubercles.
</paragraph>
<caption id="DF0A6614FFC5FFF9FF0679ADFABCF0FF" ID-Table-UUID="DF0A6614FFC5FFF9FF0679ADFABCF0FF" box="[151,1407,656,679]" httpUri="http://table.plazi.org/id/DF0A6614FFC5FFF9FF0679ADFABCF0FF" pageId="9" pageNumber="429" targetBox="[159,1290,706,1916]" targetIsTable="true" targetPageId="9">
<paragraph id="8BCA369CFFC5FFF9FF0679ADFABCF0FF" blockId="9.[151,1407,656,679]" box="[151,1407,656,679]" pageId="9" pageNumber="429">
<emphasis id="B901EA8EFFC5FFF9FF0679ADFECAF0FF" bold="true" box="[151,265,656,679]" pageId="9" pageNumber="429">TABLE 3.</emphasis>
Morphometric data for
<taxonomicName id="4C754D1FFFC5FFF9FD9B79ADFCC2F0FF" box="[522,769,656,679]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="429" phylum="Chordata" rank="species" species="peregrinator">
<emphasis id="B901EA8EFFC5FFF9FD9B79ADFCC2F0FF" box="[522,769,656,679]" italics="true" pageId="9" pageNumber="429">Exostoma peregrinator</emphasis>
</taxonomicName>
(NIFI 3831, NIFI 3762, NIFI 3764, UMMZ 250051; n=19)
</paragraph>
</caption>
<paragraph id="8BCA369CFFC5FFF9FF0E7837FAC9F524" pageId="9" pageNumber="429">
<table id="F975C43CFFC5000FFF0E79FFFAC9F524" box="[159,1290,706,1916]" gridcols="4" gridrows="10" pageId="9" pageNumber="429">
<tr id="354534DEFFC5000FFF0E79FFFAC9F178" box="[159,1290,706,800]" gridrow="0" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E79FFFE1EF178" box="[159,477,706,800]" gridcol="0" gridrow="0" pageId="9" pageNumber="429">Standard length (mm)</th>
<th id="76945DA2FFC5000FFD0C79FFFCC5F178" box="[669,774,706,800]" gridcol="1" gridrow="0" pageId="9" pageNumber="429">Holotype NIFI 3831 70.2</th>
<th id="76945DA2FFC5000FFC3179FFFBC5F178" box="[928,1030,706,800]" gridcol="2" gridrow="0" pageId="9" pageNumber="429">Range 23.370.2</th>
<th id="76945DA2FFC5000FFB3279FFFAC9F178" box="[1187,1290,706,800]" gridcol="3" gridrow="0" pageId="9" pageNumber="429">Mean±SD</th>
</tr>
<tr id="354534DEFFC5000FFF0E7809FAC9F1C4" box="[159,1290,820,924]" gridrow="1" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7809FE1EF1C4" box="[159,477,820,924]" gridcol="0" gridrow="1" pageId="9" pageNumber="429">%SL Predorsal length Preanal length</th>
<td id="76945DA2FFC5000FFD0C7809FCC5F1C4" box="[669,774,820,924]" gridcol="1" gridrow="1" pageId="9" pageNumber="429">39.5 71.4</td>
<td id="76945DA2FFC5000FFC317809FBC5F1C4" box="[928,1030,820,924]" gridcol="2" gridrow="1" pageId="9" pageNumber="429">38.943.4 68.674.2</td>
<td id="76945DA2FFC5000FFB327809FAC9F1C4" box="[1187,1290,820,924]" gridcol="3" gridrow="1" pageId="9" pageNumber="429">40.5±1.47 71.8±1.90</td>
</tr>
<tr id="354534DEFFC5000FFF0E788DFAC9F640" box="[159,1290,944,1048]" gridrow="2" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E788DFE1EF640" box="[159,477,944,1048]" gridcol="0" gridrow="2" pageId="9" pageNumber="429">Prepelvic length Prepectoral length Length of dorsal-fin base</th>
<td id="76945DA2FFC5000FFD0C788DFCC5F640" box="[669,774,944,1048]" gridcol="1" gridrow="2" pageId="9" pageNumber="429">49.7 15.2 12.5</td>
<td id="76945DA2FFC5000FFC31788DFBC5F640" box="[928,1030,944,1048]" gridcol="2" gridrow="2" pageId="9" pageNumber="429">45.150.9 14.317.5 12.313.8</td>
<td id="76945DA2FFC5000FFB32788DFAC9F640" box="[1187,1290,944,1048]" gridcol="3" gridrow="2" pageId="9" pageNumber="429">47.8±2.05 15.9±1.22 12.9±0.51</td>
</tr>
<tr id="354534DEFFC5000FFF0E7F11FAC9F6CC" box="[159,1290,1068,1172]" gridrow="3" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7F11FE1EF6CC" box="[159,477,1068,1172]" gridcol="0" gridrow="3" pageId="9" pageNumber="429">Length of anal-fin base Pelvic-fin length Pectoral-fin length</th>
<td id="76945DA2FFC5000FFD0C7F11FCC5F6CC" box="[669,774,1068,1172]" gridcol="1" gridrow="3" pageId="9" pageNumber="429">7.8 19.4 22.1</td>
<td id="76945DA2FFC5000FFC317F11FBC5F6CC" box="[928,1030,1068,1172]" gridcol="2" gridrow="3" pageId="9" pageNumber="429">6.98.3 16.520.7 19.923.8</td>
<td id="76945DA2FFC5000FFB327F11FAC9F6CC" box="[1187,1290,1068,1172]" gridcol="3" gridrow="3" pageId="9" pageNumber="429">7.6±0.49 18.2±1.35 22.2±1.18</td>
</tr>
<tr id="354534DEFFC5000FFF0E7F95FAC9F748" box="[159,1290,1192,1296]" gridrow="4" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7F95FE1EF748" box="[159,477,1192,1296]" gridcol="0" gridrow="4" pageId="9" pageNumber="429">Caudal-fin length Length of adipose-fin base Dorsal to adipose distance</th>
<td id="76945DA2FFC5000FFD0C7F95FCC5F748" box="[669,774,1192,1296]" gridcol="1" gridrow="4" pageId="9" pageNumber="429">26.4 31.8 11.3</td>
<td id="76945DA2FFC5000FFC317F95FBC5F748" box="[928,1030,1192,1296]" gridcol="2" gridrow="4" pageId="9" pageNumber="429">19.826.4 31.734.6 8.213.1</td>
<td id="76945DA2FFC5000FFB327F95FAC9F748" box="[1187,1290,1192,1296]" gridcol="3" gridrow="4" pageId="9" pageNumber="429">24.8±2.11 32.7±1.02 11.4±1.46</td>
</tr>
<tr id="354534DEFFC5000FFF0E7E19FAC9F7D4" box="[159,1290,1316,1420]" gridrow="5" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7E19FE1EF7D4" box="[159,477,1316,1420]" gridcol="0" gridrow="5" pageId="9" pageNumber="429">Length of caudal peduncle Depth of caudal peduncle Body depth at anus</th>
<td id="76945DA2FFC5000FFD0C7E19FCC5F7D4" box="[669,774,1316,1420]" gridcol="1" gridrow="5" pageId="9" pageNumber="429">17.9 8.0 12.3</td>
<td id="76945DA2FFC5000FFC317E19FBC5F7D4" box="[928,1030,1316,1420]" gridcol="2" gridrow="5" pageId="9" pageNumber="429">17.720.9 6.28.5 9.913.1</td>
<td id="76945DA2FFC5000FFB327E19FAC9F7D4" box="[1187,1290,1316,1420]" gridcol="3" gridrow="5" pageId="9" pageNumber="429">18.8±1.08 7.9±0.71 11.8±1.06</td>
</tr>
<tr id="354534DEFFC5000FFF0E7E9DFAC9F450" box="[159,1290,1440,1544]" gridrow="6" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7E9DFE1EF450" box="[159,477,1440,1544]" gridcol="0" gridrow="6" pageId="9" pageNumber="429">Pectoralpelvic distance Head length Head width</th>
<td id="76945DA2FFC5000FFD0C7E9DFCC5F450" box="[669,774,1440,1544]" gridcol="1" gridrow="6" pageId="9" pageNumber="429">34.5 21.5 18.7</td>
<td id="76945DA2FFC5000FFC317E9DFBC5F450" box="[928,1030,1440,1544]" gridcol="2" gridrow="6" pageId="9" pageNumber="429">29.935.3 21.523.9 18.020.0</td>
<td id="76945DA2FFC5000FFB327E9DFAC9F450" box="[1187,1290,1440,1544]" gridcol="3" gridrow="6" pageId="9" pageNumber="429">33.2±1.94 22.5±0.86 19.1±0.68</td>
</tr>
<tr id="354534DEFFC5000FFF0E7D21FAC9F4DC" box="[159,1290,1564,1668]" gridrow="7" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7D21FE1EF4DC" box="[159,477,1564,1668]" gridcol="0" gridrow="7" pageId="9" pageNumber="429">Head depth %HL Snout length</th>
<td id="76945DA2FFC5000FFD0C7D21FCC5F4DC" box="[669,774,1564,1668]" gridcol="1" gridrow="7" pageId="9" pageNumber="429">9.5 58.9</td>
<td id="76945DA2FFC5000FFC317D21FBC5F4DC" box="[928,1030,1564,1668]" gridcol="2" gridrow="7" pageId="9" pageNumber="429">9.110.9 56.161.2</td>
<td id="76945DA2FFC5000FFB327D21FAC9F4DC" box="[1187,1290,1564,1668]" gridcol="3" gridrow="7" pageId="9" pageNumber="429">10.3±0.68 58.8±1.85</td>
</tr>
<tr id="354534DEFFC5000FFF0E7DA5FAC9F558" box="[159,1290,1688,1792]" gridrow="8" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7DA5FE1EF558" box="[159,477,1688,1792]" gridcol="0" gridrow="8" pageId="9" pageNumber="429">Interorbital distance Eye diameter Nasal barbel length</th>
<td id="76945DA2FFC5000FFD0C7DA5FCC5F558" box="[669,774,1688,1792]" gridcol="1" gridrow="8" pageId="9" pageNumber="429">33.1 12.6 39.7</td>
<td id="76945DA2FFC5000FFC317DA5FBC5F558" box="[928,1030,1688,1792]" gridcol="2" gridrow="8" pageId="9" pageNumber="429">22.233.1 9.312.8 30.239.7</td>
<td id="76945DA2FFC5000FFB327DA5FAC9F558" box="[1187,1290,1688,1792]" gridcol="3" gridrow="8" pageId="9" pageNumber="429">29.4±3.38 11.5±1.08 34.9±3.02</td>
</tr>
<tr id="354534DEFFC5000FFF0E7C29FAC9F524" box="[159,1290,1812,1916]" gridrow="9" pageId="9" pageNumber="429">
<th id="76945DA2FFC5000FFF0E7C29FE1EF524" box="[159,477,1812,1916]" gridcol="0" gridrow="9" pageId="9" pageNumber="429">Maxillary barbel length Inner mandibular barbel length Outer mandibular barbel length</th>
<td id="76945DA2FFC5000FFD0C7C29FCC5F524" box="[669,774,1812,1916]" gridcol="1" gridrow="9" pageId="9" pageNumber="429">94.0 10.6 25.8</td>
<td id="76945DA2FFC5000FFC317C29FBC5F524" box="[928,1030,1812,1916]" gridcol="2" gridrow="9" pageId="9" pageNumber="429">69.194.0 7.712.6 21.429.3</td>
<td id="76945DA2FFC5000FFB327C29FAC9F524" box="[1187,1290,1812,1916]" gridcol="3" gridrow="9" pageId="9" pageNumber="429">82.9±8.23 9.8±1.86 25.0±2.86</td>
</tr>
</table>
</paragraph>
<caption id="DF0A6614FFC6FFFAFF067996FEB3F0B8" httpUri="https://zenodo.org/record/230837/files/figure.png" pageId="10" pageNumber="430" targetBox="[185,1396,221,590]" targetPageId="10">
<paragraph id="8BCA369CFFC6FFFAFF067996FEB3F0B8" blockId="10.[151,1436,683,736]" pageId="10" pageNumber="430">
<emphasis id="B901EA8EFFC6FFFAFF067996FED5F098" bold="true" box="[151,278,683,705]" pageId="10" pageNumber="430">FIGURE 7.</emphasis>
<taxonomicName id="4C754D1FFFC6FFFAFEB17991FDCEF099" box="[288,525,684,705]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="430" phylum="Chordata" rank="species" species="peregrinator">
<emphasis id="B901EA8EFFC6FFFAFEB17991FDCEF099" box="[288,525,684,705]" italics="true" pageId="10" pageNumber="430">Exostoma peregrinator</emphasis>
</taxonomicName>
, NIFI 3831, holotype, 70.2 mm SL; photograph taken immediately after preservation, showing color in life.
</paragraph>
</caption>
<paragraph id="8BCA369CFFC6FFFAFF567835FA8DF11C" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">Jaw teeth distally flattened, oar-shaped, in two rounded triangular patches narrowly separated at midline on upper jaw. Teeth on lower jaw situated in two well-separated, roughly triangular patches. Palate edentulous.</paragraph>
<paragraph id="8BCA369CFFC6FFFAFF56786DFB8FF1D4" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">Eyes small, dorsolaterally situated and subcutaneous. Gill openings narrow, extending from base of first pectoral-fin ray to level immediately dorsal to base of posteriormost pectoral-fin ray.</paragraph>
<paragraph id="8BCA369CFFC6FFFAFF5678AAFB81F618" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">Barbels in four pairs. Nasal barbel moderately long, reaching to middle of orbit. Maxillary barbel flattened, with flap of skin fringing posterior margin; its tip pointed; ventral surface with numerous striae; extending beyond base of pectoral fin. Inner mandibular barbel slightly flattened, very short; originating from notch on posterior margin of lower lip. Outer mandibular barbel situated lateral to inner mandibular barbel; slightly flattened, reaching to two-thirds of distance between its base and base of first pectoral-fin ray.</paragraph>
<paragraph id="8BCA369CFFC6FFFAFF567F76FB6FF764" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">Dorsal fin without spine, with i,6 (19) rays. Adipose fin with long base, posterior end separate from upper procurrent caudal-fin rays. Anal fin with ii,4,i (14) or ii,5*(5) rays. Caudal fin lunate, with i,6,7,i* (1) or i,7,7,i (18) rays; upper and lower first principal rays of approximately same length. Pelvic fin greatly enlarged, with convex distal margin and i,5 (19) rays; first ray greatly flattened, with numerous striae on ventral surface. Pectoral fin greatly enlarged, without spine and with i,8,i (2), i,9 (3), i,9,i (7) or i,l0* (7) rays; first ray greatly flattened, with numerous striae on ventral surface. Dorsal surface of pectoral-fin rays covered with sparse tubercles. Vertebrae 22+14=36 (2), 23+13=36 (2), 22+15=37* (4), 23+14=37 (4), 23+15=38 (1) or 24+14=38 (4).</paragraph>
<paragraph id="8BCA369CFFC6FFFAFF567E7AFB14F7A8" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">
<emphasis id="B901EA8EFFC6FFFAFF567E7AFE8CF738" bold="true" box="[199,335,1351,1376]" pageId="10" pageNumber="430">Coloration.</emphasis>
In 70% ethanol: brown on dorsal and lateral surfaces of the head and body, dark yellow on ventral region. Dorsal and caudal fins brown; dorsal surfaces of pectoral and pelvic fins brown, with light yellow ventral surfaces. Anal fin dark yellow. Dorsal surface of barbels brown, ventral surface dark yellow. Dorsal half of adipose fin hyaline; ventral half brown. Caudal fin with brown fin rays and hyaline interradial membranes; dark elongated elliptical spot on first to third upper and lower principal caudal rays about middle third of fin ray.
</paragraph>
<paragraph id="8BCA369CFFC6FFFAFF567EC1FCDDF557" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">
In life (
<figureCitation id="134E2A19FFC6FFFAFE8A7EC6FEA1F44C" box="[283,354,1531,1556]" captionStart="FIGURE 7" captionStartId="10.[151,250,683,705]" captionTargetBox="[185,1396,221,590]" captionTargetId="figure@10.[150,1436,171,661]" captionTargetPageId="10" captionText="FIGURE 7. Exostoma peregrinator, NIFI 3831, holotype, 70.2 mm SL; photograph taken immediately after preservation, showing color in life." httpUri="https://zenodo.org/record/230837/files/figure.png" pageId="10" pageNumber="430">Fig. 7</figureCitation>
): Olive brown on dorsal and lateral surfaces of head and body, fading to dark yellow on ventral half of flanks and to tan on ventral surfaces. Numerous pale spots densely distributed over dorsal surfaces of head and predorsal region. Dorsolateral surface of body with two indistinct tan-colored blotches: one below adipose-fin origin and second below middle of adipose-fin base. Base of adipose fin with melanophores concentrated in a dark brown, longitudinal band. Dorsal, pectoral and pelvic fins with dusky fin rays and hyaline fin membranes. Base of caudal fin with gently crescentic dark-brown bar; middle third of caudal fin with irregular, dark brown w-shaped band; whitish spots on dorsal- and ventralmost tips of caudal fin. Dorsal surface of nasal and maxillary barbels olive brown, ventral surfaces tan. Mandibular barbels tan.
</paragraph>
</subSubSection>
<subSubSection id="C36F6517FFC6FFFAFF567C26FD21F5F8" pageId="10" pageNumber="430" type="distribution">
<paragraph id="8BCA369CFFC6FFFAFF567C26FD21F5F8" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">
<emphasis id="B901EA8EFFC6FFFAFF567C26FEA3F56C" bold="true" box="[199,352,1819,1844]" pageId="10" pageNumber="430">Distribution.</emphasis>
This species is only known from the
<typeStatus id="54CE883EFFC6FFFAFC967C21FCF4F56C" box="[775,823,1820,1844]" pageId="10" pageNumber="430">type</typeStatus>
locality in the upper reaches of the Ping River (one of two main tributaries of the Chao Phraya River) drainage in northwestern
<collectingCountry id="F362760CFFC6FFFAFC707C02FB86F500" box="[993,1093,1855,1880]" name="Thailand" pageId="10" pageNumber="430">Thailand</collectingCountry>
(
<figureCitation id="134E2A19FFC6FFFAFBC77C02FB58F50F" box="[1110,1179,1855,1880]" captionStart="FIGURE 3" captionStartId="4.[151,250,1591,1613]" captionTargetBox="[151,1436,207,1569]" captionTargetId="figure@4.[151,1436,195,1569]" captionTargetPageId="4" captionText="FIGURE 3. Map showing collecting localities of Exostoma species in Thailand examined in this study. The type localities of E. effrenum and E. peregrinator are shown much further apart than they really are in order to easily distinguish between the two symbols." httpUri="https://zenodo.org/record/230833/files/figure.png" pageId="10" pageNumber="430">Fig. 3</figureCitation>
). This is also the
<typeStatus id="54CE883EFFC6FFFAFAFD7C7DFA5FF500" box="[1388,1436,1856,1880]" pageId="10" pageNumber="430">type</typeStatus>
locality of
<taxonomicName id="4C754D1FFFC6FFFAFE807C5EFE28F523" box="[273,491,1891,1915]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="430" phylum="Chordata" rank="species" species="sudarai">
<emphasis id="B901EA8EFFC6FFFAFE807C5EFE28F523" box="[273,491,1891,1915]" italics="true" pageId="10" pageNumber="430">Oreoglanis sudarai</emphasis>
</taxonomicName>
, where the coordinates were erroneously given as
<geoCoordinate id="EE41505BFFC6FFFAFBB57C5EFB43F524" box="[1060,1152,1891,1916]" direction="north" orientation="latitude" pageId="10" pageNumber="430" precision="925" value="18.583334">18°35'N</geoCoordinate>
<geoCoordinate id="EE41505BFFC6FFFAFB197C59FB22F524" box="[1160,1249,1892,1916]" direction="east" orientation="longitude" pageId="10" pageNumber="430" precision="925" value="98.166664">98°10'E</geoCoordinate>
(Vidthayanon et el., 2009), which lies in the Salween River drainage.
</paragraph>
</subSubSection>
<subSubSection id="C36F6517FFC6FFFAFF567C96FA9AF5B0" pageId="10" pageNumber="430" type="etymology">
<paragraph id="8BCA369CFFC6FFFAFF567C96FA9AF5B0" blockId="10.[151,1437,775,2024]" pageId="10" pageNumber="430">
<emphasis id="B901EA8EFFC6FFFAFF567C96FE8DF59C" bold="true" box="[199,334,1963,1988]" pageId="10" pageNumber="430">Etymology.</emphasis>
The specific epithet is the Latin noun meaning “one who travels about”, in reference to the distribution of this species, which represents the first record of the genus east of the Salween River drainage.
</paragraph>
</subSubSection>
</treatment>
</document>