144 lines
17 KiB
XML
144 lines
17 KiB
XML
<document id="EAE9F9AFAF3E08160DC193DD1B5C6C53" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE8FFEA6194FB5B073CF51E" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus compactus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="21" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="72C8884BABD32C0388677C598DD6DEF1">
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<mods:title id="08C31CDE3B7DCFEE869E964A9517EADB">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C58F99D7F854EB88C6392273F8197567" type="personal">
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<mods:role id="7ABD5EA4A1D07E11FEBBF842F1807B8E">
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<mods:roleTerm id="DC5B9190004557E4DC8489F936582F11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CA5A11F1BBFD3AE1D4CF0748C0D235E9">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="2BBFA7F03FEC9C27A3E0AB1A46BBF8C8">text</mods:typeOfResource>
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<mods:title id="43B6A2C0A502C8B7D7F6184470CD5ABD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="330B9A8C2037F4F14EE0441685E88134">
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<mods:date id="2DC96F0BC76DD1A4D2551DE064BA3410">2021</mods:date>
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<mods:number id="7A2DE82718FDFACD40F2E4EF7EF979B8">2021-06-28</mods:number>
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<mods:detail id="5CB7ADCAC87FF7D130ACB7D5F691A417" type="volume">
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<mods:number id="CD89ECF1158557274BE8F610B2FE3F40">4994</mods:number>
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</mods:detail>
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<mods:detail id="CB09052DCE84F6775E2340352518BAC0" type="issue">
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<mods:number id="1E34F00B2F33B14318D77AB57785AD9D">1</mods:number>
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<mods:start id="41A60BF98807CEA29598FECFACCC20F7">1</mods:start>
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<mods:end id="110AB1413749762840745679B969B6ED">93</mods:end>
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<mods:classification id="08AB91DE3B1F684A88D7335FBBD88BFB">journal article</mods:classification>
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<mods:identifier id="20F6A60B825A5291D6F9AE7333E1AB99" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="FF3F03DCD2D7B19F4DADB2E35BD589A8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7236474E55F2501C7934D04B3F489C1E" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="2526AC1EC8C3867DDF2908B5AA88C6A7" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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</mods:mods>
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<treatment id="03A187DEFFE8FFEA6194FB5B073CF51E" ID-DOI="http://doi.org/10.5281/zenodo.5043948" ID-Zenodo-Dep="5043948" LSID="urn:lsid:plazi:treatment:03A187DEFFE8FFEA6194FB5B073CF51E" httpUri="http://treatment.plazi.org/id/03A187DEFFE8FFEA6194FB5B073CF51E" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C3126543FFE8FFEB6194FB5B061CF3C2" box="[151,579,1195,1222]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BB736C8FFE8FFEB6194FB5B061CF3C2" blockId="20.[151,579,1195,1258]" box="[151,579,1195,1222]" pageId="20" pageNumber="21">
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<heading id="D0FF81A4FFE8FFEB6194FB5B061CF3C2" bold="true" box="[151,579,1195,1222]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
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<emphasis id="B97CEADAFFE8FFEB6194FB5B061CF3C2" bold="true" box="[151,579,1195,1222]" pageId="20" pageNumber="21">
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<taxonomicName id="4C084D4BFFE8FFEB6194FB5B05FAF3C2" ID-CoL="8Y4PP" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,421,1195,1222]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="compactus" status="sp. nov.">
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<emphasis id="B97CEADAFFE8FFEB6194FB5B05FAF3C2" bold="true" box="[151,421,1195,1222]" italics="true" pageId="20" pageNumber="21">Hydrochus compactus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE8FFEB60B7FB5B061CF3C2" box="[436,579,1195,1221]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEB6194FB3F0568F3ED" box="[151,311,1231,1258]" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BB736C8FFE8FFEB6194FB3F0568F3ED" blockId="20.[151,579,1195,1258]" box="[151,311,1231,1258]" pageId="20" pageNumber="21">
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<figureCitation id="13332A4DFFE8FFEB6194FB3F04BBF3EE" box="[151,228,1231,1257]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="20" pageNumber="21">Figs. 3</figureCitation>
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,
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<figureCitation id="13332A4DFFE8FFEB61F2FB3F0550F3EE" box="[241,271,1231,1257]" captionStart="FIGURES 34–35" captionStartId="69.[151,264,1944,1968]" captionTargetBox="[171,1426,203,1871]" captionTargetId="figure-15@69.[151,1435,181,1920]" captionTargetPageId="69" captionText="FIGURES 34–35. Habitus and male genitalia of holotypes. 34. Hydrochus compactus n. sp. 35. Hydrochus zicsii Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043318" httpUri="https://zenodo.org/record/5043318/files/figure.png" pageId="20" pageNumber="21">34</figureCitation>
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,
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<figureCitation id="13332A4DFFE8FFEB6018FB3F0568F3ED" box="[283,311,1231,1258]" captionStart="FIGURES 57–58" captionStartId="81.[151,264,1928,1952]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-15@81.[151,1435,181,1905]" captionTargetPageId="81" captionText="FIGURES 57–58. Geographical distributions. 57. Endemics and near endemics. 58. H. bituberculatus." figureDoi="http://doi.org/10.5281/zenodo.5043356" httpUri="https://zenodo.org/record/5043356/files/figure.png" pageId="20" pageNumber="21">57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEB6194FAE706B4F252" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BB736C8FFE8FFEB6194FAE706B4F252" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">
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<emphasis id="B97CEADAFFE8FFEB6194FAE70517F236" bold="true" box="[151,328,1303,1329]" pageId="20" pageNumber="21">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE8FFEB6053FAE706B4F252" collectingDate="1986-10-11" collectionCode="NMNH" collectorName="P. Spangler & R. Crombie" country="Brazil" latitude="-3.65" location="Para" longLatPrecision="1307" longitude="-52.366665" municipality="Rio Xingu Camp" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97CEADAFFE8FFEB6053FAE70646F236" bold="true" box="[336,537,1303,1329]" pageId="20" pageNumber="21">
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<typeStatus id="54B3886AFFE8FFEB6053FAE705E2F236" box="[336,445,1303,1329]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFE8FFEB6329FAE706C9F236" box="[554,662,1303,1329]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
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:
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<location id="8ED76013FFE8FFEB63A2FAE80689F235" LSID="urn:lsid:plazi:treatment:03A187DEFFE8FFEA6194FB5B073CF51E:8ED76013FFE8FFEB63A2FAE80689F235" box="[673,726,1304,1330]" country="Brazil" latitude="-3.65" longLatPrecision="1307" longitude="-52.366665" municipality="Rio Xingu Camp" name="Para" pageId="20" pageNumber="21">Para</location>
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;
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<collectingMunicipality id="6BD3ACB2FFE8FFEB63E1FAE707FAF236" box="[738,933,1303,1329]" pageId="20" pageNumber="21">Rio Xingu Camp</collectingMunicipality>
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, (
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<geoCoordinate id="EE3C500FFFE8FFEB62B9FAE70056F236" box="[954,1033,1303,1329]" degrees="3" direction="south" minutes="39" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-3.65">3°39’S</geoCoordinate>
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,
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<geoCoordinate id="EE3C500FFFE8FFEB6515FAE70023F235" box="[1046,1148,1303,1330]" degrees="52" direction="west" minutes="22" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-52.366665">52°22’W</geoCoordinate>
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), Altamira (ca.
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<quantity id="4CF09B2DFFE8FFEB642DFAE70131F236" box="[1326,1390,1303,1329]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="km" value="60.0">60km</quantity>
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S.),,
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<date id="FFB61008FFE8FFEB61A6FACB0566F252" box="[165,313,1339,1365]" pageId="20" pageNumber="21" value="1986-10-11">
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<collectingDate id="EFF2E9E0FFE8FFEB61A6FACB0566F252" box="[165,313,1339,1365]" pageId="20" pageNumber="21" value="1986-10-11">11 Oct. 1986</collectingDate>
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</date>
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,
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<collectorName id="26FD531EFFE8FFEB6047FACC059DF252" box="[324,450,1339,1366]" pageId="20" pageNumber="21">P. Spangler</collectorName>
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&
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<collectorName id="26FD531EFFE8FFEB60E6FACC0634F252" box="[485,619,1339,1366]" pageId="20" pageNumber="21">R. Crombie</collectorName>
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” (
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<collectionCode id="ED19AE0DFFE8FFEB6380FACB0680F252" box="[643,735,1339,1365]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEB61C4FAAF0593F2E2" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BB736C8FFE8FFEB61C4FAAF0593F2E2" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">
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<emphasis id="B97CEADAFFE8FFEB61C4FAAF058CF27E" bold="true" box="[199,467,1375,1401]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of small size (ca.
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<quantity id="4CF09B2DFFE8FFEB642BFAAF01CFF27D" box="[1320,1424,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="21" unit="mm" value="2.1">2.10 mm</quantity>
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), the compact body form, the black dorsum, the lack of a callus on the 5
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<superScript id="7C7D9B80FFE8FFEB629EFA7307F5F296" attach="left" box="[925,938,1411,1425]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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elytral interstria, and the male genitalia (
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<figureCitation id="13332A4DFFE8FFEB646CFA7304E9F2C6" captionStart="FIGURES 34–35" captionStartId="69.[151,264,1944,1968]" captionTargetBox="[171,1426,203,1871]" captionTargetId="figure-15@69.[151,1435,181,1920]" captionTargetPageId="69" captionText="FIGURES 34–35. Habitus and male genitalia of holotypes. 34. Hydrochus compactus n. sp. 35. Hydrochus zicsii Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043318" httpUri="https://zenodo.org/record/5043318/files/figure.png" pageId="20" pageNumber="21">Fig. 34</figureCitation>
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; described below). Differentiation from other similarly small sized species with similarly shaped pronotum will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEA61C4FA1F0728F6F2" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BB736C8FFE8FFEB61C4FA1F055DF1BA" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">
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<emphasis id="B97CEADAFFE8FFEB61C4FA1F050EF10E" bold="true" box="[199,337,1519,1545]" pageId="20" pageNumber="21">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFE8FFEB60A7FA1F0659F10E" box="[420,518,1519,1545]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.10/0.82; head width 0.56; pronotum l/w 0.53/0.56; PA 0.54; PB 0.43; elytra 1.31/0.82. Habitus as illustrated (
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<figureCitation id="13332A4DFFE8FFEB6549F9E300FFF12A" box="[1098,1184,1555,1581]" captionStart="FIGURES 34–35" captionStartId="69.[151,264,1944,1968]" captionTargetBox="[171,1426,203,1871]" captionTargetId="figure-15@69.[151,1435,181,1920]" captionTargetPageId="69" captionText="FIGURES 34–35. Habitus and male genitalia of holotypes. 34. Hydrochus compactus n. sp. 35. Hydrochus zicsii Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043318" httpUri="https://zenodo.org/record/5043318/files/figure.png" pageId="20" pageNumber="21">Fig. 34</figureCitation>
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). Body size small (ca.
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<quantity id="4CF09B2DFFE8FFEB6194F9C7055FF155" box="[151,256,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="21" unit="mm" value="2.1">2.10 mm</quantity>
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). Dorsum dark brown to black, with very weak iridescence, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 0.5x strial puncture diameter. Interstria 5
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<superScript id="7C7D9B80FFE8FFEB60EFF98F05A6F18A" attach="left" box="[492,505,1663,1677]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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raised, but lacking callus. Interstria 9
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<superScript id="7C7D9B80FFE8FFEB62A7F98F07EEF18A" attach="left" box="[932,945,1663,1677]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFE8FFEB6493F98F01C2F18A" attach="none" box="[1424,1437,1663,1677]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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interstria.
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</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4F93805BAF02E" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">Pronotum slightly wider than long (ratio ca. 32/28), widest at anterior margin, markedly narrowed at base, sides sinuate, with microdenticulations; anterior margin bisinuate; depressions shallow; punctation dense, coarse, denser in depressions than on reliefs.</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4F8C304B2F091" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">Elytra with spaces between strial punctures ca. 0.5x their diameter, or less; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without angulation; submarginal row of punctures in common groove.</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4F86F048DF0D9" blockId="20.[151,1437,1303,2014]" pageId="20" pageNumber="21">Ventral characters: Mentum and submentum densely, moderately coarsely punctate; submentum without foveae.</paragraph>
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<paragraph id="8BB736C8FFE9FFEA61C4FF6707D5F68E" blockId="21.[151,1437,151,537]" pageId="21" pageNumber="22">
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Male genitalia (
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<figureCitation id="13332A4DFFE9FFEA6076FF670599F7B6" box="[373,454,151,177]" captionStart="FIGURES 34–35" captionStartId="69.[151,264,1944,1968]" captionTargetBox="[171,1426,203,1871]" captionTargetId="figure-15@69.[151,1435,181,1920]" captionTargetPageId="69" captionText="FIGURES 34–35. Habitus and male genitalia of holotypes. 34. Hydrochus compactus n. sp. 35. Hydrochus zicsii Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043318" httpUri="https://zenodo.org/record/5043318/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
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) general characters: genitalia very narrow, elongate, length ca. 6.6x greatest width, basal piece shorter than parameres (ratio ca. 13/20); paramere tips extending well beyond aedeagus tip, medial margins beyond aedeagus tip diverging slightly one from the other, plsa absent, lateral margins of tips slightly emarginate; in lateral view parameres very narrow, slightly sinuate, sharply pointed apically; aedeagus narrow, sides arcuate, agw at ca. distal 2/3, adbl ca. same width; basal piece in dorsal/ventral views with slightly arcuate lateral margins, in lateral view arcuate, dorsal surface more arcuate than ventral, maximum width in dorsal/ventral views slightly greater than lateral width, orifice asymmetrical, margin sclerotized.
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</paragraph>
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<paragraph id="8BB736C8FFE9FFEA61C4FE630649F6D6" blockId="21.[151,1437,151,537]" pageId="21" pageNumber="22">Dorsal surface: adtl at ca. proximal 1/4; pdmm approximating alm, except underneath lateral margin of adbl; adbl large, width ca. equal to agw.</paragraph>
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<paragraph id="8BB736C8FFE9FFEA61C4FE2B0728F6F2" blockId="21.[151,1437,151,537]" box="[199,887,475,501]" pageId="21" pageNumber="22">Ventral surface: avtl at ca. distal 1/3; pvmm similar to pdmm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE9FFEA61C4FE0F073CF51E" box="[199,867,511,537]" pageId="21" pageNumber="22" type="etymology">
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<paragraph id="8BB736C8FFE9FFEA61C4FE0F073CF51E" blockId="21.[151,1437,151,537]" box="[199,867,511,537]" pageId="21" pageNumber="22">
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<emphasis id="B97CEADAFFE9FFEA61C4FE0F0517F51E" bold="true" box="[199,328,511,537]" pageId="21" pageNumber="22">Etymology</emphasis>
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. Named in reference to the compact body form.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |