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<document id="894310A848CD59898EFAC54095111C42" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE2FFE16194FF67002AF236" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus personatus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="31" updateTime="1698978858223" updateUser="plazi">
<mods:mods id="A8E7D88B7CC91FE879A532C5AA692D58" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6923913CED571568B1CF7F4039727229">
<mods:title id="BCFE292ED37B2CC35C70F0C00F2DDA8F">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="05B3EC8C36AD59EFC2FBA2D1E94F96BA">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="90BF5067101E97B2EBD25118101712C6">text</mods:typeOfResource>
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<mods:title id="469DDE9E3905A213F73CF0A7A69E4EB4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8EAC7F1BEE9373754A149400BD1F4CB4">
<mods:date id="6BD12EC51086DD3F8A0C8163C36C8AE5">2021</mods:date>
<mods:detail id="498D73F0D2636F490FFBD6D20F9D1C70" type="pubDate">
<mods:number id="B0AEE477752BEF99AA2E0310BF2B1940">2021-06-28</mods:number>
</mods:detail>
<mods:detail id="AF3C62F2D5AF8F25982889F08E8540BB" type="volume">
<mods:number id="97C2E4B0C35838197D4987A1C5662D6E">4994</mods:number>
</mods:detail>
<mods:detail id="B1C8B7861746597376BF182A9360A87F" type="issue">
<mods:number id="24855CAAE4DC7C10E46F8A696D61D841">1</mods:number>
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<mods:classification id="0EB966CA1F4438841D26BC4237FF9435">journal article</mods:classification>
<mods:identifier id="9DEF5D2B3CA55438B338F6D52041A229" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
<mods:identifier id="B509E71128C4478EC61B822FBE15CE01" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EEFAAE9A2EBD1EE6918BAC58275D0BC7" type="Zenodo-Dep">5043261</mods:identifier>
<mods:identifier id="312613F0CC59C6CC0DF3CA04011378A0" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
</mods:mods>
<treatment id="03A187DEFFE2FFE16194FF67002AF236" ID-DOI="http://doi.org/10.5281/zenodo.5043970" ID-Zenodo-Dep="5043970" LSID="urn:lsid:plazi:treatment:03A187DEFFE2FFE16194FF67002AF236" httpUri="http://treatment.plazi.org/id/03A187DEFFE2FFE16194FF67002AF236" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="C3126543FFE2FFE16194FF670618F7B6" box="[151,583,151,178]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BB736C8FFE2FFE16194FF670618F7B6" blockId="30.[151,583,151,214]" box="[151,583,151,178]" pageId="30" pageNumber="31">
<heading id="D0FF81A4FFE2FFE16194FF670618F7B6" bold="true" box="[151,583,151,178]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="B97CEADAFFE2FFE16194FF670618F7B6" bold="true" box="[151,583,151,178]" pageId="30" pageNumber="31">
<taxonomicName id="4C084D4BFFE2FFE16194FF6705F7F7B6" ID-CoL="8Y4R2" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,424,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="personatus" status="sp. nov.">
<emphasis id="B97CEADAFFE2FFE16194FF6705F7F7B6" bold="true" box="[151,424,151,178]" italics="true" pageId="30" pageNumber="31">Hydrochus personatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24F57A1FFE2FFE160B4FF670618F7B6" box="[439,583,151,177]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFE2FFE16194FF4B0568F7D1" box="[151,311,187,214]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BB736C8FFE2FFE16194FF4B0568F7D1" blockId="30.[151,583,151,214]" box="[151,311,187,214]" pageId="30" pageNumber="31">
<figureCitation id="13332A4DFFE2FFE16194FF4B04BAF7D2" box="[151,229,187,213]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="30" pageNumber="31">Figs. 2</figureCitation>
,
<figureCitation id="13332A4DFFE2FFE161F2FF4B0550F7D1" box="[241,271,187,214]" captionStart="FIGURES 14–15" captionStartId="59.[151,264,1944,1968]" captionTargetBox="[161,1428,217,1846]" captionTargetId="figure-15@59.[151,1435,181,1920]" captionTargetPageId="59" captionText="FIGURES 14–15. Habitus and male genitalia of holotypes. 14. Hydrochus reticulatus n. sp. 15. Hydrochus personatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043290" httpUri="https://zenodo.org/record/5043290/files/figure.png" pageId="30" pageNumber="31">15</figureCitation>
,
<figureCitation id="13332A4DFFE2FFE16018FF4B0568F7D1" box="[283,311,187,214]" captionStart="FIGURES 55–56" captionStartId="80.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1920]" captionTargetId="figure-28@80.[165,1422,181,1920]" captionTargetPageId="80" captionText="FIGURES 55–56. Geographical distributions. 55. Endemics and near endemics. 56. H. bicarinatus and H. merkli." figureDoi="http://doi.org/10.5281/zenodo.5043352" httpUri="https://zenodo.org/record/5043352/files/figure.png" pageId="30" pageNumber="31">55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFE2FFE16194FEF3077FF68D" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BB736C8FFE2FFE16194FEF3077FF68D" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">
<emphasis id="B97CEADAFFE2FFE16194FEF30513F61A" bold="true" box="[151,332,259,285]" pageId="30" pageNumber="31">Type Material.</emphasis>
<materialsCitation id="3B603C95FFE2FFE16054FEF304ACF645" collectingDate="1998-07-23" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" location="Dep. Cordillera" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Cordillera" typeStatus="holotype">
<emphasis id="B97CEADAFFE2FFE16054FEF3067BF61A" bold="true" box="[343,548,259,285]" pageId="30" pageNumber="31">
<typeStatus id="54B3886AFFE2FFE16054FEF3059BF61A" box="[343,452,259,285]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
“
<emphasis id="B97CEADAFFE2FFE16338FEF306F1F61A" bold="true" box="[571,686,259,285]" pageId="30" pageNumber="31">
<collectingCountry id="F31F7658FFE2FFE16338FEF306F1F61A" box="[571,686,259,285]" name="Paraguay" pageId="30" pageNumber="31">Paraguay</collectingCountry>
</emphasis>
:
<date id="FFB61008FFE2FFE163C2FEF3071CF61A" box="[705,835,259,285]" pageId="30" pageNumber="31" value="1998-07-23">
<collectingDate id="EFF2E9E0FFE2FFE163C2FEF3071CF61A" box="[705,835,259,285]" pageId="30" pageNumber="31" value="1998-07-23">23.vii.1998</collectingDate>
</date>
,
<collectingRegion id="49CCF82AFFE2FFE16250FEF40058F61A" box="[851,1031,259,286]" country="Paraguay" name="Cordillera" pageId="30" pageNumber="31">Dep. Cordillera</collectingRegion>
, San Bernadino, leg.
<collectorName id="26FD531EFFE2FFE16405FEF401CCF61A" box="[1286,1427,259,286]" pageId="30" pageNumber="31">U. Drechsel</collectorName>
” (
<collectionCode id="ED19AE0DFFE2FFE1619CFED704B7F646" box="[159,232,295,321]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="30" pageNumber="31">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B603C95FFE2FFE16000FED705BBF661" collectingDate="1959-12-27" collectingDateMax="1959-12-30" collectingDateMin="1959-12-27" collectionCode="NMNH" collectorName="Robert Cumming" country="Bolivia" location="Saavedra Exp. Sta." pageId="30" pageNumber="31" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<emphasis id="B97CEADAFFE2FFE16000FED70643F646" bold="true" box="[259,540,295,321]" pageId="30" pageNumber="31">
<typeStatus id="54B3886AFFE2FFE16000FED70524F646" box="[259,379,295,321]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
(15):
<collectingCountry id="F31F7658FFE2FFE160CBFED70643F646" box="[456,540,295,321]" name="Bolivia" pageId="30" pageNumber="31">Bolivia</collectingCountry>
</emphasis>
:
<collectingRegion id="49CCF82AFFE2FFE1632DFED706EFF646" box="[558,688,295,321]" country="Bolivia" name="Santa Cruz" pageId="30" pageNumber="31">Santa Cruz</collectingRegion>
,
<quantity id="4CF09B2DFFE2FFE163BCFED70754F646" box="[703,779,295,321]" metricMagnitude="4" metricUnit="m" metricValue="9.656064" pageId="30" pageNumber="31" unit="mi" value="60.0">60 mi.</quantity>
N.
<collectingRegion id="49CCF82AFFE2FFE16238FED707E3F646" box="[827,956,295,321]" country="Bolivia" name="Santa Cruz" pageId="30" pageNumber="31">Santa Cruz</collectingRegion>
,
<location id="8ED76013FFE2FFE162CFFED700F6F646" LSID="urn:lsid:plazi:treatment:03A187DEFFE2FFE16194FF67002AF236:8ED76013FFE2FFE162CFFED700F6F646" box="[972,1193,295,322]" country="Bolivia" name="Saavedra Exp. Sta." pageId="30" pageNumber="31" stateProvince="Santa Cruz">Saavedra Exp. Sta.</location>
,
<date id="FFB61008FFE2FFE165BBFED7013BF645" box="[1208,1380,295,322]" pageId="30" pageNumber="31" value="1959-12-27" valueMax="1959-12-30" valueMin="1959-12-27">
<collectingDate id="EFF2E9E0FFE2FFE165BBFED7013BF645" box="[1208,1380,295,322]" pageId="30" pageNumber="31" value="1959-12-27" valueMax="1959-12-30" valueMin="1959-12-27">27–30.xii.1959</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFE2FFE16194FEBB0504F662" box="[151,347,331,357]" pageId="30" pageNumber="31">Robert Cumming</collectorName>
(6
<collectionCode id="ED19AE0DFFE2FFE16081FEBC0582F661" box="[386,477,332,358]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFE2FFE160F3FEBB07ACF662" box="[496,1011,331,358]" collectingDate="1958-06-06" collectionCode="CAS" collectorName="E. Pinckert" country="Bolivia" location="Okinawa" pageId="30" pageNumber="31" specimenCount="1" typeStatus="paratype">
<location id="8ED76013FFE2FFE160F3FEBB0607F662" LSID="urn:lsid:plazi:treatment:03A187DEFFE2FFE16194FF67002AF236:8ED76013FFE2FFE160F3FEBB0607F662" box="[496,600,331,357]" country="Bolivia" name="Okinawa" pageId="30" pageNumber="31">Okinawa</location>
,
<date id="FFB61008FFE2FFE16367FEBB068FF661" box="[612,720,331,358]" pageId="30" pageNumber="31" value="1958-06-06">
<collectingDate id="EFF2E9E0FFE2FFE16367FEBB068FF661" box="[612,720,331,358]" pageId="30" pageNumber="31" value="1958-06-06">6.vi.1958</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFE2FFE1620EFEBC07D5F662" box="[781,906,331,358]" pageId="30" pageNumber="31">E. Pinckert</collectorName>
(6
<collectionCode id="ED19AE0DFFE2FFE162B3FEBB07B4F662" box="[944,1003,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="30" pageNumber="31" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFE2FFE16503FEBC0742F68D" collectingDate="1969-02-21" collectionCode="NMNH" collectorName="A. Martinez &amp; R. E. Woodruff" country="Bolivia" location="Prov. Sara" pageId="30" pageNumber="31" specimenCount="1" typeStatus="paratype">
<location id="8ED76013FFE2FFE16503FEBC002AF662" LSID="urn:lsid:plazi:treatment:03A187DEFFE2FFE16194FF67002AF236:8ED76013FFE2FFE16503FEBC002AF662" box="[1024,1141,331,358]" country="Bolivia" name="Prov. Sara" pageId="30" pageNumber="31">Prov. Sara</location>
,
<locationDeviation id="9A052FEAFFE2FFE16581FEBB01C7F661" box="[1154,1432,331,358]" location="Santa Rosa" pageId="30" pageNumber="31">
<quantity id="4CF09B2DFFE2FFE16581FEBB00E4F662" box="[1154,1211,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="km" value="2.0">2 km</quantity>
NW of Santa Rosa
</locationDeviation>
,
<date id="FFB61008FFE2FFE16194FE9F0554F68E" box="[151,267,367,393]" pageId="30" pageNumber="31" value="1969-02-21">
<collectingDate id="EFF2E9E0FFE2FFE16194FE9F0554F68E" box="[151,267,367,393]" pageId="30" pageNumber="31" value="1969-02-21">21.ii.1969</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFE2FFE16046FE9F0592F68E" box="[325,461,367,393]" pageId="30" pageNumber="31">A. Martinez</collectorName>
&amp;
<collectorName id="26FD531EFFE2FFE160F3FE8006CAF68E" box="[496,661,367,394]" pageId="30" pageNumber="31">R.E. Woodruff</collectorName>
(3
<collectionCode id="ED19AE0DFFE2FFE163B9FE80074AF68D" box="[698,789,368,394]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFE2FFE161C4FE6305A1F53A" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BB736C8FFE2FFE161C4FE6305A1F53A" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">
<emphasis id="B97CEADAFFE2FFE161C4FE63058BF6AA" bold="true" box="[199,468,403,429]" pageId="30" pageNumber="31">Differential Diagnosis.</emphasis>
TO MODIFY: Among Bolivian species, recognized by the combination of moderate size (ca. 3.18), the grey dorsum with a large black area on each elytron, the pronotum with depressions shallow, reliefs very weakly raised, if at all, depressions moderately densely punctate, reliefs very sparsely punctate, and the male genitalia (
<figureCitation id="13332A4DFFE2FFE16041FE0F05CBF51D" box="[322,404,511,538]" captionStart="FIGURES 14–15" captionStartId="59.[151,264,1944,1968]" captionTargetBox="[161,1428,217,1846]" captionTargetId="figure-15@59.[151,1435,181,1920]" captionTargetPageId="59" captionText="FIGURES 14–15. Habitus and male genitalia of holotypes. 14. Hydrochus reticulatus n. sp. 15. Hydrochus personatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043290" httpUri="https://zenodo.org/record/5043290/files/figure.png" pageId="30" pageNumber="31">Fig. 15</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C084D4BFFE2FFE162BDFE0F0067F51E" authorityName="Leach" authorityYear="1817" box="[958,1080,511,537]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEADAFFE2FFE162BDFE0F0067F51E" box="[958,1080,511,537]" italics="true" pageId="30" pageNumber="31">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFE2FFE161C4FDB70786F20A" pageId="30" pageNumber="31" type="description">
<paragraph id="8BB736C8FFE2FFE161C4FDB70674F412" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">
<emphasis id="B97CEADAFFE2FFE161C4FDB7050EF566" bold="true" box="[199,337,583,609]" pageId="30" pageNumber="31">Description</emphasis>
. Size:
<typeStatus id="54B3886AFFE2FFE160A7FDB70659F566" box="[420,518,583,609]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.18/1.08; head width 0.73; pronotum l/w 0.70/0.73; PA 0.70; PB 0.63; elytra 1.98/1.08. Habitus as illustrated (
<figureCitation id="13332A4DFFE2FFE1652CFD9B00DEF581" box="[1071,1153,619,646]" captionStart="FIGURES 14–15" captionStartId="59.[151,264,1944,1968]" captionTargetBox="[161,1428,217,1846]" captionTargetId="figure-15@59.[151,1435,181,1920]" captionTargetPageId="59" captionText="FIGURES 14–15. Habitus and male genitalia of holotypes. 14. Hydrochus reticulatus n. sp. 15. Hydrochus personatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043290" httpUri="https://zenodo.org/record/5043290/files/figure.png" pageId="30" pageNumber="31">Fig. 15</figureCitation>
). Dorsum grey, with very slight to moderate iridescence, elytra with large medial mask-like black maculae. Legs brown to testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 1–2x strial puncture diameter. Interstria 9
<superScript id="7C7D9B80FFE2FFE16087FD2705CEF5E2" attach="left" box="[388,401,727,741]" fontSize="6" pageId="30" pageNumber="31">th</superScript>
more convex than others, overhanging 10
<superScript id="7C7D9B80FFE2FFE16274FD2707DBF5E2" attach="left" box="[887,900,727,741]" fontSize="6" pageId="30" pageNumber="31">th</superScript>
interstria. Callus on usual area of 5
<superScript id="7C7D9B80FFE2FFE16420FD27016FF5E2" attach="left" box="[1315,1328,727,741]" fontSize="6" pageId="30" pageNumber="31">th</superScript>
interstria moderately raised, moderately long.
</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FCD006EFF485" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">Pronotum length and width ca. equal, widest at about anterior 1/3, narrowed at base, sides sinuate, smooth; anterior margin very slightly bisinuate; depressions shallow, reliefs very weakly raised, if at all, depressions moderately densely punctate, reliefs very sparsely punctate.</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FC7B0691F4CE" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">Elytra with spaces between strial punctures ca. 1.0–1.5xpd; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with angulation.</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FC2304B6F316" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">Ventral characters: Mentum punctate, with central depression; submentum with two large, narrowly separated foveae.</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FBEB01C8F3C2" blockId="30.[151,1437,259,1330]" pageId="30" pageNumber="31">
Male genitalia (
<figureCitation id="13332A4DFFE2FFE1607CFBEB058BF331" box="[383,468,1051,1078]" captionStart="FIGURES 14–15" captionStartId="59.[151,264,1944,1968]" captionTargetBox="[161,1428,217,1846]" captionTargetId="figure-15@59.[151,1435,181,1920]" captionTargetPageId="59" captionText="FIGURES 14–15. Habitus and male genitalia of holotypes. 14. Hydrochus reticulatus n. sp. 15. Hydrochus personatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043290" httpUri="https://zenodo.org/record/5043290/files/figure.png" pageId="30" pageNumber="31">Fig. 15</figureCitation>
) general characters: genitalia narrow (l/w ratio ca. 49/9); paramere tips distal ends pointed, lateral margins emarginate from rounded plsa to distal tip, medial margins of tips parallel, distal end of tips extending slightly beyond aedeagus, in lateral view parameres ca. as wide as basal piece; aedeagus lanceolate in dorsal/ventral views, widest at ca. midlength; basal piece slightly longer than parameres (ratio ca. 27/22), in dorsal/ ventral views nearly parallel-sided for most of length, tapering near orifice, with rim of orifice weakly sclerotized.
</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FB3F0117F3EE" blockId="30.[151,1437,259,1330]" box="[199,1352,1231,1258]" pageId="30" pageNumber="31">Dorsal surface: adtl near distal end; pdmm widely overlapping alm except at paramere tips; adbl small.</paragraph>
<paragraph id="8BB736C8FFE2FFE161C4FB030786F20A" blockId="30.[151,1437,259,1330]" box="[199,985,1267,1294]" pageId="30" pageNumber="31">Ventral surface: pvmm approximating alm; avtl at about proximal 1/4.</paragraph>
</subSubSection>
<subSubSection id="C3126543FFE2FFE161C4FAE7002AF236" box="[199,1141,1303,1330]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BB736C8FFE2FFE161C4FAE7002AF236" blockId="30.[151,1437,259,1330]" box="[199,1141,1303,1330]" pageId="30" pageNumber="31">
<emphasis id="B97CEADAFFE2FFE161C4FAE70511F236" bold="true" box="[199,334,1303,1329]" pageId="30" pageNumber="31">Etymology.</emphasis>
Named in reference to the large, mask-like black maculae on the elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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