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<mods:title id="59274541CE872797B4C925AD511962A7">On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<emphasis id="B97CEACEFFDA94371FEAFCB09025FC37" bold="true" box="[178,610,888,910]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFCB093AAFC34" ID-CoL="4R3BW" authority="ERICHSON, 1839" box="[178,493,888,910]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus" subGenus="Raphirus">
<emphasis id="B97CEACEFFDA94371FEAFCB0934EFC34" bold="true" box="[178,265,888,910]" italics="true" pageId="2" pageNumber="151">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDA94371E40FCB0933EFC34" bold="true" box="[280,377,888,910]" italics="true" pageId="2" pageNumber="151">Raphirus</emphasis>
) 
<emphasis id="B97CEACEFFDA94371ED0FCB093AAFC34" bold="true" box="[392,493,888,910]" italics="true" pageId="2" pageNumber="151">umbrinus</emphasis>
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ERICHSON
</emphasis>
, 
<emphasis id="B97CEACEFFDA94371D35FCB090E7FC37" bold="true" box="[621,672,888,909]" pageId="2" pageNumber="151">1839</emphasis>
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</paragraph>
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<paragraph id="8BB736DCFFDA94371FEAFC6C91E3FC03" blockId="2.[178,1086,931,1586]" box="[178,932,931,954]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFC6C9063FC03" ID-CoL="4R3BW" authority="ERICHSON, 1839: 491" authorityName="ERICHSON" authorityPageNumber="491" authorityYear="1839" box="[178,548,932,953]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371FEAFC6C931EFC03" box="[178,345,932,953]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
ERICHSON, 1839: 
<specimenCount id="9D0EFD55FFDA94371D5AFC6C9063FC03" box="[514,548,932,953]" pageId="2" pageNumber="151" type="generic">491</specimenCount>
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f. [
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locality: &quot;Neustadt-Eberswalde&quot;].
</paragraph>
</subSubSection>
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<paragraph id="8BB736DCFFDA94371FEAFC7691B5FC6E" blockId="2.[178,1086,931,1586]" box="[178,1010,957,980]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFC7691B5FC6E" box="[178,1010,957,980]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFC7693ADFC6E" ID-CoL="4RKPN" authority="J. SAHLBERG, 1876" box="[178,490,958,980]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="variety" species="umbrinus" variety="maritimus">
<emphasis id="B97CEACEFFDA94371FEAFC769327FC69" box="[178,352,958,979]" italics="true" pageId="2" pageNumber="151">Raphirus umbrinus</emphasis>
var. 
<emphasis id="B97CEACEFFDA94371ED6FC7793ADFC6E" box="[398,490,959,980]" italics="true" pageId="2" pageNumber="151">maritimus</emphasis>
</taxonomicName>
J. 
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<bibRefCitation id="EF994B2DFFDA94371D5BFC769084FC69" author="SAHLBERG J" box="[515,707,958,980]" pageId="2" pageNumber="151" pagination="1 - 248" refId="ref21331" refString="SAHLBERG J. R. (1876): Enumeratio coleopterorum brachelytrorum Fenniae. Systematisk forteckning ofver de inom Finlands naturalhistoriska omrade hittills funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och beskrifningar af nya och mindre kanda species. I. Staphylinidae. - Acta Societatis pro Fauna Fl ora Fennica 1: 1 - 248." type="journal article" year="1876">SAHLBERG, 1876: 29</bibRefCitation>
</treatmentCitation>
[first synonymy: 
<bibRefCitation id="EF994B2DFFDA94371C3CFC7691AFFC6E" author="FAUVEL A" box="[868,1000,957,980]" pageId="2" pageNumber="151" pagination="215 - 253" refId="ref20718" refString="FAUVEL A. (1900): Staphylinides palearctiques nouveaux. - Revue d'Ento mologie 19: 215 - 253." type="journal article" year="1900">Fauvel (1900)</bibRefCitation>
].
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</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFC109117FC57" blockId="2.[178,1086,931,1586]" box="[178,848,984,1006]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFC109324FC57" ID-CoL="8XS9M" authority="" box="[178,355,984,1005]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="subGenus" subGenus="Sauridus">
<emphasis id="B97CEACEFFDA94371FEAFC1092B9FC57" box="[178,254,984,1005]" italics="true" pageId="2" pageNumber="151">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDA94371E53FC10931CFC57" box="[267,347,984,1005]" italics="true" pageId="2" pageNumber="151">Sauridus</emphasis>
)
</taxonomicName>
<emphasis id="B97CEACEFFDA94371E31FC119388FC54" box="[361,463,985,1006]" italics="true" pageId="2" pageNumber="151">cyanescens</emphasis>
MULSANT &amp; REY, 1876: 727 ff.; 
<emphasis id="B97CEACEFFDA94371C59FC109117FC57" bold="true" box="[769,848,984,1005]" pageId="2" pageNumber="151">nov.syn.</emphasis>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFC3A90F3FBBD" blockId="2.[178,1086,931,1586]" box="[178,692,1010,1032]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFC3A90F3FBBD" box="[178,692,1010,1032]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFC3A93A9FBB2" ID-CoL="4R3WY" authority="LEINBERG, 1900" box="[178,494,1010,1032]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="variety" species="umbrinus" variety="tetrastigma">
<emphasis id="B97CEACEFFDA94371FEAFC3A931EFBBD" box="[178,345,1010,1031]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
var. 
<emphasis id="B97CEACEFFDA94371EDFFC3B93A9FBB2" box="[391,494,1011,1032]" italics="true" pageId="2" pageNumber="151">tetrastigma</emphasis>
</taxonomicName>
<treatmentCitation id="0AA910CDFFDA94371EACFC3B90E8FBBD" author="LEINBERG A" box="[500,687,1010,1032]" page="79" pageId="2" pageNumber="151" year="1900">
<bibRefCitation id="EF994B2DFFDA94371EACFC3B90E8FBBD" author="LEINBERG A" box="[500,687,1010,1032]" pageId="2" pageNumber="151" pagination="79 - 80" refId="ref21138" refString="LEINBERG A. (1900): Einige neue Staphyliniden-Varietaten. - Meddelanden af Societas pro Fauna et Flora Fennica 26: 79 - 80." type="journal article" year="1900">LEINBERG, 1900: 79</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFBC491BEFB98" blockId="2.[178,1086,931,1586]" box="[178,1017,1036,1058]" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFBC49398FB9B" ID-CoL="4R3XD" box="[178,479,1036,1058]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="variety" species="dubius" variety="umbripennis">
<emphasis id="B97CEACEFFDA94371FEAFBC49307FB9B" box="[178,320,1036,1057]" italics="true" pageId="2" pageNumber="151">Quedius dubius</emphasis>
var. 
<emphasis id="B97CEACEFFDA94371E36FBC49398FB9B" box="[366,479,1036,1057]" italics="true" pageId="2" pageNumber="151">umbripennis</emphasis>
</taxonomicName>
BERNHAUER &amp; SCHUBERT, 1916: 79; see comment below.
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFBEE9080FB86" blockId="2.[178,1086,931,1586]" box="[178,711,1062,1084]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFBEE9080FB86" box="[178,711,1062,1084]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFBEE9323FB81" box="[178,356,1062,1083]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFBEE92B9FB81" ID-CoL="8VS2M" box="[178,254,1062,1083]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
bulgaricus
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371E32FBEF9016FB86" author="SCHEERPELTZ O" box="[362,593,1063,1084]" page="219" pageId="2" pageNumber="151" year="1937">
<bibRefCitation id="EF994B2DFFDA94371E32FBEF9016FB86" author="SCHEERPELTZ O" box="[362,593,1063,1084]" pageId="2" pageNumber="151" pagination="185 - 246" refId="ref21464" refString="SCHEERPELTZ O. (1937): Wissenschaftliche Ergebnisse einer von Herrn Hofrat F. Schubert, seinem Sohne Herrn cand. phil. F. Schubert und Herrn Prof. Ing. K. Mandl im Sommer 1935 (1936) nach Bulgarien unternommenen Studienreise. Coleoptera: I. Staphylinidae. - Mitteilungen aus den Konigliche n Naturwissenschaftlichen Instituten in Sofia - Bulgarien 10: 185 - 246." type="journal article" year="1937">SCHEERPELTZ, 1937: 219</bibRefCitation>
</treatmentCitation>
ff.; 
<emphasis id="B97CEACEFFDA94371D20FBEF9080FB86" bold="true" box="[632,711,1063,1084]" pageId="2" pageNumber="151">nov.syn.</emphasis>
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</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFB889030FBEC" blockId="2.[178,1086,931,1586]" box="[178,631,1088,1110]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFB889030FBEC" box="[178,631,1088,1110]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFB88932AFBEC" box="[178,365,1088,1110]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFB8892B9FBEF" ID-CoL="8VS2M" box="[178,254,1088,1109]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
cyprusensis
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371E2BFB89904DFBEC" author="LAST H" box="[371,522,1088,1110]" page="251" pageId="2" pageNumber="151" year="1955">
<bibRefCitation id="EF994B2DFFDA94371E2BFB89904DFBEC" author="LAST H" box="[371,522,1088,1110]" pageId="2" pageNumber="151" pagination="251 - 252" refId="ref21105" refString="LAST H. R. (1955): A new Palaearctic species of Quedius (Col., Staphylinidae). - The Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1955">
LAST, 1955: 
<specimenCount id="9D0EFD55FFDA94371EB0FB88904DFBEC" box="[488,522,1088,1110]" pageId="2" pageNumber="151" type="generic">251</specimenCount>
</bibRefCitation>
</treatmentCitation>
f.; 
<emphasis id="B97CEACEFFDA94371D70FB899030FBEC" bold="true" box="[552,631,1089,1110]" pageId="2" pageNumber="151">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFB9390DCFBCA" blockId="2.[178,1086,931,1586]" box="[178,667,1115,1137]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFB9390DCFBCA" box="[178,667,1115,1137]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFB93936AFBCA" box="[178,301,1115,1136]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFB9392B9FBCA" ID-CoL="8VS2M" box="[178,254,1115,1136]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
freyi
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371E6AFB939062FBCA" author="SCHEERPELTZ O" box="[306,549,1115,1137]" page="1102" pageId="2" pageNumber="151" year="1956">
<bibRefCitation id="EF994B2DFFDA94371E6AFB939062FBCA" author="SCHEERPELTZ O" box="[306,549,1115,1137]" pageId="2" pageNumber="151" pagination="1092 - 1117" refId="ref21567" refString="SCHEERPELTZ O. (1956): Die von Herrn Dr. E. Haaf vom Museum Frey-Tutzing gelegentlich seiner Studienreise 1955 in Griechenland und auf der Insel Cypern aufgesammelten Staphylinidae (Col.). - Entomologische Arbeiten aus dem Museum G. Frey 7: 1092 - 1117." type="journal article" year="1956">SCHEERPELTZ, 1956: 1102</bibRefCitation>
</treatmentCitation>
ff.; 
<emphasis id="B97CEACEFFDA94371D14FB9390DCFBCA" bold="true" box="[588,667,1115,1136]" pageId="2" pageNumber="151">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFBBD91B6FB31" blockId="2.[178,1086,931,1586]" box="[178,1009,1141,1163]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFBBD91B6FB31" box="[178,1009,1141,1163]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFBBD93DEFB30" box="[178,409,1141,1162]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFBBD92B9FB30" ID-CoL="8VS2M" box="[178,254,1141,1162]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
pseudoumbrinus
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371EC7FBBD907DFB31" author="LOHSE G" box="[415,570,1141,1163]" page="59" pageId="2" pageNumber="151" year="1958">
<bibRefCitation id="EF994B2DFFDA94371EC7FBBD907DFB31" author="LOHSE G" box="[415,570,1141,1163]" pageId="2" pageNumber="151" pagination="59 - 60" refId="ref21164" refString="LOHSE G. A. (1958): Quedius umbrinus ERICHSON und der Quedius umbrinus der Autoren. - Entomologische Blatter 54: 59 - 60." type="journal article" year="1958">LOHSE, 1958: 59</bibRefCitation>
</treatmentCitation>
; synonymy by 
<bibRefCitation id="EF994B2DFFDA94371D9CFBBD9170FB31" author="POPE R" box="[708,823,1141,1163]" pageId="2" pageNumber="151" refId="ref21240" refString="POPE R. D. (1977): Part 3. XXV. Coleoptera and Strepsiptera. - In: K LOET G. S. &amp; W. D. HINCKS, A check list of British insects. 2 nd ed. Handbooks for the identification of British insects. Vol. XI. - Royal Entom ological Society, London: xiv + 105 pp." type="book" year="1977">POPE (1977)</bibRefCitation>
and 
<bibRefCitation id="EF994B2DFFDA94371C3AFBBD91A9FB31" author="ASSING V" box="[866,1006,1141,1163]" pageId="2" pageNumber="151" pagination="35 - 46" refId="ref19493" refString="ASSING V. (1999): Zur Kenntnis und Synonymie einiger mitteleuropaischer Arten der Gattung Quedius STEPHENS (Col., Staphylinidae). - Entomologische Blatter 95 (1): 35 - 46." type="journal article" year="1999">ASSING (1999)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFB47903BFB1E" blockId="2.[178,1086,931,1586]" box="[178,636,1167,1189]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFB47903BFB1E" box="[178,636,1167,1189]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFB479318FB1F" box="[178,351,1167,1189]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFB4792B9FB1E" ID-CoL="8VS2M" box="[178,254,1167,1188]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
maronitus
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371E3DFB479063FB1E" author="COIFFAIT H" box="[357,548,1167,1189]" page="410" pageId="2" pageNumber="151" year="1963">
<bibRefCitation id="EF994B2DFFDA94371E3DFB479063FB1E" author="COIFFAIT H" box="[357,548,1167,1189]" pageId="2" pageNumber="151" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT, 1963: 410</bibRefCitation>
</treatmentCitation>
; 
<emphasis id="B97CEACEFFDA94371D75FB47903BFB1E" bold="true" box="[557,636,1167,1188]" pageId="2" pageNumber="151">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFB6190D8FB04" blockId="2.[178,1086,931,1586]" box="[178,671,1193,1215]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="AB1811F2FFDA94371FEAFB6190D8FB04" box="[178,671,1193,1215]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFB619328FB05" box="[178,367,1193,1215]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371FEAFB6192B9FB04" ID-CoL="8VS2M" box="[178,254,1193,1214]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
gueorguievi
</emphasis>
<treatmentCitation id="0AA910CDFFDA94371E2DFB619076FB04" author="COIFFAIT H" box="[373,561,1193,1215]" page="399" pageId="2" pageNumber="151" year="1967">
<bibRefCitation id="EF994B2DFFDA94371E2DFB619076FB04" author="COIFFAIT H" box="[373,561,1193,1215]" pageId="2" pageNumber="151" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">
COIFFAIT, 1967: 
<specimenCount id="9D0EFD55FFDA94371D57FB619076FB04" box="[527,561,1193,1214]" pageId="2" pageNumber="151" type="generic">399</specimenCount>
</bibRefCitation>
</treatmentCitation>
f.; 
<emphasis id="B97CEACEFFDA94371D08FB6190D8FB04" bold="true" box="[592,671,1193,1214]" pageId="2" pageNumber="151">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDA94371FEAFB009090FA29" blockId="2.[178,1086,931,1586]" pageId="2" pageNumber="151">
<materialsCitation id="3B603C81FFDA94371FEAFB009303FADC" ID-GBIF-Occurrence="2846796370" collectionCode="MHNL" collectorName="D. Drugmand Rev. &amp; V. Assing" determinerName="V. Assing" pageId="2" pageNumber="151" specimenCount="5" specimenCount-male="5" typeStatus="lectotype">
T y p e m a t e r i a l e x a m i n e d: 
<emphasis id="B97CEACEFFDA94371D38FB0090AEFB64" box="[608,745,1224,1246]" italics="true" pageId="2" pageNumber="151">Q. cyanescens</emphasis>
: 
<typeStatus id="54B3887EFFDA94371DA4FB009110FB67" box="[764,855,1224,1245]" pageId="2" pageNumber="151" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D0EFD55FFDA94371C3BFB009131FB67" box="[867,886,1224,1245]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
, present designation [dissected prior to present study]: &quot; 
<specimenCount id="9D0EFD55FFDA94371EACFB179041FB4E" box="[500,518,1247,1268]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
14 / 
<typeStatus id="54B3887EFFDA94371D6DFB1790C8FB4E" box="[565,655,1247,1268]" pageId="2" pageNumber="151" type="lectotype">Lectotype</typeStatus>
/ 
<collectorName id="26FD530AFFDA94371DFBFB179115FB4E" box="[675,850,1247,1269]" pageId="2" pageNumber="151">D. Drugmand Rev.</collectorName>
1995 
<taxonomicName id="4C084D5FFFDA94371CD3FB17967AFB4E" ID-CoL="8XS9M" authority="" box="[907,1085,1247,1268]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="subGenus" subGenus="Sauridus">
<emphasis id="B97CEACEFFDA94371CD3FB179191FB4E" box="[907,982,1247,1268]" italics="true" pageId="2" pageNumber="151">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDA94371CBDFB179671FB4E" box="[997,1078,1247,1268]" italics="true" pageId="2" pageNumber="151">Sauridus</emphasis>
)
</taxonomicName>
<emphasis id="B97CEACEFFDA94371FEAFB3E935FFAB1" box="[178,280,1270,1291]" italics="true" pageId="2" pageNumber="151">cyanescens</emphasis>
Muls. Rey / 
<typeStatus id="54B3887EFFDA94371ED7FB3E93B3FAB1" box="[399,500,1270,1291]" pageId="2" pageNumber="151" type="lectotype">Lectotypus</typeStatus>
<specimenCount id="9D0EFD55FFDA94371EA2FB3E904CFAB1" box="[506,523,1270,1291]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
<emphasis id="B97CEACEFFDA94371D49FB3D908FFAB1" box="[529,712,1269,1291]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371D49FB3D901AFAB0" ID-CoL="8VS2M" box="[529,605,1269,1290]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
cyanescens
</emphasis>
Mulsant &amp; Rey, desig. 
<collectorName id="26FD530AFFDA94371CFAFB3E91BAFAB1" box="[930,1021,1269,1291]" pageId="2" pageNumber="151">V. Assing</collectorName>
2017 / 
<taxonomicName id="4C084D5FFFDA94371FEAFAC493F4FA98" ID-CoL="4R3BW" authority="Erichson" authorityName="Erichson" box="[178,435,1292,1314]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371FEAFAC4931EFA9B" box="[178,345,1292,1313]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
Erichson
</taxonomicName>
, det. 
<determinerName id="7E6BDA18FFDA94371EBCFAC49078FA98" box="[484,575,1292,1314]" pageId="2" pageNumber="151">V. Assing</determinerName>
2017&quot; (
<collectionCode id="ED19AE19FFDA94371DD2FAC49096FA9B" box="[650,721,1292,1313]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="2" pageNumber="151">MHNL</collectionCode>
). 
<typeStatus id="54B3887EFFDA94371DB9FAC4912EFA98" box="[737,873,1292,1314]" pageId="2" pageNumber="151" type="paralectotype">Paralectotypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFDA94371C29FAC491D7FA9B" box="[881,912,1292,1313]" pageId="2" pageNumber="151" type="male">1♂</specimenCount>
: &quot;1 / 
<typeStatus id="54B3887EFFDA94371C9AFAC4967AFA98" box="[962,1085,1292,1314]" pageId="2" pageNumber="151" type="paralectotype">Paralectotype</typeStatus>
/ 
<collectorName id="26FD530AFFDA94371F98FAEB9335FA82" box="[192,370,1315,1337]" pageId="2" pageNumber="151">D. Drugmand Rev.</collectorName>
1995 
<taxonomicName id="4C084D5FFFDA94371EF5FAEB9026FA82" ID-CoL="8XS9M" authority="" box="[429,609,1315,1336]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="subGenus" subGenus="Sauridus">
<emphasis id="B97CEACEFFDA94371EF5FAEB93BEFA82" box="[429,505,1315,1336]" italics="true" pageId="2" pageNumber="151">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDA94371D50FAEB901EFA82" box="[520,601,1315,1336]" italics="true" pageId="2" pageNumber="151">Sauridus</emphasis>
)
</taxonomicName>
<emphasis id="B97CEACEFFDA94371D30FAEB9089FA82" box="[616,718,1315,1336]" italics="true" pageId="2" pageNumber="151">cyanescens</emphasis>
Muls. Rey / 
<typeStatus id="54B3887EFFDA94371C14FAEB9196FA83" box="[844,977,1315,1337]" pageId="2" pageNumber="151" type="paralectotype">Paralectotypus</typeStatus>
<specimenCount id="9D0EFD55FFDA94371C81FAEB91ADFA82" box="[985,1002,1315,1336]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
<emphasis id="B97CEACEFFDA94371CAAFAEB935FFAF5" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371CAAFAEB967AFA82" ID-CoL="8VS2M" box="[1010,1085,1315,1336]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
cyanescens
</emphasis>
Mulsant &amp; Rey, desig. 
<collectorName id="26FD530AFFDA94371EA5FAF2901DFAF5" box="[509,602,1337,1359]" pageId="2" pageNumber="151">V. Assing</collectorName>
2017 / 
<taxonomicName id="4C084D5FFFDA94371DFEFAF191ECFAF5" ID-CoL="4R3BW" authority="Erichson" authorityName="Erichson" box="[678,939,1337,1359]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371DFEFAF19117FAF4" box="[678,848,1337,1358]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
Erichson
</taxonomicName>
, det. 
<determinerName id="7E6BDA18FFDA94371CB8FAF2967AFAF5" box="[992,1085,1337,1359]" pageId="2" pageNumber="151">V. Assing</determinerName>
2017&quot; (
<collectionCode id="ED19AE19FFDA94371FAEFA989379FADC" box="[246,318,1360,1382]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="2" pageNumber="151">MHNL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDA94371E16FA989095FA29" ID-GBIF-Occurrence="2846796387" collectionCode="MHNL" collectorName="D. Drugmand Rev. &amp; V. Assing" determinerName="V. Assing" isTailEnumeration="true" pageId="2" pageNumber="151" specimenCount="3" specimenCount-female="3" typeStatus="paralectotype">
<specimenCount id="9D0EFD55FFDA94371E16FA98932DFAD2" box="[334,362,1360,1384]" pageId="2" pageNumber="151" type="female">1♀</specimenCount>
: &quot; 
<specimenCount id="9D0EFD55FFDA94371ED9FA9B93D5FAD2" box="[385,402,1363,1384]" pageId="2" pageNumber="151" type="female">♀</specimenCount>
/ 
<typeStatus id="54B3887EFFDA94371EFEFA989066FADC" box="[422,545,1360,1382]" pageId="2" pageNumber="151" type="paralectotype">Paralectotype</typeStatus>
/ 
<collectorName id="26FD530AFFDA94371D6DFA9890A3FADF" box="[565,740,1360,1382]" pageId="2" pageNumber="151">D. Drugmand Rev.</collectorName>
1995 
<taxonomicName id="4C084D5FFFDA94371C46FA989196FADF" ID-CoL="8XS9M" authority="" box="[798,977,1360,1381]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="subGenus" subGenus="Sauridus">
<emphasis id="B97CEACEFFDA94371C46FA98912DFADF" box="[798,874,1360,1381]" italics="true" pageId="2" pageNumber="151">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDA94371C20FA98918EFADF" box="[888,969,1360,1381]" italics="true" pageId="2" pageNumber="151">Sauridus</emphasis>
)
</taxonomicName>
<emphasis id="B97CEACEFFDA94371C8FFA99967AFADC" box="[983,1085,1361,1382]" italics="true" pageId="2" pageNumber="151">cyanescens</emphasis>
Muls. Rey / 
<typeStatus id="54B3887EFFDA94371E6BFAAF93FFFAC7" box="[307,440,1383,1405]" pageId="2" pageNumber="151" type="paralectotype">Paralectotypus</typeStatus>
<specimenCount id="9D0EFD55FFDA94371E9CFAA29392FAC5" box="[452,469,1386,1407]" pageId="2" pageNumber="151" type="female">♀</specimenCount>
<emphasis id="B97CEACEFFDA94371EB8FAAF90D9FAC6" box="[480,670,1383,1404]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371EB8FAAF906BFAC6" ID-CoL="8VS2M" box="[480,556,1383,1404]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
cyanescens
</emphasis>
Mulsant &amp; Rey, desig. 
<collectorName id="26FD530AFFDA94371CCAFAAF91B5FAC7" box="[914,1010,1383,1405]" pageId="2" pageNumber="151">V. Assing</collectorName>
2017 / 
<taxonomicName id="4C084D5FFFDA94371FEAFAB593F6FA29" ID-CoL="4R3BW" authority="Erichson" authorityName="Erichson" box="[178,433,1405,1427]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371FEAFAB5931EFA28" box="[178,345,1405,1426]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
Erichson
</taxonomicName>
, det. 
<determinerName id="7E6BDA18FFDA94371EB9FAB6907BFA29" box="[481,572,1405,1427]" pageId="2" pageNumber="151">V. Assing</determinerName>
2017&quot; (
<collectionCode id="ED19AE19FFDA94371DDCFAB5908BFA29" box="[644,716,1405,1427]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="2" pageNumber="151">MHNL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDA94361FEAFA5C93B2FCDA" lastPageId="3" lastPageNumber="152" pageId="2" pageNumber="151" type="description">
<paragraph id="8BB736DCFFDA94371FEAFA5C90CAF988" blockId="2.[178,1086,931,1586]" pageId="2" pageNumber="151">
<emphasis id="B97CEACEFFDA94371FEAFA5C9376FA13" box="[178,305,1428,1449]" italics="true" pageId="2" pageNumber="151">Q. bulgaricus</emphasis>
: 
<materialsCitation id="3B603C81FFDA94371E67FA5C9073FA54" ID-GBIF-Occurrence="2846796358" collectionCode="NHMW" collectorName="Scheerpeltz &amp; O. Scheerpeltz &amp; V. Assing" determinerName="V. Assing" elevation="1900" pageId="2" pageNumber="151" specimenCount="3" specimenCount-male="3" typeStatus="lectotype">
<typeStatus id="54B3887EFFDA94371E67FA5C93DDFA13" box="[319,410,1428,1449]" pageId="2" pageNumber="151" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D0EFD55FFDA94371EF9FA5C93F3FA13" box="[417,436,1428,1449]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
, present designation [dissected prior to present study]: &quot; 
<specimenCount id="9D0EFD55FFDA94371C9AFA5C9193FA13" box="[962,980,1428,1449]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
/ Rosalitopolje, Schipka-Balkan, ca. 
<quantity id="4CF09B39FFDA94371EFEFA6393ABFA7A" box="[422,492,1451,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="151" unit="m" value="1900.0">
<elevation id="0025D1EFFFDA94371EFEFA6393ABFA7A" box="[422,492,1451,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="151" unit="m" value="1900.0">1900 m</elevation>
</quantity>
/ ex coll. 
<collectorName id="26FD530AFFDA94371D1DFA6390E8FA7B" box="[581,687,1451,1473]" pageId="2" pageNumber="151">Scheerpeltz</collectorName>
/ Typus 
<emphasis id="B97CEACEFFDA94371DA7FA6391F5FA7A" box="[767,946,1451,1472]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371DA7FA63910DFA7A" box="[767,842,1451,1472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
bulgaricus
</emphasis>
<collectorName id="26FD530AFFDA94371CEFFA63967AFA7B" box="[951,1085,1451,1473]" pageId="2" pageNumber="151">O. Scheerpeltz</collectorName>
/ 
<typeStatus id="54B3887EFFDA94371F9DFA0A936DFA6D" box="[197,298,1474,1495]" pageId="2" pageNumber="151" type="lectotype">Lectotypus</typeStatus>
<specimenCount id="9D0EFD55FFDA94371E6EFA0A9300FA6D" box="[310,327,1474,1495]" pageId="2" pageNumber="151" type="male">♂</specimenCount>
<emphasis id="B97CEACEFFDA94371E0BFA09904BFA6C" box="[339,524,1473,1494]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371E0BFA0993D9FA6C" box="[339,414,1473,1494]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
bulgaricus
</emphasis>
Scheerpeltz, desig. 
<collectorName id="26FD530AFFDA94371D8EFA0A9170FA6D" box="[726,823,1473,1495]" pageId="2" pageNumber="151">V. Assing</collectorName>
2017 / 
<taxonomicName id="4C084D5FFFDA94371CD7FA099342FA54" authority="Erichson" authorityName="Erichson" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371CD7FA09967AFA6C" box="[911,1085,1473,1494]" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
Erichson
</taxonomicName>
, det. 
<determinerName id="7E6BDA18FFDA94371E6FFA1093D4FA54" box="[311,403,1496,1518]" pageId="2" pageNumber="151">V. Assing</determinerName>
2017&quot; (
<collectionCode id="ED19AE19FFDA94371E86FA109069FA54" box="[478,558,1496,1518]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="151" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B603C81FFDA94371D65FA10967EFA54" ID-GBIF-Occurrence="2846796334" box="[573,1081,1496,1520]" collectionCode="NHMW" pageId="2" pageNumber="151" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="54B3887EFFDA94371D65FA109082FA54" box="[573,709,1496,1518]" pageId="2" pageNumber="151" type="paralectotype">Paralectotypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFDA94371D97FA1090ACFA4A" box="[719,747,1496,1520]" pageId="2" pageNumber="151" type="female">1♀</specimenCount>
: same labels as 
<typeStatus id="54B3887EFFDA94371CDCFA109190FA54" box="[900,983,1496,1518]" pageId="2" pageNumber="151" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED19AE19FFDA94371CBCFA109674FA54" box="[996,1075,1496,1518]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="151" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDA94371FEAFA2790F7F9BF" ID-GBIF-Occurrence="2846796316" box="[178,688,1519,1541]" collectionCode="NHMW" collectorName="Scheerpeltz &amp; O. Scheerpeltz &amp; V. Assing" determinerName="V. Assing" elevation="1900" pageId="2" pageNumber="151" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="9D0EFD55FFDA94371FEAFA279296F9BE" box="[178,209,1519,1540]" pageId="2" pageNumber="151" type="male">1♂</specimenCount>
: same data as lectotype, but &quot;Cotypus&quot; (
<collectionCode id="ED19AE19FFDA94371D02FA2790EDF9BF" box="[602,682,1519,1541]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="151" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDA94371DE6FA2790CEF988" ID-GBIF-Occurrence="2846796353" collectionCode="NHMW" collectorName="Scheerpeltz &amp; O. Scheerpeltz" determinerName="V. Assing" elevation="1600" pageId="2" pageNumber="151" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<specimenCount id="9D0EFD55FFDA94371DE6FA27909DF9BD" box="[702,730,1519,1543]" pageId="2" pageNumber="151" type="female">1♀</specimenCount>
: &quot; 
<specimenCount id="9D0EFD55FFDA94371DABFA3A9143F9BD" box="[755,772,1522,1543]" pageId="2" pageNumber="151" type="female">♀</specimenCount>
/ Jumruktschal, Schipka-Balkan, 
<quantity id="4CF09B39FFDA94371FEAF9CD9376F9A1" box="[178,305,1541,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="2" pageNumber="151" unit="m" value="1600.0" valueMax="1800.0" valueMin="1400.0">
<elevation id="0025D1EFFFDA94371FEAF9CD9376F9A1" box="[178,305,1541,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="2" pageNumber="151" unit="m" value="1600.0" valueMax="1800.0" valueMin="1400.0">1400-1800 m</elevation>
</quantity>
/ ex coll. 
<collectorName id="26FD530AFFDA94371EFDF9CD9048F9A1" box="[421,527,1541,1563]" pageId="2" pageNumber="151">Scheerpeltz</collectorName>
/ Cotypus 
<emphasis id="B97CEACEFFDA94371DDCF9CD917BF9A0" box="[644,828,1541,1562]" italics="true" pageId="2" pageNumber="151">
<taxonomicName id="4C084D5FFFDA94371DDCF9CD9088F9A0" box="[644,719,1541,1562]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
bulgaricus
</emphasis>
<collectorName id="26FD530AFFDA94371C10F9CD9193F9A1" box="[840,980,1541,1563]" pageId="2" pageNumber="151">O. Scheerpeltz</collectorName>
/ 
<taxonomicName id="4C084D5FFFDA94371CAAF9CD9327F988" authority="Erichson" authorityName="Erichson" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDA94371CAAF9CD934FF98B" italics="true" pageId="2" pageNumber="151">Quedius umbrinus</emphasis>
Erichson
</taxonomicName>
, det. 
<determinerName id="7E6BDA18FFDA94371EC8F9D493ACF988" box="[400,491,1564,1586]" pageId="2" pageNumber="151">V. Assing</determinerName>
2017&quot; (
<collectionCode id="ED19AE19FFDA94371D6BF9D490C4F988" box="[563,643,1564,1586]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="151" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB736DCFFDB94361FEAFF1F91F0FEA6" blockId="3.[178,1086,214,285]" pageId="3" pageNumber="152">
A d d i t i o n a l m a t e r i a l e x a m i n e d: Numerous specimens from 
<collectingCountry id="F31F764CFFDB94361CBEFF1F967EFF56" box="[998,1081,215,236]" name="Germany" pageId="3" pageNumber="152">Germany</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361F87FF269359FEBE" box="[223,286,238,260]" name="Poland" pageId="3" pageNumber="152">Poland</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361E76FF269310FEBE" box="[302,343,238,260]" name="Italy" pageId="3" pageNumber="152">Italy</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361E30FF2693EDFEBE" box="[360,426,238,260]" name="Austria" pageId="3" pageNumber="152">Austria</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361EE2FF26904FFEBE" box="[442,520,238,260]" name="Slovakia" pageId="3" pageNumber="152">Slovakia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361D41FF26902DFEBE" box="[537,618,238,260]" name="Romania" pageId="3" pageNumber="152">Romania</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361D22FF26908FFEBE" box="[634,712,238,260]" name="Slovenia" pageId="3" pageNumber="152">Slovenia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361D80FF269155FEBE" box="[728,786,238,260]" name="Serbia" pageId="3" pageNumber="152">Serbia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361C7AFF26912EFEBE" box="[802,873,238,260]" name="Albania" pageId="3" pageNumber="152">Albania</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361C21FF26919BFEBE" box="[889,988,238,260]" name="Macedonia" pageId="3" pageNumber="152">Macedonia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361CB5FF26967DFEBE" box="[1005,1082,238,260]" name="Bulgaria" pageId="3" pageNumber="152">Bulgaria</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361F87FECF9358FEA6" box="[223,287,263,284]" name="Greece" pageId="3" pageNumber="152">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361E7FFECE932FFEA6" box="[295,360,262,284]" name="Turkey" pageId="3" pageNumber="152">Turkey</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361E29FECF93F5FEA6" box="[369,434,263,284]" name="Cyprus" pageId="3" pageNumber="152">Cyprus</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361EE3FECE904DFEA6" box="[443,522,262,284]" name="Lebanon" pageId="3" pageNumber="152">Lebanon</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361D4AFECE900BFEA6" box="[530,588,262,284]" name="Russia" pageId="3" pageNumber="152">Russia</collectingCountry>
(West Caucasus), 
<collectingCountry id="F31F764CFFDB94361DB7FECE9170FEA6" box="[751,823,262,284]" name="Georgia" pageId="3" pageNumber="152">Georgia</collectingCountry>
, and 
<collectingCountry id="F31F764CFFDB94361C3CFECE91F4FEA6" box="[868,947,262,284]" name="Armenia" pageId="3" pageNumber="152">Armenia</collectingCountry>
.
</paragraph>
<paragraph id="8BB736DCFFDB94361FEAFEE39127FEE4" blockId="3.[178,1085,299,350]" pageId="3" pageNumber="152">
C o m m e n t: 
<taxonomicName id="4C084D5FFFDB94361E3FFEE39062FEFA" box="[359,549,299,320]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361E3FFEE39062FEFA" box="[359,549,299,320]" italics="true" pageId="3" pageNumber="152">Quedius umbrinus</emphasis>
</taxonomicName>
was described based on an unspecified number of 
<typeStatus id="54B3887EFFDB94361FEAFE80934CFEE7" box="[178,267,328,349]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
from &quot;Neustadt-Eberswalde&quot;, 
<collectingCountry id="F31F764CFFDB94361D1AFE8090E6FEE7" box="[578,673,328,349]" name="Germany" pageId="3" pageNumber="152">Germany</collectingCountry>
(ERICHSON 1839).
</paragraph>
<paragraph id="8BB736DCFFDB94361FEAFEA59163FD89" blockId="3.[178,1086,365,563]" pageId="3" pageNumber="152">
The original description of 
<emphasis id="B97CEACEFFDB94361E91FEA6901DFE39" box="[457,602,366,387]" italics="true" pageId="3" pageNumber="152">Q. cyanescens</emphasis>
is based on an unspecified number of 
<typeStatus id="54B3887EFFDB94361CBCFEA6967AFE39" box="[996,1085,366,387]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
from &quot;les environs d'Hyères&quot; (MULSANT &amp; REY 1876). The three 
<typeStatus id="54B3887EFFDB94361C33FE439183FE1A" box="[875,964,395,416]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
in the Rey collection had been examined, partly dissected, and labelled as 
<typeStatus id="54B3887EFFDB94361C1EFE6091E4FE04" box="[838,931,424,446]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
and 
<typeStatus id="54B3887EFFDB94361C80FE6092ADFE61" pageId="3" pageNumber="152" type="paralectotype">paralectotypes</typeStatus>
, respectively, by Didier Drugmand. A designation, however, was never published. In order to establish a formal designation, the male in better condition is designated as the 
<typeStatus id="54B3887EFFDB94361F80FDC8937FFDAC" box="[216,312,512,534]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
. The aedeagus of the 
<typeStatus id="54B3887EFFDB94361D40FDC89032FDAC" box="[536,629,512,534]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
is within the range of intraspecific variation of that of 
<taxonomicName id="4C084D5FFFDB94361E4DFDD693D4FD88" box="[277,403,541,563]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361E4DFDD693D4FD88" box="[277,403,541,563]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
. Hence the synonymy proposed above.
</paragraph>
<paragraph id="8BB736DCFFDB94361FEAFD8B93B2FCDA" blockId="3.[178,1085,577,865]" pageId="3" pageNumber="152">
The first and second editions of the Palaearctic Catalogue (
<bibRefCitation id="EF994B2DFFDB94361C4FFD8B91FFFDE2" author="SMETANA A" box="[791,952,579,600]" pageId="3" pageNumber="152" pagination="505 - 698" refId="ref21710" refString="SMETANA A. (2004): Staphylinidae, subfamilies Omaliinae-Dasycerinae, Phloeocharinae- Apaticinae, Piestinae-Staphylininae, pp. 237 - 272, 329 - 495, 505 - 698. - In: L OBL I. &amp; A. SMETANA (eds), Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea - Histeroidea - Staphylinoidea. - Apollo B ooks, Stenstrup: 942 pp." type="book chapter" year="2004">SMETANA 2004</bibRefCitation>
, SCHÜLKE &amp; SMETANA 2015) list 
<emphasis id="B97CEACEFFDB94361EC9FDA89028FDCF" box="[401,623,608,629]" italics="true" pageId="3" pageNumber="152">
<taxonomicName id="4C084D5FFFDB94361EC9FDA893A2FDCF" box="[401,485,608,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
umbripennis
</emphasis>
<bibRefCitation id="EF994B2DFFDB94361D22FDA89153FDCF" author="ROUBAL J" box="[634,788,608,629]" pageId="3" pageNumber="152" pagination="111 - 112" refId="ref21310" refString="ROUBAL J. (1913): Verschiedene koleopterologische Notizen. - Coleopterolo gische Rundschau 2: 111 - 112." type="journal article" year="1913">ROUBAL, 1913</bibRefCitation>
as a junior synonym of 
<taxonomicName id="4C084D5FFFDB94361B7CFDA89355FD28" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361B7CFDA89355FD28" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EF994B2DFFDB94361E47FDB69387FD2E" author="ROUBAL J" box="[287,448,638,660]" pageId="3" pageNumber="152" pagination="111 - 112" refId="ref21310" refString="ROUBAL J. (1913): Verschiedene koleopterologische Notizen. - Coleopterolo gische Rundschau 2: 111 - 112." type="journal article" year="1913">ROUBAL (1913)</bibRefCitation>
described 
<emphasis id="B97CEACEFFDB94361D68FDB69097FD28" box="[560,720,637,659]" italics="true" pageId="3" pageNumber="152">Q. umbripennis</emphasis>
as an aberration of 
<taxonomicName id="4C084D5FFFDB94361CC6FDB5967AFD28" baseAuthorityName="Heer" baseAuthorityYear="1839" box="[926,1085,637,658]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B97CEACEFFDB94361CC6FDB5967AFD28" box="[926,1085,637,658]" italics="true" pageId="3" pageNumber="152">Quedius dubius</emphasis>
</taxonomicName>
from the Caucasus, i.e., as an infrasubspecific and consequently unavailable name (see also 
<bibRefCitation id="EF994B2DFFDB94361FBFFD7093C2FD74" author="HERMAN L" box="[231,389,696,718]" pageId="3" pageNumber="152" refId="ref20779" refString="HERMAN L. H. (2001): Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII. - Bulletin of the America n Museum of Natural History 265: 4218 pp." type="book" year="2001">HERMAN 2001</bibRefCitation>
). In treating the name as a variety of 
<taxonomicName id="4C084D5FFFDB94361C6BFD7091DCFD77" baseAuthorityName="Heer" baseAuthorityYear="1839" box="[819,923,696,717]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B97CEACEFFDB94361C6BFD7091DCFD77" box="[819,923,696,717]" italics="true" pageId="3" pageNumber="152">Q. dubius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF994B2DFFDB94361CF7FD709676FD74" author="HEER O" box="[943,1073,696,718]" pageId="3" pageNumber="152" pagination="361 - 542" refId="ref20739" refString="HEER O. (1839): Fascicule III. Pp. 361 - 542. - In: F auna Coleopterorum Helvetica Pars I. - Oreliii, Fuesslini et Sociorum, Turici: xii + 652 pp." type="book chapter" year="1839">HEER, 1839</bibRefCitation>
), BERNHAUER &amp; SCHUBERT (1916) made the name available and, according to the Code, they must be considered the authors of the taxon. In more recent articles (
<bibRefCitation id="EF994B2DFFDB94361CC7FD3B967EFCB3" author="COIFFAIT H" box="[927,1081,755,777]" pageId="3" pageNumber="152" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT 1963</bibRefCitation>
, 1968, 
<bibRefCitation id="EF994B2DFFDB94361FABFCD8933FFC9C" author="KORGE H" box="[243,376,784,806]" pageId="3" pageNumber="152" pagination="105 - 126" refId="ref20886" refString="KORGE H. (1964): Carabiden- und Staphylinidenfunde in den Pontischen Gebirgen Kleinasiens und in Mazedonien (Coleoptera). - Reichenbachia 4 (14): 105 - 126." type="journal article" year="1964">KORGE 1964</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFDB94361EDCFCD8905DFC9C" author="HERMAN L" box="[388,538,784,806]" pageId="3" pageNumber="152" refId="ref20779" refString="HERMAN L. H. (2001): Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII. - Bulletin of the America n Museum of Natural History 265: 4218 pp." type="book" year="2001">HERMAN 2001</bibRefCitation>
) it is treated as a distinct species. Based on the illustrations of the aedeagus provided by 
<bibRefCitation id="EF994B2DFFDB94361D77FCE690A2FCFE" author="COIFFAIT H" box="[559,741,814,836]" pageId="3" pageNumber="152" refId="ref20382" refString="COIFFAIT H. (1978 a): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvell e Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">COIFFAIT (1978a)</bibRefCitation>
, however, it was correctly placed in synonymy with 
<taxonomicName id="4C084D5FFFDB94361E37FC8393AAFCDA" box="[367,493,843,864]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361E37FC8393AAFCDA" box="[367,493,843,864]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDB94311FEAFCB8901AFD89" lastPageId="4" lastPageNumber="153" pageId="3" pageNumber="152" type="materials_examined">
<paragraph id="8BB736DCFFDB94361FEAFCB89370FA6A" blockId="3.[178,1087,880,1488]" pageId="3" pageNumber="152">
<bibRefCitation id="EF994B2DFFDB94361FEAFCB89371FC3C" author="LAST H" box="[178,310,880,902]" pageId="3" pageNumber="152" pagination="251 - 252" refId="ref21105" refString="LAST H. R. (1955): A new Palaearctic species of Quedius (Col., Staphylinidae). - The Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1955">LAST (1955)</bibRefCitation>
described 
<emphasis id="B97CEACEFFDB94361EF6FCB8900FFC3C" box="[430,584,880,902]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
based on a male 
<typeStatus id="54B3887EFFDB94361C49FCB8912DFC3C" box="[785,874,880,902]" pageId="3" pageNumber="152" type="holotype">holotype</typeStatus>
and an unspecified number of 
<typeStatus id="54B3887EFFDB94361E75FC4693C9FC19" box="[301,398,910,931]" pageId="3" pageNumber="152" type="paratype">paratypes</typeStatus>
from &quot;Mount Troodus&quot;, 
<collectingCountry id="F31F764CFFDB94361DFAFC4690A9FC19" box="[674,750,910,931]" name="Cyprus" pageId="3" pageNumber="152">Cyprus</collectingCountry>
. 
<emphasis id="B97CEACEFFDB94361DA6FC4591CBFC19" box="[766,908,909,931]" italics="true" pageId="3" pageNumber="152">
<taxonomicName id="4C084D5FFFDB94361DA6FC459115FC18" box="[766,850,909,930]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
freyi
</emphasis>
, which was described based on 
<specimenCount id="9D0EFD55FFDB94361E3BFC6393F7FC7A" box="[355,432,939,961]" pageId="3" pageNumber="152" type="generic">16 type</specimenCount>
specimens from &quot;Pedhoulas&quot; only one year later (
<bibRefCitation id="EF994B2DFFDB94361CE9FC6392A1FC64" author="SCHEERPELTZ O" pageId="3" pageNumber="152" pagination="1092 - 1117" refId="ref21567" refString="SCHEERPELTZ O. (1956): Die von Herrn Dr. E. Haaf vom Museum Frey-Tutzing gelegentlich seiner Studienreise 1955 in Griechenland und auf der Insel Cypern aufgesammelten Staphylinidae (Col.). - Entomologische Arbeiten aus dem Museum G. Frey 7: 1092 - 1117." type="journal article" year="1956">SCHEERPELTZ 1956</bibRefCitation>
) was subsequently synonymized with 
<emphasis id="B97CEACEFFDB94361DD3FC009161FC64" box="[651,806,968,990]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
by 
<bibRefCitation id="EF994B2DFFDB94361C00FC009653FC64" author="COIFFAIT H" box="[856,1044,968,990]" pageId="3" pageNumber="152" refId="ref20382" refString="COIFFAIT H. (1978 a): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvell e Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">COIFFAIT (1978a)</bibRefCitation>
. At present, 
<emphasis id="B97CEACEFFDB94361E53FC2E93E4FC41" box="[267,419,998,1019]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
is known from 
<collectingCountry id="F31F764CFFDB94361D0AFC2E90D9FC41" box="[594,670,998,1019]" name="Cyprus" pageId="3" pageNumber="152">Cyprus</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361DF3FC2D9141FC41" box="[683,774,997,1019]" name="Lebanon" pageId="3" pageNumber="152">Lebanon</collectingCountry>
, and 
<collectingCountry id="F31F764CFFDB94361C1AFC2D91CCFC41" box="[834,907,997,1019]" name="Turkey" pageId="3" pageNumber="152">Turkey</collectingCountry>
(ANLAŞ &amp; ROSE 2009, ASSING &amp; WUNDERLE 2001, 
<bibRefCitation id="EF994B2DFFDB94361D67FBCB90A6FBA2" author="BORDONI A" box="[575,737,1027,1048]" pageId="3" pageNumber="152" pagination="331 - 345" refId="ref20178" refString="BORDONI A. (1984): Note su alcuni stafilinidi del Libano. - Fragmenta Ento mologica, Roma 17 (2): 331 - 345." type="journal article" year="1984">BORDONI 1984</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFDB94361DACFBCB91D1FBA3" author="COIFFAIT H" box="[756,918,1027,1049]" pageId="3" pageNumber="152" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT 1963</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFDB94361CF0FBCB919AFBA2" author="COIFFAIT H" box="[936,989,1027,1048]" pageId="3" pageNumber="152" pagination="61 - 68" refId="ref20324" refString="COIFFAIT H. (1970): Staphylinides egeidiens. - L'Entomologiste 26 (3): 61 - 68." type="journal article" year="1970">1970</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFDB94361CA8FBCB9675FBA3" author="COIFFAIT H" box="[1008,1074,1027,1049]" pageId="3" pageNumber="152" refId="ref20382" refString="COIFFAIT H. (1978 a): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvell e Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">1978a</bibRefCitation>
). According to 
<bibRefCitation id="EF994B2DFFDB94361E66FBE893FAFB8C" author="LAST H" box="[318,445,1056,1078]" pageId="3" pageNumber="152" pagination="251 - 252" refId="ref21105" refString="LAST H. R. (1955): A new Palaearctic species of Quedius (Col., Staphylinidae). - The Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1955">LAST (1955)</bibRefCitation>
and 
<bibRefCitation id="EF994B2DFFDB94361EB7FBE8908FFB8C" author="SCHEERPELTZ O" box="[495,712,1056,1078]" pageId="3" pageNumber="152" pagination="1092 - 1117" refId="ref21567" refString="SCHEERPELTZ O. (1956): Die von Herrn Dr. E. Haaf vom Museum Frey-Tutzing gelegentlich seiner Studienreise 1955 in Griechenland und auf der Insel Cypern aufgesammelten Staphylinidae (Col.). - Entomologische Arbeiten aus dem Museum G. Frey 7: 1092 - 1117." type="journal article" year="1956">SCHEERPELTZ (1956)</bibRefCitation>
, the species is distinguished from 
<taxonomicName id="4C084D5FFFDB94361B7CFBE89355FBE8" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361B7CFBE89355FBE8" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
by darker coloration of the legs, a larger and more robust body, differently shaped antennae and pronotum, and by the shape of the paramere. 
<bibRefCitation id="EF994B2DFFDB94361C05FB9391ACFBCB" author="LOHSE G" box="[861,1003,1115,1137]" pageId="3" pageNumber="152" pagination="59 - 60" refId="ref21164" refString="LOHSE G. A. (1958): Quedius umbrinus ERICHSON und der Quedius umbrinus der Autoren. - Entomologische Blatter 54: 59 - 60." type="journal article" year="1958">LOHSE (1958)</bibRefCitation>
already noted that in both external and the male sexual characters 
<emphasis id="B97CEACEFFDB94361C4EFBB091E9FB34" box="[790,942,1144,1166]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
much resembled 
<taxonomicName id="4C084D5FFFDB94361FBFFB5E932EFB10" box="[231,361,1173,1195]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361FBFFB5E932EFB10" box="[231,361,1173,1195]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
. A comparison of specimens previously identified as 
<emphasis id="B97CEACEFFDB94361CFCFB5E967AFB11" box="[932,1085,1174,1195]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
(see ASSING &amp; WUNDERLE 2001) with material of 
<taxonomicName id="4C084D5FFFDB94361DEEFB7B9173FB72" box="[694,820,1203,1224]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361DEEFB7B9173FB72" box="[694,820,1203,1224]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
from other regions yield- ed no discrete characters suggesting that the former should represent a distinct species. Neither is the aedeagus of significantly different shape (males of 
<emphasis id="B97CEACEFFDB94361C31FB269645FAB9" box="[873,1026,1262,1283]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
from 
<collectingCountry id="F31F764CFFDB94361FEAFAC392BCFA9A" box="[178,251,1291,1312]" name="Cyprus" pageId="3" pageNumber="152">Cyprus</collectingCountry>
and 
<collectingCountry id="F31F764CFFDB94361E77FAC393C0FA9B" box="[303,391,1291,1313]" name="Lebanon" pageId="3" pageNumber="152">Lebanon</collectingCountry>
examined), nor are the specimens significantly larger than material of 
<taxonomicName id="4C084D5FFFDB94361F88FAE09317FA87" box="[208,336,1320,1341]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361F88FAE09317FA87" box="[208,336,1320,1341]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
seen, e.g., from 
<collectingCountry id="F31F764CFFDB94361D5AFAE0900BFA87" box="[514,588,1320,1341]" name="Greece" pageId="3" pageNumber="152">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361D01FAE090ECFA84" box="[601,683,1320,1342]" name="Georgia" pageId="3" pageNumber="152">Georgia</collectingCountry>
and 
<collectingCountry id="F31F764CFFDB94361DB8FAE0917CFA84" box="[736,827,1320,1342]" name="Armenia" pageId="3" pageNumber="152">Armenia</collectingCountry>
, even though they are at the upper end of the size range. Specimens examined from 
<collectingCountry id="F31F764CFFDB94361C42FA8E9124FAE1" box="[794,867,1350,1371]" name="Cyprus" pageId="3" pageNumber="152">Cyprus</collectingCountry>
and 
<collectingCountry id="F31F764CFFDB94361CCFFA8D91A8FAE1" box="[919,1007,1349,1371]" name="Lebanon" pageId="3" pageNumber="152">Lebanon</collectingCountry>
belong to the macropterous morph (hence the long elytra). I have seen macropterous and similarly dark-coloured specimens also from South 
<collectingCountry id="F31F764CFFDB94361DC1FA4890A4FA2F" box="[665,739,1408,1429]" name="Greece" pageId="3" pageNumber="152">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFDB94361DB6FA48910EFA2C" box="[750,841,1408,1430]" name="Armenia" pageId="3" pageNumber="152">Armenia</collectingCountry>
, and 
<collectingCountry id="F31F764CFFDB94361C27FA489193FA2C" box="[895,980,1408,1430]" name="Georgia" pageId="3" pageNumber="152">Georgia</collectingCountry>
. In consequence, both 
<emphasis id="B97CEACEFFDB94361E62FA569388FA09" box="[314,463,1438,1459]" italics="true" pageId="3" pageNumber="152">Q. cyprusensis</emphasis>
and its junior synonym 
<emphasis id="B97CEACEFFDB94361D9CFA569156FA09" box="[708,785,1438,1459]" italics="true" pageId="3" pageNumber="152">Q. freyi</emphasis>
are placed in synonymy with 
<taxonomicName id="4C084D5FFFDB94361FEAFA739376FA6A" box="[178,305,1467,1488]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDB94361FEAFA739376FA6A" box="[178,305,1467,1488]" italics="true" pageId="3" pageNumber="152">Q. umbrinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736DCFFDB94311FEAFA28901AFD89" blockId="3.[178,1086,1504,1587]" lastBlockId="4.[178,1085,219,563]" lastPageId="4" lastPageNumber="153" pageId="3" pageNumber="152">
The original description of 
<emphasis id="B97CEACEFFDB94361E8AFA289027FA4F" box="[466,608,1504,1525]" italics="true" pageId="3" pageNumber="152">Q. bulgaricus</emphasis>
is based on &quot; 
<specimenCount id="9D0EFD55FFDB94361DAAFA289161FA4F" box="[754,806,1504,1525]" pageId="3" pageNumber="152" type="male">2♂♂</specimenCount>
(
<specimenCount id="9D0EFD55FFDB94361C60FA2891C8FA4F" box="[824,911,1504,1525]" pageId="3" pageNumber="152" type="generic">1 Typus</specimenCount>
, 1 Cotypus) und 
<specimenCount id="9D0EFD55FFDB94361FEAFA3692A2F9AC" box="[178,229,1534,1558]" pageId="3" pageNumber="152" type="female">2♀♀</specimenCount>
(
<specimenCount id="9D0EFD55FFDB94361FA7FA36931BF9A9" box="[255,348,1534,1555]" pageId="3" pageNumber="152" type="generic">1 Typus</specimenCount>
, 1 Cotypus) vom Rosalito-polje, Schipka Balkan&quot; and &quot; 
<specimenCount id="9D0EFD55FFDB94361CB8FA369647F9AC" box="[992,1024,1534,1558]" pageId="3" pageNumber="152" type="female">1♀</specimenCount>
vom Jumruktchal, Zentral-Balkan... und 
<specimenCount id="9D0EFD55FFDB94361D47F9D39079F989" box="[543,574,1563,1587]" pageId="3" pageNumber="152" type="female">1♀</specimenCount>
von ebendort... (Cotypen)&quot; (
<bibRefCitation id="EF994B2DFFDB94361C30F9D39675F98B" author="SCHEERPELTZ O" box="[872,1074,1563,1585]" pageId="3" pageNumber="152" pagination="185 - 246" refId="ref21464" refString="SCHEERPELTZ O. (1937): Wissenschaftliche Ergebnisse einer von Herrn Hofrat F. Schubert, seinem Sohne Herrn cand. phil. F. Schubert und Herrn Prof. Ing. K. Mandl im Sommer 1935 (1936) nach Bulgarien unternommenen Studienreise. Coleoptera: I. Staphylinidae. - Mitteilungen aus den Konigliche n Naturwissenschaftlichen Instituten in Sofia - Bulgarien 10: 185 - 246." type="journal article" year="1937">SCHEERPELTZ 1937</bibRefCitation>
). Since a 
<typeStatus id="54B3887EFFDC94311E5FFF139318FF4B" box="[263,351,219,241]" pageId="4" pageNumber="153" type="holotype">holotype</typeStatus>
is not specified, all the type specimens have 
<typeStatus id="54B3887EFFDC94311C64FF1391CCFF4A" box="[828,907,219,240]" pageId="4" pageNumber="153" type="syntype">syntype</typeStatus>
status. The male labelled by Scheerpeltz as &quot;Typus&quot; is designated as the 
<typeStatus id="54B3887EFFDC94311DAEFF309111FEB4" box="[758,854,248,270]" pageId="4" pageNumber="153" type="lectotype">lectotype</typeStatus>
. According to the key provided by 
<bibRefCitation id="EF994B2DFFDC94311E1BFEDE9063FE96" author="SCHEERPELTZ O" box="[323,548,278,300]" pageId="4" pageNumber="153" pagination="185 - 246" refId="ref21464" refString="SCHEERPELTZ O. (1937): Wissenschaftliche Ergebnisse einer von Herrn Hofrat F. Schubert, seinem Sohne Herrn cand. phil. F. Schubert und Herrn Prof. Ing. K. Mandl im Sommer 1935 (1936) nach Bulgarien unternommenen Studienreise. Coleoptera: I. Staphylinidae. - Mitteilungen aus den Konigliche n Naturwissenschaftlichen Instituten in Sofia - Bulgarien 10: 185 - 246." type="journal article" year="1937">SCHEERPELTZ (1937)</bibRefCitation>
, 
<emphasis id="B97CEACEFFDC94311D6EFEDE908DFE90" box="[566,714,277,299]" italics="true" pageId="4" pageNumber="153">Q. bulgaricus</emphasis>
is mainly distinguished from 
<taxonomicName id="4C084D5FFFDC94311B7CFEDE9355FEF2" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311B7CFEDE9355FEF2" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
by the shape of the head, shorter hind wings, and the absence of a palisade fringe at the posterior margin of the abdominal tergite VII. An examination of the type specimens revealed, however, that the shape of the head, the length of the hind wings, and the shape of the aedeagus are within the range of intraspecific variation of 
<taxonomicName id="4C084D5FFFDC94311B7CFE439355FE07" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311B7CFE439355FE07" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
and that the palisade fringe at the posterior margin of tergite VII is indeed present. The median lobe of the aedeagus is of similar shape as that of material seen from 
<collectingCountry id="F31F764CFFDC94311FEAFE2B92A7FE43" box="[178,224,483,505]" name="Italy" pageId="4" pageNumber="153">Italy</collectingCountry>
and 
<collectingCountry id="F31F764CFFDC94311E44FE2B9336FE43" box="[284,369,483,505]" name="Georgia" pageId="4" pageNumber="153">Georgia</collectingCountry>
. Hence the synonymy proposed above. Externally, the populations from the Stara Planina are characterized by rather extensive yellowish coloration of the lateral and posterior margins of the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDC94311FEAFD8B9308FA64" pageId="4" pageNumber="153" type="reference_group">
<paragraph id="8BB736DCFFDC94311FEAFD8B9314FD51" blockId="4.[178,1085,579,747]" pageId="4" pageNumber="153">
<emphasis id="B97CEACEFFDC94311FEAFD8B9331FDE2" box="[178,374,579,600]" italics="true" pageId="4" pageNumber="153">
<taxonomicName id="4C084D5FFFDC94311FEAFD8B9341FDE2" ID-CoL="8VS2M" box="[178,262,579,600]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
maronitus
</emphasis>
was described from a unique male collected in &quot; 
<collectingCountry id="F31F764CFFDC94311C2EFD8B91F2FDE3" box="[886,949,579,601]" name="Lebanon" pageId="4" pageNumber="153">Liban</collectingCountry>
, Aïn Batara&quot; (
<bibRefCitation id="EF994B2DFFDC94311FE2FDA89310FDCC" author="COIFFAIT H" box="[186,343,608,630]" pageId="4" pageNumber="153" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT 1963</bibRefCitation>
). The 
<typeStatus id="54B3887EFFDC94311EC4FDA893B2FDCC" box="[412,501,608,630]" pageId="4" pageNumber="153" type="holotype">holotype</typeStatus>
is deposited in the Coiffait collection and consequently unavailable for taxonomic revision (see introduction). However, based on the descriptions and illustrations of the aedeagus provided by 
<bibRefCitation id="EF994B2DFFDC94311DE2FD53911CFD0B" author="COIFFAIT H" box="[698,859,667,689]" pageId="4" pageNumber="153" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT (1963</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFDC94311C3EFD5391E0FD0B" author="COIFFAIT H" box="[870,935,667,689]" pageId="4" pageNumber="153" refId="ref20382" refString="COIFFAIT H. (1978 a): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvell e Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">1978a</bibRefCitation>
), as well as on additional material from 
<collectingCountry id="F31F764CFFDC94311EE8FD70904CFD74" box="[432,523,696,718]" name="Lebanon" pageId="4" pageNumber="153">Lebanon</collectingCountry>
, the 
<typeStatus id="54B3887EFFDC94311D64FD7090D2FD74" box="[572,661,696,718]" pageId="4" pageNumber="153" type="holotype">holotype</typeStatus>
is undoubtedly a macropterous specimen of 
<taxonomicName id="4C084D5FFFDC94311F96FD1E930BFD50" ID-CoL="4R3BW" box="[206,332,725,747]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311F96FD1E930BFD50" box="[206,332,725,747]" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736DCFFDC94311FEAFD339088FC64" blockId="4.[178,1085,763,990]" pageId="4" pageNumber="153">
<emphasis id="B97CEACEFFDC94311FEAFD3393CEFCAA" box="[178,393,763,784]" italics="true" pageId="4" pageNumber="153">
<taxonomicName id="4C084D5FFFDC94311FEAFD339341FCAA" ID-CoL="8VS2M" box="[178,262,763,784]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
gueorguievi
</emphasis>
was described based on a male 
<typeStatus id="54B3887EFFDC94311DBAFD33917CFCAB" box="[738,827,763,785]" pageId="4" pageNumber="153" type="holotype">holotype</typeStatus>
and two 
<typeStatus id="54B3887EFFDC94311CFAFD339644FCAA" box="[930,1027,763,784]" pageId="4" pageNumber="153" type="paratype">paratypes</typeStatus>
from &quot;Grotte Porojnata Dupka, 
<collectingCountry id="F31F764CFFDC94311E94FCD09062FC94" box="[460,549,792,814]" name="Bulgaria" pageId="4" pageNumber="153">Bulgarie</collectingCountry>
&quot; (
<bibRefCitation id="EF994B2DFFDC94311D1AFCD090A4FC94" author="COIFFAIT H" box="[578,739,792,814]" pageId="4" pageNumber="153" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">COIFFAIT 1967</bibRefCitation>
). The type material is currently inaccessible for scientific study (see introduction). However, neither the illustrations of the aedeagus nor the description provided by 
<bibRefCitation id="EF994B2DFFDC94311DFAFC9B9109FCD3" author="COIFFAIT H" box="[674,846,851,873]" pageId="4" pageNumber="153" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">COIFFAIT (1967)</bibRefCitation>
provide any evidence suggesting that 
<emphasis id="B97CEACEFFDC94311E0FFCB893B4FC3C" box="[343,499,880,902]" italics="true" pageId="4" pageNumber="153">Q. gueorguievi</emphasis>
should be distinct from 
<taxonomicName id="4C084D5FFFDC94311DA1FCB8913BFC3F" ID-CoL="4R3BW" box="[761,892,880,901]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311DA1FCB8913BFC3F" box="[761,892,880,901]" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
. The shape of the median lobe of the aedeagus is identical to that of 
<taxonomicName id="4C084D5FFFDC94311DE9FC469168FC18" ID-CoL="4R3BW" box="[689,815,909,931]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311DE9FC469168FC18" box="[689,815,909,931]" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
and the shape and chaetotaxy of the paramere (figure 2L in 
<bibRefCitation id="EF994B2DFFDC94311D7AFC6390FBFC7B" author="COIFFAIT H" box="[546,700,939,961]" pageId="4" pageNumber="153" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">COIFFAIT 1967</bibRefCitation>
) are within the range of its intraspecific variation. Hence the synonymy proposed above.
</paragraph>
<paragraph id="8BB736DCFFDC94311FEAFC259308FA64" blockId="4.[178,1085,1004,1502]" pageId="4" pageNumber="153">
I n t r a - a n d i n t e r s p e c i f i c v a r i a t i o n: 
<taxonomicName id="4C084D5FFFDC94311C71FC2491AEFBBB" ID-CoL="4R3BW" box="[809,1001,1004,1025]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311C71FC2491AEFBBB" box="[809,1001,1004,1025]" italics="true" pageId="4" pageNumber="153">Quedius umbrinus</emphasis>
</taxonomicName>
is subject to enormous variation not only of external characters such as size, coloration, and the length of the hind wings and the elytra, but also of the shape of the aedeagus. The macropterous morph, which is usually of more robust habitus, darker coloration, and larger size, was observed in populations from South 
<collectingCountry id="F31F764CFFDC94311D87FB95916EFBC8" box="[735,809,1117,1138]" name="Greece" pageId="4" pageNumber="153">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFDC94311C6FFB9591C4FBC8" box="[823,899,1117,1138]" name="Cyprus" pageId="4" pageNumber="153">Cyprus</collectingCountry>
, the Middle East, and the Caucasus region (
<collectingCountry id="F31F764CFFDC94311E9EFBB0905BFB34" box="[454,540,1144,1166]" name="Georgia" pageId="4" pageNumber="153">Georgia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDC94311D72FBB090C0FB34" box="[554,647,1144,1166]" name="Armenia" pageId="4" pageNumber="153">Armenia</collectingCountry>
). In 
<collectingCountry id="F31F764CFFDC94311DE3FBB09151FB34" box="[699,790,1144,1166]" name="Armenia" pageId="4" pageNumber="153">Armenia</collectingCountry>
, it was repeatedly collected together with the brachypterous morph. The coloration of the body is mostly dark-brown to blackish-brown, but ranges from nearly uniformly reddish (some specimens from North 
<collectingCountry id="F31F764CFFDC94311FADFB059314FB58" box="[245,339,1229,1250]" name="Germany" pageId="4" pageNumber="153">Germany</collectingCountry>
seen) to partly or completely black. The elytra may be bicoloured (dark with more or less extensively paler margins and suture) or unicolorous, and the colour of the legs ranges from yellowish-red (usually with the meso- and metatibiae more or less distinctly infuscate) to blackish. The position of the subapical tooth of the median lobe of the aedeagus may be very close to the apex or more distant (with all transitions). The paramere varies in breadth and, to a lesser extent, also in length; it may be basally constricted or not. The apex of the paramere may be smoothly rounded or obtusely pointed; the sensory peg setae may be arranged in two distinct lateral rows or occupy nearly all of the apical portion of the paramere. Some of the aedeagal variation is illustrated by 
<bibRefCitation id="EF994B2DFFDC94311FEAFA01930CFA64" author="ASSING V" box="[178,331,1480,1502]" pageId="4" pageNumber="153" pagination="35 - 46" refId="ref19493" refString="ASSING V. (1999): Zur Kenntnis und Synonymie einiger mitteleuropaischer Arten der Gattung Quedius STEPHENS (Col., Staphylinidae). - Entomologische Blatter 95 (1): 35 - 46." type="journal article" year="1999">ASSING (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDC94301FEAFA2590E0FE91" lastPageId="5" lastPageNumber="154" pageId="4" pageNumber="153" type="description">
<paragraph id="8BB736DCFFDC94301FEAFA2590E0FE91" blockId="4.[178,1085,1517,1598]" lastBlockId="5.[178,1085,219,299]" lastPageId="5" lastPageNumber="154" pageId="4" pageNumber="153">
<emphasis id="B97CEACEFFDC94311FEAFA259318F9B8" box="[178,351,1517,1538]" italics="true" pageId="4" pageNumber="153">
<taxonomicName id="4C084D5FFFDC94311FEAFA259341F9B8" box="[178,262,1517,1538]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
sigwalti
</emphasis>
COIFFAIT, 1972, an endemic of 
<collectingRegion id="49CCF83EFFDC94311DF7FA2690A1F9B9" box="[687,742,1518,1539]" country="Greece" name="Kriti" pageId="4" pageNumber="153">Crete</collectingRegion>
and probably the closest relative of 
<taxonomicName id="4C084D5FFFDC94311F97F9C39308F99A" box="[207,335,1547,1568]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311F97F9C39308F99A" box="[207,335,1547,1568]" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
, resembles 
<taxonomicName id="4C084D5FFFDC94311E93F9C3900BF99A" box="[459,588,1547,1568]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDC94311E93F9C3900BF99A" box="[459,588,1547,1568]" italics="true" pageId="4" pageNumber="153">Q. umbrinus</emphasis>
</taxonomicName>
both in external and in sexual characters, but is distinguished by a much broader paramere. Since transitional conditions have not been observed, this difference is interpreted as interspecific variation. It seems likely that 
<collectingRegion id="49CCF83EFFDD94301FEAFF3092AEFEB7" box="[178,233,248,269]" country="Greece" name="Kriti" pageId="5" pageNumber="154">Crete</collectingRegion>
was colonized by the ancestors of 
<emphasis id="B97CEACEFFDD94301D07FF309096FEB7" box="[607,721,248,269]" italics="true" pageId="5" pageNumber="154">Q. sigwalti</emphasis>
millions of years ago and that the Cretan populations have been isolated ever since.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDD94301FEAFEF390E9FE7C" pageId="5" pageNumber="154" type="materials_examined">
<paragraph id="8BB736DCFFDD94301FEAFEF390E9FE7C" blockId="5.[178,1085,315,454]" pageId="5" pageNumber="154">
<taxonomicName id="4C084D5FFFDD94301FEAFEF39329FEEA" box="[178,366,315,336]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDD94301FEAFEF39329FEEA" box="[178,366,315,336]" italics="true" pageId="5" pageNumber="154">Quedius umbrinus</emphasis>
</taxonomicName>
is widespread and common in practically all of Europe (eastwards to the Caucasus region), 
<collectingCountry id="F31F764CFFDD94301EF9FE9093AAFED4" box="[417,493,344,366]" name="Turkey" pageId="5" pageNumber="154">Turkey</collectingCountry>
, and the Middle East (material from 
<collectingCountry id="F31F764CFFDD94301CDAFE91918DFED4" box="[898,970,345,366]" name="France" pageId="5" pageNumber="154">France</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301C80FE90967EFED7" box="[984,1081,344,365]" name="Germany" pageId="5" pageNumber="154">Germany</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301FEAFEBD934AFE31" box="[178,269,373,395]" name="Slovakia" pageId="5" pageNumber="154">Slovakia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301E42FEBD9320FE31" box="[282,359,373,395]" name="Austria" pageId="5" pageNumber="154">Austria</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301E2BFEBD93E3FE31" box="[371,420,373,395]" name="Italy" pageId="5" pageNumber="154">Italy</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301EE8FEBD904CFE31" box="[432,523,373,395]" name="Slovenia" pageId="5" pageNumber="154">Slovenia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301D40FEBD9031FE31" box="[536,630,373,395]" name="Romania" pageId="5" pageNumber="154">Romania</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301DDAFEBD9082FE31" box="[642,709,373,395]" name="Serbia" pageId="5" pageNumber="154">Serbia</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301D8AFEBD9162FE31" box="[722,805,373,395]" name="Albania" pageId="5" pageNumber="154">Albania</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301C69FEBD91CDFE31" box="[817,906,373,395]" name="Bulgaria" pageId="5" pageNumber="154">Bulgaria</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301CCFFEBE91A6FE31" box="[919,993,374,395]" name="Greece" pageId="5" pageNumber="154">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301CB5FEBE967EFE31" box="[1005,1081,374,395]" name="Cyprus" pageId="5" pageNumber="154">Cyprus</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301FEAFE5B92B9FE13" box="[178,254,403,425]" name="Turkey" pageId="5" pageNumber="154">Turkey</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301E53FE5B9321FE13" box="[267,358,403,425]" name="Lebanon" pageId="5" pageNumber="154">Lebanon</collectingCountry>
, 
<collectingCountry id="F31F764CFFDD94301E2BFE5B93F1FE13" box="[371,438,403,425]" name="Russia" pageId="5" pageNumber="154">Russia</collectingCountry>
(West Caucasus), and 
<collectingCountry id="F31F764CFFDD94301DF3FE5B9143FE13" box="[683,772,403,425]" name="Armenia" pageId="5" pageNumber="154">Armenia</collectingCountry>
examined). The species is an epigeic and eurytopic inhabitant of moist habitats.
</paragraph>
</subSubSection>
</treatment>
</document>