564 lines
111 KiB
XML
564 lines
111 KiB
XML
<document id="2F1D5399C19C06D9D85938A39C32DDB1" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Perkovsky, Evgeny" docDate="2016" docId="03A187A92930FFFFFF03C1359BEDF992" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rovnoleptochromus Jałoszyński & Perkovsky, 2016, gen. n." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="10" updateTime="1698382676775" updateUser="plazi">
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<mods:mods id="B7C7EFB6CD82F92A6EFDF34DC17932DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CFCE1DFD4008E7F6FCF9803201ABFBC4">
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<mods:title id="D70FBF24C537F883340C74D2D7C4224E">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:name id="29CDF5F622A0583153EE24DFC105ADB7" type="personal">
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<mods:namePart id="3A56E3A96B434C87BC67E3C762D7BDCB">Jałoszyński, Paweł</mods:namePart>
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<mods:roleTerm id="2E4A065ED3B690E5F580395B226A4EFC">Author</mods:roleTerm>
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<mods:namePart id="841D44098E178E75BB8E3B21BB29CC58">Perkovsky, Evgeny</mods:namePart>
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<mods:typeOfResource id="8781D4186B6B01F44FC91FB5D0B1FF8D">text</mods:typeOfResource>
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<mods:title id="79761E9B806D00B32DEDEB3AEB94DE2F">Zootaxa</mods:title>
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<mods:part id="17FF61ABC3FAEDA193740A1DC7300688">
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<mods:date id="2D9FD9241C2FACD415337C0982DEEE90">2016</mods:date>
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<mods:number id="5B4B5EC478720E604D33FACB530D80B0">4157</mods:number>
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<mods:detail id="3A18BE179AB09E4404863016C7BAC8EA" type="issue">
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<mods:number id="1291BBAE1550494A2ED128F96B64E5D6">1</mods:number>
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<mods:classification id="80A41D71773DEF09AD764A8D24E4674C">journal article</mods:classification>
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<mods:identifier id="6F6750628BB0A367AF077F122D0CD8E6" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<mods:identifier id="4F3B037BA8B4F6C4092A468F64963E41" type="GBIF-Dataset">2389f078-1812-4b4e-826a-561cd7e46654</mods:identifier>
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<mods:identifier id="18108BEECEE65B8288AAB9BCC2909332" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A187A92930FFFFFF03C1359BEDF992" ID-DOI="http://doi.org/10.5281/zenodo.5681731" ID-GBIF-Taxon="124407836" ID-Zenodo-Dep="5681731" LSID="urn:lsid:plazi:treatment:03A187A92930FFFFFF03C1359BEDF992" httpUri="http://treatment.plazi.org/id/03A187A92930FFFFFF03C1359BEDF992" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10">
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<subSubSection id="C31265342930FFF8FF03C1359CA6FBE3" box="[151,498,1047,1074]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BB736BF2930FFF8FF03C1359CA6FBE3" blockId="9.[151,498,1047,1074]" box="[151,498,1047,1074]" pageId="9" pageNumber="10">
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<heading id="D0FF81D32930FFF8FF03C1359CA6FBE3" bold="true" box="[151,498,1047,1074]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<emphasis id="B97CEAAD2930FFF8FF03C1359CA6FBE3" bold="true" box="[151,498,1047,1074]" pageId="9" pageNumber="10">
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†
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<taxonomicName id="4C084D3C2930FFF8FF31C13A9CC3FBE0" ID-CoL="8XPPJ" box="[165,407,1047,1073]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97CEAAD2930FFF8FF31C13A9CC3FBE0" bold="true" box="[165,407,1047,1073]" italics="true" pageId="9" pageNumber="10">Rovnoleptochromus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24F57D62930FFF8FE0BC1359CA6FBE3" box="[415,498,1048,1074]" pageId="9" pageNumber="10" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31265342930FFF8FF03C1729E14FBA5" box="[151,832,1118,1140]" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BB736BF2930FFF8FF03C1729E14FBA5" blockId="9.[151,832,1118,1140]" box="[151,832,1118,1140]" pageId="9" pageNumber="10">
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Type species:
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<taxonomicName id="4C084D3C2930FFF8FEB2C1739FD0FBA5" box="[294,644,1118,1140]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ableptonoides">
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<emphasis id="B97CEAAD2930FFF8FEB2C1739FD0FBA5" box="[294,644,1118,1140]" italics="true" pageId="9" pageNumber="10">Rovnoleptochromus ableptonoides</emphasis>
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</taxonomicName>
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(here designated).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31265342930FFF8FF03C18C9C0FFA43" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="8BB736BF2930FFF8FF03C18C9C0FFA43" blockId="9.[151,1437,1185,2002]" pageId="9" pageNumber="10">
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<emphasis id="B97CEAAD2930FFF8FF03C18C9C45FB6B" bold="true" box="[151,273,1185,1210]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
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A genus of Clidicini with postgenae bearing long bristles but lacking cuticular postgenal processes; vertex lacking posterolateral bristles; maxillary palpomere II lacking cuticular projection but bearing one long bristle; maxillary palpomere
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<date id="FFB6107F2930FFF8FE7AC1C79F72FAD3" box="[494,550,1258,1282]" pageId="9" pageNumber="10">III 3</date>
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× as long as broad, with strongly oblique apex; maxillary palpomere IV subtriangular, broadest at base; pedicel distinctly longer than antennomere III; mesoventral intercoxal process narrowly subtriangular and pointed posteriorly; mesocoxal cavities carinate posteriorly; metaventrite with median longitudinal carina; abdominal sternite III (i.e., the first visible sternite) with broadly inversely V-shaped anterior transverse carina.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31265342930FFFCFF53C0B09CBDFF04" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BB736BF2930FFF8FF53C0B099FAFA08" blockId="9.[151,1437,1185,2002]" pageId="9" pageNumber="10">
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<emphasis id="B97CEAAD2930FFF8FF53C0B09C0DFA67" bold="true" box="[199,345,1437,1462]" pageId="9" pageNumber="10">Description.</emphasis>
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Body (
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<figureCitation id="13332A3A2930FFF8FE2DC0B09F15FA67" box="[441,577,1437,1462]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="9" pageNumber="10">Figs 39–42</figureCitation>
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,
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<figureCitation id="13332A3A2930FFF8FDC0C0B09FC8FA67" box="[596,668,1437,1462]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="9" pageNumber="10">44–45</figureCitation>
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,
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<figureCitation id="13332A3A2930FFF8FD3BC0B39FA2FA67" box="[687,758,1438,1462]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">47–48</figureCitation>
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) strongly elongate and flattened, with distinctly marked constrictions between head and pronotum and between pronotum and elytra, BL about
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<quantity id="4CF09B5A2930FFF8FBF1C0EF99FDFA08" box="[1125,1193,1474,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="2.0">2 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BB736BF2930FFF8FF53C0C89841F800" blockId="9.[151,1437,1185,2002]" pageId="9" pageNumber="10">
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Head capsule (
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<figureCitation id="13332A3A2930FFF8FEE5C0C89CA3FA2F" box="[369,503,1509,1534]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="9" pageNumber="10">Figs 39, 43</figureCitation>
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,
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<figureCitation id="13332A3A2930FFF8FD92C0C89F2DFA2F" box="[518,633,1509,1534]" captionStart-0="FIGURES 44 – 45" captionStart-1="FIGURES 46 – 48" captionStartId-0="12.[151,264,1157,1180]" captionStartId-1="15.[151,264,1833,1856]" captionTargetBox-0="[156,1428,201,1132]" captionTargetBox-1="[159,1435,886,1804]" captionTargetId-0="figure@12.[151,1436,193,1137]" captionTargetId-1="figure@15.[151,1436,876,1813]" captionTargetPageId-0="12" captionTargetPageId-1="15" captionText-0="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." captionText-1="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri-0="https://zenodo.org/record/262418/files/figure.png" httpUri-1="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">44–46, 48</figureCitation>
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) with anterior part (in front of occipital constriction) transverse; eyes located anteriorly, large, strongly convex, oval and finely faceted; occipital constriction much narrower than vertex just behind eyes; vertex lacking posterolateral bristles; mouthparts only partly visible, labrum (
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<figureCitation id="13332A3A2930FFF8FA93C3009834F997" box="[1287,1376,1581,1606]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 48</figureCitation>
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;
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<emphasis id="B97CEAAD2930FFF8FAE6C30098C7F997" box="[1394,1427,1581,1606]" italics="true" pageId="9" pageNumber="10">lbr</emphasis>
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) transverse and anteriorly emarginated; mandibles (
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<figureCitation id="13332A3A2930FFF8FD7CC37C9E15F9BB" box="[744,833,1617,1642]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
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;
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<emphasis id="B97CEAAD2930FFF8FCC6C37C9E20F9BB" box="[850,884,1617,1642]" italics="true" pageId="9" pageNumber="10">md</emphasis>
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) each with at least two preapical mesal teeth. Maxillary palps longer than head and slender, each composed of small, elongate palpomere I (
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<figureCitation id="13332A3A2930FFF8FB7CC358986BF95F" box="[1256,1343,1653,1678]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
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;
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<emphasis id="B97CEAAD2930FFF8FADAC35B98DFF95F" box="[1358,1419,1654,1678]" italics="true" pageId="9" pageNumber="10">mxp1</emphasis>
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); evenly curved palpomere II (
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<figureCitation id="13332A3A2930FFF8FE7FC3B49F16F963" box="[491,578,1689,1714]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
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;
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<emphasis id="B97CEAAD2930FFF8FDC6C3B79FDAF963" box="[594,654,1690,1714]" italics="true" pageId="9" pageNumber="10">mxp2</emphasis>
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) which is about 4.5 × as long as broad, gradually and moderately broadening distally, bearing one long bristle in proximal third of dorsal margin; strongly elongate palpomere III (
|
||
<figureCitation id="13332A3A2930FFF8FF0BC3CC9DA0F92B" box="[159,244,1761,1786]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FE95C3CF9C69F92B" box="[257,317,1762,1786]" italics="true" pageId="9" pageNumber="10">mxp3</emphasis>
|
||
) 3 × as long as broad, strongly and gradually broadening distally and with strongly oblique apical margin; and short, subtriangular palpomere IV (
|
||
<figureCitation id="13332A3A2930FFF8FD26C2289E52F8CF" box="[690,774,1797,1822]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FC86C22B9E1BF8CF" box="[786,847,1798,1822]" italics="true" pageId="9" pageNumber="10">mxp4</emphasis>
|
||
; the shape of palpomeres III and IV is visible in the left palp, which is shown with palpomeres III and IV in strictly ventral view), about twice as long as broad, broadest at base. Constriction between maxillary palpomeres III and IV weakly marked. Labial palps (
|
||
<figureCitation id="13332A3A2930FFF8FA88C2609825F8B7" box="[1308,1393,1869,1894]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FAE9C26098C7F8B7" box="[1405,1427,1869,1894]" italics="true" pageId="9" pageNumber="10">lp</emphasis>
|
||
) with large and elongate palpomere II and small, elongate and pointed palpomere III. Posterior tentorial pits (
|
||
<figureCitation id="13332A3A2930FFF8FAFBC25C9DE2F87F" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FF50C2BB9DBCF87C" box="[196,232,1942,1965]" italics="true" pageId="9" pageNumber="10">ptp</emphasis>
|
||
) elongate and well-visible, each connected with outer margin of postcardinal region by hypomeral ridge (
|
||
<figureCitation id="13332A3A2930FFF8FF0BC2949DA7F803" box="[159,243,1977,2002]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FF6AC2949C43F803" box="[254,279,1977,2002]" italics="true" pageId="9" pageNumber="10">hr</emphasis>
|
||
). Each postgena with two long postgenal bristles (
|
||
<figureCitation id="13332A3A2930FFF8FCD9C2949EF4F803" box="[845,928,1977,2002]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="9" pageNumber="10">Fig. 46</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2930FFF8FC38C2949E82F803" box="[940,982,1977,2002]" italics="true" pageId="9" pageNumber="10">pgb</emphasis>
|
||
), lacking postgenal process.
|
||
</paragraph>
|
||
<caption id="DF7766372933FFFBFF03C2B99E31F818" httpUri="https://zenodo.org/record/262416/files/figure.png" pageId="10" pageNumber="11" targetBox="[181,1396,193,1909]" targetPageId="10">
|
||
<paragraph id="8BB736BF2933FFFBFF03C2B99E31F818" blockId="10.[151,1436,1940,1993]" pageId="10" pageNumber="11">
|
||
<emphasis id="B97CEAAD2933FFFBFF03C2B99C03F87B" bold="true" box="[151,343,1940,1963]" pageId="10" pageNumber="11">FIGURES 35–38.</emphasis>
|
||
Location of the Rovno Region (35−36) and the Klesov, Dubrovitsa and Vyrka amber mines (37) in the present-day Europe; location of the Rovno Region during Eocene (38).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7766372932FFFAFF03C3409983F973" httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,193,1623]" targetPageId="11">
|
||
<paragraph id="8BB736BF2932FFFAFF03C3409983F973" blockId="11.[151,1436,1645,1698]" pageId="11" pageNumber="12">
|
||
<emphasis id="B97CEAAD2932FFFAFF03C3409C00F952" bold="true" box="[151,340,1645,1668]" pageId="11" pageNumber="12">FIGURES 39–43.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C084D3C2932FFFAFE4DC3439E63F955" box="[473,823,1646,1668]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ableptonoides" status="sp. nov.">
|
||
<emphasis id="B97CEAAD2932FFFAFE4DC3439E63F955" box="[473,823,1646,1668]" italics="true" pageId="11" pageNumber="12">Rovnoleptochromus ableptonoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24F57D62932FFFAFCA9C3439E22F952" box="[829,886,1646,1667]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B97CEAAD2932FFFAFCA9C3439E22F952" bold="true" box="[829,886,1646,1667]" pageId="11" pageNumber="12">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(K-27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB736BF2932FFFAFF53C3E79F1EF8A2" blockId="11.[151,1436,1738,2014]" pageId="11" pageNumber="12">
|
||
Antennae (
|
||
<figureCitation id="13332A3A2932FFFAFED5C3E79CBFF933" box="[321,491,1738,1763]" captionStart-0="FIGURES 39 – 43" captionStart-1="FIGURES 44 – 45" captionStart-2="FIGURES 46 – 48" captionStartId-0="11.[151,264,1645,1668]" captionStartId-1="12.[151,264,1157,1180]" captionStartId-2="15.[151,264,1833,1856]" captionTargetBox-0="[151,1436,193,1623]" captionTargetBox-1="[156,1428,201,1132]" captionTargetBox-2="[159,1435,886,1804]" captionTargetId-0="figure@11.[151,1436,193,1624]" captionTargetId-1="figure@12.[151,1436,193,1137]" captionTargetId-2="figure@15.[151,1436,876,1813]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="15" captionText-0="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." captionText-1="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." captionText-2="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri-0="https://zenodo.org/record/262417/files/figure.png" httpUri-1="https://zenodo.org/record/262418/files/figure.png" httpUri-2="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Figs 39–46, 48</figureCitation>
|
||
) long and slender, slightly shorter than half BL, scape (
|
||
<figureCitation id="13332A3A2932FFFAFBC9C3E799E4F933" box="[1117,1200,1738,1763]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2932FFFAFB28C3E799B3F933" box="[1212,1255,1738,1762]" italics="true" pageId="11" pageNumber="12">an1</emphasis>
|
||
) about 5–6 × as long as broad, nearly straight, distinctly but weakly broadening distally, with deep apical notch; pedicel (
|
||
<figureCitation id="13332A3A2932FFFAFAD7C3C098C3F8D7" box="[1347,1431,1773,1798]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2932FFFAFF03C23F9D94F8FB" box="[151,192,1810,1834]" italics="true" pageId="11" pageNumber="12">an2</emphasis>
|
||
) much narrower and shorter than scape, slightly broadened distally, distinctly longer than antennomere III; flagellomeres (i.e., antennomeres III–XI) gradually broadening, each except XI broadest distally to middle, antennomere XI suboval, symmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2932FFFAFF53C25299BEF80F" blockId="11.[151,1436,1738,2014]" pageId="11" pageNumber="12">
|
||
Pronotum (
|
||
<figureCitation id="13332A3A2932FFFAFED1C2509CA7F847" box="[325,499,1917,1942]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="11" pageNumber="12">Figs 39–41, 43</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2932FFFAFE6BC2509F13F847" box="[511,583,1917,1942]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">46, 47</figureCitation>
|
||
) in dorsal view (
|
||
<figureCitation id="13332A3A2932FFFAFC9CC2509E0FF847" box="[776,859,1917,1942]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Fig. 47</figureCitation>
|
||
) strongly elongate and broadest near anterior fifth, convex, with short posterior pronotal collar (
|
||
<figureCitation id="13332A3A2932FFFAFD0FC28F9FA5F86B" box="[667,753,1954,1979]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Fig. 47</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2932FFFAFC94C28E9E7CF86B" box="[768,808,1955,1978]" italics="true" pageId="11" pageNumber="12">pnc</emphasis>
|
||
) distinctly demarcated from disc by transverse row of several (at least four, poorly visible in specimen) large antebasal pits or impressions (
|
||
<figureCitation id="13332A3A2932FFFAFBC5C2E899F0F80F" box="[1105,1188,1989,2014]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="11" pageNumber="12">Fig. 47</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2932FFFAFB24C2E8998EF80F" box="[1200,1242,1989,2014]" italics="true" pageId="11" pageNumber="12">abp</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7766372935FFFDFF03C1A89F20FB08" httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13" targetBox="[156,1428,201,1132]" targetPageId="12">
|
||
<paragraph id="8BB736BF2935FFFDFF03C1A89F20FB08" blockId="12.[151,1437,1157,1241]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97CEAAD2935FFFDFF03C1A89C02FB4A" bold="true" box="[151,342,1157,1180]" pageId="12" pageNumber="13">FIGURES 44–45.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C084D3C2935FFFDFE77C1AB9E17FB4D" box="[483,835,1158,1180]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ableptonoides" status="sp. nov.">
|
||
<emphasis id="B97CEAAD2935FFFDFE77C1AB9E17FB4D" box="[483,835,1158,1180]" italics="true" pageId="12" pageNumber="13">Rovnoleptochromus ableptonoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24F57D62935FFFDFCD8C1AA9ED3FB4D" box="[844,903,1159,1180]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B97CEAAD2935FFFDFCD8C1AA9ED3FB4D" bold="true" box="[844,903,1159,1180]" pageId="12" pageNumber="13">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(K-27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest3, metanepisternum; epm3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB736BF2935FFFDFF53C02C980CFAEC" blockId="12.[151,1437,1281,2026]" pageId="12" pageNumber="13">
|
||
Ventral structures of prothorax (
|
||
<figureCitation id="13332A3A2935FFFDFDA7C02C9FD8FAC8" box="[563,652,1281,1306]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="12" pageNumber="13">Figs 42</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFD0DC02C9FE2FAC8" box="[665,694,1281,1305]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">48</figureCitation>
|
||
) poorly visible; basisternal part of prosternum seems moderately long, procoxae contiguous; shape of mesal margins of prothoracic hypomera indicate open procoxal sockets.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C0679985FAB3" blockId="12.[151,1437,1281,2026]" box="[199,1233,1353,1378]" pageId="12" pageNumber="13">
|
||
Mesonotum only partly and poorly visible (
|
||
<figureCitation id="13332A3A2935FFFDFD3AC0649E57FAB0" box="[686,771,1353,1378]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 47</figureCitation>
|
||
); mesoscutellum (
|
||
<figureCitation id="13332A3A2935FFFDFC5AC0649975FAB0" box="[974,1057,1353,1378]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 47</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFBB9C064990EFAB3" box="[1069,1114,1353,1378]" italics="true" pageId="12" pageNumber="13">scl3</emphasis>
|
||
) elongate.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C0419E05FA78" blockId="12.[151,1437,1281,2026]" pageId="12" pageNumber="13">
|
||
Mesoventrite (
|
||
<figureCitation id="13332A3A2935FFFDFEF8C0419CEBFA54" box="[364,447,1388,1413]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
) with carinate, very narrow and subtriangular mesoventral intercoxal process (
|
||
<figureCitation id="13332A3A2935FFFDFAD6C04198C3FA54" box="[1346,1431,1388,1413]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFF03C0BF9D85FA78" box="[151,209,1426,1449]" italics="true" pageId="12" pageNumber="13">msvp</emphasis>
|
||
), distinctly narrowing posteriorly and with pointed apex.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C0999E2AF977" blockId="12.[151,1437,1281,2026]" pageId="12" pageNumber="13">
|
||
Metaventrite (
|
||
<figureCitation id="13332A3A2935FFFDFEF2C0999CEBFA1C" box="[358,447,1460,1485]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="12" pageNumber="13">Figs 42</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFE5FC0999CBDFA1C" box="[459,489,1460,1485]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">45</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFE62C0989F47FA1C" box="[502,531,1461,1485]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">48</figureCitation>
|
||
) subrectangular with sides rounded and in posterior fourth narrowing; posterior margins of mesocoxal cavities with carinae (
|
||
<figureCitation id="13332A3A2935FFFDFD3AC0F49E53FA20" box="[686,775,1497,1522]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFC8DC0F79E00FA20" box="[793,852,1498,1521]" italics="true" pageId="12" pageNumber="13">pcmc</emphasis>
|
||
); posterior margin of ventrite deeply concave in admetacoxal regions and forming very short and broad metaventral intercoxal process (
|
||
<figureCitation id="13332A3A2935FFFDFB1BC0D199B1F9C4" box="[1167,1253,1532,1557]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFB60C0D3987EF9C4" box="[1268,1322,1534,1557]" italics="true" pageId="12" pageNumber="13">mtvp</emphasis>
|
||
) between metacoxae, with weakly concave posterior margin. Metaventrite with narrow but distinct median longitudinal carina (
|
||
<figureCitation id="13332A3A2935FFFDFF67C3689C1AF98C" box="[243,334,1605,1630]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFEF6C36B9CDBF98C" box="[354,399,1606,1629]" italics="true" pageId="12" pageNumber="13">mvc</emphasis>
|
||
) extending from mesoventral intercoxal process to near posterior margin of ventrite. Metanepisterna (
|
||
<figureCitation id="13332A3A2935FFFDFEC1C3449CFEF953" box="[341,426,1641,1666]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">Fig. 45</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFE23C3449CA6F950" box="[439,498,1641,1665]" italics="true" pageId="12" pageNumber="13">aest3</emphasis>
|
||
) broadening posteriorly, partly visible in ventral view (
|
||
<figureCitation id="13332A3A2935FFFDFBF9C3449997F950" box="[1133,1219,1641,1666]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
); metepimera (
|
||
<figureCitation id="13332A3A2935FFFDFAFAC3449DE2F977" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">Fig. 45</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFF56C3A09DABF974" box="[194,255,1677,1701]" italics="true" pageId="12" pageNumber="13">epm3</emphasis>
|
||
) narrow, nearly parallel-sided and visible in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C39C9911F8C3" blockId="12.[151,1437,1281,2026]" pageId="12" pageNumber="13">
|
||
Elytra (
|
||
<figureCitation id="13332A3A2935FFFDFE89C39C9CF5F918" box="[285,417,1713,1738]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="12" pageNumber="13">Figs 39–41</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFE3AC39C9CA3F91B" box="[430,503,1713,1738]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">44–45</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFD91C39C9F76F918" box="[517,546,1713,1737]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">47</figureCitation>
|
||
) strongly elongate and flattened; humeral calli weakly marked, lateral elytral margins nearly parallel in anterior half; elytral apices rounded; each elytron with five impressed rows of large pits visible in dorsal view (
|
||
<figureCitation id="13332A3A2935FFFDFE03C3D49CBEF8C0" box="[407,490,1785,1810]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 47</figureCitation>
|
||
), striae reducing in depth in posterior fourth of elytra.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C2319EECF8E4" blockId="12.[151,1437,1281,2026]" box="[199,952,1820,1845]" pageId="12" pageNumber="13">
|
||
Hind wings (
|
||
<figureCitation id="13332A3A2935FFFDFECCC2319CF8F8E4" box="[344,428,1820,1845]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">Fig. 45</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFE23C2319C8CF8E4" box="[439,472,1820,1845]" italics="true" pageId="12" pageNumber="13">hw</emphasis>
|
||
) present and protruding from under elytra.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFDFF53C26F99B9F817" blockId="12.[151,1437,1281,2026]" pageId="12" pageNumber="13">
|
||
Legs (
|
||
<figureCitation id="13332A3A2935FFFDFE9AC26C9CE8F888" box="[270,444,1857,1882]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1645,1668]" captionTargetBox="[151,1436,193,1623]" captionTargetId="figure@11.[151,1436,193,1624]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (39), lateral (40), dorsal (41) and ventral (42) view of whole specimen; anterior portion of body showing head in ventrolateral view (43)." httpUri="https://zenodo.org/record/262417/files/figure.png" pageId="12" pageNumber="13">Figs 39–40, 42</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFE5EC26C9F46F88B" box="[458,530,1857,1882]" captionStart="FIGURES 44 – 45" captionStartId="12.[151,264,1157,1180]" captionTargetBox="[156,1428,201,1132]" captionTargetId="figure@12.[151,1436,193,1137]" captionTargetPageId="12" captionText="FIGURES 44 – 45. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Laterodorsal (44) and lateral 45) view. Abbreviations: aest 3, metanepisternum; epm 3, metepimeron; hw, hind wing; mcp, mesocoxal projection; pcxb, procoxal bristles; pgb; postgenal bristles; pre, prepectus." httpUri="https://zenodo.org/record/262418/files/figure.png" pageId="12" pageNumber="13">44–45</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2935FFFDFDB4C26C9F69F888" box="[544,573,1857,1881]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">48</figureCitation>
|
||
) long and slender; pro- and mesocoxae subconical, metacoxae with weakly transverse proximal portion and subconical distal part; all trochanters short and subtriangular, protrochanter with weakly visible distal bristles; femora gradually clavate, profemora lacking ventral bristles; tibiae long and slender, weakly curved or nearly straight; tarsi slender and long, all tarsomeres elongate and subcylindrical.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2935FFFCFF53C2FC9CBDFF04" blockId="12.[151,1437,1281,2026]" lastBlockId="13.[151,1437,151,1941]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
Abdomen (
|
||
<figureCitation id="13332A3A2935FFFDFED0C2FC9CC1F838" box="[324,405,2001,2026]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
) covered by elytra, six sternites visible in ventral view (
|
||
<figureCitation id="13332A3A2935FFFDFB92C2FC990CF838" box="[1030,1112,2001,2026]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="12" pageNumber="13">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2935FFFDFBF0C2FC99F4F838" box="[1124,1184,2001,2025]" italics="true" pageId="12" pageNumber="13">st3–8</emphasis>
|
||
); sternite III distinctly longer than IV, with broadly and inversely V-shaped anterior transverse carina (
|
||
<figureCitation id="13332A3A2934FFFCFBD9C5BA99F2FF61" box="[1101,1190,151,176]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="13" pageNumber="14">Fig. 48</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2934FFFCFB23C5B4999BFF61" box="[1207,1231,153,176]" italics="true" pageId="13" pageNumber="14">vc</emphasis>
|
||
); suture between sternites VII and VIII distinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31265342934FFFCFF53C5F29DADFE91" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BB736BF2934FFFCFF53C5F29DADFE91" blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97CEAAD2934FFFCFF53C5F29F62FF29" bold="true" box="[199,566,223,248]" pageId="13" pageNumber="14">Distribution and composition.</emphasis>
|
||
<taxonomicName id="4C084D3C2934FFFCFDD5C5F29E74FF29" box="[577,800,223,248]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFDD5C5F29E74FF29" box="[577,800,223,248]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
is represented by only one species,
|
||
<taxonomicName id="4C084D3C2934FFFCFB40C5CC98C1FF29" box="[1236,1429,223,248]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ableptonoides">
|
||
<emphasis id="B97CEAAD2934FFFCFB40C5CC98C1FF29" box="[1236,1429,223,248]" italics="true" pageId="13" pageNumber="14">R. ableptonoides</emphasis>
|
||
</taxonomicName>
|
||
, described below and known from the Late Eocene of Europe (
|
||
<figureCitation id="13332A3A2934FFFCFC1FC4299EB0FECD" box="[907,996,260,285]" captionStart="FIGURES 35 – 38" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[181,1396,193,1909]" captionTargetId="figure@10.[181,1396,193,1910]" captionTargetPageId="10" captionText="FIGURES 35 – 38. Location of the Rovno Region (35 − 36) and the Klesov, Dubrovitsa and Vyrka amber mines (37) in the present-day Europe; location of the Rovno Region during Eocene (38)." httpUri="https://zenodo.org/record/262416/files/figure.png" pageId="13" pageNumber="14">Fig. 38</figureCitation>
|
||
), area currently within north-western
|
||
<collectingCountry id="F31F762F2934FFFCFF03C40A9DA1FE91" box="[151,245,295,320]" name="Ukraine" pageId="13" pageNumber="14">Ukraine</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31265342934FFFCFF53C4619C96FE59" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BB736BF2934FFFCFF53C4619C96FE59" blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97CEAAD2934FFFCFF53C4619C1AFEB4" bold="true" box="[199,334,332,357]" pageId="13" pageNumber="14">Etymology.</emphasis>
|
||
<taxonomicName id="4C084D3C2934FFFCFECDC4619F6CFEB4" box="[345,568,332,357]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFECDC4619F6CFEB4" box="[345,568,332,357]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
is derived from the genus name
|
||
<taxonomicName id="4C084D3C2934FFFCFC5DC461993CFEB4" box="[969,1128,332,357]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC5DC461993CFEB4" box="[969,1128,332,357]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
and the geographic name
|
||
<collectingRegion id="49CCF85D2934FFFCFF03C45D9DB1FE56" box="[151,229,368,391]" country="Ukraine" name="Rivne" pageId="13" pageNumber="14">Rovno</collectingRegion>
|
||
. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31265342934FFFCFF53C4B99DA5F844" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="8BB736BF2934FFFCFF53C4B99E73FCC4" blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97CEAAD2934FFFCFF53C4B99C6FFE7C" bold="true" box="[199,315,404,429]" pageId="13" pageNumber="14">Remarks.</emphasis>
|
||
Mastigitae are most likely a paraphyletic group in relation to
|
||
<taxonomicName id="4C084D3C2934FFFCFBAEC4B99998FE7C" box="[1082,1228,404,429]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
|
||
of Scydmaenitae (
|
||
<bibRefCitation id="EF994B4E2934FFFCFF0AC49A9C24FE01" author="Jaloszynski" box="[158,368,439,464]" pageId="13" pageNumber="14" refString="Jaloszynski, P. (2012 c) Beetles with ' trochantelli': phylogeny of Cephenniini (Coleoptera: Staphylinidae: Scydmaeninae) with focus on Neotropical genera. Systematic Entomology, 37, 448 - 477. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2012.00627. x" type="journal article" year="2012" yearSuffix="c">Jałoszyński 2012c</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF994B4E2934FFFCFEEFC49A9C91FE01" author="Jaloszynski" box="[379,453,439,464]" pageId="13" pageNumber="14" refString="Jaloszynski, P. (2014 b) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. http: // dx. doi. org / 10.1111 / syen. 12044" type="journal article" year="2014" yearSuffix="b">2014b</bibRefCitation>
|
||
); beetles currently classified in these taxa share the geniculate antennae (the apex of the scape is notched, so that the antennae can bend between the scape and pedicel), the prosternum laterally fused with prothoracic hypomera, and the metacoxae broadly separated.
|
||
<taxonomicName id="4C084D3C2934FFFCFCF1C4D29EAEFDC9" box="[869,1018,511,536]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
|
||
, as currently defined, can be easily distinguished from all species of Mastigitae on the basis of the maxillary palpomere IV, which is short, domeshaped and symmetrical, and its base is clearly narrower than the apex of palpomere III. In Mastigitae, palpomere IV is either large, elongate subtriangular or suboval (Mastigini and Clidicini), or short, but then strongly asymmetrical (with strongly oblique base) and at base as broad as the apex of palpomere III (Leptomastacini). Based on these characters,
|
||
<taxonomicName id="4C084D3C2934FFFCFE5CC7999FF3FD1C" box="[456,679,692,717]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFE5CC7999FF3FD1C" box="[456,679,692,717]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
can be unambiguously recognized as a member of Mastigitae. The structure of maxillary palps (palpomere IV elongate), the pedicel short and thin and the scape and pedicel lacking two rows of long bristles place this genus within Clidicini.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2934FFFCFF53C6329E6DF8D5" blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="14">
|
||
Clidicini include three extant genera,
|
||
<taxonomicName id="4C084D3C2934FFFCFDF9C6329E2DFCE9" authority="Laporte, 1832" authorityName="Laporte" authorityYear="1832" box="[621,889,799,824]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFDF9C6329F9EFCE9" box="[621,714,799,824]" italics="true" pageId="13" pageNumber="14">Clidicus</emphasis>
|
||
,
|
||
<bibRefCitation id="EF994B4E2934FFFCFD4DC60D9E2DFCE9" author="Laporte" box="[729,889,800,824]" pageId="13" pageNumber="14" refString="Laporte, F. L. (1832) Memoire sur cinquante especes nouvelles ou peu connues d'insectes. Annales de la Societe Entomologique de France, 1, 386 - 415." type="journal article" year="1832">Laporte, 1832</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFC10C63299ABFCE9" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[900,1279,799,824]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC10C6329977FCE9" box="[900,1059,799,824]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
<bibRefCitation id="EF994B4E2934FFFCFBBEC63299ABFCE9" author="Motschulsky" box="[1066,1279,799,824]" pageId="13" pageNumber="14" refString="Motschulsky, V. (1855) Voyages. Lettre de M. de Motschulsky a M. Menetries. No. 2. A bord du bateau a vapeur United-States, 20 Mars 1854. Etudes Entomologiques, 4, 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C084D3C2934FFFCFAA2C60C9C70FC8D" authority="Casey, 1897" authorityName="Casey" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFAA2C60C98C1FCE9" box="[1334,1429,801,824]" italics="true" pageId="13" pageNumber="14">Papusus</emphasis>
|
||
,
|
||
<bibRefCitation id="EF994B4E2934FFFCFF03C6699C70FC8D" author="Casey" box="[151,292,836,860]" pageId="13" pageNumber="14" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Science, 9, 285 - 684. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" type="journal article" year="1897">Casey, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, and three extinct genera,
|
||
<taxonomicName id="4C084D3C2934FFFCFDC5C6699EB0FC8D" authority="O'Keefe, 1997" authorityName="O'Keefe" authorityYear="1997" box="[593,996,836,861]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFDC5C6699E6CFC8C" box="[593,824,836,861]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
<bibRefCitation id="EF994B4E2934FFFCFCD4C6699EB0FC8D" author="O'Keefe" box="[832,996,836,861]" pageId="13" pageNumber="14" refString="O'Keefe, S. T., Pike, T. & Poinar, G. (1997) Palaeoleptochromus schaufussi (gen. n., sp. nov.), a new antlike stone beetle (Coleoptera: Scydmaenidae) from Canadian Cretaceous amber. The Canadian Entomologist, 129, 379 - 385. http: // dx. doi. org / 10.4039 / Ent 129379 - 3" type="journal article" year="1997">O'Keefe, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="EF994B4E2934FFFCFB80C669983FFC8D" author="O'Keefe" box="[1044,1387,836,861]" pageId="13" pageNumber="14" refString="O'Keefe, S. T., Pike, T. & Poinar, G. (1997) Palaeoleptochromus schaufussi (gen. n., sp. nov.), a new antlike stone beetle (Coleoptera: Scydmaenidae) from Canadian Cretaceous amber. The Canadian Entomologist, 129, 379 - 385. http: // dx. doi. org / 10.4039 / Ent 129379 - 3" type="journal article" year="1997">
|
||
O'Keefe,
|
||
<bibRefCitation id="EF994B4E2934FFFCFBEBC669983FFC8D" author="Pike" box="[1151,1387,836,861]" pageId="13" pageNumber="14" refString="O'Keefe, S. T., Pike, T. & Poinar, G. (1997) Palaeoleptochromus schaufussi (gen. n., sp. nov.), a new antlike stone beetle (Coleoptera: Scydmaenidae) from Canadian Cretaceous amber. The Canadian Entomologist, 129, 379 - 385. http: // dx. doi. org / 10.4039 / Ent 129379 - 3" type="journal article" year="1997">Pike & Poinar (1997</bibRefCitation>
|
||
</bibRefCitation>
|
||
)) in Canadian Campanian amber,
|
||
<taxonomicName id="4C084D3C2934FFFCFDB5C64A99A1FC51" authority="Jaloszynski, 2012a in Eocene Baltic" authorityName="Jaloszynski, 2012a in Eocene Baltic" box="[545,1269,871,896]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFDB5C64A9FBBFC51" box="[545,751,871,896]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
<bibRefCitation id="EF994B4E2934FFFCFC98C64A9EA1FC51" author="Jaloszynski" box="[780,1013,871,896]" pageId="13" pageNumber="14" refString="Jaloszynski, P. (2012 a) Description of Euroleptochromus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae) from Baltic amber, with discussion of biogeography and mouthpart evolution within Clidicini. Systematic Entomology, 37, 346 - 359. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2012.00623. x" type="journal article" year="2012" yearSuffix="a">Jałoszyński, 2012a</bibRefCitation>
|
||
in Eocene Baltic
|
||
</taxonomicName>
|
||
amber and
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C6A19C22FC74" box="[151,374,908,933]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C6A19C22FC74" box="[151,374,908,933]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
. The systematic position of
|
||
<taxonomicName id="4C084D3C2934FFFCFD55C6A09E74FC75" box="[705,800,909,932]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFD55C6A09E74FC75" box="[705,800,909,932]" italics="true" pageId="13" pageNumber="14">Papusus</emphasis>
|
||
</taxonomicName>
|
||
remains problematic and unsolved (
|
||
<bibRefCitation id="EF994B4E2934FFFCFB50C6A198C3FC75" author="Jaloszynski" box="[1220,1431,908,933]" pageId="13" pageNumber="14" refString="Jaloszynski, P. (2012 a) Description of Euroleptochromus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae) from Baltic amber, with discussion of biogeography and mouthpart evolution within Clidicini. Systematic Entomology, 37, 346 - 359. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2012.00623. x" type="journal article" year="2012" yearSuffix="a">Jałoszyński 2012a</bibRefCitation>
|
||
, 2015); this genus, associated with unusual for
|
||
<taxonomicName id="4C084D3C2934FFFCFD57C6829E32FC19" box="[707,870,943,968]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||
arid habitats of Nearctic deserts, shows unique characters and may in future be excluded from Clidicini.
|
||
<taxonomicName id="4C084D3C2934FFFCFCEBC6F99E88FC3C" box="[895,988,980,1005]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFCEBC6F99E88FC3C" box="[895,988,980,1005]" italics="true" pageId="13" pageNumber="14">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFC61C6F999C0FC3C" box="[1013,1172,980,1005]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC61C6F999C0FC3C" box="[1013,1172,980,1005]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFB3AC6F998C1FC3C" box="[1198,1429,980,1005]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFB3AC6F998C1FC3C" box="[1198,1429,980,1005]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C6DA9C31FBC1" box="[151,357,1015,1040]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C6DA9C31FBC1" box="[151,357,1015,1040]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C084D3C2934FFFCFE09C6DA9F2FFBC1" box="[413,635,1015,1040]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFE09C6DA9F2FFBC1" box="[413,635,1015,1040]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
share the strongly elongate scape, which is more than twice as long as the pedicel, and distinct rows of punctures on the elytra. All these genera, except
|
||
<taxonomicName id="4C084D3C2934FFFCFBE9C131998EFBE4" box="[1149,1242,1052,1077]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFBE9C131998EFBE4" box="[1149,1242,1052,1077]" italics="true" pageId="13" pageNumber="14">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
, have postgenal ('subocular' of
|
||
<bibRefCitation id="EF994B4E2934FFFCFEDBC1129F19FB89" author="O'Keefe" box="[335,589,1087,1112]" pageId="13" pageNumber="14" refString="O'Keefe, S. T., Pike, T. & Poinar, G. (1997) Palaeoleptochromus schaufussi (gen. n., sp. nov.), a new antlike stone beetle (Coleoptera: Scydmaenidae) from Canadian Cretaceous amber. The Canadian Entomologist, 129, 379 - 385. http: // dx. doi. org / 10.4039 / Ent 129379 - 3" type="journal article" year="1997">O'Keefe et al. (1997</bibRefCitation>
|
||
)) bristles on the head and bristles on protrochanters;
|
||
<taxonomicName id="4C084D3C2934FFFCFB62C11298C1FB89" box="[1270,1429,1087,1112]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFB62C11298C1FB89" box="[1270,1429,1087,1112]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C1499C2AFBAC" box="[151,382,1124,1149]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C1499C2AFBAC" box="[151,382,1124,1149]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C084D3C2934FFFCFE56C1499FC4FBAC" box="[450,656,1124,1149]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFE56C1499FC4FBAC" box="[450,656,1124,1149]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
additionally have bristles along the ventral margin of profemur.
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C1AA9C62FB71" box="[151,310,1159,1184]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C1AA9C62FB71" box="[151,310,1159,1184]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
is defined (among other characters) by an elongate cuticular projection on the maxillary palpomere II bearing two thick bristles at apex, and a strongly elongate (at least four times as long as broad) maxillary palpomere IV, distinctly broadest in its distal half (
|
||
<bibRefCitation id="EF994B4E2934FFFCFD7CC1E29ED9FB39" author="O'Keefe" box="[744,909,1231,1256]" pageId="13" pageNumber="14" refString="O'Keefe, S. T. (2002) Revision of the Neotropical genus Leptochromus Motschulsky (Coleoptera: Scydmaenidae): Systematic Entomology, 27, 211 - 234. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.2002.00179. x" type="journal article" year="2002">O'Keefe 2002</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C084D3C2934FFFCFC37C1E299DDFB39" box="[931,1161,1231,1256]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC37C1E299DDFB39" box="[931,1161,1231,1256]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFB0FC1E2983DFB39" box="[1179,1385,1231,1256]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFB0FC1E2983DFB39" box="[1179,1385,1231,1256]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C1D99C22FADC" box="[151,374,1268,1293]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C1D99C22FADC" box="[151,374,1268,1293]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
lack the cuticular projection of palpomere III and have the palpomere IV less elongate (less than three times as long as broad), broadest either at base or near middle.
|
||
<taxonomicName id="4C084D3C2934FFFCFC7CC03A999BFAE1" box="[1000,1231,1303,1328]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC7CC03A999BFAE1" box="[1000,1231,1303,1328]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
has the maxillary palpomere IV clearly subtriangular, broadest at base; the maxillary palpomere II evenly curved; and a pair of long bristles on the posterior margin of the vertex (
|
||
<bibRefCitation id="EF994B4E2934FFFCFD03C0729E21FAA9" author="O'Keefe" box="[663,885,1375,1400]" pageId="13" pageNumber="14" refString="O'Keefe, S. T., Pike, T. & Poinar, G. (1997) Palaeoleptochromus schaufussi (gen. n., sp. nov.), a new antlike stone beetle (Coleoptera: Scydmaenidae) from Canadian Cretaceous amber. The Canadian Entomologist, 129, 379 - 385. http: // dx. doi. org / 10.4039 / Ent 129379 - 3" type="journal article" year="1997">O'Keefe et al. 1997</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C084D3C2934FFFCFC1DC0729903FAA9" box="[905,1111,1375,1400]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC1DC0729903FAA9" box="[905,1111,1375,1400]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
has the maxillary palpomere IV suboval, broadening from base to middle and then narrowing distally; the maxillary palpomere II with its mesal margin with an angulate protuberance bearing a group of bristles; it lacks the pair of the posterior vertexal bristles; and has a slender and strongly elongate postgenal projection with two apical bristles (
|
||
<bibRefCitation id="EF994B4E2934FFFCFBD8C0E1984BFA35" author="Jaloszynski" box="[1100,1311,1484,1509]" pageId="13" pageNumber="14" refString="Jaloszynski, P. (2012 a) Description of Euroleptochromus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae) from Baltic amber, with discussion of biogeography and mouthpart evolution within Clidicini. Systematic Entomology, 37, 346 - 359. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2012.00623. x" type="journal article" year="2012" yearSuffix="a">Jałoszyński 2012a</bibRefCitation>
|
||
). The latter character was recently found in a newly described extant species of
|
||
<taxonomicName id="4C084D3C2934FFFCFC55C0C29D8FF9FD" authority="Lord, Carlton & Leschen 2014" authorityName="Lord, Carlton & Leschen" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC55C0C29934F9D9" box="[961,1120,1519,1544]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF994B4E2934FFFCFBE7C0C29D87F9FD" author="Lord" pageId="13" pageNumber="14" refString="Lord, N. P., Carlton, C. E. & Leschen, R. A. B. (2014) A new species of Leptochromus Motschulsky (Coleoptera: Staphylinidae: Scydmaeninae) from Costa Rica, with notes on morphology and natural history. The Coleopterists Bulletin, 68 (1), 119 - 125. http: // dx. doi. org / 10.1649 / 0010 - 065 X- 68.1.119" type="journal article" year="2014">Lord, Carlton & Leschen 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, all remaining species of
|
||
<taxonomicName id="4C084D3C2934FFFCFE61C3399FC0F9FC" box="[501,660,1556,1581]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFE61C3399FC0F9FC" box="[501,660,1556,1581]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
has the postgenae either unmodified or with a weakly raised angulate or tuberculate protuberance bearing the postgenal bristles (
|
||
<bibRefCitation id="EF994B4E2934FFFCFC24C31A9932F981" author="O'Keefe" box="[944,1126,1591,1616]" pageId="13" pageNumber="14" refString="O'Keefe, S. T. (2002) Revision of the Neotropical genus Leptochromus Motschulsky (Coleoptera: Scydmaenidae): Systematic Entomology, 27, 211 - 234. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.2002.00179. x" type="journal article" year="2002">O'Keefe 2002</bibRefCitation>
|
||
). The newly described
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C3719C22F9A4" box="[151,374,1628,1653]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C3719C22F9A4" box="[151,374,1628,1653]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
has the maxillary palpomere IV approximately subtriangular, broadest at base (see the left palp in
|
||
<figureCitation id="13332A3A2934FFFCFF22C3529C58F949" box="[182,268,1663,1688]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1833,1856]" captionTargetBox="[159,1435,886,1804]" captionTargetId="figure@15.[151,1436,876,1813]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Holotype of Rovnoleptochromus ableptonoides sp. n. (K- 27418). Head in ventrolateral view (46), pronotum and elytra in dorsal view (47), body in ventral view (48). Abbreviations: abp, antebasal pit; aest 3, metanepisternum; an 1 − 11, antennomere I − XI; fr, frons; hr, hypostomal ridge; lbr, labrum; lp 2 − 3, labial palpomere II − III; md, mandible; mn, mentum; msvp, mesoventral process; mtvp, metaventral process; mvc, metaventral carina; mxp 1 − 4, maxillary palpomere I − IV; pcmc, posterior carina of mesocoxal cavity; pgb, postgenal bristles; pn, pronotum; pnc, pronotal collar; ptp, posterior tentorial pit; slc 2, mesoscutellum; smn, submentum; st 3 − 8; abdominal sternite III − VIII; v 3, metaventrite; vc, V-shaped carina." httpUri="https://zenodo.org/record/262419/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
|
||
); the maxillary palpomere II evenly curved; and lacks the vertexal bristles and the postgenal process. Additionally,
|
||
<taxonomicName id="4C084D3C2934FFFCFEA6C3899F45F96C" box="[306,529,1700,1725]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFEA6C3899F45F96C" box="[306,529,1700,1725]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
has the longitudinal metaventral carina, a character not known in any extant or extinct Clidicini, but present in
|
||
<taxonomicName id="4C084D3C2934FFFCFE63C3EA9E7EF931" authority="Frivaldszky, 1877" authorityName="Frivaldszky" authorityYear="1877" box="[503,810,1735,1760]" class="Insecta" family="Staphylinidae" genus="Ablepton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFE63C3EA9F08F931" box="[503,604,1735,1760]" italics="true" pageId="13" pageNumber="14">Ablepton</emphasis>
|
||
<bibRefCitation id="EF994B4E2934FFFCFDF6C3EA9E7EF931" author="Frivaldszky" box="[610,810,1735,1760]" pageId="13" pageNumber="14" refString="Frivaldszky, J. (1877) Coleoptera nova e Hungaria meridionale. Termeszetrajzi Fuzetek, 1, 17 - 22." type="journal article" year="1877">Frivaldszky, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
of Leptomastacini (Jałoszyński, unpublished obs.). Key characters of all genera of Clidicini are compiled in
|
||
<tableCitation id="C68A03042934FFFCFD4BC3C19E67F8D5" box="[735,819,1772,1797]" captionStart="TABLE 1" captionStartId="14.[119,203,575,600]" captionText="TABLE 1. Comparison of key morphological characters of the genera of Cliđicini." pageId="13" pageNumber="14">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2934FFFCFF53C2229DA5F844" blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C084D3C2934FFFCFF53C2229CF2F8F9" box="[199,422,1807,1832]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2934FFFCFF53C2229CF2F8F9" box="[199,422,1807,1832]" italics="true" pageId="13" pageNumber="14">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
in a preliminary phylogenetic analysis was placed as a component of the '
|
||
<taxonomicName id="4C084D3C2934FFFCFB69C22298C8F8F9" box="[1277,1436,1807,1832]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFB69C22298C8F8F9" box="[1277,1436,1807,1832]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
lineage' of Clidicini, i.e., a member of a group that includes
|
||
<taxonomicName id="4C084D3C2934FFFCFC54C219990BF89C" box="[960,1119,1844,1869]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFC54C219990BF89C" box="[960,1119,1844,1869]" italics="true" pageId="13" pageNumber="14">Leptochromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C2934FFFCFBEDC2199834F89C" box="[1145,1376,1844,1869]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFBEDC2199834F89C" box="[1145,1376,1844,1869]" italics="true" pageId="13" pageNumber="14">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C084D3C2934FFFCFF03C27A9C31F8A1" box="[151,357,1879,1904]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2934FFFCFF03C27A9C31F8A1" box="[151,357,1879,1904]" italics="true" pageId="13" pageNumber="14">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
, and as a sister group to a clade composed of these three genera (Jałoszyński, unpublished results).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7766372937FFFFFFE3C7129EE4FD89" box="[119,944,575,601]" pageId="14" pageNumber="15">
|
||
<paragraph id="8BB736BF2937FFFFFFE3C7129EE4FD89" blockId="14.[119,944,575,601]" box="[119,944,575,601]" pageId="14" pageNumber="15">
|
||
<emphasis id="B97CEAAD2937FFFFFFE3C7129DB0FD89" bold="true" box="[119,228,575,600]" pageId="14" pageNumber="15">TABLE 1.</emphasis>
|
||
Comparison of key morphological characters of the genera of Cliđicini.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31265342937FFFFFEF0C7AC9BEDF992" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8BB736BF2937FFFFFEF0C7AC9A07FD4B" blockId="14.[356,1875,641,666]" box="[356,1875,641,666]" pageId="14" pageNumber="15">
|
||
<emphasis id="B97CEAAD2937FFFFFEF0C7AC9A07FD4B" bold="true" box="[356,1875,641,666]" pageId="14" pageNumber="15">
|
||
<emphasis id="B97CEAAD2937FFFFFEF0C7AC9955FD4B" bold="true" box="[356,1025,641,666]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="4C084D3C2937FFFFFEF0C7AC9CECFD4B" box="[356,440,641,666]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Clidicus</taxonomicName>
|
||
<taxonomicName id="4C084D3C2937FFFFFDFDC7AF9F94FD48" box="[617,704,642,665]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Papusus</taxonomicName>
|
||
<taxonomicName id="4C084D3C2937FFFFFCF9C7AC9955FD4B" box="[877,1025,641,666]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Leptochromus</taxonomicName>
|
||
</emphasis>
|
||
†
|
||
<taxonomicName id="4C084D3C2937FFFFFBEAC7AC9804FD4B" box="[1150,1360,641,666]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2937FFFFFBEAC7AC9804FD4B" bold="true" box="[1150,1360,641,666]" italics="true" pageId="14" pageNumber="15">Palaeoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
†
|
||
<taxonomicName id="4C084D3C2937FFFFFA16C7AC9B16FD4B" authority="Rovnoleptochromus" authorityName="Rovnoleptochromus" box="[1410,1602,641,666]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2937FFFFFA16C7AC9B16FD4B" bold="true" box="[1410,1602,641,666]" italics="true" pageId="14" pageNumber="15">Euroleptochromus</emphasis>
|
||
</taxonomicName>
|
||
†
|
||
<taxonomicName id="4C084D3C2937FFFFF912C7AC9A07FD4B" box="[1670,1875,641,666]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97CEAAD2937FFFFF912C7AC9A07FD4B" bold="true" box="[1670,1875,641,666]" italics="true" pageId="14" pageNumber="15">Rovnoleptochromus</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C7EF9BEDFD0A" blockId="14.[119,1885,706,1603]" box="[119,1721,706,731]" pageId="14" pageNumber="15">Cuticular projection absent absent long anđ bearing two absent tuberculate, weakly absent</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFF06C7CF9B0BFD2A" blockId="14.[119,1885,706,1603]" box="[146,1631,738,763]" pageId="14" pageNumber="15">maxillary apical bristles raiseđ, with 2-3 bristles</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C62F9DA3FCCB" blockId="14.[119,1885,706,1603]" box="[119,247,770,795]" pageId="14" pageNumber="15">palpomere II</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C60C9EE7FC4B" blockId="14.[119,1885,706,1603]" pageId="14" pageNumber="15">Maxillary palpomere 2$3 × as long as broađ, at least 4 ×as long as at least 4 × as long as about 3 × as long as> 5 × as long as broađ, 3 × as long as broađ, with oblique apical broađ, with transverse broađ, with nearly broađ, with oblique with transverse apical with oblique apical margin apical margin transverse apical apical margin margin margin margin</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C68C9A54FC2B" blockId="14.[119,1885,706,1603]" pageId="14" pageNumber="15">Maxillary palpomere subtriangular, broađest subtriangular, broađest suboval, broađest in subtriangular, broađest suboval, broađest near subtriangular, broađest at base, 2$3 × as long at base, about 2 × as đistal half, at least 4 × at base, about 2 × as miđđle, 2.5 × as long at base, about 2 × as as broađ long as broađ as long as broađ long as broađ as broađ long as broađ</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C12D9BEDFBC8" blockId="14.[119,1885,706,1603]" box="[119,1721,1024,1049]" pageId="14" pageNumber="15">Posterolateral absent absent absent present absent absent</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C10D9DAAFBE9" blockId="14.[119,1885,706,1603]" box="[119,254,1056,1081]" pageId="14" pageNumber="15">vertexal setae</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C16D9918FBA8" blockId="14.[119,1885,706,1603]" pageId="14" pageNumber="15">Postgenal process absent absent absent or present, from absent rođ-like absent weakly raiseđ tubercle</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFCF9C1AC9EA1FB48" blockId="14.[119,1885,706,1603]" box="[877,1013,1152,1177]" pageId="14" pageNumber="15">to slenđer rođ</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C18D9B95FB69" blockId="14.[119,1885,706,1603]" box="[119,1729,1184,1209]" pageId="14" pageNumber="15">Poastgenal absent absent present present present present</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFEAC1929C62FB09" blockId="14.[119,1885,706,1603]" box="[126,310,1215,1240]" pageId="14" pageNumber="15">subocular) bristles</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C1F29B9CFB29" blockId="14.[119,1885,706,1603]" box="[119,1736,1247,1272]" pageId="14" pageNumber="15">Posterior margins of carinate carinate non-carinate unknown carinate carinate</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C1D29C67FAC9" blockId="14.[119,1885,706,1603]" box="[119,307,1279,1304]" pageId="14" pageNumber="15">mesocoxal cavities</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0329A05FAE9" blockId="14.[119,1885,706,1603]" box="[119,1873,1311,1336]" pageId="14" pageNumber="15">Mesoventral parallel-siđeđ or parallel-siđeđ parallel-siđeđ unknown parallel-siđeđ narrowing posteriorly</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0129F70FA89" blockId="14.[119,1885,706,1603]" box="[119,548,1343,1368]" pageId="14" pageNumber="15">intercoxal process slightly broađening</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFEF0C0729C9BFAA9" blockId="14.[119,1885,706,1603]" box="[356,463,1375,1400]" pageId="14" pageNumber="15">posteriorly</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0529B97FA49" blockId="14.[119,1885,706,1603]" box="[119,1731,1407,1432]" pageId="14" pageNumber="15">Elytral striae đistinct inđistinct or absent đistinct đistinct đistinct đistinct</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0879B95FA13" blockId="14.[119,1885,706,1603]" box="[119,1729,1450,1475]" pageId="14" pageNumber="15">Međian longituđinal absent absent absent unknown absent present</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0E79C7BFA32" blockId="14.[119,1885,706,1603]" box="[119,303,1482,1507]" pageId="14" pageNumber="15">metaventral carina</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C0C79B95F9D3" blockId="14.[119,1885,706,1603]" box="[119,1729,1514,1539]" pageId="14" pageNumber="15">Protrochanteral absent absent present present present present</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C3279DEAF9F2" blockId="14.[119,1885,706,1603]" box="[119,190,1546,1571]" pageId="14" pageNumber="15">bristles</paragraph>
|
||
<paragraph id="8BB736BF2937FFFFFFE3C3079BEDF992" blockId="14.[119,1885,706,1603]" box="[119,1721,1578,1603]" pageId="14" pageNumber="15">Profemoral bristles absent absent present present present absent</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |