446 lines
58 KiB
XML
446 lines
58 KiB
XML
<document id="E1F0EE5EC115B55C49A6A161496DF2CB" ID-DOI="10.5281/zenodo.5385668" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385668" ID-ZooBank="A7DA557E-5E0D-4629-B715-6BE88124DCB7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630302160134" checkinUser="valdenar" docAuthor="Stasolla, G., Abbarchi, A. & Innocenti, G." docDate="2015" docId="0398FD3B9706FFDBFEEBFEA90192F8C7" docLanguage="en" docName="63rbz448-453.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Sundathelphusa spelaeophila Stasolla & Abbarchi & Innocenti 2015, n. sp." docType="treatment" docUuid="DAB6C5CD-5C5A-4291-8389-CA7FB3E88626" docUuidSource="ZooBank" docVersion="5" lastPageNumber="451" masterDocId="FFA185439707FFD8FFEAFF950331FFEE" masterDocTitle="Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae)" masterLastPageNumber="453" masterPageNumber="448" pageNumber="449" updateTime="1699220854849" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4B998A3D254FF0A24288820ECD27EBAD">Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae)</mods:title>
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<mods:name id="8C17B2ACBA4301F360A95BE495F16C7E" type="personal">
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<mods:roleTerm id="5322B997ACBA8066B58817D3C9F0A581">Author</mods:roleTerm>
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<mods:namePart id="A8E15DE0602AC39E834737B6738CC5A6">Stasolla, G.</mods:namePart>
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<mods:roleTerm id="80CBC589F5DAFE2B744A94C6D716014C">Author</mods:roleTerm>
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<mods:namePart id="DA3A81936437C7A4D92B04A2376B4657">Abbarchi, A.</mods:namePart>
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<mods:roleTerm id="1B2BB1111E692E841DD0276C2848D9E5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="51C99AF5FE163CA0FBB6602D8BAAAF48">Innocenti, G.</mods:namePart>
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<mods:typeOfResource id="8C252DB469F3557656EBE1F1D3E9FDAF">text</mods:typeOfResource>
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<mods:title id="A3475C303A606DDB043C961CB7210AD5">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="D5182744AF9E84481CB3CA69B0EF19A3">
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<mods:date id="0B8D54D1EEDEE2CADA09D8B24E72B322">2015</mods:date>
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<mods:number id="85028A68902DCABD447F822EDDC3CF18">2015-10-09</mods:number>
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<mods:number id="9BF219A87F13FB6E96ED11F7C20240C3">63</mods:number>
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<mods:classification id="DACC159D51AF675FFE17B96044A8B18E">journal article</mods:classification>
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<mods:identifier id="9190285C15AE044925EF020375DD4107" type="DOI">10.5281/zenodo.5385668</mods:identifier>
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<mods:identifier id="28232F0253774D46A7F3F81EBFDE850A" type="ISSN">2345-7600</mods:identifier>
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<treatment id="0398FD3B9706FFDBFEEBFEA90192F8C7" ID-DOI="http://doi.org/10.5281/zenodo.5466957" ID-Zenodo-Dep="5466957" LSID="urn:lsid:zoobank.org:act:DAB6C5CD-5C5A-4291-8389-CA7FB3E88626" httpUri="http://treatment.plazi.org/id/0398FD3B9706FFDBFEEBFEA90192F8C7" lastPageId="3" lastPageNumber="451" pageId="1" pageNumber="449">
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<subSubSection id="C32B1FA69706FFD9FEEBFEA901BCFEBB" box="[257,653,316,341]" pageId="1" pageNumber="449" type="nomenclature">
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<paragraph id="8B8E4C2D9706FFD9FEEBFEA901BCFEBB" blockId="1.[257,653,316,373]" box="[257,653,316,341]" pageId="1" pageNumber="449">
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<heading id="D0C6FB419706FFD9FEEBFEA901BCFEBB" bold="true" box="[257,653,316,341]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="449" reason="2">
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<emphasis id="B945903F9706FFD9FEEBFEA901BCFEBB" bold="true" box="[257,653,316,341]" pageId="1" pageNumber="449">
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<taxonomicName id="4C3137AE9706FFD9FEEBFEA90176FEBA" ID-CoL="53GC6" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[257,583,316,340]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
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<emphasis id="B945903F9706FFD9FEEBFEA90176FEBA" bold="true" box="[257,583,316,340]" italics="true" pageId="1" pageNumber="449">Sundathelphusa spelaeophila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2762D449706FFD9FDA5FEA801BCFEBB" box="[591,653,317,341]" pageId="1" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B1FA69706FFD9FEB3FEC80104FE9B" box="[345,565,349,373]" pageId="1" pageNumber="449" type="description">
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<paragraph id="8B8E4C2D9706FFD9FEB3FEC80104FE9B" blockId="1.[257,653,316,373]" box="[345,565,349,373]" pageId="1" pageNumber="449">
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(
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<figureCitation id="130A50A89706FFD9FE8BFEC802D0FE9B" box="[353,481,349,373]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Figs. 1A–C</figureCitation>
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||
,
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||
<figureCitation id="130A50A89706FFD9FE04FEC8011FFE9B" box="[494,558,349,373]" captionStart="Fig" captionStartId="2.[113,148,1317,1339]" captionTargetBox="[116,1451,190,1302]" captionTargetId="figure-317@2.[95,1469,173,1305]" captionTargetPageId="2" captionText="Fig. 2. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="1" pageNumber="449">2A–E</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32B1FA69706FFD9FF62FE0807EBFCFB" pageId="1" pageNumber="449" type="materials_examined">
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<paragraph id="8B8E4C2D9706FFD9FF62FE080617FEFB" blockId="1.[135,776,413,725]" lastBlockId="1.[834,1474,189,278]" pageId="1" pageNumber="449">
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||
<emphasis id="B945903F9706FFD9FF62FE080242FE5B" bold="true" box="[136,371,413,437]" pageId="1" pageNumber="449">Material examined.</emphasis>
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||
<materialsCitation id="3B5946709706FFD9FE94FE080298FDFB" collectingDate="1991-03-19" collectionCode="MZUF" collectorName="C. Ferron" country="Philippines" location="Can Gortio Cave" pageId="1" pageNumber="449" specimenCode="MZUF 3920" specimenCount="1" stateProvince="Samar" typeStatus="holotype">
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||
<typeStatus id="548AF28F9706FFD9FE94FE0802DBFE5B" box="[382,490,413,437]" pageId="1" pageNumber="449" type="holotype">Holotype</typeStatus>
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||
: male (15.9 ×
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||
<quantity id="4CC9E1C89706FFD9FD7DFE0801CFFE5B" box="[663,766,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="449" unit="mm" value="14.1">14.1 mm</quantity>
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||
) (
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||
<specimenCode id="DB97E4569706FFD9FF65FE280217FE3B" box="[143,294,445,469]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="1" pageNumber="449">MZUF 3920</specimenCode>
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);
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||
<collectingCountry id="F3260CBD9706FFD9FED7FE28028EFE3B" box="[317,447,445,469]" name="Philippines" pageId="1" pageNumber="449">Philippines</collectingCountry>
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||
,
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||
<collectingRegion id="49F582CF9706FFD9FE25FE2801B7FE3B" box="[463,646,445,469]" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar province</collectingRegion>
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||
, San Jorge municipality, Barangay Matalud,
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||
<location id="8EEE1AF69706FFD9FDEEFE4801F9FE1B" LSID="urn:lsid:plazi:treatment:0398FD3B9706FFDBFEEBFEA90192F8C7:8EEE1AF69706FFD9FDEEFE4801F9FE1B" box="[516,712,477,501]" country="Philippines" name="Can Gortio Cave" pageId="1" pageNumber="449" stateProvince="Samar">Can Gortio Cave</location>
|
||
, coll.
|
||
<collectorName id="26C429FB9706FFD9FF62FE6803C2FDFA" box="[136,243,509,533]" pageId="1" pageNumber="449">C. Ferron</collectorName>
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||
,
|
||
<date id="FF8F6AED9706FFD9FF15FE680298FDFB" box="[255,425,509,533]" pageId="1" pageNumber="449" value="1991-03-19">
|
||
<collectingDate id="EFCB93059706FFD9FF15FE680298FDFB" box="[255,425,509,533]" pageId="1" pageNumber="449" value="1991-03-19">19 March 1991</collectingDate>
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||
</date>
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||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B5946709706FFD9FE5CFE68023BFD9B" collectingDate="1991-03-19" collectionCode="MZUF" collectorName="C. Ferron" country="Philippines" location="Can Gortio Cave" pageId="1" pageNumber="449" specimenCode="MZUF 4273" specimenCount="1" specimenCount-male="1" stateProvince="Samar" typeStatus="paratype">
|
||
<typeStatus id="548AF28F9706FFD9FE5CFE680113FDFB" box="[438,546,509,533]" pageId="1" pageNumber="449" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D3787A49706FFD9FDC7FE680146FDFB" box="[557,631,509,533]" pageId="1" pageNumber="449" type="male">1 male</specimenCount>
|
||
(11.7 ×
|
||
<quantity id="4CC9E1C89706FFD9FD3DFE680384FDDB" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="449" unit="mm" value="10.6">10.6 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB97E4569706FFD9FF23FD880260FDDB" box="[201,337,541,565]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="1" pageNumber="449">MZUF 4273</specimenCode>
|
||
)
|
||
<collectingRegion id="49F582CF9706FFD9FEB7FD880137FDDB" box="[349,518,541,565]" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar province</collectingRegion>
|
||
, San Jorge municipality, Barangay Matalud,
|
||
<location id="8EEE1AF69706FFD9FE8EFDA80114FDBB" LSID="urn:lsid:plazi:treatment:0398FD3B9706FFDBFEEBFEA90192F8C7:8EEE1AF69706FFD9FE8EFDA80114FDBB" box="[356,549,573,597]" country="Philippines" name="Can Gortio Cave" pageId="1" pageNumber="449" stateProvince="Samar">Can Gortio Cave</location>
|
||
, coll.
|
||
<collectorName id="26C429FB9706FFD9FD84FDA801EDFDBA" box="[622,732,573,597]" pageId="1" pageNumber="449">C. Ferron</collectorName>
|
||
,
|
||
<date id="FF8F6AED9706FFD9FD00FDA8023BFD9B" pageId="1" pageNumber="449" value="1991-03-19">
|
||
<collectingDate id="EFCB93059706FFD9FD00FDA8023BFD9B" pageId="1" pageNumber="449" value="1991-03-19">19 March 1991</collectingDate>
|
||
</date>
|
||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B5946709706FFD9FEF9FDC8016CFD5B" collectingDate="1991-03-20" collectionCode="NMCR" collectorName="P. Marcel" country="Philippines" location="Can Gortio Cave" pageId="1" pageNumber="449" specimenCode="NMCR 40102" specimenCount="1" specimenCount-male="1" stateProvince="Samar" typeStatus="paratype">
|
||
<specimenCount id="9D3787A49706FFD9FEF9FDC80269FD9B" box="[275,344,605,629]" pageId="1" pageNumber="449" type="male">1 male</specimenCount>
|
||
(15.4 ×
|
||
<quantity id="4CC9E1C89706FFD9FE47FDC8013AFD9B" box="[429,523,605,629]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="449" unit="mm" value="12.9">12.9 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB97E4569706FFD9FDF4FDC80186FD9B" box="[542,695,605,629]" collectionCode="NMCR" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="1" pageNumber="449" type="Herbarium">NMCR 40102</specimenCode>
|
||
)
|
||
<collectingRegion id="49F582CF9706FFD9FD28FDC803D6FD7B" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar province</collectingRegion>
|
||
, San Jorge municipality, Barangay Matalud,
|
||
<location id="8EEE1AF69706FFD9FD30FDE8023DFD5B" LSID="urn:lsid:plazi:treatment:0398FD3B9706FFDBFEEBFEA90192F8C7:8EEE1AF69706FFD9FD30FDE8023DFD5B" country="Philippines" name="Can Gortio Cave" pageId="1" pageNumber="449" stateProvince="Samar">Can Gortio Cave</location>
|
||
, coll.
|
||
<collectorName id="26C429FB9706FFD9FEA3FD080280FD5B" box="[329,433,669,693]" pageId="1" pageNumber="449">P. Marcel</collectorName>
|
||
,
|
||
<date id="FF8F6AED9706FFD9FE51FD08016CFD5B" box="[443,605,669,693]" pageId="1" pageNumber="449" value="1991-03-20">
|
||
<collectingDate id="EFCB93059706FFD9FE51FD08016CFD5B" box="[443,605,669,693]" pageId="1" pageNumber="449" value="1991-03-20">20 March 1991</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B5946709706FFD9FD8CFD080612FEFB" collectingDate="1993-05-26" collectionCode="MZUF, SNAZ" collectorName="Cave & Zootechnics & C. Ferron" country="Philippines" location="National School of Agriculture" pageId="1" pageNumber="449" specimenCode="MZUF 3927, SNAZ 1" specimenCount="1" specimenCount-male="1" stateProvince="Samar">
|
||
Other material:
|
||
<specimenCount id="9D3787A49706FFD9FF6DFD2803E0FD3B" box="[135,209,701,725]" pageId="1" pageNumber="449" type="male">1 male</specimenCount>
|
||
(16.8 ×
|
||
<quantity id="4CC9E1C89706FFD9FEDEFD2802A6FD3B" box="[308,407,701,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="449" unit="mm" value="14.7">14.7 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB97E4569706FFD9FE45FD28010FFD3B" box="[431,574,701,725]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="1" pageNumber="449">MZUF 3927</specimenCode>
|
||
),
|
||
<collectingRegion id="49F582CF9706FFD9FDB9FD280030FD3B" box="[595,769,701,725]" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar province</collectingRegion>
|
||
, San Jorge municipality, Barangay Matalud, “
|
||
<specimenCode id="DB97E4569706FFD9FAF5FF280646FF3B" box="[1311,1399,189,213]" collectionCode="SNAZ" pageId="1" pageNumber="449">SNAZ 1</specimenCode>
|
||
”
|
||
<collectorName id="26C429FB9706FFD9FA61FF2806F3FF3B" box="[1419,1474,189,213]" pageId="1" pageNumber="449">Cave</collectorName>
|
||
(
|
||
<quantity id="4CC9E1C89706FFD9FCA7FF4800B4FF1B" box="[845,901,221,245]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="449" unit="km" value="1.0">1 km</quantity>
|
||
from the local
|
||
<location id="8EEE1AF69706FFD9FBD2FF4806A1FF1B" LSID="urn:lsid:plazi:treatment:0398FD3B9706FFDBFEEBFEA90192F8C7:8EEE1AF69706FFD9FBD2FF4806A1FF1B" box="[1080,1424,221,245]" country="Philippines" name="National School of Agriculture" pageId="1" pageNumber="449" stateProvince="Samar">National School of Agriculture</location>
|
||
and
|
||
<collectorName id="26C429FB9706FFD9FCA9FF6800FBFEFB" box="[835,970,253,277]" pageId="1" pageNumber="449">Zootechnics</collectorName>
|
||
), coll.
|
||
<collectorName id="26C429FB9706FFD9FBFFFF6807B1FEFB" box="[1045,1152,253,277]" pageId="1" pageNumber="449">C. Ferron</collectorName>
|
||
,
|
||
<date id="FF8F6AED9706FFD9FB67FF680612FEFB" box="[1165,1315,253,278]" pageId="1" pageNumber="449" value="1993-05-26">
|
||
<collectingDate id="EFCB93059706FFD9FB67FF680612FEFB" box="[1165,1315,253,278]" pageId="1" pageNumber="449" value="1993-05-26">26 May 1993</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9706FFD9FCA9FEA807EBFCFB" blockId="1.[834,1475,317,789]" pageId="1" pageNumber="449">
|
||
<emphasis id="B945903F9706FFD9FCA9FEA80775FEBB" bold="true" box="[835,1092,317,341]" pageId="1" pageNumber="449">Comparative material.</emphasis>
|
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The specimens were compared with the descriptions of
|
||
<taxonomicName id="4C3137AE9706FFD9FBFDFEC800EDFE7B" authority="Husana, Naruse & Kase, 2009" authorityName="Husana, Naruse & Kase" authorityYear="2009" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="waray">
|
||
<emphasis id="B945903F9706FFD9FBFDFEC8063FFE9B" box="[1047,1294,349,373]" italics="true" pageId="1" pageNumber="449">Sundathelphusa waray</emphasis>
|
||
<bibRefCitation id="EFA031DC9706FFD9FAFDFECB00EDFE7B" author="Husana DEM & Naruse T & Kase T" pageId="1" pageNumber="449" pagination="419 - 427" refId="ref3400" refString="Husana DEM, Naruse T & Kase T (2009) Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3): 419 - 427." type="journal article" year="2009">Husana, Naruse & Kase, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3137AE9706FFD9FBFCFEEB068FFE7B" authority="Husana, Naruse & Kase, 2009" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[1046,1470,381,405]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="lobo">
|
||
<emphasis id="B945903F9706FFD9FBFCFEEB0752FE7B" box="[1046,1123,381,405]" italics="true" pageId="1" pageNumber="449">S. lobo</emphasis>
|
||
<bibRefCitation id="EFA031DC9706FFD9FB87FEEB068FFE7B" author="Husana DEM & Naruse T & Kase T" box="[1133,1470,381,405]" pageId="1" pageNumber="449" pagination="419 - 427" refId="ref3400" refString="Husana DEM, Naruse T & Kase T (2009) Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3): 419 - 427." type="journal article" year="2009">Husana, Naruse & Kase, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, as well as with the direct observation of the specimens of
|
||
<taxonomicName id="4C3137AE9706FFD9FCA9FE2B079FFE3B" authority="(von Martens, 1868)" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[835,1198,445,469]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9706FFD9FCA9FE2B00FDFE3B" box="[835,972,445,469]" italics="true" pageId="1" pageNumber="449">S. philippina</emphasis>
|
||
(von
|
||
<bibRefCitation id="EFA031DC9706FFD9FBE0FE2B0797FE3B" author="Martens E von" box="[1034,1190,445,469]" pageId="1" pageNumber="449" pagination="608 - 615" refId="ref3637" refString="Martens E von (1868) Uber einige neue Crustaceen. Monatsberichte der Koniglichen Preussischen Akademien der Wissenschaften zu Berlin, 1868: 608 - 615." type="journal article" year="1868">Martens, 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
deposited in the Museum für Naturkunde, Berlin (
|
||
<collectionCode id="ED20D4E89706FFD9FBB9FE4807A3FE1B" box="[1107,1170,477,501]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="449" type="Museum">ZMB</collectionCode>
|
||
), a
|
||
<taxonomicName id="4C3137AE9706FFD9FB54FE4806F3FE1B" box="[1214,1474,477,501]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B945903F9706FFD9FB54FE48065BFE1B" box="[1214,1386,477,501]" italics="true" pageId="1" pageNumber="449">Sundathelphusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
collected in
|
||
<collectingRegion id="49F582CF9706FFD9FC28FE680737FDFB" box="[962,1030,509,533]" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar</collectingRegion>
|
||
Island. These last specimens are presently catalogued as
|
||
<taxonomicName id="4C3137AE9706FFD9FC3CFD8B0755FDDB" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[982,1124,541,565]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9706FFD9FC3CFD8B0755FDDB" box="[982,1124,541,565]" italics="true" pageId="1" pageNumber="449">S. grapsoides</emphasis>
|
||
</taxonomicName>
|
||
, possibly following
|
||
<bibRefCitation id="EFA031DC9706FFD9FAD6FD8806F3FDDB" author="Balss H" box="[1340,1474,541,565]" pageId="1" pageNumber="449" pagination="143 - 197" refId="ref2987" refString="Balss H (1937) Potamoniden (Dekapoda Brachyura) der Philippinen und des Malayischen Archipels. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34: 143 - 197." type="journal article" year="1937">Balss (1937)</bibRefCitation>
|
||
(O. Coleman pers. comm.).
|
||
<taxonomicName id="4C3137AE9706FFD9FB80FDA806B9FDBB" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1130,1416,573,597]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9706FFD9FB80FDA806B9FDBB" box="[1130,1416,573,597]" italics="true" pageId="1" pageNumber="449">Sundathelphusa philippina</emphasis>
|
||
</taxonomicName>
|
||
were collected in
|
||
<collectingRegion id="49F582CF9706FFD9FC2DFDC8073AFD9B" box="[967,1035,605,629]" country="Philippines" name="Samar" pageId="1" pageNumber="449">Samar</collectingRegion>
|
||
in the following localities: Calbiga river and Loquilocum (= Ulut) river (
|
||
<bibRefCitation id="EFA031DC9706FFD9FB43FDE80613FD7B" author="Bott R" box="[1193,1314,637,661]" pageId="1" pageNumber="449" pagination="1 - 338" refId="ref3044" refString="Bott R (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526: 1 - 338." type="journal article" year="1970">Bott, 1970</bibRefCitation>
|
||
). As the
|
||
<collectionCode id="ED20D4E89706FFD9FA63FDEB06F2FD7B" box="[1417,1475,638,661]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="449" type="Museum">ZMB</collectionCode>
|
||
specimens were presently identified as
|
||
<taxonomicName id="4C3137AE9706FFD9FB15FD0B0702FD3B" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9706FFD9FB15FD0B06A5FD5B" box="[1279,1428,669,693]" italics="true" pageId="1" pageNumber="449">S. grapsoides</emphasis>
|
||
(H.
|
||
<bibRefCitation id="EFA031DC9706FFD9FCA9FD28071DFD3B" author="Milne Edwards H" box="[835,1068,701,725]" pageId="1" pageNumber="449" pagination="163 - 228" refId="ref3750" refString="Milne Edwards H (1853) Memoire sur la famille des Ocypodides. Annales des Sciences Naturelles, (3) 20: 163 - 228." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the original descriptions of this latter species were also carefully compared, following A. Milne Edwards (1869) and
|
||
<bibRefCitation id="EFA031DC9706FFD9FBC2FD6807E7FCFB" author="Rathbun MJ" box="[1064,1238,765,789]" pageId="1" pageNumber="449" pagination="225 - 312" refId="ref4075" refString="Rathbun MJ (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 4 e serie, 6: 225 - 312." type="journal article" year="1904">Rathbun (1904)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B1FA69706FFDAFCA9FCA802EFF7EA" lastPageId="2" lastPageNumber="450" pageId="1" pageNumber="449" type="description">
|
||
<paragraph id="8B8E4C2D9706FFD9FCA9FCA806F2FA9B" blockId="1.[835,1476,829,1398]" pageId="1" pageNumber="449">
|
||
<emphasis id="B945903F9706FFD9FCA9FCA800FFFCBB" bold="true" box="[835,974,829,853]" pageId="1" pageNumber="449">Description.</emphasis>
|
||
Carapace (
|
||
<figureCitation id="130A50A89706FFD9FBA2FCA807AAFCBB" box="[1096,1179,829,853]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Fig. 1A</figureCitation>
|
||
) subquadrate, broader than long (width-to-length ratio ca. 1.1) quite flat; dorsal surface with distinct rugosity in frontal, hepatic regions; branchial region rugose, striated; regions poorly defined, cervical, gastric grooves shallow; epigastric cristae low but distinct, continuous with postorbital cristae; postorbital cristae low but distinct, gently curving toward anterior, converging toward and connecting epibranchial teeth; frontal margin slightly convex; anterolateral margins slightly convex, granular; external orbital angle not very broad; single epibranchial tooth acutely triangular, developed, anteriorly directed, separated from exorbital tooth by V-shaped notch; posterolateral margins slightly concave before converging towards posterior carapace margin; supraorbital margin smooth, parallel with frontal margin; infraorbital margins granular, complete, congruent with anterolateral margin; suborbital region rugose, extending to sub-branchial region. Antennules slender. Eyes well-developed, cornea pigmented.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9706FFD9FCA9FA0807DCF9DB" blockId="1.[834,1475,1437,1589]" pageId="1" pageNumber="449">
|
||
Epistome narrow (
|
||
<figureCitation id="130A50A89706FFD9FBECFA080766FA5B" box="[1030,1111,1437,1461]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Fig. 1C</figureCitation>
|
||
), posterior margin with one lateral cleft, median part sub-triangular, margin smooth. Third maxilliped (
|
||
<figureCitation id="130A50A89706FFD9FC21FA48071CFA1B" box="[971,1069,1501,1525]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="130A50A89706FFD9FBD7FA480751FA1B" box="[1085,1120,1501,1525]" captionStart="Fig" captionStartId="2.[113,148,1317,1339]" captionTargetBox="[116,1451,190,1302]" captionTargetId="figure-317@2.[95,1469,173,1305]" captionTargetPageId="2" captionText="Fig. 2. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="1" pageNumber="449">2A</figureCitation>
|
||
) with slender, sinuous exopod, narrower than ischium, flagellum well developed; merus broad, antero-external margin rounded.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9706FFD9FCA9F9C80617F91B" blockId="1.[834,1475,1629,1781]" pageId="1" pageNumber="449">
|
||
Male thoracic sternum (
|
||
<figureCitation id="130A50A89706FFD9FBD6F9C807BAF99B" box="[1084,1163,1629,1653]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Fig. 1B</figureCitation>
|
||
) broad and generally smooth, with scattered granularities, lateral margins setose; sternites 1–4 fused, with traces of sutures between 3–4 sternites; sterno-abdominal cavity deep; press button on sternite 5, near to the suture between sternites 4 and 5.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9706FFD9FCA9F88807E1F83B" blockId="1.[834,1474,1821,2005]" pageId="1" pageNumber="449">
|
||
Male chelae (
|
||
<figureCitation id="130A50A89706FFD9FC36F8880703F8DB" box="[988,1074,1821,1845]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Fig. 1A</figureCitation>
|
||
) distinctly unequal, relatively long, surface punctuated, with scattered rugosities, fingers shorter than palm; distal region of fingers with pointed, chitinous tooth; margin of merus irregularly granular. Carpus rounded, rugose, inner distal angle with well-developed tooth and with 1 or 2 smaller teeth in inner margin.
|
||
</paragraph>
|
||
<caption id="DF4E1CA59706FFD9FF6CF86602E5F7AD" ID-DOI="http://doi.org/10.5281/zenodo.5385670" ID-Zenodo-Dep="5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449" startId="1.[134,169,2035,2057]" targetBox="[135,774,786,2020]" targetPageId="1">
|
||
<paragraph id="8B8E4C2D9706FFD9FF6CF86602E5F7AD" blockId="1.[134,774,2035,2115]" pageId="1" pageNumber="449">
|
||
Fig. 1.
|
||
<taxonomicName id="4C3137AE9706FFD9FF3AF86602DCF7E6" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[208,493,2035,2056]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
|
||
<emphasis id="B945903F9706FFD9FF3AF86602DCF7E6" box="[208,493,2035,2056]" italics="true" pageId="1" pageNumber="449">Sundathelphusa spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2762D449706FFD9FE1FF861011CF7E7" box="[501,557,2036,2057]" pageId="1" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E4C2D9706FFDAFCA9F86801C1F9AA" blockId="1.[835,1474,2045,2101]" lastBlockId="2.[112,753,1452,1604]" lastPageId="2" lastPageNumber="450" pageId="1" pageNumber="449">
|
||
Ambulatory legs (
|
||
<figureCitation id="130A50A89706FFD9FBE0F868076EF7FB" box="[1034,1119,2045,2069]" captionStart="Fig" captionStartId="1.[134,169,2035,2057]" captionTargetBox="[135,774,786,2020]" captionTargetId="figure-528@1.[135,774,786,2020]" captionTargetPageId="1" captionText="Fig. 1. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920). A, dorsal view (Scale bar = 10 mm); B, ventral view; C, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5385670" httpUri="https://zenodo.org/record/5385670/files/figure.png" pageId="1" pageNumber="449">Fig. 1A</figureCitation>
|
||
) sub-triangular in cross section; second, third ambulatory legs longest; anterior dorsal margin of merus (M) smooth; propodi (P) with spines, all dactyli (D) bearing spines; 3M long (length to width ratio 3.3), 3P long (length to width ratio 2.3), 3D long (length to width ratio 6.4); 4M short (length to width ratio 2.9), 4P long (length to width ratio 2), 4D relatively short (length to width ratio 3.8).
|
||
</paragraph>
|
||
<caption id="DF4E1CA59705FFDAFF9BFAB002DFFA9B" ID-DOI="http://doi.org/10.5281/zenodo.5385672" ID-Zenodo-Dep="5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="2" pageNumber="450" startId="2.[113,148,1317,1339]" targetBox="[116,1451,190,1302]" targetPageId="2">
|
||
<paragraph id="8B8E4C2D9705FFDAFF9BFAB002DFFA9B" blockId="2.[113,1452,1317,1397]" pageId="2" pageNumber="450">
|
||
Fig. 2.
|
||
<taxonomicName id="4C3137AE9705FFDAFF50FAB002E6FAD4" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[186,471,1317,1338]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
|
||
<emphasis id="B945903F9705FFDAFF50FAB002E6FAD4" box="[186,471,1317,1338]" italics="true" pageId="2" pageNumber="450">Sundathelphusa spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2762D449705FFDAFE35FAB30126FAD5" box="[479,535,1318,1339]" pageId="2" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E4C2D9705FFDAFF9BF9F901D4F92A" blockId="2.[112,753,1644,1957]" pageId="2" pageNumber="450">
|
||
Male abdomen (
|
||
<figureCitation id="130A50A89705FFDAFECBF9F90242F96A" box="[289,371,1644,1668]" captionStart="Fig" captionStartId="2.[113,148,1317,1339]" captionTargetBox="[116,1451,190,1302]" captionTargetId="figure-317@2.[95,1469,173,1305]" captionTargetPageId="2" captionText="Fig. 2. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="2" pageNumber="450">Fig. 2B</figureCitation>
|
||
) T-shaped; lateral margins covered with short setae; Telson slightly shorter than somite 6, narrow, with lateral margins converging to rounded distal margin.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9705FFDAFF9BF95902D9F84B" blockId="2.[112,753,1644,1957]" pageId="2" pageNumber="450">
|
||
G1 (
|
||
<figureCitation id="130A50A89705FFDAFF75F959023AF90A" box="[159,267,1740,1764]" captionStart="Fig" captionStartId="2.[113,148,1317,1339]" captionTargetBox="[116,1451,190,1302]" captionTargetId="figure-317@2.[95,1469,173,1305]" captionTargetPageId="2" captionText="Fig. 2. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="2" pageNumber="450">Fig. 2D, E</figureCitation>
|
||
) relatively stout, basal part swollen, terminally curved, subterminal segment straight with outer margin distinctly concave; proximal part evenly cylindrical, terminal part occupying about 0.2 times total length, nearly straight, conical, with fairly fluted tip. G2 (
|
||
<figureCitation id="130A50A89705FFDAFE00F8D9010FF88A" box="[490,574,1868,1892]" captionStart="Fig" captionStartId="2.[113,148,1317,1339]" captionTargetBox="[116,1451,190,1302]" captionTargetId="figure-317@2.[95,1469,173,1305]" captionTargetPageId="2" captionText="Fig. 2. Sundathelphusa spelaeophila n. sp., holotype, male (15.9 × 14.1 mm) (MZUF 3920): A, right third maxilliped; B, abdomen; C, right G2, respectively ventral and dorsal view; D, right G1, respectively ventral and dorsal view; E, tip of right G1, respectively ventral and dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5385672" httpUri="https://zenodo.org/record/5385672/files/figure.png" pageId="2" pageNumber="450">Fig. 2C</figureCitation>
|
||
) relatively long, slightly longer than G1, distal segment long, sinuous, about 0.5 times length of basal segment.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9705FFDAFF9BF85802EFF7EA" blockId="2.[113,752,1996,2053]" pageId="2" pageNumber="450">No females were collected, thus it is not possible to give any detail of female morphology.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B1FA69705FFDAFCC7FA3907AEF9EA" pageId="2" pageNumber="450" type="etymology">
|
||
<paragraph id="8B8E4C2D9705FFDAFCC7FA3907AEF9EA" blockId="2.[813,1451,1452,1540]" pageId="2" pageNumber="450">
|
||
<emphasis id="B945903F9705FFDAFCC7FA390089FA2A" bold="true" box="[813,952,1452,1476]" pageId="2" pageNumber="450">Etymology.</emphasis>
|
||
From the composition of
|
||
<emphasis id="B945903F9705FFDAFB13FA390660FA2A" box="[1273,1361,1452,1476]" italics="true" pageId="2" pageNumber="450">spelaeo</emphasis>
|
||
(Latin: spēlaeum, i) cavern and
|
||
<emphasis id="B945903F9705FFDAFBDCFA59075CFA0A" box="[1078,1133,1484,1508]" italics="true" pageId="2" pageNumber="450">phila</emphasis>
|
||
(Latin: -philus, -phila) lover, as it has been collected in a cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B1FA69705FFDBFCC7F9B90192F8C7" lastPageId="3" lastPageNumber="451" pageId="2" pageNumber="450" type="discussion">
|
||
<paragraph id="8B8E4C2D9705FFDAFCC7F9B90696F98A" blockId="2.[813,1451,1580,1636]" pageId="2" pageNumber="450">
|
||
<emphasis id="B945903F9705FFDAFCC7F9B900AAF9AA" bold="true" box="[813,923,1580,1604]" pageId="2" pageNumber="450">Remarks.</emphasis>
|
||
At present, only three species of
|
||
<taxonomicName id="4C3137AE9705FFDAFAEBF9B9069AF9AA" authorityName="Bott" authorityYear="1969" box="[1281,1451,1580,1604]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945903F9705FFDAFAEBF9B9069AF9AA" box="[1281,1451,1580,1604]" italics="true" pageId="2" pageNumber="450">Sundathelphusa</emphasis>
|
||
</taxonomicName>
|
||
are known for
|
||
<collectingRegion id="49F582CF9705FFDAFC25F9D90726F98A" box="[975,1047,1612,1636]" country="Philippines" name="Samar" pageId="2" pageNumber="450">Samar</collectingRegion>
|
||
:
|
||
<taxonomicName id="4C3137AE9705FFDAFBCFF9D8079FF98A" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1061,1198,1612,1636]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9705FFDAFBCFF9D8079FF98A" box="[1061,1198,1612,1636]" italics="true" pageId="2" pageNumber="450">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3137AE9705FFDAFB57F9D80639F98A" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[1213,1288,1612,1636]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="lobo">
|
||
<emphasis id="B945903F9705FFDAFB57F9D80639F98A" box="[1213,1288,1612,1636]" italics="true" pageId="2" pageNumber="450">S. lobo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3137AE9705FFDAFAA9F9D80691F98A" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[1347,1440,1613,1636]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="waray">
|
||
<emphasis id="B945903F9705FFDAFAA9F9D8067FF98A" box="[1347,1358,1613,1636]" italics="true" pageId="2" pageNumber="450">S</emphasis>
|
||
.
|
||
<emphasis id="B945903F9705FFDAFAB7F9D80691F98A" box="[1373,1440,1613,1636]" italics="true" pageId="2" pageNumber="450">waray</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9705FFDAFCC7F9180666F88A" blockId="2.[813,1452,1676,1892]" pageId="2" pageNumber="450">
|
||
However
|
||
<taxonomicName id="4C3137AE9705FFDAFC7DF918070DF94A" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[919,1084,1676,1700]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="spelaeophila">
|
||
<emphasis id="B945903F9705FFDAFC7DF918070DF94A" box="[919,1084,1676,1700]" italics="true" pageId="2" pageNumber="450">S. spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from
|
||
<taxonomicName id="4C3137AE9705FFDAFAC9F918069DF94A" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1315,1452,1676,1700]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9705FFDAFAC9F918069DF94A" box="[1315,1452,1676,1700]" italics="true" pageId="2" pageNumber="450">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
in: (a) the subquadrate shape of the carapace (versus subexagonal, inflated and broader in
|
||
<taxonomicName id="4C3137AE9705FFDAFB37F958065FF90A" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1245,1390,1740,1764]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9705FFDAFB37F958065FF90A" box="[1245,1390,1740,1764]" italics="true" pageId="2" pageNumber="450">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
); (b) poorly defined regions in the carapace (versus well defined regions in
|
||
<taxonomicName id="4C3137AE9705FFDAFC49F898071EF8CA" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[931,1071,1804,1828]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9705FFDAFC49F898071EF8CA" box="[931,1071,1804,1828]" italics="true" pageId="2" pageNumber="450">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
); and (c) G1 subterminal segment almost straight with outer margin distinctly concave (versus curved with outer margin convex in
|
||
<taxonomicName id="4C3137AE9705FFDAFB55F8D80676F88A" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1215,1351,1868,1892]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="B945903F9705FFDAFB55F8D80676F88A" box="[1215,1351,1868,1892]" italics="true" pageId="2" pageNumber="450">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E4C2D9705FFDBFCC7F8190036FAA7" blockId="2.[813,1453,1932,2052]" lastBlockId="3.[136,775,1265,1353]" lastPageId="3" lastPageNumber="451" pageId="2" pageNumber="450">
|
||
The other two species, described from caves,
|
||
<taxonomicName id="4C3137AE9705FFDAFAF5F818064EF84A" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[1311,1407,1933,1956]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="waray">
|
||
<emphasis id="B945903F9705FFDAFAF5F818064EF84A" box="[1311,1407,1933,1956]" italics="true" pageId="2" pageNumber="450">S. waray</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3137AE9705FFDAFCC7F838004BF82A" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[813,890,1964,1988]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="lobo">
|
||
<emphasis id="B945903F9705FFDAFCC7F838004BF82A" box="[813,890,1964,1988]" italics="true" pageId="2" pageNumber="450">S. lobo</emphasis>
|
||
</taxonomicName>
|
||
, are both regarded as obligate cave inhabitants by
|
||
<bibRefCitation id="EFA031DC9705FFDAFCC7F858073AF80A" author="Husana DEM & Naruse T & Kase T" box="[813,1035,1996,2020]" pageId="2" pageNumber="450" pagination="419 - 427" refId="ref3400" refString="Husana DEM, Naruse T & Kase T (2009) Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3): 419 - 427." type="journal article" year="2009">Husana et al. (2009)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C3137AE9705FFDAFBFCF859067EF80A" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[1046,1359,1996,2020]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
|
||
<emphasis id="B945903F9705FFDAFBFCF859067EF80A" box="[1046,1359,1996,2020]" italics="true" pageId="2" pageNumber="450">Sundathelphusa spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2762D449705FFDAFABDF858069DF80A" box="[1367,1452,1997,2020]" pageId="2" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from these two species by the absence of evident troglobitic traits (e.g., reduced cornea pigment; absence of body pigment; length of legs and spines, see
|
||
<bibRefCitation id="EFA031DC9704FFDBFF62FAA70266FAA7" author="Husana DEM & Naruse T & Kase T" box="[136,343,1329,1353]" pageId="3" pageNumber="451" pagination="419 - 427" refId="ref3400" refString="Husana DEM, Naruse T & Kase T (2009) Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3): 419 - 427." type="journal article" year="2009">Husana et al., 2009</bibRefCitation>
|
||
) clearly present in
|
||
<taxonomicName id="4C3137AE9704FFDBFDCEFAA701B3FAA7" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[548,642,1330,1353]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="waray">
|
||
<emphasis id="B945903F9704FFDBFDCEFAA701B3FAA7" box="[548,642,1330,1353]" italics="true" pageId="3" pageNumber="451">S. waray</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B945903F9704FFDBFD5CFAA70036FAA7" box="[694,775,1329,1353]" italics="true" pageId="3" pageNumber="451">
|
||
<taxonomicName id="4C3137AE9704FFDBFD5CFAA70035FAA7" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[694,772,1329,1353]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="lobo">S. lobo</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DF4E1CA59704FFDBFF62FBED01BBFB44" ID-DOI="http://doi.org/10.5281/zenodo.5385674" ID-Zenodo-Dep="5385674" httpUri="https://zenodo.org/record/5385674/files/figure.png" pageId="3" pageNumber="451" startId="3.[136,171,1144,1166]" targetBox="[136,1475,188,1128]" targetPageId="3">
|
||
<paragraph id="8B8E4C2D9704FFDBFF62FBED01BBFB44" blockId="3.[136,1474,1144,1195]" pageId="3" pageNumber="451">
|
||
Fig. 3. Map of the localities of the four
|
||
<taxonomicName id="4C3137AE9704FFDBFDF2FBED01CEFB60" box="[536,767,1144,1166]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B945903F9704FFDBFDF2FBED0180FB63" box="[536,689,1144,1165]" italics="true" pageId="3" pageNumber="451">Sundathelphusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
collected in western Samar: ▲
|
||
<taxonomicName id="4C3137AE9704FFDBFBD5FBED07E5FB63" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[1087,1236,1144,1165]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
|
||
<emphasis id="B945903F9704FFDBFBD5FBED07E5FB63" box="[1087,1236,1144,1165]" italics="true" pageId="3" pageNumber="451">S. spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2762D449704FFDBFB36FBEC0625FB60" box="[1244,1300,1145,1166]" pageId="3" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="B945903F9704FFDBFACBFBED068DFB63" box="[1313,1468,1144,1165]" italics="true" pageId="3" pageNumber="451">
|
||
<emphasis id="B945903F9704FFDBFACBFBED0607FB63" bold="true" box="[1313,1334,1144,1165]" italics="true" pageId="3" pageNumber="451">«</emphasis>
|
||
<taxonomicName id="4C3137AE9704FFDBFAD4FBED068DFB63" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1342,1468,1144,1165]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="philippina">S. philippina</taxonomicName>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B945903F9704FFDBFF62FB0103ACFB44" bold="true" box="[136,157,1172,1194]" pageId="3" pageNumber="451">È</emphasis>
|
||
<taxonomicName id="4C3137AE9704FFDBFF4FFB0003CAFB45" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[165,251,1173,1195]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="waray">
|
||
<emphasis id="B945903F9704FFDBFF4FFB0003CAFB45" box="[165,251,1173,1195]" italics="true" pageId="3" pageNumber="451">S. waray</emphasis>
|
||
</taxonomicName>
|
||
(Langun cave); ■
|
||
<taxonomicName id="4C3137AE9704FFDBFE2AFB000134FB44" authorityName="Husana, Naruse & Kase" authorityYear="2009" box="[448,517,1173,1194]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="lobo">
|
||
<emphasis id="B945903F9704FFDBFE2AFB000134FB44" box="[448,517,1173,1194]" italics="true" pageId="3" pageNumber="451">S. lobo</emphasis>
|
||
</taxonomicName>
|
||
(Lobo cave).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E4C2D9704FFDBFF62FAE40192F8C7" blockId="3.[134,775,1393,1833]" pageId="3" pageNumber="451">
|
||
<taxonomicName id="4C3137AE9704FFDBFF62FAE402D3FA67" authority="Stasolla & Abbarchi & Innocenti, 2015" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[136,482,1393,1417]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="spelaeophila" status="sp. nov.">
|
||
<emphasis id="B945903F9704FFDBFF62FAE402D3FA67" box="[136,482,1393,1417]" italics="true" pageId="3" pageNumber="451">Sundathelphusa spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2762D449704FFDBFE04FAE7017CFA67" box="[494,589,1394,1417]" pageId="3" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can apparently resemble
|
||
<taxonomicName id="4C3137AE9704FFDBFF1BFA07019BFA47" authority="(H Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[241,682,1425,1449]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9704FFDBFF1BFA0702B5FA47" box="[241,388,1425,1449]" italics="true" pageId="3" pageNumber="451">S. grapsoides</emphasis>
|
||
(H
|
||
<bibRefCitation id="EFA031DC9704FFDBFE58FA040190FA47" author="Milne Edwards H" box="[434,673,1425,1449]" pageId="3" pageNumber="451" pagination="163 - 228" refId="ref3750" refString="Milne Edwards H (1853) Memoire sur la famille des Ocypodides. Annales des Sciences Naturelles, (3) 20: 163 - 228." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Even if this species is reported from Luzon (
|
||
<collectingRegion id="49F582CF9704FFDBFDECFA240146FA27" box="[518,631,1457,1481]" country="Philippines" name="Pampanga" pageId="3" pageNumber="451">Pampanga</collectingRegion>
|
||
and
|
||
<collectingRegion id="49F582CF9704FFDBFD42FA240032FA27" box="[680,771,1457,1481]" country="Philippines" name="Bulacan" pageId="3" pageNumber="451">Bulacan</collectingRegion>
|
||
;
|
||
<bibRefCitation id="EFA031DC9704FFDBFF6DFA4402A4FA07" author="Mendoza JCE & Naruse T" box="[135,405,1489,1513]" pageId="3" pageNumber="451" pagination="61 - 70" refId="ref3664" refString="Mendoza JCE & Naruse T (2010) A new species of riverine crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from Northeastern Luzon, Philippines. Philippine Journal of Science, 139 (1): 61 - 70." type="journal article" year="2010">Mendoza & Naruse, 2010</bibRefCitation>
|
||
), an island quite distant from
|
||
<collectingRegion id="49F582CF9704FFDBFD28FA440034FA07" box="[706,773,1489,1513]" country="Philippines" name="Samar" pageId="3" pageNumber="451">Samar</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFA031DC9704FFDBFF64FA640266F9E7" author="Ng PKL & Sket B" box="[142,343,1521,1545]" pageId="3" pageNumber="451" pagination="695 - 706" refId="ref3925" refString="Ng PKL & Sket B (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On a collection of Parathelphusidae from Bohol. Proceedings of the Biological Society of Washington, 109 (4): 695 - 706." type="journal article" year="1996">Ng & Sket, 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA031DC9704FFDBFE8CFA6702E5F9E7" author="Ng PKL" box="[358,468,1521,1546]" pageId="3" pageNumber="451" pagination="561 - 571" refId="ref3818" refString="Ng PKL (2010) On the identity of Para-Bary-Thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). In: Fransen CHJM, De Grave S & Ng PKL (eds.) Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume, Crustaceana Monographs, Vol. 14. Brill, Netherlands. Pp. 561 - 571." type="book chapter" year="2010">Ng, 2010</bibRefCitation>
|
||
), it also shows remarkable morphological differences.
|
||
<taxonomicName id="4C3137AE9704FFDBFE2AF9840037F9C7" authorityName="Stasolla & Abbarchi & Innocenti" authorityYear="2015" box="[448,774,1553,1577]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="spelaeophila">
|
||
<emphasis id="B945903F9704FFDBFE2AF9840037F9C7" box="[448,774,1553,1577]" italics="true" pageId="3" pageNumber="451">Sundathelphusa spelaeophila</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3137AE9704FFDBFEE4F9A702AFF9A7" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[270,414,1585,1609]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9704FFDBFEE4F9A702AFF9A7" box="[270,414,1585,1609]" italics="true" pageId="3" pageNumber="451">S. grapsoides</emphasis>
|
||
</taxonomicName>
|
||
descriptions reported in A. Milne Edwards (1869) and
|
||
<bibRefCitation id="EFA031DC9704FFDBFE8FF9C4013CF987" author="Rathbun MJ" box="[357,525,1617,1641]" pageId="3" pageNumber="451" pagination="225 - 312" refId="ref4075" refString="Rathbun MJ (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 4 e serie, 6: 225 - 312." type="journal article" year="1904">Rathbun (1904)</bibRefCitation>
|
||
by: (a) the subquadrate shape of the carapace (versus rounded shape of the carapace in
|
||
<taxonomicName id="4C3137AE9704FFDBFF4CF9070209F947" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[166,312,1681,1705]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9704FFDBFF4CF9070209F947" box="[166,312,1681,1705]" italics="true" pageId="3" pageNumber="451">S. grapsoides</emphasis>
|
||
</taxonomicName>
|
||
); (b) the anterior side of the merus of the ambulatory legs smooth (versus numerous teeth in the anterior side of the merus of the ambulatory legs in
|
||
<taxonomicName id="4C3137AE9704FFDBFD8DF94701C7F907" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[615,758,1745,1769]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9704FFDBFD8DF94701C7F907" box="[615,758,1745,1769]" italics="true" pageId="3" pageNumber="451">S. grapsoides</emphasis>
|
||
</taxonomicName>
|
||
); and (c) poorly defined regions of the carapace (versus well defined regions of the carapace in
|
||
<taxonomicName id="4C3137AE9704FFDBFDE8F88701A2F8C7" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[514,659,1809,1833]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="grapsoides">
|
||
<emphasis id="B945903F9704FFDBFDE8F88701A2F8C7" box="[514,659,1809,1833]" italics="true" pageId="3" pageNumber="451">S. grapsoides</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |