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<document id="B18B3B9C2EBCE32D1DB19824E88CAC5D" ID-DOI="10.11646/zootaxa.5415.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10695722" ID-ZooBank="DFD45E44-A4BA-4F42-B859-3C1F35BF4695" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708678264697" checkinUser="plazi" docAuthor="Lü, Liang" docDate="2024" docId="03FBCB12FFA63155FF47F803FB4A6B00" docLanguage="en" docName="zootaxa.5415.4.9.pdf" docOrigin="Zootaxa 5415 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxytelus acriculiclypeatus Lü 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="597" masterDocId="FFC2B36AFFA63151FFD0FFACFF896A1D" masterDocTitle="Oxytelus acriculiclypeatus, a New Species of Oxytelus Gravenhorst, 1802 (Coleoptera: Staphylinidae: Oxytelinae) from Borneo" masterLastPageNumber="597" masterPageNumber="593" pageNumber="593" updateTime="1708678633768" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EEDA378C8D0A5251171DA049876F6A07" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2A9FF527D440FB5FAAB89F777F716EBB">
<mods:title id="069D9862BC6AA146C9C8F1B2EB48F73B">Oxytelus acriculiclypeatus, a New Species of Oxytelus Gravenhorst, 1802 (Coleoptera: Staphylinidae: Oxytelinae) from Borneo</mods:title>
</mods:titleInfo>
<mods:name id="3F42D50138A7CE01A25E2DE33B8F6160" type="personal">
<mods:role id="8A5BF84CC3E48D24043B67194BDD4EC1">
<mods:roleTerm id="CF80400CF92134865CECF762E646D300">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2530F64559D02C1614790C86EDA6F622">Lü, Liang</mods:namePart>
<mods:nameIdentifier id="086CD9A60D949D0095AB5594964DC973" type="ORCID">0000-0001-7533-6244</mods:nameIdentifier>
<mods:affiliation id="E7145965F7FD3B52E6A7614D961AF9BA">Hebei Collaborative Innovation Center for Eco-Environment, College of Life Sciences, Hebei Normal University, Shijiazhuang, Hebei Province 050024, China.</mods:affiliation>
<mods:nameIdentifier id="8A54EB1C917DB02F7800A4553D9A143E" type="email">lianges.luex@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="DA5CC8C9A1DBA18AF93AE480EB0536F0">text</mods:typeOfResource>
<mods:relatedItem id="C54C0ACDE30F6C05B167B76F05BFF043" type="host">
<mods:titleInfo id="3A3C2CCC7B35AD75D99782F89224F15D">
<mods:title id="1E1718DA228CFACE9D33FA2856C9896D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="64FFB658615C54282F469DA8759028B4">
<mods:date id="6BF4C7E67DBD53553046B000C69FE557">2024</mods:date>
<mods:detail id="3D537B9557D8B1316BC533EC897874C4" type="pubDate">
<mods:number id="56870905557D26A19BEC03A3EB1DB38F">2024-02-23</mods:number>
</mods:detail>
<mods:detail id="44924CEC84DB70B60271D113D43993A8" type="volume">
<mods:number id="C50860D06EC10CBB4BFC070137A2E93F">5415</mods:number>
</mods:detail>
<mods:detail id="20248349AC76B53D310B44CEABD897E4" type="issue">
<mods:number id="A3D4B6BAA76030D64C9E9599FDDF5C14">4</mods:number>
</mods:detail>
<mods:extent id="25A79F9CB7125D0B45FE5536CA5F8A2C" unit="page">
<mods:start id="E890A7080356E9F68BB2051105075EEE">593</mods:start>
<mods:end id="A02AAF804EAC99DCFF2350574A98126A">597</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="41C54503D6E4ADC4515EE9D5E8F0FB22">
<mods:url id="8F006172DE748DF70BDF96C0C1FAF617">http://dx.doi.org/10.11646/zootaxa.5415.4.9</mods:url>
</mods:location>
<mods:classification id="CE8A409D6849564C97F83E561E905C2A">journal article</mods:classification>
<mods:identifier id="D76B3B68654B4148968407BCAF07FAAC" type="DOI">10.11646/zootaxa.5415.4.9</mods:identifier>
<mods:identifier id="E2E5ED5DC8EDBF38BA917C6BB61A6384" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="577E4462EB34D0966041CC5E4619E50A" type="Zenodo-Dep">10695722</mods:identifier>
<mods:identifier id="49020D19721F490F110DA3E75860C755" type="ZooBank">DFD45E44-A4BA-4F42-B859-3C1F35BF4695</mods:identifier>
</mods:mods>
<treatment id="03FBCB12FFA63155FF47F803FB4A6B00" LSID="urn:lsid:plazi:treatment:03FBCB12FFA63155FF47F803FB4A6B00" httpUri="http://treatment.plazi.org/id/03FBCB12FFA63155FF47F803FB4A6B00" lastPageId="4" lastPageNumber="597" pageId="0" pageNumber="593">
<subSubSection id="C348298FFFA63151FF47F803FDB26DD4" box="[151,571,1967,1994]" pageId="0" pageNumber="593" type="nomenclature">
<paragraph id="8BED7A04FFA63151FF47F803FDB26DD4" blockId="0.[151,571,1967,2029]" box="[151,571,1967,1994]" pageId="0" pageNumber="593">
<heading id="D0A5CD68FFA63151FF47F803FDB26DD4" bold="true" box="[151,571,1967,1994]" fontSize="11" level="1" pageId="0" pageNumber="593" reason="1">
<emphasis id="B926A616FFA63151FF47F803FDB26DD4" bold="true" box="[151,571,1967,1994]" pageId="0" pageNumber="593">
<taxonomicName id="4C520187FFA63151FF47F803FE5C6DD7" authority="Lü, 2024" authorityName="Lü" authorityYear="2024" box="[151,469,1967,1994]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="593" phylum="Arthropoda" rank="species" species="acriculiclypeatus" status="sp. nov.">
<emphasis id="B926A616FFA63151FF47F803FE5C6DD7" bold="true" box="[151,469,1967,1994]" italics="true" pageId="0" pageNumber="593">Oxytelus acriculiclypeatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2151B6DFFA63151FE0DF803FDB26DD4" box="[477,571,1967,1993]" pageId="0" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA63151FF47F87FFE9B6DF0" box="[151,274,2003,2029]" pageId="0" pageNumber="593" type="description">
<paragraph id="8BED7A04FFA63151FF47F87FFE9B6DF0" blockId="0.[151,571,1967,2029]" box="[151,274,2003,2029]" pageId="0" pageNumber="593">
(
<figureCitation id="13696681FFA63151FF4FF87FFE806DF0" box="[159,265,2003,2029]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1922,1946]" captionStartId-1="3.[151,250,1626,1650]" captionTargetBox-0="[163,1426,190,1885]" captionTargetBox-1="[163,1423,193,1590]" captionTargetId-0="figure-26@2.[151,1435,179,1896]" captionTargetId-1="figure-16@3.[151,1435,181,1602]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." captionText-1="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10695724" figureDoi-1="http://doi.org/10.5281/zenodo.10695726" httpUri-0="https://zenodo.org/record/10695724/files/figure.png" httpUri-1="https://zenodo.org/record/10695726/files/figure.png" pageId="0" pageNumber="593">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA73150FF47FF3AFAD56B01" pageId="1" pageNumber="594" type="materials_examined">
<paragraph id="8BED7A04FFA73150FF47FF3AFAD56B01" blockId="1.[151,1436,150,717]" pageId="1" pageNumber="594">
<emphasis id="B926A616FFA73150FF47FF3AFECC6AAD" bold="true" box="[151,325,150,176]" pageId="1" pageNumber="594">Type material.</emphasis>
<materialsCitation id="3B3A7059FFA73150FE9BFF3AFC626AC9" collectingDate="1999-06" collectionCode="BMNH" collectorName="H. Mendel" country="Malaysia" latitude="4.9666667" location="Danum Valley" longLatPrecision="1306" longitude="117.78333" municipality="Borneo" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B926A616FFA73150FE9BFF3AFE316AAD" bold="true" box="[331,440,150,176]" pageId="1" pageNumber="594">
<typeStatus id="54E9C4A6FFA73150FE9BFF3AFE316AAD" box="[331,440,150,176]" pageId="1" pageNumber="594" type="holotype">Holotype</typeStatus>
</emphasis>
: ♁,
<emphasis id="B926A616FFA73150FE39FF3AFDF66AAD" bold="true" box="[489,639,150,176]" pageId="1" pageNumber="594">
<collectingCountry id="F3453A94FFA73150FE39FF3AFDF66AAD" box="[489,639,150,176]" name="Malaysia" pageId="1" pageNumber="594">MALAYSIA</collectingCountry>
</emphasis>
,
<collectingMunicipality id="6B89E07EFFA73150FD5CFF3BFD576AAC" box="[652,734,151,177]" pageId="1" pageNumber="594">Borneo</collectingMunicipality>
<collectingRegion id="4996B4E6FFA73150FD35FF3AFCA56AAD" box="[741,812,150,176]" country="Malaysia" name="Sabah" pageId="1" pageNumber="594">Sabah</collectingRegion>
,
<location id="8E8D2CDFFFA73150FCE7FF3BFC506AAD" LSID="urn:lsid:plazi:treatment:03FBCB12FFA63155FF47F803FB4A6B00:8E8D2CDFFFA73150FCE7FF3BFC506AAD" box="[823,985,150,177]" country="Malaysia" latitude="4.9666667" longLatPrecision="1306" longitude="117.78333" municipality="Borneo" name="Danum Valley" pageId="1" pageNumber="594" stateProvince="Sabah">Danum Valley</location>
,
<geoCoordinate id="EE661CC3FFA73150FC33FF34FBBA6AAE" box="[995,1075,152,179]" degrees="4" direction="north" minutes="58" orientation="latitude" pageId="1" pageNumber="594" precision="925" value="4.9666667">4°58′N</geoCoordinate>
<geoCoordinate id="EE661CC3FFA73150FBE9FF34FB2A6AAE" box="[1081,1187,152,179]" degrees="117" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="594" precision="925" value="117.78333">117°47′E</geoCoordinate>
,
<collectingMethod id="52130213FFA73150FB7EFF3AFA116AAC" box="[1198,1432,150,177]" pageId="1" pageNumber="594">Flight Intercept Trap</collectingMethod>
,
<date id="FFEC5CC4FFA73150FF47FF17FE836AC9" box="[151,266,186,213]" pageId="1" pageNumber="594" value="1999-06">
<collectingDate id="EFA8A52CFFA73150FF47FF17FE836AC9" box="[151,266,186,213]" pageId="1" pageNumber="594" value="1999-06">June 1999</collectingDate>
</date>
//
<collectionCode id="ED43E2C1FFA73150FEF6FF16FEF66AC9" box="[294,383,186,212]" pageId="1" pageNumber="594">BMNH</collectionCode>
(E), 2005-177,
<collectorName id="26A71FD2FFA73150FDFAFF17FD2A6AC9" box="[554,675,186,213]" pageId="1" pageNumber="594">H. Mendel</collectorName>
//
<taxonomicName id="4C520187FFA73150FD6EFF16FC626AC9" authorityName="Lü" authorityYear="2024" box="[702,1003,186,212]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="acriculiclypeatus">Oxytelus acriculiclypeatus</taxonomicName>
</materialsCitation>
♁
<materialsCitation id="3B3A7059FFA73150FBC3FF16FE886AE5" collectionCode="LU, BMNH" country="Malaysia" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54E9C4A6FFA73150FBC3FF16FB2E6AC9" box="[1043,1191,186,212]" pageId="1" pageNumber="594" type="holotype">HOLOTYPE</typeStatus>
det.
<collectionCode id="ED43E2C1FFA73150FB33FF10FA816ACA" box="[1251,1288,188,215]" country="Russia" name="St. Petersburg University" pageId="1" pageNumber="594">LÜ</collectionCode>
Liang, 2024 (
<collectionCode id="ED43E2C1FFA73150FF4FFF72FF706AE5" box="[159,249,222,248]" pageId="1" pageNumber="594">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3A7059FFA73150FEDFFF72FCC66B01" collectingDate="1999-06" collectionCode="BMNH" collectorName="H. Mendel" country="Malaysia" latitude="4.9666667" location="Danum Valley" longLatPrecision="1306" longitude="117.78333" municipality="Borneo" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B926A616FFA73150FEDFFF72FE0E6AE5" bold="true" box="[271,391,222,248]" pageId="1" pageNumber="594">
<typeStatus id="54E9C4A6FFA73150FEDFFF72FE0E6AE5" box="[271,391,222,248]" pageId="1" pageNumber="594" type="paratype">Paratypes</typeStatus>
</emphasis>
: 2♁♁,
<collectingMunicipality id="6B89E07EFFA73150FE32FF73FDBD6AE4" box="[482,564,223,249]" pageId="1" pageNumber="594">Borneo</collectingMunicipality>
<collectingRegion id="4996B4E6FFA73150FDEEFF72FD0C6AE5" box="[574,645,222,248]" country="Malaysia" name="Sabah" pageId="1" pageNumber="594">Sabah</collectingRegion>
,
<location id="8E8D2CDFFFA73150FD44FF73FCB16AE5" LSID="urn:lsid:plazi:treatment:03FBCB12FFA63155FF47F803FB4A6B00:8E8D2CDFFFA73150FD44FF73FCB16AE5" box="[660,824,222,249]" country="Malaysia" latitude="4.9666667" longLatPrecision="1306" longitude="117.78333" municipality="Borneo" name="Danum Valley" pageId="1" pageNumber="594" stateProvince="Sabah">Danum Valley</location>
,
<geoCoordinate id="EE661CC3FFA73150FC96FF4CFC1F6AE6" box="[838,918,224,251]" degrees="4" direction="north" minutes="58" orientation="latitude" pageId="1" pageNumber="594" precision="925" value="4.9666667">4°58′N</geoCoordinate>
<geoCoordinate id="EE661CC3FFA73150FC70FF4CFB806AE6" box="[928,1033,224,251]" degrees="117" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="594" precision="925" value="117.78333">117°47′E</geoCoordinate>
,
<collectingMethod id="52130213FFA73150FBC8FF72FA806AE4" box="[1048,1289,222,249]" pageId="1" pageNumber="594">Flight Intercept Trap</collectingMethod>
,
<date id="FFEC5CC4FFA73150FAC7FF73FA026AE5" box="[1303,1419,222,249]" pageId="1" pageNumber="594" value="1999-06">
<collectingDate id="EFA8A52CFFA73150FAC7FF73FA026AE5" box="[1303,1419,222,249]" pageId="1" pageNumber="594" value="1999-06">June 1999</collectingDate>
</date>
//
<collectionCode id="ED43E2C1FFA73150FF47FEAEFF796B01" box="[151,240,258,284]" pageId="1" pageNumber="594">BMNH</collectionCode>
(E), 2005-177,
<collectorName id="26A71FD2FFA73150FE46FEAFFD856B01" box="[406,524,258,285]" pageId="1" pageNumber="594">H. Mendel</collectorName>
//
<taxonomicName id="4C520187FFA73150FDF5FEAEFCC66B01" authorityName="Lü" authorityYear="2024" box="[549,847,258,284]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="acriculiclypeatus">Oxytelus acriculiclypeatus</taxonomicName>
</materialsCitation>
♁
<materialsCitation id="3B3A7059FFA73150FCA1FEAEFBD06B02" box="[881,1113,258,287]" collectionCode="LU" country="Malaysia" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54E9C4A6FFA73150FCA1FEAEFC756B01" box="[881,1020,258,284]" pageId="1" pageNumber="594" type="paratype">PARATYPE</typeStatus>
det.
<collectionCode id="ED43E2C1FFA73150FBE4FEA8FBD06B02" box="[1076,1113,260,287]" country="Russia" name="St. Petersburg University" pageId="1" pageNumber="594">LÜ</collectionCode>
</materialsCitation>
Liang, 2024 (
<collectionCode id="ED43E2C1FFA73150FB26FEAEFAD96B01" box="[1270,1360,258,284]" pageId="1" pageNumber="594">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA73152FF17FE8AFE166D3A" lastPageId="3" lastPageNumber="596" pageId="1" pageNumber="594" type="description">
<paragraph id="8BED7A04FFA73150FF17FE8AFA9A6B78" blockId="1.[151,1436,150,717]" pageId="1" pageNumber="594">
<emphasis id="B926A616FFA73150FF17FE8AFED06B5D" bold="true" box="[199,345,294,320]" pageId="1" pageNumber="594">Description.</emphasis>
Body (
<figureCitation id="13696681FFA73150FE7EFE8AFE666B5D" box="[430,495,294,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1922,1946]" captionTargetBox="[163,1426,190,1885]" captionTargetId="figure-26@2.[151,1435,179,1896]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="1" pageNumber="594">Fig. 1</figureCitation>
) medium-sized, glossy, dark brown to blackish. First two antennomeres, maxillary palpi, and legs yellowish brown and elytra brown and vaguely translucent. Body length about
<quantity id="4CAAD7E1FFA73150FB66FEE6FA866B78" box="[1206,1295,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="594" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
<paragraph id="8BED7A04FFA73150FF17FEC2FCB568D0" blockId="1.[151,1436,150,717]" pageId="1" pageNumber="594">
<emphasis id="B926A616FFA73150FF17FEC2FE886B95" box="[199,257,366,392]" italics="true" pageId="1" pageNumber="594">Male</emphasis>
. Head (
<figureCitation id="13696681FFA73150FE8EFEC2FE276B95" box="[350,430,366,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1922,1946]" captionTargetBox="[163,1426,190,1885]" captionTargetId="figure-26@2.[151,1435,179,1896]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="1" pageNumber="594">Figs. 1</figureCitation>
&amp;
<figureCitation id="13696681FFA73150FE07FEC2FE726B95" box="[471,507,366,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1626,1650]" captionTargetBox="[163,1423,193,1590]" captionTargetId="figure-16@3.[151,1435,181,1602]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="1" pageNumber="594">2A</figureCitation>
) sub-pentagonal, widest at eyes or temples (accurately at temples). Disc bearing scarce setae and covered with large punctures. Clypeus elongated triangularly, protruding beyond anterior margin of supra-antennal ridges, as long as 1/4–1/3 head length, slightly depressed in central part, surface glabrous and occasionally and slightly coriaceous; anterior margin narrow and pointed in middle. Epistomal suture with lateral portions lightly incurved and running posteriorly to level of anterior margin of eyes. Vertex slightly convex but flat and feebly coriaceous near clypeus, posterior part limited (
<typeStatus id="54E9C4A6FFA73150FCEDFD8EFC2B6821" box="[829,930,546,572]" pageId="1" pageNumber="594" type="holotype">holotype</typeStatus>
) or not (
<typeStatus id="54E9C4A6FFA73150FBDAFD8EFBF06821" box="[1034,1145,546,572]" pageId="1" pageNumber="594" type="paratype">paratypes</typeStatus>
) in median portion; midlongitudinal suture indistinct (
<typeStatus id="54E9C4A6FFA73150FE20FDEAFDDC687D" box="[496,597,582,608]" pageId="1" pageNumber="594" type="holotype">holotype</typeStatus>
) or missing (
<typeStatus id="54E9C4A6FFA73150FD20FDEAFCEB687D" box="[752,866,582,608]" pageId="1" pageNumber="594" type="paratype">paratypes</typeStatus>
); paralateral sutures vague. Eyes with fine facets, convex, and shorter than temples. Temples slightly rugose, posterolaterally dilated. Occipital suture with middle portion distinct (in
<typeStatus id="54E9C4A6FFA73150FEBCFD22FE5B68B5" box="[364,466,654,680]" pageId="1" pageNumber="594" type="holotype">holotype</typeStatus>
), punctately connected (in
<specimenCount id="9D54B18DFFA73150FD2BFD23FC0668B5" box="[763,911,654,681]" count="1" pageId="1" pageNumber="594" type="generic" typeStatus="paratype">one paratype</specimenCount>
), or completely missing (in the other
<typeStatus id="54E9C4A6FFA73150FAFBFD22FA1868B5" box="[1323,1425,654,680]" pageId="1" pageNumber="594" type="paratype">paratype</typeStatus>
); nuchal ridge interrupted in middle, dorsobasal ridge present.
</paragraph>
<caption id="DF2D2A8CFFA73150FF47FD56FBD96908" box="[151,1104,762,789]" pageId="1" pageNumber="594" startId="1.[151,244,762,788]" targetType="table">
<paragraph id="8BED7A04FFA73150FF47FD56FBD96908" blockId="1.[151,1104,762,789]" box="[151,1104,762,789]" pageId="1" pageNumber="594">
<emphasis id="B926A616FFA73150FF47FD56FE986909" bold="true" box="[151,273,762,788]" pageId="1" pageNumber="594">TABLE 1.</emphasis>
Measurements (in millimeters) of the types of
<taxonomicName id="4C520187FFA73150FCCFFD56FBC06909" authorityName="Lü" authorityYear="2024" box="[799,1097,762,788]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="acriculiclypeatus">
<emphasis id="B926A616FFA73150FCCFFD56FBC06909" box="[799,1097,762,788]" italics="true" pageId="1" pageNumber="594">Oxytelus acriculiclypeatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BED7A04FFA73150FCC4FC88FABC6F6D" pageId="1" pageNumber="594">
<table id="F95288A4FFA7CEAEFF73FC88FAC66F6D" box="[163,1359,804,1392]" gridcols="4" gridrows="15" pageId="1" pageNumber="594">
<tr id="35627846FFA7CEAEFF73FC88FAC66920" box="[163,1359,804,829]" gridrow="0" pageId="1" pageNumber="594" rowspan-0="1">
<th id="76B3113AFFA7CEAEFCC4FC88FCFB6920" box="[788,882,804,829]" gridcol="1" gridrow="0" pageId="1" pageNumber="594">Holotype</th>
<th id="76B3113AFFA7CEAEFC3EFC88FBD36920" box="[1006,1114,804,829]" gridcol="2" gridrow="0" pageId="1" pageNumber="594">Paratype 1</th>
<th id="76B3113AFFA7CEAEFB33FC88FAC66920" box="[1251,1359,804,829]" gridcol="3" gridrow="0" pageId="1" pageNumber="594">Paratype 2</th>
</tr>
<tr id="35627846FFA7CEAEFF73FCE0FAC66978" box="[163,1359,844,869]" gridrow="1" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FCE0FD9C6978" box="[163,533,844,869]" gridcol="0" gridrow="1" pageId="1" pageNumber="594">Body length</th>
<td id="76B3113AFFA7CEAEFCC4FCE0FCFB6978" box="[788,882,844,869]" gridcol="1" gridrow="1" pageId="1" pageNumber="594">4.594</td>
<td id="76B3113AFFA7CEAEFC3EFCE0FBD36978" box="[1006,1114,844,869]" gridcol="2" gridrow="1" pageId="1" pageNumber="594">4.307</td>
<td id="76B3113AFFA7CEAEFB33FCE0FAC66978" box="[1251,1359,844,869]" gridcol="3" gridrow="1" pageId="1" pageNumber="594">4.452</td>
</tr>
<tr id="35627846FFA7CEAEFF73FCD9FAC66990" box="[163,1359,885,909]" gridrow="2" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FCD9FD9C6990" box="[163,533,885,909]" gridcol="0" gridrow="2" pageId="1" pageNumber="594">Forebody length</th>
<td id="76B3113AFFA7CEAEFCC4FCD9FCFB6990" box="[788,882,885,909]" gridcol="1" gridrow="2" pageId="1" pageNumber="594">2.183</td>
<td id="76B3113AFFA7CEAEFC3EFCD9FBD36990" box="[1006,1114,885,909]" gridcol="2" gridrow="2" pageId="1" pageNumber="594">2.140</td>
<td id="76B3113AFFA7CEAEFB33FCD9FAC66990" box="[1251,1359,885,909]" gridcol="3" gridrow="2" pageId="1" pageNumber="594">2.337</td>
</tr>
<tr id="35627846FFA7CEAEFF73FC31FAC669A8" box="[163,1359,925,949]" gridrow="3" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FC31FD9C69A8" box="[163,533,925,949]" gridcol="0" gridrow="3" pageId="1" pageNumber="594">Head length</th>
<td id="76B3113AFFA7CEAEFCC4FC31FCFB69A8" box="[788,882,925,949]" gridcol="1" gridrow="3" pageId="1" pageNumber="594">0.625</td>
<td id="76B3113AFFA7CEAEFC3EFC31FBD369A8" box="[1006,1114,925,949]" gridcol="2" gridrow="3" pageId="1" pageNumber="594">0.800</td>
<td id="76B3113AFFA7CEAEFB33FC31FAC669A8" box="[1251,1359,925,949]" gridcol="3" gridrow="3" pageId="1" pageNumber="594">0.862</td>
</tr>
<tr id="35627846FFA7CEAEFF73FC69FAC669C0" box="[163,1359,965,989]" gridrow="4" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FC69FD9C69C0" box="[163,533,965,989]" gridcol="0" gridrow="4" pageId="1" pageNumber="594">Head width</th>
<td id="76B3113AFFA7CEAEFCC4FC69FCFB69C0" box="[788,882,965,989]" gridcol="1" gridrow="4" pageId="1" pageNumber="594">0.849</td>
<td id="76B3113AFFA7CEAEFC3EFC69FBD369C0" box="[1006,1114,965,989]" gridcol="2" gridrow="4" pageId="1" pageNumber="594">0.824</td>
<td id="76B3113AFFA7CEAEFB33FC69FAC669C0" box="[1251,1359,965,989]" gridcol="3" gridrow="4" pageId="1" pageNumber="594">0.871</td>
</tr>
<tr id="35627846FFA7CEAEFF73FC41FAC66E1B" box="[163,1359,1005,1030]" gridrow="5" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FC41FD9C6E1B" box="[163,533,1005,1030]" gridcol="0" gridrow="5" pageId="1" pageNumber="594">Eye length</th>
<td id="76B3113AFFA7CEAEFCC4FC41FCFB6E1B" box="[788,882,1005,1030]" gridcol="1" gridrow="5" pageId="1" pageNumber="594">0.232</td>
<td id="76B3113AFFA7CEAEFC3EFC41FBD36E1B" box="[1006,1114,1005,1030]" gridcol="2" gridrow="5" pageId="1" pageNumber="594">0.222</td>
<td id="76B3113AFFA7CEAEFB33FC41FAC66E1B" box="[1251,1359,1005,1030]" gridcol="3" gridrow="5" pageId="1" pageNumber="594">0.234</td>
</tr>
<tr id="35627846FFA7CEAEFF73FBBAFAC66E33" box="[163,1359,1046,1070]" gridrow="6" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FBBAFD9C6E33" box="[163,533,1046,1070]" gridcol="0" gridrow="6" pageId="1" pageNumber="594">Temple length</th>
<td id="76B3113AFFA7CEAEFCC4FBBAFCFB6E33" box="[788,882,1046,1070]" gridcol="1" gridrow="6" pageId="1" pageNumber="594">0.294</td>
<td id="76B3113AFFA7CEAEFC3EFBBAFBD36E33" box="[1006,1114,1046,1070]" gridcol="2" gridrow="6" pageId="1" pageNumber="594">0.294</td>
<td id="76B3113AFFA7CEAEFB33FBBAFAC66E33" box="[1251,1359,1046,1070]" gridcol="3" gridrow="6" pageId="1" pageNumber="594">0.318</td>
</tr>
<tr id="35627846FFA7CEAEFF73FB92FAC66E4B" box="[163,1359,1086,1110]" gridrow="7" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FB92FD9C6E4B" box="[163,533,1086,1110]" gridcol="0" gridrow="7" pageId="1" pageNumber="594">Supra-antennal ridge vertex distance</th>
<td id="76B3113AFFA7CEAEFCC4FB92FCFB6E4B" box="[788,882,1086,1110]" gridcol="1" gridrow="7" pageId="1" pageNumber="594">0.482</td>
<td id="76B3113AFFA7CEAEFC3EFB92FBD36E4B" box="[1006,1114,1086,1110]" gridcol="2" gridrow="7" pageId="1" pageNumber="594">0.449</td>
<td id="76B3113AFFA7CEAEFB33FB92FAC66E4B" box="[1251,1359,1086,1110]" gridcol="3" gridrow="7" pageId="1" pageNumber="594">0.480</td>
</tr>
<tr id="35627846FFA7CEAEFF73FBCAFAC66E63" box="[163,1359,1126,1150]" gridrow="8" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FBCAFD9C6E63" box="[163,533,1126,1150]" gridcol="0" gridrow="8" pageId="1" pageNumber="594">Neck width</th>
<td id="76B3113AFFA7CEAEFCC4FBCAFCFB6E63" box="[788,882,1126,1150]" gridcol="1" gridrow="8" pageId="1" pageNumber="594">0.498</td>
<td id="76B3113AFFA7CEAEFC3EFBCAFBD36E63" box="[1006,1114,1126,1150]" gridcol="2" gridrow="8" pageId="1" pageNumber="594">0.490</td>
<td id="76B3113AFFA7CEAEFB33FBCAFAC66E63" box="[1251,1359,1126,1150]" gridcol="3" gridrow="8" pageId="1" pageNumber="594">0.510</td>
</tr>
<tr id="35627846FFA7CEAEFF73FB22FAC66EBA" box="[163,1359,1166,1191]" gridrow="9" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FB22FD9C6EBA" box="[163,533,1166,1191]" gridcol="0" gridrow="9" pageId="1" pageNumber="594">Pronotum length</th>
<td id="76B3113AFFA7CEAEFCC4FB22FCFB6EBA" box="[788,882,1166,1191]" gridcol="1" gridrow="9" pageId="1" pageNumber="594">0.824</td>
<td id="76B3113AFFA7CEAEFC3EFB22FBD36EBA" box="[1006,1114,1166,1191]" gridcol="2" gridrow="9" pageId="1" pageNumber="594">0.627</td>
<td id="76B3113AFFA7CEAEFB33FB22FAC66EBA" box="[1251,1359,1166,1191]" gridcol="3" gridrow="9" pageId="1" pageNumber="594">0.633</td>
</tr>
<tr id="35627846FFA7CEAEFF73FB1BFAC66ED2" box="[163,1359,1207,1231]" gridrow="10" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FB1BFD9C6ED2" box="[163,533,1207,1231]" gridcol="0" gridrow="10" pageId="1" pageNumber="594">Pronotum width</th>
<td id="76B3113AFFA7CEAEFCC4FB1BFCFB6ED2" box="[788,882,1207,1231]" gridcol="1" gridrow="10" pageId="1" pageNumber="594">0.809</td>
<td id="76B3113AFFA7CEAEFC3EFB1BFBD36ED2" box="[1006,1114,1207,1231]" gridcol="2" gridrow="10" pageId="1" pageNumber="594">0.862</td>
<td id="76B3113AFFA7CEAEFB33FB1BFAC66ED2" box="[1251,1359,1207,1231]" gridcol="3" gridrow="10" pageId="1" pageNumber="594">0.896</td>
</tr>
<tr id="35627846FFA7CEAEFF73FB73FAC66EEA" box="[163,1359,1247,1271]" gridrow="11" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FB73FD9C6EEA" box="[163,533,1247,1271]" gridcol="0" gridrow="11" pageId="1" pageNumber="594">Elytra length</th>
<td id="76B3113AFFA7CEAEFCC4FB73FCFB6EEA" box="[788,882,1247,1271]" gridcol="1" gridrow="11" pageId="1" pageNumber="594">0.766</td>
<td id="76B3113AFFA7CEAEFC3EFB73FBD36EEA" box="[1006,1114,1247,1271]" gridcol="2" gridrow="11" pageId="1" pageNumber="594">0.755</td>
<td id="76B3113AFFA7CEAEFB33FB73FAC66EEA" box="[1251,1359,1247,1271]" gridcol="3" gridrow="11" pageId="1" pageNumber="594">0.782</td>
</tr>
<tr id="35627846FFA7CEAEFF73FAABFAC66F02" box="[163,1359,1287,1311]" gridrow="12" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FAABFD9C6F02" box="[163,533,1287,1311]" gridcol="0" gridrow="12" pageId="1" pageNumber="594">Left elytra width</th>
<td id="76B3113AFFA7CEAEFCC4FAABFCFB6F02" box="[788,882,1287,1311]" gridcol="1" gridrow="12" pageId="1" pageNumber="594">0.538</td>
<td id="76B3113AFFA7CEAEFC3EFAABFBD36F02" box="[1006,1114,1287,1311]" gridcol="2" gridrow="12" pageId="1" pageNumber="594">0.528</td>
<td id="76B3113AFFA7CEAEFB33FAABFAC66F02" box="[1251,1359,1287,1311]" gridcol="3" gridrow="12" pageId="1" pageNumber="594">0.534</td>
</tr>
<tr id="35627846FFA7CEAEFF73FA83FAC66F55" box="[163,1359,1327,1352]" gridrow="13" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FA83FD9C6F55" box="[163,533,1327,1352]" gridcol="0" gridrow="13" pageId="1" pageNumber="594">Right elytra width</th>
<td id="76B3113AFFA7CEAEFCC4FA83FCFB6F55" box="[788,882,1327,1352]" gridcol="1" gridrow="13" pageId="1" pageNumber="594">0.508</td>
<td id="76B3113AFFA7CEAEFC3EFA83FBD36F55" box="[1006,1114,1327,1352]" gridcol="2" gridrow="13" pageId="1" pageNumber="594">NA</td>
<td id="76B3113AFFA7CEAEFB33FA83FAC66F55" box="[1251,1359,1327,1352]" gridcol="3" gridrow="13" pageId="1" pageNumber="594">0.555</td>
</tr>
<tr id="35627846FFA7CEAEFF73FAFBFAC66F6D" box="[163,1359,1367,1392]" gridrow="14" pageId="1" pageNumber="594">
<th id="76B3113AFFA7CEAEFF73FAFBFD9C6F6D" box="[163,533,1367,1392]" gridcol="0" gridrow="14" pageId="1" pageNumber="594">Abdomen width</th>
<td id="76B3113AFFA7CEAEFCC4FAFBFCFB6F6D" box="[788,882,1367,1392]" gridcol="1" gridrow="14" pageId="1" pageNumber="594">1.050</td>
<td id="76B3113AFFA7CEAEFC3EFAFBFBD36F6D" box="[1006,1114,1367,1392]" gridcol="2" gridrow="14" pageId="1" pageNumber="594">1.093</td>
<td id="76B3113AFFA7CEAEFB33FAFBFAC66F6D" box="[1251,1359,1367,1392]" gridcol="3" gridrow="14" pageId="1" pageNumber="594">1.128</td>
</tr>
</table>
</paragraph>
<paragraph id="8BED7A04FFA73150FF17FA33FD3E6C1C" blockId="1.[151,1437,1439,2042]" pageId="1" pageNumber="594">
Mandible stout, falciform, strongly incurved; apex sharply pointed. Antenna (
<typeStatus id="54E9C4A6FFA73150FB96FA33FBF16FA4" box="[1094,1144,1439,1465]" pageId="1" pageNumber="594">type</typeStatus>
II) as long as or slightly longer than head and pronotum together, with apical antennomere slightly shorter than two preceding together, antennomeres 4–11 with basal dish (
<figureCitation id="13696681FFA73150FDFFFA4BFD236C1C" box="[559,682,1511,1537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1922,1946]" captionTargetBox="[163,1426,190,1885]" captionTargetId="figure-26@2.[151,1435,179,1896]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="1" pageNumber="594">Fig. 1A–C</figureCitation>
).
</paragraph>
<paragraph id="8BED7A04FFA73150FF17F9A0FBBD6C8C" blockId="1.[151,1437,1439,2042]" pageId="1" pageNumber="594">
Pronotum (
<figureCitation id="13696681FFA73150FE96F9A7FE4F6C38" box="[326,454,1547,1573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1922,1946]" captionTargetBox="[163,1426,190,1885]" captionTargetId="figure-26@2.[151,1435,179,1896]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="1" pageNumber="594">Figs. 1D–F</figureCitation>
,
<figureCitation id="13696681FFA73150FE02F9A7FE7F6C38" box="[466,502,1547,1573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1626,1650]" captionTargetBox="[163,1423,193,1590]" captionTargetId="figure-16@3.[151,1435,181,1602]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="1" pageNumber="594">2A</figureCitation>
) transverse, broadest at near anterior 1/4, as wide as head or slightly broader. Disc 3-sulcate, median sulcus and two curved paramedial sulci deep and punctate, two paralateral depressions punctate; lateral margin even in front and indistinctly crenulate behind, posterolateral angles rounded. Elytra (
<figureCitation id="13696681FFA73150FACCF9FFFA1D6C70" box="[1308,1428,1619,1645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1922,1946]" captionTargetBox="[163,1426,190,1885]" captionTargetId="figure-26@2.[151,1435,179,1896]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–C) and forebody (D–F) of the types of O. acriculiclypeatus sp. nov. A, D: holotype; B, E: paratype 1; C, F: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="1" pageNumber="594">Fig. 1D–F</figureCitation>
) punctate and sparsely pubescent, but not rugose, without lateral longitudinal ridge.
</paragraph>
<paragraph id="8BED7A04FFA73150FF17F937FC596D3C" blockId="1.[151,1437,1439,2042]" pageId="1" pageNumber="594">
Abdomen barely coriaceous and sparsely pubescent, broadest at segment VI. Sternite VII (
<figureCitation id="13696681FFA73150FB71F937FB7C6CA8" box="[1185,1269,1691,1717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1626,1650]" captionTargetBox="[163,1423,193,1590]" captionTargetId="figure-16@3.[151,1435,181,1602]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="1" pageNumber="594">Fig. 2B</figureCitation>
) with posterior margin straight and with densely haired patch in middle. Sternite VIII (
<figureCitation id="13696681FFA73150FC7EF913FB8D6CC4" box="[942,1028,1727,1753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1626,1650]" captionTargetBox="[163,1423,193,1590]" captionTargetId="figure-16@3.[151,1435,181,1602]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="1" pageNumber="594">Fig. 2C</figureCitation>
) with subbasal ridge continuous and straight; posterior margin shallowly and narrowly bi-emarginate near middle; central plate short, broadly and shortly protruding in middle, with apical margin weakly emarginate and carinate.
</paragraph>
<paragraph id="8BED7A04FFA73150FF17F887FBB96DE7" blockId="1.[151,1437,1439,2042]" pageId="1" pageNumber="594">
Aedeagus (
<figureCitation id="13696681FFA73150FE96F887FE4C6D58" box="[326,453,1835,1861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1626,1650]" captionTargetBox="[163,1423,193,1590]" captionTargetId="figure-16@3.[151,1435,181,1602]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of O. acriculiclypeatus sp. nov. A: head and pronotum in dorsal view; B: sternite VII in ventral view; C: sternite VIII in ventral view; D: aedeagus in ventral view; E: aedeagus in dorsal view; F: median lobe of aedeagus in lateral view; G: right paramere of aedeagus in lateral view; H: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="1" pageNumber="594">Fig. 2D–H</figureCitation>
). Median lobe elongate and slightly broader basally, ventral edge of apical orifice with two pointed processes near middle, with sclerite-like or membranous structures in apical part of median lobe; apico-medial hook with apical tip pointed and downcurved and with transverse carina behind tip, basal process short; dorsal membranous area covering most of apical part of dorsum and nearly across length of median lobe. Parameres arm-like, with apex flattened, a little broadened and rounded, with ridge on ventral and lateral surface of each piece and screw-tracked at elbow bend, at near 1/2 of apical arm situated with long seta.
</paragraph>
<caption id="DF2D2A8CFFA43153FF47F82EFD656DA3" ID-DOI="http://doi.org/10.5281/zenodo.10695724" ID-Zenodo-Dep="10695724" httpUri="https://zenodo.org/record/10695724/files/figure.png" pageId="2" pageNumber="595" startId="2.[151,250,1922,1946]" targetBox="[163,1426,190,1885]" targetPageId="2" targetType="figure">
<paragraph id="8BED7A04FFA43153FF47F82EFD656DA3" blockId="2.[151,1436,1922,1983]" pageId="2" pageNumber="595">
<emphasis id="B926A616FFA43153FF47F82EFE9A6D87" bold="true" box="[151,275,1922,1946]" pageId="2" pageNumber="595">FIGURE 1.</emphasis>
Habitus (
<emphasis id="B926A616FFA43153FEA7F82EFE216D87" bold="true" box="[375,424,1922,1946]" pageId="2" pageNumber="595">A–C</emphasis>
) and forebody (
<emphasis id="B926A616FFA43153FD9CF82EFDF36D87" bold="true" box="[588,634,1922,1946]" pageId="2" pageNumber="595">D–F</emphasis>
) of the types of
<taxonomicName id="4C520187FFA43153FCF0F82FFC646D87" authority="Lü, 2024" authorityName="Lü" authorityYear="2024" box="[800,1005,1922,1946]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="acriculiclypeatus" status="sp. nov.">
<emphasis id="B926A616FFA43153FCF0F82FFC646D87" box="[800,1005,1922,1946]" italics="true" pageId="2" pageNumber="595">O. acriculiclypeatus</emphasis>
</taxonomicName>
<emphasis id="B926A616FFA43153FC24F82EFBD46D87" bold="true" box="[1012,1117,1922,1946]" pageId="2" pageNumber="595">
<taxonomicNameLabel id="A2151B6DFFA43153FC24F82EFBCD6D87" box="[1012,1092,1922,1946]" pageId="2" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
,
<emphasis id="B926A616FFA43153FBBAF82EFBF56D87" bold="true" box="[1130,1148,1922,1946]" pageId="2" pageNumber="595">D</emphasis>
: holotype;
<emphasis id="B926A616FFA43153FB3FF82EFA896D87" bold="true" box="[1263,1280,1922,1946]" pageId="2" pageNumber="595">B</emphasis>
,
<emphasis id="B926A616FFA43153FADDF82EFA976D87" bold="true" box="[1293,1310,1922,1946]" pageId="2" pageNumber="595">E</emphasis>
: paratype 1;
<emphasis id="B926A616FFA43153FF47F80AFF206DA3" bold="true" box="[151,169,1958,1982]" pageId="2" pageNumber="595">C</emphasis>
,
<emphasis id="B926A616FFA43153FF66F80AFF4C6DA3" bold="true" box="[182,197,1958,1982]" pageId="2" pageNumber="595">F</emphasis>
: paratype 2. Scale bars: A–C = 1 mm, D–F = 0.5 mm.
</paragraph>
</caption>
<caption id="DF2D2A8CFFA53152FF47F9F6FCDF6CC3" ID-DOI="http://doi.org/10.5281/zenodo.10695726" ID-Zenodo-Dep="10695726" httpUri="https://zenodo.org/record/10695726/files/figure.png" pageId="3" pageNumber="596" startId="3.[151,250,1626,1650]" targetBox="[163,1423,193,1590]" targetPageId="3" targetType="figure">
<paragraph id="8BED7A04FFA53152FF47F9F6FCDF6CC3" blockId="3.[151,1436,1626,1759]" pageId="3" pageNumber="596">
<emphasis id="B926A616FFA53152FF47F9F6FE9D6C6F" bold="true" box="[151,276,1626,1650]" pageId="3" pageNumber="596">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C520187FFA53152FE4FF9F7FDE46C6E" authority="Lü, 2024" authorityName="Lü" authorityYear="2024" box="[415,621,1627,1651]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="acriculiclypeatus" status="sp. nov.">
<emphasis id="B926A616FFA53152FE4FF9F7FDE46C6E" box="[415,621,1627,1651]" italics="true" pageId="3" pageNumber="596">O. acriculiclypeatus</emphasis>
</taxonomicName>
<emphasis id="B926A616FFA53152FDA5F9F7FD686C6F" bold="true" box="[629,737,1626,1651]" pageId="3" pageNumber="596">
<taxonomicNameLabel id="A2151B6DFFA53152FDA5F9F7FD4E6C6E" box="[629,711,1627,1651]" pageId="3" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
: head and pronotum in dorsal view;
<emphasis id="B926A616FFA53152FB8FF9F7FBF96C6E" bold="true" box="[1119,1136,1627,1651]" pageId="3" pageNumber="596">B</emphasis>
: sternite VII in ventral view;
<emphasis id="B926A616FFA53152FF47F9D2FF206C8B" bold="true" box="[151,169,1662,1686]" pageId="3" pageNumber="596">C</emphasis>
: sternite VIII in ventral view;
<emphasis id="B926A616FFA53152FE0EF9D3FE796C8A" bold="true" box="[478,496,1663,1687]" pageId="3" pageNumber="596">D</emphasis>
: aedeagus in ventral view;
<emphasis id="B926A616FFA53152FCD6F9D3FC9E6C8A" bold="true" box="[774,791,1663,1687]" pageId="3" pageNumber="596">E</emphasis>
: aedeagus in dorsal view;
<emphasis id="B926A616FFA53152FBF5F9D3FBBD6C8A" bold="true" box="[1061,1076,1663,1687]" pageId="3" pageNumber="596">F</emphasis>
: median lobe of aedeagus in lateral view;
<emphasis id="B926A616FFA53152FF05F90EFF606CA7" bold="true" box="[213,233,1698,1722]" pageId="3" pageNumber="596">G</emphasis>
: right paramere of aedeagus in lateral view;
<emphasis id="B926A616FFA53152FD7AF90FFD376CA6" bold="true" box="[682,702,1699,1723]" pageId="3" pageNumber="596">H</emphasis>
: line-drawing showing details of apical orifice and apico-medial hook of median lobe in ventral view. Scale bars: A = 0.5 mm, B–H = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BED7A04FFA53152FF17F8A1FE166D3A" blockId="3.[151,1437,1805,2011]" box="[199,415,1805,1831]" pageId="3" pageNumber="596">
<emphasis id="B926A616FFA53152FF17F8A1FE926D3A" box="[199,283,1805,1831]" italics="true" pageId="3" pageNumber="596">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA53152FF17F89CFCC96D56" box="[199,832,1840,1867]" pageId="3" pageNumber="596" type="distribution">
<paragraph id="8BED7A04FFA53152FF17F89CFCC96D56" blockId="3.[151,1437,1805,2011]" box="[199,832,1840,1867]" pageId="3" pageNumber="596">
<emphasis id="B926A616FFA53152FF17F89CFD856D57" bold="true" box="[199,524,1840,1866]" pageId="3" pageNumber="596">
Distribution (
<typeStatus id="54E9C4A6FFA53152FEB8F89CFE146D57" box="[360,413,1840,1866]" pageId="3" pageNumber="596">type</typeStatus>
locality).
</emphasis>
<collectingCountry id="F3453A94FFA53152FDC2F89DFDF06D56" box="[530,633,1841,1867]" name="Malaysia" pageId="3" pageNumber="596">Malaysia</collectingCountry>
(Borneo:
<collectingRegion id="4996B4E6FFA53152FD3BF89DFCBD6D56" box="[747,820,1841,1867]" country="Malaysia" name="Sabah" pageId="3" pageNumber="596">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA53152FF17F8F9FED16D8E" pageId="3" pageNumber="596" type="etymology">
<paragraph id="8BED7A04FFA53152FF17F8F9FED16D8E" blockId="3.[151,1437,1805,2011]" pageId="3" pageNumber="596">
<emphasis id="B926A616FFA53152FF17F8F9FEC76D72" bold="true" box="[199,334,1877,1903]" pageId="3" pageNumber="596">Etymology.</emphasis>
The specific epithet is derived from two Latin adjectives
<emphasis id="B926A616FFA53152FC0BF8F9FBCA6D72" box="[987,1091,1877,1903]" italics="true" pageId="3" pageNumber="596">acriculus</emphasis>
and
<taxonomicName id="4C520187FFA53152FBABF8F9FB6C6D72" authorityName="Motschulsky" authorityYear="1860" box="[1147,1253,1877,1903]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B926A616FFA53152FBABF8F9FB6C6D72" box="[1147,1253,1877,1903]" italics="true" pageId="3" pageNumber="596">clypeatus</emphasis>
</taxonomicName>
, referring to the sharp clypeal tip.
</paragraph>
</subSubSection>
<subSubSection id="C348298FFFA53155FF17F831FB4A6B00" lastPageId="4" lastPageNumber="597" pageId="3" pageNumber="596" type="discussion">
<paragraph id="8BED7A04FFA53152FF17F831FA156DC6" blockId="3.[151,1437,1805,2011]" pageId="3" pageNumber="596">
<emphasis id="B926A616FFA53152FF17F831FEB26DAA" bold="true" box="[199,315,1949,1975]" pageId="3" pageNumber="596">Remarks.</emphasis>
This species closely resembles
<taxonomicName id="4C520187FFA53152FD4BF831FCBD6DAA" authorityName="Fauvel" authorityYear="1895" box="[667,820,1949,1975]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="megaceros">
<emphasis id="B926A616FFA53152FD4BF831FCBD6DAA" box="[667,820,1949,1975]" italics="true" pageId="3" pageNumber="596">O. megaceros</emphasis>
</taxonomicName>
and was found in overlapping areas (
<bibRefCitation id="EFC307F5FFA53152FB00F831FA186DA5" author="Lu, L. &amp; Zhou, H. - Z." box="[1232,1425,1949,1976]" pageId="3" pageNumber="596" pagination="1 - 94" refId="ref1782" refString="Lu, L. &amp; Zhou, H. - Z. (2015) Revision of the genus Oxytelus Gravenhorst (Staphylinidae: Oxytelinae) in Southeast Asia. Zootaxa, 3992 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 3992.1.1" type="journal article" year="2015">Lü &amp; Zhou 2015</bibRefCitation>
). However, it can be distinguished from the latter by several male morphological features (cf. Fig.
<quantity id="4CAAD7E1FFA53152FB0DF86DFA9F6DC6" box="[1245,1302,1985,2011]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="3" pageNumber="596" unit="in" value="11.0">11 in</quantity>
<bibRefCitation id="EFC307F5FFA53152FACDF86DFA156DC6" author="Lu, L. &amp; Zhou, H. - Z." box="[1309,1436,1985,2011]" pageId="3" pageNumber="596" pagination="1 - 63" refId="ref1728" refString="Lu, L. &amp; Zhou, H. - Z. (2012) Taxonomy of the genus Oxytelus Gravenhorst (Coleoptera: Staphylinidae: Oxytelinae) from China. Zootaxa, 3576 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3576.1.1" type="journal article" year="2012">Lü &amp; Zhou</bibRefCitation>
</paragraph>
<paragraph id="8BED7A04FFA23155FF47FF3BFB4A6B00" blockId="4.[151,1436,150,285]" pageId="4" pageNumber="597">
2012, Fig.
<quantity id="4CAAD7E1FFA23155FEDEFF3BFECF6AAC" box="[270,326,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="597" unit="in" value="34.0">34 in</quantity>
2015): the anterior clypeal margin with a sharper tip than that of
<taxonomicName id="4C520187FFA23155FBC5FF3BFB266AAD" authorityName="Fauvel" authorityYear="1895" box="[1045,1199,151,177]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="megaceros">
<emphasis id="B926A616FFA23155FBC5FF3BFB266AAD" box="[1045,1199,151,177]" italics="true" pageId="4" pageNumber="597">O. megaceros</emphasis>
</taxonomicName>
, the posterior margin of sternite VIII with a broader central plate and a carina lying across the whole width of the posterior margin of central plate, the sharper and more slender processes on the apical orifice, and the apico-medial hook with a smaller apical tip but without the connecting scape that attaches it to the body of aedeagal median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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