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<document id="0F053437899F58ABF3A9EA8B2C30CD50" ID-DOI="10.11646/zootaxa.5497.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13617972" ID-ZooBank="42D8CC88-4245-4DFF-AEAE-C86D74F7F7A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725025211448" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2024" docId="03AF87EAFF926621FF0CF5303476F9EA" docLanguage="en" docName="zootaxa.5497.1.2.pdf" docOrigin="Zootaxa 5497 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Taphurini Distant 1905" docType="treatment" docVersion="2" lastPageNumber="50" masterDocId="FF96FF92FF836630FF9BF64F3637FFC8" masterDocTitle="The cicada (Hemiptera: Cicadidae) fauna of the Lesser Antilles sensu lato with the description of two new species, two new combinations, and a key to species" masterLastPageNumber="69" masterPageNumber="33" pageNumber="50" updateTime="1725026498551" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0ED132E268A01C53E58991DF2A599946">The cicada (Hemiptera: Cicadidae) fauna of the Lesser Antilles sensu lato with the description of two new species, two new combinations, and a key to species</mods:title>
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<mods:date id="6E582FEF795FF49748FC3A7A31B116AB">2024</mods:date>
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<subSubSection id="C31C6577FF926621FF0CF5303438FC29" pageId="17" pageNumber="50" type="nomenclature">
<paragraph id="8BB936FCFF926621FF0CF530342FFC51" blockId="17.[151,536,895,921]" box="[151,536,895,921]" pageId="17" pageNumber="50">
<heading id="D0F18190FF926621FF0CF530342FFC51" bold="true" box="[151,536,895,921]" fontSize="11" level="1" pageId="17" pageNumber="50" reason="1">
<emphasis id="B972EAEEFF926621FF0CF530342FFC51" bold="true" box="[151,536,895,921]" pageId="17" pageNumber="50">
Tribe
<taxonomicName id="4C064D7FFF926621FF79F530342FFC51" ID-CoL="8HC2H" authority="Distant, 1905 e" authorityName="Distant" authorityYear="1905" box="[226,536,895,921]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Taphurini">
Taphurini
<bibRefCitation id="EF974B0DFF926621FEFCF530342FFC51" author="Distant, W. L." box="[359,536,895,921]" pageId="17" pageNumber="50" pagination="22 - 35" refId="ref31418" refString="Distant, W. L. (1905 e) Rhynchotal notes XXXIII. Annals and Magazine of Natural History, Series 7, 16, 22 - 35. https: // doi. org / 10.1080 / 03745480509443650" type="journal article" year="1905">Distant, 1905e</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936FCFF926621FF0CF5883438FC29" blockId="17.[151,527,967,993]" box="[151,527,967,993]" pageId="17" pageNumber="50">
<heading id="D0F18190FF926621FF0CF5883438FC29" bold="true" box="[151,527,967,993]" fontSize="11" level="1" pageId="17" pageNumber="50" reason="1">
<emphasis id="B972EAEEFF926621FF0CF5883438FC29" bold="true" box="[151,527,967,993]" pageId="17" pageNumber="50">
Taphuraria
<bibRefCitation id="EF974B0DFF926621FEB4F588343EFC29" author="Distant, W. L." box="[303,521,967,993]" pageId="17" pageNumber="50" pagination="22 - 35" refId="ref31418" refString="Distant, W. L. (1905 e) Rhynchotal notes XXXIII. Annals and Magazine of Natural History, Series 7, 16, 22 - 35. https: // doi. org / 10.1080 / 03745480509443650" type="journal article" year="1905">Distant 1905e: 25</bibRefCitation>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C6577FF926621FF0CF25F34A0FBE2" box="[151,663,1039,1066]" pageId="17" pageNumber="50" type="type_taxon">
<paragraph id="8BB936FCFF926621FF0CF25F34A0FBE2" blockId="17.[151,1437,1039,1570]" box="[151,663,1039,1066]" pageId="17" pageNumber="50">
<emphasis id="B972EAEEFF926621FF0CF25F3713FBE2" bold="true" box="[151,292,1040,1066]" pageId="17" pageNumber="50">
<typeStatus id="54BD885EFF926621FF0CF25F36E5FBE2" box="[151,210,1040,1066]" pageId="17" pageNumber="50">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="4C064D7FFF926621FEB0F24034A4FBE2" ID-CoL="8KLCR" authority="Stal 1862: 20" authorityName="Stal" authorityPageNumber="20" authorityYear="1862" box="[299,659,1039,1066]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="subGenus" subGenus="Taphura">
<emphasis id="B972EAEEFF926621FEB0F24037C3FBE1" box="[299,500,1039,1065]" italics="true" pageId="17" pageNumber="50">Cicada (Taphura)</emphasis>
<bibRefCitation id="EF974B0DFF926621FE60F24034A4FBE2" author="Stal, C." box="[507,659,1039,1066]" pageId="17" pageNumber="50" pagination="1 - 75" refId="ref35843" refString="Stal, C. (1862) Bidrag till Rio Janeiro - traktens Hemipter-fauna. II. Handlingar. Kongliga Svenska Vetenskaps Akademien, Stockholm, 3, 1 - 75." type="journal article" year="1862">Stål 1862: 20</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C6577FF926621FF5CF27B32F4FAC9" pageId="17" pageNumber="50" type="discussion">
<paragraph id="8BB936FCFF926621FF5CF27B32F4FAC9" blockId="17.[151,1437,1039,1570]" pageId="17" pageNumber="50">
<emphasis id="B972EAEEFF926621FF5CF27B370CFB86" bold="true" box="[199,315,1076,1102]" pageId="17" pageNumber="50">Remarks.</emphasis>
The tribe has been a repository for small species of uncertain classification historically as illustrated by the distribution of genera previously assigned to the
<taxonomicName id="4C064D7FFF926621FCB1F21835A1FBB9" authorityName="Distant" authorityYear="1905" box="[810,918,1111,1137]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Taphurini">Taphurini</taxonomicName>
being reassigned recently to seven tribes in four subfamilies (
<bibRefCitation id="EF974B0DFF926621FEF9F2343464FB5E" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." box="[354,595,1147,1174]" pageId="17" pageNumber="50" pagination="1 - 64" refId="ref32908" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B972EAEEFF926621FE4AF233343BFB5D" box="[465,524,1147,1173]" italics="true" pageId="17" pageNumber="50">et al.</emphasis>
2018
</bibRefCitation>
). The reassignment of all the genera previously assigned to the
<taxonomicName id="4C064D7FFF926621FAACF2343705FB72" authority="Moulds, 2005" authorityName="Moulds" authorityYear="2005" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="subTribe" subTribe="Tryellina">
Tryellina
<bibRefCitation id="EF974B0DFF926621FF0CF2D03705FB72" author="Moulds, M. S." box="[151,306,1183,1210]" pageId="17" pageNumber="50" pagination="375 - 446" refId="ref33404" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds, 2005</bibRefCitation>
</taxonomicName>
to the
<taxonomicName id="4C064D7FFF926621FEE5F2D03482FB71" authority="Boulard, 1976" authorityName="Boulard" authorityYear="1976" box="[382,693,1183,1209]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Lamotialnini">Lamotialnini Boulard, 1976</taxonomicName>
(
<bibRefCitation id="EF974B0DFF926621FD58F2D0359FFB72" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." box="[707,936,1183,1210]" pageId="17" pageNumber="50" pagination="1 - 64" refId="ref32908" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B972EAEEFF926621FCB4F2EF3550FB71" box="[815,871,1183,1209]" italics="true" pageId="17" pageNumber="50">et al.</emphasis>
2018
</bibRefCitation>
) and the Anopercalnina Boulard, 2008 to the
<taxonomicName id="4C064D7FFF926621FF0CF28C37D9FB16" authority="Distant, 1905 c" authorityName="Distant" authorityYear="1905" box="[151,494,1219,1246]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Tettigomyiinae">
Tettigomyiinae
<bibRefCitation id="EF974B0DFF926621FED2F28C37D9FB16" author="Distant, W. L." box="[329,494,1219,1246]" pageId="17" pageNumber="50" pagination="265 - 281" refId="ref31338" refString="Distant, W. L. (1905 c) Rhynchotal notes XXXV. Annals and Magazine of Natural History, Series 7, 16, 265 - 281. https: // doi. org / 10.1080 / 03745480509442862" type="journal article" year="1905">Distant, 1905c</bibRefCitation>
</taxonomicName>
left only Neotropical genera in the tribe (
<bibRefCitation id="EF974B0DFF926621FC55F28C3245FB16" author="Sanborn, A. F." box="[974,1138,1219,1246]" pageId="17" pageNumber="50" pagination="1 - 79" refId="ref35121" refString="Sanborn, A. F. (2021) The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species. Zootaxa, 4937 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 4937.1.1" type="journal article" year="2021">Sanborn 2021</bibRefCitation>
). Eight of the nine genera currently assigned to the tribe are monospecific with only
<taxonomicName id="4C064D7FFF926621FCBFF2A835B5FAC9" authorityName="Stal" authorityYear="1862" box="[804,898,1255,1281]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B972EAEEFF926621FCBFF2A835B5FAC9" box="[804,898,1255,1281]" italics="true" pageId="17" pageNumber="50">Taphura</emphasis>
</taxonomicName>
containing multiple species.
</paragraph>
</subSubSection>
<subSubSection id="C31C6577FF926621FF5CF3443476F9EA" pageId="17" pageNumber="50" type="diagnosis">
<paragraph id="8BB936FCFF926621FF5CF3443476F9EA" blockId="17.[151,1437,1039,1570]" pageId="17" pageNumber="50">
<emphasis id="B972EAEEFF926621FF5CF3443775FAED" bold="true" box="[199,322,1291,1317]" pageId="17" pageNumber="50">Diagnosis.</emphasis>
Members of the tribe are characterized by a head that is wider than the maximum width of the pronotum, a subquadrate pronotum not distinctly narrowed anteriorly, pronotal lateral margins that are not developed or dilated, male opercula that curve towards the abdominal midline but remain widely separated medially, opercula that are small, narrow, and do not cover the tympanal cavity, the abdomen length is about as long as the distance between the apex of the postclypeus and the posterior cruciform elevation, timbals do not extend ventral to the wing bases, the uncus is absent but the anal tube may possess lateral lobes, the upper pygofer lobes are small and unsclerotized, and the basal lobes of the pygofer are large and ornamented, especially in
<taxonomicName id="4C064D7FFF926621FB15F3AC32DBFA35" authorityName="Stal" authorityYear="1862" box="[1166,1260,1507,1533]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B972EAEEFF926621FB15F3AC32DBFA35" box="[1166,1260,1507,1533]" italics="true" pageId="17" pageNumber="50">Taphura</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF974B0DFF926621FB66F3AC33A0FA36" author="Moulds, M. S." box="[1277,1431,1507,1534]" pageId="17" pageNumber="50" pagination="375 - 446" refId="ref33404" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds 2005</bibRefCitation>
;
<bibRefCitation id="EF974B0DFF926621FF0CF0483773F9EA" author="Sanborn, A. F." box="[151,324,1543,1570]" pageId="17" pageNumber="50" pagination="451 - 481" refId="ref34525" refString="Sanborn, A. F. (2017 a) Generic redescription, seven new species and a key to the Taphura Stal, 1862 (Hemiptera: Cicadidae: Cicadettinae: Taphurini). Zootaxa, 4324 (3), 451 - 481. https: // doi. org / 10.11646 / zootaxa. 4324.3.3" type="journal article" year="2017">Sanborn 2017a</bibRefCitation>
;
<bibRefCitation id="EF974B0DFF926621FED4F0483402F9EA" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." box="[335,565,1543,1570]" pageId="17" pageNumber="50" pagination="1 - 64" refId="ref32908" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B972EAEEFF926621FE22F04737C5F9E9" box="[441,498,1543,1569]" italics="true" pageId="17" pageNumber="50">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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