treatments-xml/data/AE/75/87/AE7587E3ED2EBA4213F6EC14A5DF1D32.xml
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<mods:title id="9206A218A36463CCB4395F9293F33302">Multi-locus phylogeny and taxonomic revision of Uperoleia toadlets (Anura: Myobatrachidae) from the western arid zone of Australia, with a description of a new species</mods:title>
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<subSubSection id="6EC6657EED2EBA4213F6EC14A7F31F06" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="266336F5ED2EBA4213F6EC14A7991E2C" blockId="14.[151,413,1027,1054]" box="[151,413,1027,1054]" pageId="14" pageNumber="15">
<heading id="7D2B8199ED2EBA4213F6EC14A7991E2C" bold="true" box="[151,413,1027,1054]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="E1DC4D76ED2EBA4213F6EC14A7991E2C" ID-CoL="84ZK" authority="Gray, 1841" authorityName="Gray" authorityYear="1841" box="[151,413,1027,1054]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4213F6EC14A7991E2C" bold="true" box="[151,413,1027,1054]" pageId="14" pageNumber="15">
<emphasis id="14A8EAE7ED2EBA4213F6EC14A70A1E2F" bold="true" box="[151,270,1028,1054]" italics="true" pageId="14" pageNumber="15">Uperoleia</emphasis>
Gray, 1841
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="266336F5ED2EBA4213F6EC5AA4E61E51" blockId="14.[151,1437,1098,1366]" box="[151,738,1098,1120]" pageId="14" pageNumber="15">
<taxonomicName id="E1DC4D76ED2EBA4213F6EC5AA7711E51" authority="Gray, 1841" authorityName="Gray" authorityYear="1841" box="[151,373,1098,1120]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4213F6EC5AA6F91E51" box="[151,253,1098,1120]" italics="true" pageId="14" pageNumber="15">Uperoleia</emphasis>
Gray, 1841
</taxonomicName>
, Ann. Mag. Nat. Hist., Ser. 1, 7: 90.
</paragraph>
<paragraph id="266336F5ED2EBA4213F6EC79A5E41E4E" blockId="14.[151,1437,1098,1366]" box="[151,992,1129,1151]" pageId="14" pageNumber="15">
<emphasis id="14A8EAE7ED2EBA4213F6EC79A70C1E4E" box="[151,264,1129,1151]" italics="true" pageId="14" pageNumber="15">Hyperoleia</emphasis>
Agassiz, 1846, Nomencl. Zool., Fasc. 12: 384. Unjustified emendation.
</paragraph>
<paragraph id="266336F5ED2EBA4213F6EC98A2051E8D" blockId="14.[151,1437,1098,1366]" pageId="14" pageNumber="15">
<taxonomicName id="E1DC4D76ED2EBA4213F6EC98A7A91EAC" authority="Loveridge, 1933" authorityName="Loveridge" authorityYear="1933" box="[151,429,1160,1182]" class="Amphibia" family="Myobatrachidae" genus="Glauertia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4213F6EC98A6FD1EAF" box="[151,249,1160,1182]" italics="true" pageId="14" pageNumber="15">Glauertia</emphasis>
Loveridge, 1933
</taxonomicName>
, Occas. Pap. Boston Soc. Nat. Hist., 8: 89.
<typeStatus id="F9678857ED2EBA421019EC99A5AE1EAF" box="[888,938,1161,1182]" pageId="14" pageNumber="15">Type</typeStatus>
species:
<taxonomicName id="E1DC4D76ED2EBA42176DEC98A3721EAC" authority="Loveridge, 1933" authorityName="Loveridge" authorityYear="1933" box="[1036,1398,1160,1182]" class="Amphibia" family="Myobatrachidae" genus="Glauertia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="russelli">
<emphasis id="14A8EAE7ED2EBA42176DEC98A2C51EAF" box="[1036,1217,1160,1182]" italics="true" pageId="14" pageNumber="15">Glauertia russelli</emphasis>
Loveridge, 1933
</taxonomicName>
, by monotypy. Synonymy by
<bibRefCitation id="424D4B04ED2EBA4212ADECB6A4781E8D" author="Tyler" box="[460,636,1190,1213]" pageId="14" pageNumber="15" refString="Tyler, M. J., Davies, M. &amp; Martin, A. A. (1981) Australian frogs of the Leptodactylid Genus Uperoleia Gray. Australian Journal of Zoology Supplementary Series, 79, 1 - 64." type="journal article" year="1981">
Tyler
<emphasis id="14A8EAE7ED2EBA421166ECB8A4451E8C" box="[519,577,1191,1213]" italics="true" pageId="14" pageNumber="15">et. al.</emphasis>
1981
</bibRefCitation>
, Aust. J. Zool., Suppl. Ser., 29 (79): 9.
</paragraph>
<paragraph id="266336F5ED2EBA4213F6ECD6A4691EC8" blockId="14.[151,1437,1098,1366]" pageId="14" pageNumber="15">
<taxonomicName id="E1DC4D76ED2EBA4213F6ECD6A42E1EEA" authority="Wells and Wellington, 1985" authorityName="Wells and Wellington" authorityYear="1985" box="[151,554,1221,1243]" class="Amphibia" family="Myobatrachidae" genus="Hosmeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4213F6ECD6A6FF1EEA" box="[151,251,1222,1243]" italics="true" pageId="14" pageNumber="15">Hosmeria</emphasis>
Wells and Wellington, 1985
</taxonomicName>
, Aust. J. Herpetol., Suppl. Ser., 1: 2.
<typeStatus id="F9678857ED2EBA4210DCECD6A5EB1EEA" box="[957,1007,1222,1243]" pageId="14" pageNumber="15">Type</typeStatus>
species:
<taxonomicName id="E1DC4D76ED2EBA421732ECD5A7761EC8" authority="Keferstein, 1867" authorityName="Keferstein" authorityYear="1867" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="marmorata" subSpecies="laevigata">
<emphasis id="14A8EAE7ED2EBA421732ECD5A3981EEA" box="[1107,1436,1221,1243]" italics="true" pageId="14" pageNumber="15">Uperoleia marmorata laevigata</emphasis>
Keferstein, 1867
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="266336F5ED2EBA4213F6ED12A7F31F06" blockId="14.[151,1437,1098,1366]" pageId="14" pageNumber="15">
<taxonomicName id="E1DC4D76ED2EBA4213F6ED12A4221F29" authority="Wells and Wellington, 1985" authorityName="Wells and Wellington" authorityYear="1985" box="[151,550,1282,1304]" class="Amphibia" family="Myobatrachidae" genus="Prohartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4213F6ED12A6FE1F29" box="[151,250,1282,1304]" italics="true" pageId="14" pageNumber="15">Prohartia</emphasis>
Wells and Wellington, 1985
</taxonomicName>
, Aust. J. Herpetol., Suppl. Ser., 1: 3.
<typeStatus id="F9678857ED2EBA4210D0ED13A5E71F29" box="[945,995,1283,1304]" pageId="14" pageNumber="15">Type</typeStatus>
species:
<taxonomicName id="E1DC4D76ED2EBA421725ED12A6F81F06" authority="Parker, 1926" authorityName="Parker" authorityYear="1926" class="Amphibia" family="Myobatrachidae" genus="Pseudophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="fimbrianus">
<emphasis id="14A8EAE7ED2EBA421725ED12A3481F29" box="[1092,1356,1282,1304]" italics="true" pageId="14" pageNumber="15">Pseudophryne fimbrianus</emphasis>
Parker, 1926
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="6EC6657EED2EBA4213F6ED50A4511F67" box="[151,597,1343,1366]" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="266336F5ED2EBA4213F6ED50A4511F67" blockId="14.[151,1437,1098,1366]" box="[151,597,1343,1366]" pageId="14" pageNumber="15">
<emphasis id="14A8EAE7ED2EBA4213F6ED50A7221F67" bold="true" box="[151,294,1343,1366]" pageId="14" pageNumber="15">
<typeStatus id="F9678857ED2EBA4213F6ED50A6C81F64" box="[151,204,1344,1365]" pageId="14" pageNumber="15">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="E1DC4D76ED2EBA42124AED51A7B81F67" box="[299,444,1345,1366]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="marmorata">
<emphasis id="14A8EAE7ED2EBA42124AED51A7B81F67" box="[299,444,1345,1366]" italics="true" pageId="14" pageNumber="15">U. marmorata</emphasis>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="6EC6657EED2EBA4213F6ED92A5DF1D32" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="266336F5ED2EBA4213F6ED92A6F11C7F" blockId="14.[151,1437,1410,1795]" pageId="14" pageNumber="15">
<emphasis id="14A8EAE7ED2EBA4213F6ED92A7151FAA" bold="true" box="[151,273,1410,1435]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
A group of small-bodied (to
<quantity id="E1249B10ED2EBA42113DED92A4A81FAB" box="[604,684,1410,1434]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="14" pageNumber="15" unit="mm" value="41.0">41 mm</quantity>
) terrestrial frogs with short limbs, no webbing on fingers, first finger shorter than second, inner metatarsal tubercle not compressed, lacking vomerine teeth, covered tympana, pupil rhomboidal, parotoid, dorsolateral/inguinal and coccygeal glands usually present, males with unilobular submandibular vocal sac and unpigmented nuptial pads, aquatic reproduction with small pigmented eggs and free-swimming larvae (
<bibRefCitation id="424D4B04ED2EBA42124EEE02A7F11C1B" author="Tyler" box="[303,501,1554,1579]" pageId="14" pageNumber="15" refString="Tyler, M. J., Davies, M. &amp; Martin, A. A. (1981) Australian frogs of the Leptodactylid Genus Uperoleia Gray. Australian Journal of Zoology Supplementary Series, 79, 1 - 64." type="journal article" year="1981">
Tyler
<emphasis id="14A8EAE7ED2EBA421214EE04A7B41C1A" box="[373,432,1555,1579]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
1981
</bibRefCitation>
). Cloacal flap present, moderately fimbriated in males and strongly fimbriated in females.
</paragraph>
<paragraph id="266336F5ED2EBA4213A6EE4AA5DF1D32" blockId="14.[151,1437,1410,1795]" pageId="14" pageNumber="15">
<emphasis id="14A8EAE7ED2EBA4213A6EE4AA4161C42" bold="true" box="[199,530,1626,1651]" pageId="14" pageNumber="15">Other genera in synonymy.</emphasis>
<taxonomicName id="E1DC4D76ED2EBA42117AEE4BA4881C42" box="[539,652,1627,1651]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA42117AEE4BA4881C42" box="[539,652,1627,1651]" italics="true" pageId="14" pageNumber="15">Uperoleia</emphasis>
</taxonomicName>
are morphologically conservative and monophyly is not in question based on morphological (
<bibRefCitation id="424D4B04ED2EBA4212D8EE6EA4871CA7" author="Tyler" box="[441,643,1662,1687]" pageId="14" pageNumber="15" refString="Tyler, M. J., Davies, M. &amp; Martin, A. A. (1981) Australian frogs of the Leptodactylid Genus Uperoleia Gray. Australian Journal of Zoology Supplementary Series, 79, 1 - 64." type="journal article" year="1981">
Tyler
<emphasis id="14A8EAE7ED2EBA42129EEE90A4371CA7" box="[511,563,1662,1687]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 1981
</bibRefCitation>
) or molecular data (
<bibRefCitation id="424D4B04ED2EBA42100FEE6EA23D1CA7" author="Read" box="[878,1081,1662,1687]" pageId="14" pageNumber="15" refString="Read, K., Keogh, J. S., Scott, I. A. W., Roberts, J. D. &amp; Doughty, P. (2001) Molecular phylogeny of the Australian frog genera Crinia, Geocrinia, and allied taxa (Anura: Myobatrachidae). Molecular Phylogenetics and Evolution, 21, 294 - 308." type="journal article" year="2001">
Read
<emphasis id="14A8EAE7ED2EBA4210D2EE90A5E21CA7" box="[947,998,1662,1687]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 2001
</bibRefCitation>
;
<bibRefCitation id="424D4B04ED2EBA421726EE6FA30A1CA7" author="Frost" box="[1095,1294,1662,1687]" pageId="14" pageNumber="15" refString="Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., De Sa, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drews, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. &amp; Wheeler, W. C. (2006) The Amphibian tree of life. Bulletin of the American Museum of Natural History, 297, 1 - 370." type="journal article" year="2006">
Frost
<emphasis id="14A8EAE7ED2EBA4217EBEE90A2B91CA7" box="[1162,1213,1662,1687]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 2006
</bibRefCitation>
, this study).
<bibRefCitation id="424D4B04ED2EBA4213F6EEB2A7361C8B" author="Parker" box="[151,306,1698,1722]" pageId="14" pageNumber="15" refString="Parker, H. W. (1940) The Australasian frogs of the family Leptodactylidae. Novitates Zoologicae, 42, 1 - 106." type="journal article" year="1940">Parker (1940)</bibRefCitation>
indicated
<emphasis id="14A8EAE7ED2EBA4212C8EEB3A4221C8A" box="[425,550,1699,1723]" italics="true" pageId="14" pageNumber="15">Hyperoleia</emphasis>
was an unjustified emendation of
<taxonomicName id="E1DC4D76ED2EBA4210CDEEB3A2191C8A" box="[940,1053,1699,1723]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA4210CDEEB3A2191C8A" box="[940,1053,1699,1723]" italics="true" pageId="14" pageNumber="15">Uperoleia</emphasis>
</taxonomicName>
. There are no compelling reasons to recognize further genera within this clade. Therefore we maintain the synonymy of
<taxonomicName id="E1DC4D76ED2EBA42171AEED6A2EC1CEF" box="[1147,1256,1734,1758]" class="Amphibia" family="Myobatrachidae" genus="Glauertia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA42171AEED6A2EC1CEF" box="[1147,1256,1734,1758]" italics="true" pageId="14" pageNumber="15">Glauertia</emphasis>
</taxonomicName>
by
<bibRefCitation id="424D4B04ED2EBA421677EED6A6E51D33" pageId="14" pageNumber="15" refString="Tyler, M. J., Davies, M. &amp; Martin, A. A. (1981) Australian frogs of the Leptodactylid Genus Uperoleia Gray. Australian Journal of Zoology Supplementary Series, 79, 1 - 64." type="journal article">
Tyler
<emphasis id="14A8EAE7ED2EBA42163AEED8A3981CEF" box="[1371,1436,1734,1759]" italics="true" pageId="14" pageNumber="15">et. al.</emphasis>
(1981)
</bibRefCitation>
and tranfer
<taxonomicName id="E1DC4D76ED2EBA421208EEFCA7D31D32" box="[361,471,1772,1795]" class="Amphibia" family="Myobatrachidae" genus="Hosmeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA421208EEFCA7D31D32" box="[361,471,1772,1795]" italics="true" pageId="14" pageNumber="15">Hosmeria</emphasis>
</taxonomicName>
and
<taxonomicName id="E1DC4D76ED2EBA42116FEEFBA47F1D32" box="[526,635,1771,1795]" class="Amphibia" family="Myobatrachidae" genus="Prohartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="14A8EAE7ED2EBA42116FEEFBA47F1D32" box="[526,635,1771,1795]" italics="true" pageId="14" pageNumber="15">Prohartia</emphasis>
</taxonomicName>
to the synonymy of
<emphasis id="14A8EAE7ED2EBA421002EEFBA5DF1D32" box="[867,987,1771,1795]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="E1DC4D76ED2EBA421002EEFBA5D31D32" box="[867,983,1771,1795]" class="Amphibia" family="Myobatrachidae" genus="Uperoleia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">Uperoleia</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>