195 lines
25 KiB
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195 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.70.762" ID-GBIF-Dataset="b8fceb5a-9f78-4dc4-8fa2-92fe20bb2c42" ID-PMC="PMC3088446" ID-Pensoft-Pub="1313-2970-70-1" ID-PubMed="21594041" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-70-1" ModsDocOrigin="ZooKeys 70" ModsDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an “oceanic” island" checkinTime="1451250880545" checkinUser="pensoft" docAuthor="Rowson, B, Warren, B. H. & Ngereza, C. F." docDate="2010" docId="A625690629B57583AEB09D98A0844A72" docLanguage="en" docName="ZooKeys 70: 1-39" docOrigin="ZooKeys 70" docSource="http://dx.doi.org/10.3897/zookeys.70.762" docTitle=""Dendrolimax" vangoethemi Rowson, sp. n." docType="treatment" docVersion="6" lastPageNumber="11" masterDocId="FFB2FFB6591BFFBAFFFF0472FFE6A56D" masterDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an " oceanic " island" masterLastPageNumber="39" masterPageNumber="1" pageNumber="8" updateTime="1668164835731" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an " oceanic " island</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rowson, B</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Warren, B. H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ngereza, C. F.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>70</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>39</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.70.762</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.70.762</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-70-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360803" LSID="urn:lsid:zoobank.org:act:40EB98A1-E93A-47FA-B81C-C4F545FFD298" httpUri="http://treatment.plazi.org/id/A625690629B57583AEB09D98A0844A72" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph pageId="7" pageNumber="8">
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2.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:40EB98A1-E93A-47FA-B81C-C4F545FFD298" authority="Rowson" lsidName=""Dendrolimax" vangoethemi" pageId="7" pageNumber="8" rank="species" species="vangoethemi">
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<normalizedToken originalValue="“Dendrolimax”">"Dendrolimax"</normalizedToken>
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vangoethemi Rowson
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</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
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Figs 11
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<normalizedToken originalValue="–1264–">-1264-</normalizedToken>
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75
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="type material">
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<paragraph pageId="7" pageNumber="8">Type material</paragraph>
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<paragraph pageId="7" pageNumber="8">(all from TANZANIA: Zanzibar: Pemba Island). Holotype (NMW.Z.2009.013.00211): slug 30.0 mm long in 80% ethanol, on understorey foliage during day, Ngezi FR (Locality 2 in Fig. 1 and Table 1), 8 February 2009, leg. B. Rowson, B. H. Warren, C. F. Ngereza & local collectors.</paragraph>
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<paragraph pageId="7" pageNumber="8">Paratype 1 (NMW.Z.2009.013.00212): slug 37.5 mm long in 80% ethanol; other data as holotype. Paratype 2 (NMW.Z.2009.013.00213): slug 31.0 mm long in 80% ethanol; other data as holotype. Paratype 3 (NMW.Z.2009.013.00214): slug 25.0 mm long in 80% ethanol; other data as holotype. Paratype 4 (IRSNB.IG.31599/MT2317): slug 18.0 mm long in 80% ethanol; other data as holotype. Paratype 5 (NMT): slug 19.0 mm long in 80% ethanol, on understorey foliage during day, Ngezi FR (Locality 6 in Fig. 1 and Table 1), 8 February 2009, leg. B. Rowson, B. H. Warren, C. F. Ngereza & local collectors. Paratype 6 (NMW.Z.2009.013.00215): slug 18.5 mm long in 80% ethanol; leaf litter during day, Ras Kiuyu FR (Locality 7 in Fig. 1 and Table 1), leg. B. Rowson, B. H. Warren, C. F. Ngereza & local collectors.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="diagnosis">
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<paragraph pageId="7" pageNumber="8">Diagnosis:</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Medium-sized slug (to at least 55mm in life) with strong keel prolonged into long caudal appendage, with mantle completely covering shell. Pale to colourless, with dorsum covered in pustules. Viscera not extending far into tail, shell mineralised, jaw with no or weak projection. Radula unique in having up to 280 tiny, tricuspid teeth per half-row. Genitalia broadly similar to other
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
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<paragraph pageId="7" pageNumber="8">Description:</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Note: points of agreement with an unnamed East Usambara species as discussed by
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History (Special supplement)" pageId="24" pageNumber="25" pagination="1 - 36" title="East African slugs of the family Urocyclidae, III & IV. The genus Trichotoxon." volume="7" year="1961">Verdcourt and Polhill (1961)</bibRefCitation>
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(see below) are marked with
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<normalizedToken originalValue="“*”">"*"</normalizedToken>
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.
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</paragraph>
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<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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External features: Medium-sized slug (extended length to at least 55mm in life, or 37.5mm in 80% ethanol)*. Tail strongly keeled* and hollowed out behind mantle*; keel
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<pageBreakToken pageId="8" pageNumber="9" start="start">prolonged</pageBreakToken>
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into long straight caudal appendage* above small caudal pore*. Viscera extending little more than half-way into tail*. Body bell-shaped in cross-section when extended, but able to flatten body considerably. Sole not narrow, evenly tripartite*. Peripodial grooves clear, from tail to genital orifice and head. Mantle fully attached posteriorly, free anteriorly, not grooved, subangulate rather than rounded behind, completely covering shell, lacking a dorsal pore or slit*. Pneumostome in posterior third of mantle*. Genital orifice far forward, near right lower tentacle. No head wart or similar structure detected. Integumental tubercles barely detectable on mantle, tail or cephalopodium; instead, whole dorsum rather densely and regularly covered in hemispherical (rather than prickly) pustules*. Dorsum largely colourless and translucent*, with green, grey or pink tinge*, acquiring a green cast when on foliage; keel white; sole colourless. Diffuse, slate-grey pigment on caudal appendage, bordering keel* and/or in obscure blotches or bands on mantle in some specimens*, absent in others; remains on preservation. Pustules conspicuously white*, remaining so on preservation. Ommatophore retractors grey-ochre on preservation.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Jaw and radula: Jaw solid, semi-lunate, lacking median projection (holotype), or projection very weak (paratype). Radula of holotype broad (3.65 mm wide
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<normalizedToken originalValue="×">x</normalizedToken>
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2.20 mm long), of 155 rows (over 50 angular rows per mm length). Teeth extremely small* and extremely numerous*, to nearly 450 in each half-row*, with a central tooth. All teeth (including central tooth) tricuspid*, with very little change across the row, perhaps becoming more s-shaped laterally. Ectocones larger (or at least projecting further) than mesocones in all teeth except central tooth. This is unlike any radula figured in
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">van Goethem (1977)</bibRefCitation>
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where mesocones are always the largest cusps, and the maximum number of teeth per half-row is around 280, and only in radulae over 5.00 mm wide. The radula form may suggest a microphagous, rather than phytophagous diet.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Shell and pallial complex: Shell unguiform, bilaterally symmetrical, to at least 4.10 mm long, infilled, mineralised and white* (i.e. not fragile as in other
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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). Pallial area well vascularised.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Genitalia: Right ommatophore retractor passes between penis and vagina. No atrial diverticulum or stimulator*. Long flagellum present in place of calc sac*. Epiphallus long, stiff, not spiralling around penis*. Penial retractor short, attaching well below flagellum, perhaps obtaining from diaphragm. Penis with basal sheath, contiguous with penis wall apically, internally with longitudinal pilasters and a basal papilla; similar to that of several
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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(see
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">van Goethem 1977</bibRefCitation>
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). Bursa copulatrix duct arising low on vagina, bound to it by sheath-like circular muscle fibres. Bursa copulatrix long, weakly clavate, reaching upper part of spermoviduct*. Vagina and free oviduct with a clear, thick-walled sheath*. Hermaphroditic duct extremely short*, barely perceptible between spermoviduct and large yellow ovotestis, which lies near rear of mantle. Albumen gland small, hook-shaped*. No spermatophores were recovered from the Pemba material which may not be fully adult.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="remarks">
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<paragraph pageId="8" pageNumber="9">Remarks:</paragraph>
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<paragraph pageId="8" pageNumber="9">This distinctive species was found only in FRs. At Ngezi, the slugs were found on the underside of large understorey leaves, up to 2m above ground. The body was held flattened with one optic tentacle protruding (Fig. 12). At Ras Kiuyu FR the species was found in litter. It appears to be undescribed although it (or a similar species) may occur in the East Usambara Mts. Beyond that its affinities are less certain.</paragraph>
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<paragraph pageId="9" pageNumber="10">
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">
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<pageBreakToken pageId="9" pageNumber="10" start="start">Van</pageBreakToken>
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Goethem (1977)
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</bibRefCitation>
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thoroughly revised the known urocyclid slugs of Africa and Madagascar and provided keys to internal characters. The Pemba species keys readily out to Dendrolimacini (sole genus
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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Heynemann, 1868). The only East African record of the mainly Central-West African
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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is an unnamed and incompletely described species collected by
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History" pageId="24" pageNumber="25" pagination="200 - 209" title="East African slugs of the family Urocyclidae." volume="23" year="1960">Verdcourt (1960)</bibRefCitation>
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from Thika Gorge, Kenya, who indicated a swollen lower oviduct not present in the Pemba species. Moreover the Pemba species differs from all other
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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in the radula, body form, and shell, although the genitalia are similar. It shows much clearer similarities to a taxon referred to as "Genus et species nov." by
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History (Special supplement)" pageId="24" pageNumber="25" pagination="1 - 36" title="East African slugs of the family Urocyclidae, III & IV. The genus Trichotoxon." volume="7" year="1961">Verdcourt and Polhill (1961, p. 32-33, fig. 42)</bibRefCitation>
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from Sigi in the East Usambara Mts. (1961). They said, "This mollusc does not belong to the family Urocyclidae judging by [the] radula but to an isolated subfamily of the Helicarionidae near to the Durgellinae".
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">Van Goethem (1977)</bibRefCitation>
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could not obtain material of this taxon for his revision but considered it a urocyclid. He treated it as "Species E", incertae sedis after Dendrolimacini and Upembellini along with a "Species D" from Grand Comore to which he noted a similarity in the genitalia, but not the radula. Neither
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History (Special supplement)" pageId="24" pageNumber="25" pagination="1 - 36" title="East African slugs of the family Urocyclidae, III & IV. The genus Trichotoxon." volume="7" year="1961">Verdcourt and Polhill (1961)</bibRefCitation>
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nor
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">van Goethem (1977)</bibRefCitation>
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were certain whether the specimens of "Species E" or "Species D" were adult. This is an important consideration since slugs may change in appearance as they grow. However, neither Verdcourt and Polhill, van Goethem with his experience of growth series of many taxa, nor other slug workers (e.g.
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<bibRefCitation author="Forcart, L" journalOrPublisher="Annals of the Natal Museum" pageId="22" pageNumber="23" pagination="505 - 570" title="Studies on the Veronicellidae, Aperidae and Urocyclidae (Mollusca) of South Africa." volume="18" year="1967">Forcart 1967</bibRefCitation>
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) could attribute these forms to any known species or genus.
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History (Special supplement)" pageId="24" pageNumber="25" pagination="1 - 36" title="East African slugs of the family Urocyclidae, III & IV. The genus Trichotoxon." volume="7" year="1961">Verdcourt and Polhill (1961)</bibRefCitation>
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noted that there was no absolute criterion, e.g. concerning the size of the albumen gland, for recognising adulthood in urocyclid slugs. Since 1977 there has been little further work on the group in East Africa. The Pemba species, which will probably prove to include also the East Usambara species, is here described provisionally in
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
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. Consideration was given to erecting a new genus but owing to a lack of unique features in the genitalia, and the large number of available genus-group names, is avoided until more data are available.
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</paragraph>
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<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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Although this species is fully limacised, there are also similarities in the body form and genitalia to numerous Afrotropical semi-slug genera, among them
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Verrucarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Verrucarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Verrucarion</taxonomicName>
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van Mol, 1970 of West and central Africa (
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<bibRefCitation author="van Mol, JJ" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="23" pageNumber="24" pagination="1 - 234" title="Revision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie-Systematique-Zoogeographie." volume="180" year="1970">van Mol 1970</bibRefCitation>
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) and
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Malagarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malagarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Malagarion</taxonomicName>
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Tillier, 1979 of Madagascar. The resemblance to the latter extends to the radula and white pustules (see
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<bibRefCitation author="Tillier, S" journalOrPublisher="The Veliger" pageId="24" pageNumber="25" pagination="361 - 368" title="Malagarion paenelimax gen. nov., spec. nov., a new slug-like helicarionid from Madagascar." volume="21" year="1979">Tillier 1979</bibRefCitation>
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,
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<bibRefCitation author="Emberton, KC" journalOrPublisher="Proceedings of the National Academy of Sciences of Philadelphia" pageId="22" pageNumber="23" pagination="147 - 189" title="Thirty new species of Madagascan land snails (Mollusca: Gastropoda)." volume="145" year="1994">Emberton 1994</bibRefCitation>
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).
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<bibRefCitation author="van Mol, JJ" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="23" pageNumber="24" pagination="1 - 234" title="Revision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie-Systematique-Zoogeographie." volume="180" year="1970">Van Mol (1970)</bibRefCitation>
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treated all African genera in Urocyclidae: Urocyclinae or Gymnarioninae.
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<bibRefCitation author="Tillier, S" journalOrPublisher="The Veliger" pageId="24" pageNumber="25" pagination="361 - 368" title="Malagarion paenelimax gen. nov., spec. nov., a new slug-like helicarionid from Madagascar." volume="21" year="1979">Tillier (1979)</bibRefCitation>
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considered
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Malagarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malagarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Malagarion</taxonomicName>
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to belong, with the Mascarene
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Colparion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colparion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Colparion</taxonomicName>
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Laidlaw, 1938 in Helicarionidae: Ariophantinae and not Urocyclinae(/idae). He noted that the radula, but not genitalia, of
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Malagarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malagarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Malagarion</taxonomicName>
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was similar to the Asian Durgellini (founded on the Burmese semi-slug
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<taxonomicName class="Gastropoda" family="Helicarionidae" genus="Durgella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Durgella" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Durgella</taxonomicName>
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Blanford, 1863).
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<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of East African Natural History (Special supplement)" pageId="24" pageNumber="25" pagination="1 - 36" title="East African slugs of the family Urocyclidae, III & IV. The genus Trichotoxon." volume="7" year="1961">Verdcourt and Polhill (1961)</bibRefCitation>
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had also noted a radula similarity between their East Usambara species and the Durgellinae, although this was not discussed by
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">van Goethem (1977)</bibRefCitation>
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. Certainly the radula would be unique in Urocyclinae sensu
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<bibRefCitation author="van Goethem, JL" journalOrPublisher="Annales Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques" pageId="22" pageNumber="23" pagination="1 - 355" title="Revision systematique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)." volume="218" year="1977">van Goethem (1977)</bibRefCitation>
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, and resembles
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<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Malagarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malagarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Malagarion</taxonomicName>
|
||
in the size and number of teeth, and the large ectocones (at least on the more marginal teeth of
|
||
<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Malagarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malagarion" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Malagarion</taxonomicName>
|
||
). The monophyly of these major groups is questionable while the systematics of trop
|
||
<pageBreakToken pageId="10" pageNumber="11" start="start">ical</pageBreakToken>
|
||
Limacoidea is still far from resolved (e.g.
|
||
<bibRefCitation author="Tillier, S" journalOrPublisher="The Veliger" pageId="24" pageNumber="25" pagination="361 - 368" title="Malagarion paenelimax gen. nov., spec. nov., a new slug-like helicarionid from Madagascar." volume="21" year="1979">Tillier 1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Hausdorf, B" journalOrPublisher="Journal of Molluscan Studies" pageId="23" pageNumber="24" pagination="35 - 66" title="Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora)." volume="64" year="1998">Hausdorf 1998</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="24" pageNumber="25" pagination="1166 - 1307" title="Treatise on recent terrestrial pulmonate molluscs. 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae." volume="2" year="2002">Schileyko 2002</bibRefCitation>
|
||
), but it remains possible that this species is related to one of them rather than other
|
||
<taxonomicName class="Gastropoda" family="Urocyclidae" genus="Dendrolimax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dendrolimax" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">Dendrolimax</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph pageId="10" pageNumber="11">Distribution:</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Pemba island; probably also East Usambara Mts.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph pageId="10" pageNumber="11">Etymology:</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">vangoethemi, a noun in the generative case, for Dr. J. L. van Goethem of IRSNB, in recognition of his thorough and highly accessible monograph on Afrotropical urocyclid slugs.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |