treatments-xml/data/86/2D/C0/862DC000FFB2FFE50D2EFEE6FB30FB5D.xml

<document id="E36BCB00E18568B142ED6F20B658EDE9" ID-CLB-Dataset="53406" ID-DOI="10.1093/zoolinnean/zlac098" ID-GBIF-Dataset="5d7fb159-154d-4918-9611-642b07af490e" ID-ISSN="0024-4082" ID-Zenodo-Dep="7981596" ID-ZooBank="142F33C8-72A5-4BDAAEAE-B2F0DBAB5413" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="admin" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1685085173754" checkinUser="plazi" docAuthor="Marick, Jit, Brabec, Jan, Choudhury, Anindo, Scholz, Tomáš &amp; Ash, Anirban" docDate="2023" docId="862DC000FFB2FFE50D2EFEE6FB30FB5D" docLanguage="en" docName="zlac098.pdf" docOrigin="Zoological Journal of the Linnean Society 198" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Gangesia mukutmanipurensis Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash 2023, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="522" masterDocId="7A14B878FFB5FFE80E58FF95FF84FFBD" masterDocTitle="The evolution of an ancient tapeworm lineage in its catfish hosts: vicariance, dispersal and diversification in Gangesiinae (Cestoda: Proteocephalidae)" masterLastPageNumber="533" masterPageNumber="509" pageNumber="516" updateTime="1699509586099" updateUser="plazi">
<mods:mods id="75D504DE29E8597528236E8CB7B3EBA4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="712958FA2E610501B85CE0D6D618F1C7">
<mods:title id="FF3F0D733B3A7A76EBA17E96C362E0A4">The evolution of an ancient tapeworm lineage in its catfish hosts: vicariance, dispersal and diversification in Gangesiinae (Cestoda: Proteocephalidae)</mods:title>
</mods:titleInfo>
<mods:name id="054AE8F351BAC35D9ADBA0B6323146EB" type="personal">
<mods:role id="11108452B91A136F8825790CB28C0BCD">
<mods:roleTerm id="B33447561A96762006EDC4D8216F17B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E97B57035571B63BA06D1A873378998">Marick, Jit</mods:namePart>
</mods:name>
<mods:name id="2311B08F3F566849C0098250ED2EFE46" type="personal">
<mods:role id="E0014163D87E1B6D90AB3EF9768F0E16">
<mods:roleTerm id="D98345A50DAA284ABFE8608085C96CE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B13C1E48494B67F1D627B49C52F0073F">Brabec, Jan</mods:namePart>
</mods:name>
<mods:name id="220918A437C6E0C76EB42D8464800F60" type="personal">
<mods:role id="4410D6F995E2322ED83BAD163FF92AF2">
<mods:roleTerm id="8CF96DE397D8A44D851A20AF03EFDAFD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="964B459917C97754B3BAD03AF59CC5A7">Choudhury, Anindo</mods:namePart>
</mods:name>
<mods:name id="322F0F1402C650F9D942D81F9802C137" type="personal">
<mods:role id="5D2146718BA3120383534324B7385ED9">
<mods:roleTerm id="294FCD2F8331F11997A91E5FEE6E3AF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6A12023D5B7734412BF2ECA93B48823">Scholz, Tomáš</mods:namePart>
</mods:name>
<mods:name id="E8B88E2F95B3BDBBA8E1A523A0AD20E9" type="personal">
<mods:role id="325A9E312C36C7F7F74D4A1E239CEC0D">
<mods:roleTerm id="7C1583A23F7128314535221700848601">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A21B1ADCA0438FAD2EDE74C94E49F195">Ash, Anirban</mods:namePart>
</mods:name>
<mods:typeOfResource id="C392670E17DC8F8CF72C7ABE6806893E">text</mods:typeOfResource>
<mods:relatedItem id="63E1F5493EC2E9FEA63BAF73DF068965" type="host">
<mods:titleInfo id="2CB8D92A9ECD870B8DFAEC6AD61B539C">
<mods:title id="55DA310F41EF1F5347D0FEA5F33AF9BC">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="95E740EA77D2D27AC0CA5D440093B736">
<mods:date id="E5A03C050CAD2A1C73D1053D8A9CBE17">2023</mods:date>
<mods:detail id="4D7EC76163CA7D8583F4436D44B424A5" type="pubDate">
<mods:number id="6B841774E8A8458302BC6375C261CF7B">2023-03-20</mods:number>
</mods:detail>
<mods:detail id="5D9D4D7711BEB3095112CF2B1F4EEB8C" type="volume">
<mods:number id="430A618FC02FC68E33422780847AEA85">198</mods:number>
</mods:detail>
<mods:extent id="9590786D20083947E2FF4534155AFB5C" unit="page">
<mods:start id="3184FC4A4CD8C4B62497DCB7B3CAA822">509</mods:start>
<mods:end id="01DC3240E6B1B82026AFB60336139948">533</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EBD54E8073E656F4BE2206601E687E01">journal article</mods:classification>
<mods:identifier id="C3F33AD6E41C2E3161F97F4A9F99C070" type="CLB-Dataset">53406</mods:identifier>
<mods:identifier id="CC505C70C9756A3F258E1304BEE2FBBA" type="DOI">10.1093/zoolinnean/zlac098</mods:identifier>
<mods:identifier id="7A8AA701E4F439F1D13C4472AF162142" type="GBIF-Dataset">5d7fb159-154d-4918-9611-642b07af490e</mods:identifier>
<mods:identifier id="09C5348573667680244A155B1586D99E" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="AB916A5D405B1BC13CE74B2EC569F507" type="Zenodo-Dep">7981596</mods:identifier>
<mods:identifier id="00A86D82E51496F61D60CFCFA2AAAE4F" type="ZooBank">142F33C8-72A5-4BDAAEAE-B2F0DBAB5413</mods:identifier>
</mods:mods>
<treatment id="862DC000FFB2FFE50D2EFEE6FB30FB5D" ID-DOI="http://doi.org/10.5281/zenodo.7972992" ID-GBIF-Taxon="209633978" ID-Zenodo-Dep="7972992" LSID="urn:lsid:plazi:treatment:862DC000FFB2FFE50D2EFEE6FB30FB5D" httpUri="http://treatment.plazi.org/id/862DC000FFB2FFE50D2EFEE6FB30FB5D" lastPageId="13" lastPageNumber="522" pageId="7" pageNumber="516">
<subSubSection id="469E229DFFB2FFEF0D2EFEE6FAE3FE36" box="[886,1383,371,395]" pageId="7" pageNumber="516" type="nomenclature">
<paragraph id="0E3B7116FFB2FFEF0D2EFEE6FAE3FE36" blockId="7.[886,1383,371,395]" box="[886,1383,371,395]" pageId="7" pageNumber="516">
<heading id="5573C67AFFB2FFEF0D2EFEE6FAE3FE36" box="[886,1383,371,395]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="516" reason="2">
<taxonomicName id="C9840A95FFB2FFEF0D2EFEE6FB79FE37" ID-CoL="ceec21fa-1122-4b28-a719-560a62b22c85" authority="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash, 2023" authorityName="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash" authorityYear="2023" box="[886,1277,371,395]" class="Cestoda" family="Proteocephalidae" genus="Gangesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="7" pageNumber="528" phylum="Platyhelminthes" rank="species" species="mukutmanipurensis" status="sp. nov.">
<smallCapsWord id="08DDE7CAFFB2FFEF0D2EFEE6FC75FE37" baselines="389,389" box="[886,1009,371,395]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gangesia" pageId="7" pageNumber="516">GANGESIA</smallCapsWord>
<smallCapsWord id="08DDE7CAFFB2FFEF0DA0FEE2FB79FE37" baselines="389" box="[1016,1277,375,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="mukutmanipurensis" pageId="7" pageNumber="516">MUKUTMANIPURENSIS</smallCapsWord>
</taxonomicName>
<taxonomicNameLabel id="27C3107FFFB2FFEF0B5CFEE2FAE3FE36" box="[1284,1383,371,395]" pageId="7" pageNumber="516" rank="species">
<emphasis id="3CF0AD04FFB2FFEF0B5CFEE2FAE3FE36" bold="true" box="[1284,1383,371,395]" pageId="7" pageNumber="516">
<smallCapsWord id="08DDE7CAFFB2FFEF0B5CFEE2FAA5FE37" baselines="389" box="[1284,1313,375,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="7" pageNumber="516">SP</smallCapsWord>
. 
<smallCapsWord id="08DDE7CAFFB2FFEF0B77FEE2FAE4FE37" baselines="389" box="[1327,1376,375,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="7" pageNumber="516">NOV</smallCapsWord>
.
</emphasis>
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB2FFE50A6BFE0FFDFBFB8A" lastPageId="13" lastPageNumber="522" pageId="7" pageNumber="516" type="description">
<paragraph id="0E3B7116FFB2FFEF0A6BFE0FFB2EFE0E" blockId="7.[1075,1194,410,435]" box="[1075,1194,410,435]" pageId="7" pageNumber="516">
(
<figureCitation id="96BF6D93FFB2FFEF0A63FE0FFB26FE0E" box="[1083,1186,410,435]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="10.[145,223,1284,1306]" captionStartId-1="11.[163,245,1700,1722]" captionStartId-2="12.[147,227,1474,1496]" captionTargetBox-0="[147,1420,197,1249]" captionTargetBox-1="[232,1377,199,1669]" captionTargetBox-2="[148,1425,196,1444]" captionTargetId-0="figure-280@10.[143,1426,195,1254]" captionTargetId-1="figure-24@11.[227,1378,196,1671]" captionTargetId-2="figure-194@12.[147,1427,195,1445]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." captionText-1="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." captionText-2="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi-0="http://doi.org/10.5281/zenodo.7981608" figureDoi-1="http://doi.org/10.5281/zenodo.7981610" figureDoi-2="http://doi.org/10.5281/zenodo.7981612" httpUri-0="https://zenodo.org/record/7981608/files/figure.png" httpUri-1="https://zenodo.org/record/7981610/files/figure.png" httpUri-2="https://zenodo.org/record/7981612/files/figure.png" pageId="7" pageNumber="516">
<smallCapsWord id="08DDE7CAFFB2FFEF0A63FE0FFBF5FE0F" baselines="429,429" box="[1083,1137,410,435]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="7" pageNumber="516">FIGS</smallCapsWord>
4–6
</figureCitation>
)
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D63FE56FA27FE4A" blockId="7.[827,1443,450,503]" pageId="7" pageNumber="516">
<emphasis id="3CF0AD04FFB2FFEF0D63FE56FB0DFE65" box="[827,1161,450,472]" italics="true" pageId="7" pageNumber="516">Z o o B a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act: 
<uuid id="7A224BC3FFB2FFEF0DDEFE74FA27FE4A" box="[902,1443,481,503]" pageId="7" pageNumber="516">142F33C8-72A5-4BDA-AEAE-B2F0DBAB5413</uuid>
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D63FDD6FC43FDE7" blockId="7.[827,967,579,602]" box="[827,967,579,602]" pageId="7" pageNumber="516">
<emphasis id="3CF0AD04FFB2FFEF0D63FDD6FC43FDE7" box="[827,967,579,602]" italics="true" pageId="7" pageNumber="516">Description</emphasis>
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D63FDFFFC19FBB3" blockId="7.[827,1443,618,1898]" pageId="7" pageNumber="516">
<materialsCitation id="BEEC7B4BFFB2FFEF0D63FDFFFC1DFBB3" ID-GBIF-Occurrence="4118152302" country="India" location="West Bengal" pageId="7" pageNumber="516" specimenCount="110" specimenCount-immature="94" stateProvince="West Bengal">
Based on 
<specimenCount id="1882BA9FFFB2FFEF0DF0FDFFFBBBFD3D" box="[936,1087,618,640]" pageId="7" pageNumber="516" type="generic">16 specimens</specimenCount>
from 
<taxonomicName id="C9840A95FFB2FFEF0ADAFDFFFAE3FDC2" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[1154,1383,618,639]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="516" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="3CF0AD04FFB2FFEF0ADAFDFFFAE3FDC2" box="[1154,1383,618,639]" italics="true" pageId="7" pageNumber="516">Ompok bimaculatus</emphasis>
</taxonomicName>
from 
<collectingRegion id="CC40BFF4FFB2FFEF0D63FD1FFC49FD22" box="[827,973,649,671]" country="India" name="West Bengal" pageId="7" pageNumber="516">West Bengal</collectingRegion>
, 
<collectingCountry id="76933186FFB2FFEF0D81FD1CFB9CFD22" box="[985,1048,649,671]" name="India" pageId="7" pageNumber="516">India</collectingCountry>
, including three scoleces observed using SEM: strobila more than 
<quantity id="C97CDCF3FFB2FFEF0AF5FD3DFA8FFD00" box="[1197,1291,680,702]" metricMagnitude="-1" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="516" unit="mm" value="158.0">158 mm</quantity>
long (
<emphasis id="3CF0AD04FFB2FFEF0B0DFD3CFAEDFD03" box="[1365,1385,681,702]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 3), with acraspedote proglottids, up to 1.96 mm wide (
<emphasis id="3CF0AD04FFB2FFEF0D1BFD73FCD3FD46" box="[835,855,742,763]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 3), consisting of more than 143 proglottids (
<emphasis id="3CF0AD04FFB2FFEF0B0FFD73FAEFFD46" box="[1367,1387,742,763]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 3): 40– 
<specimenCount id="1882BA9FFFB2FFEF0D3DFC91FC7DFCA7" box="[869,1017,772,794]" pageId="7" pageNumber="516" type="immature">94 immature</specimenCount>
(up to appearance of spermatozoa in vas deferens), four to five mature (up to appearance of eggs in uterus), 67–74 pregravid (up to appearance of hooks in oncospheres) and gravid. Proglottids variable in shape, usually slightly wider than long in mature and pregravid and longer than wide in gravid towards terminal end; mature proglottids 575–1038 (733) long by 725–1413 (990) wide, L/W = 0.43–1.59 (0.79) (
<emphasis id="3CF0AD04FFB2FFEF0D1BFC6FFCD3FBB2" box="[835,855,1018,1039]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 18)
</materialsCitation>
.
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D0BFB8DFC30FA20" blockId="7.[827,1443,618,1898]" pageId="7" pageNumber="516">
Inner longitudinal musculature well-developed, anastomosed, forming isolated bundles of muscle fibres (
<figureCitation id="96BF6D93FFB2FFEF0D1BFBC0FC3DFBD6" box="[835,953,1109,1131]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="7" pageNumber="516">Fig. 4C, D</figureCitation>
). Subtegumental muscles well-developed (
<figureCitation id="96BF6D93FFB2FFEF0D1CFBE1FC3FFB34" box="[836,955,1140,1162]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="7" pageNumber="516">Fig. 4C, D</figureCitation>
). Osmoregulatory canals running alongside vitelline follicles. Ventral osmoregulatory canals thin-walled, width representing 4–7% of width of mature proglottids, overlapping lateral-most testes, form anastomoses in some proglottids. Dorsal osmoregulatory canal narrow, thick-walled, prominent in immature proglottids but gradually disintegrating with maturation of proglottids, not observable in gravid proglottids, canal may reach lateral-most testes (
<figureCitation id="96BF6D93FFB2FFEF0B2AFAFCFCFCFA20" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="7" pageNumber="516">Figs 4A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFB2FFEF0DDDFA1DFC21FA23" box="[901,933,1416,1438]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="7" pageNumber="516">5E</figureCitation>
).
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D0BFA33FAC6F9E8" blockId="7.[827,1443,618,1898]" pageId="7" pageNumber="516">
Scolex wider than neck, 134–203 (157) long by 144–203 wide (169) (
<emphasis id="3CF0AD04FFB2FFEF0A19FA53FBD1FA66" box="[1089,1109,1478,1499]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 7). Rostellum-like organ 34–61 (51) long and 53–89 (71) wide, devoid of hooks. Suckers four, uniloculate, devoid of hooklets on rims (
<figureCitation id="96BF6D93FFB2FFEF0D1BF9B4FC3BF98A" box="[835,959,1569,1591]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="7" pageNumber="516">Figs 5A–C</figureCitation>
, 
<figureCitation id="96BF6D93FFB2FFEF0D95F9B4FB8AF98B" box="[973,1038,1569,1591]" captionStart="Figure 6" captionStartId="12.[147,227,1474,1496]" captionTargetBox="[148,1425,196,1444]" captionTargetId="figure-194@12.[147,1427,195,1445]" captionTargetPageId="12" captionText="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi="http://doi.org/10.5281/zenodo.7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="7" pageNumber="516">6A, B</figureCitation>
), 59–89 (76) in diameter (
<emphasis id="3CF0AD04FFB2FFEF0B1EF9B7FADEF98A" box="[1350,1370,1570,1591]" italics="true" pageId="7" pageNumber="516">N</emphasis>
= 17); R/S = 0.75–1.14 (0.93), R/X = 0.35–0.50 (0.41).
</paragraph>
<paragraph id="0E3B7116FFB2FFEF0D0BF9CBFB97F972" blockId="7.[827,1443,618,1898]" pageId="7" pageNumber="516">
Prominent retractor muscle bundles surrounding rostellum-like organ, joining at level of suckers, forming a wide band of muscles (retractors) in neck region (
<figureCitation id="96BF6D93FFB2FFEF0DCAF92FFB87F96D" box="[914,1027,1722,1744]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="7" pageNumber="516">Fig. 5B–D</figureCitation>
).
</paragraph>
<paragraph id="0E3B7116FFB2FFE20D0BF94CFE66F9AE" blockId="7.[827,1443,618,1898]" lastBlockId="10.[145,762,1473,1893]" lastPageId="10" lastPageNumber="519" pageId="7" pageNumber="516">
Rostellum-like organ covered with capilliform filitriches (
<figureCitation id="96BF6D93FFB2FFEF0DE0F96DFBA8F8B3" box="[952,1068,1784,1806]" captionStart="Figure 6" captionStartId="12.[147,227,1474,1496]" captionTargetBox="[148,1425,196,1444]" captionTargetId="figure-194@12.[147,1427,195,1445]" captionTargetPageId="12" captionText="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi="http://doi.org/10.5281/zenodo.7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="7" pageNumber="516">Fig. 6C–G</figureCitation>
), apical part additionally armed with several rows of large, coniform spinitriches (
<figureCitation id="96BF6D93FFB2FFEF0D1BF8A0FC5CF8F7" box="[835,984,1845,1867]" captionStart="Figure 6" captionStartId="12.[147,227,1474,1496]" captionTargetBox="[148,1425,196,1444]" captionTargetId="figure-194@12.[147,1427,195,1445]" captionTargetPageId="12" captionText="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi="http://doi.org/10.5281/zenodo.7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="7" pageNumber="516">Fig. 6A–C, H</figureCitation>
). Suckers also covered with capilliform filitriches, rim of suckers with coniform spinitriches that are smaller than those on rostellum (
<figureCitation id="96BF6D93FFBFFFE20C20FA54FD6FFA6A" box="[632,747,1473,1495]" captionStart="Figure 6" captionStartId="12.[147,227,1474,1496]" captionTargetBox="[148,1425,196,1444]" captionTargetId="figure-194@12.[147,1427,195,1445]" captionTargetPageId="12" captionText="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi="http://doi.org/10.5281/zenodo.7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="10" pageNumber="519">Fig. 6D–F</figureCitation>
). Surface of scolex between suckers bearing interspersed gladiate spinitriches (
<figureCitation id="96BF6D93FFBFFFE20FD1FA6BFE57F9A9" box="[393,467,1534,1556]" captionStart="Figure 6" captionStartId="12.[147,227,1474,1496]" captionTargetBox="[148,1425,196,1444]" captionTargetId="figure-194@12.[147,1427,195,1445]" captionTargetPageId="12" captionText="Figure 6. Scanning electron micrographs of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus. A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region." figureDoi="http://doi.org/10.5281/zenodo.7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="10" pageNumber="519">Fig. 6I</figureCitation>
).
</paragraph>
<caption id="5AFB219EFFBDFFE00ECBF976FC60F8D2" ID-DOI="http://doi.org/10.5281/zenodo.7981600" ID-Zenodo-Dep="7981600" httpUri="https://zenodo.org/record/7981600/files/figure.png" pageId="8" pageNumber="517" startId="8.[147,229,1763,1785]" targetBox="[244,1330,198,1732]" targetPageId="8" targetType="figure">
<paragraph id="0E3B7116FFBDFFE00ECBF976FC60F8D2" blockId="8.[147,1427,1763,1903]" pageId="8" pageNumber="517">
<emphasis id="3CF0AD04FFBDFFE00ECBF976FE85F945" bold="true" box="[147,257,1763,1785]" pageId="8" pageNumber="517">Figure 1.</emphasis>
Phylogenetic interrelationships of basal groups of 
<taxonomicName id="C9840A95FFBDFFE00D19F971FBC6F947" authorityName="Caira, Jensen, Waeschenbach, Olson &amp; Littlewood" authorityYear="2014" box="[833,1090,1764,1786]" class="Cestoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="8" pageNumber="517" phylum="Platyhelminthes" rank="order">Onchoproteocephalidea</taxonomicName>
with focus on the subfamilies 
<taxonomicName id="C9840A95FFBDFFE00ECBF894FE91F8AA" authorityName="Mola" authorityYear="1929" box="[147,277,1793,1815]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Onchoproteocephalidea" pageId="8" pageNumber="524" phylum="Platyhelminthes" rank="subFamily" subFamily="Gangesiinae">Gangesiinae</taxonomicName>
and 
<taxonomicName id="C9840A95FFBDFFE00F11F894FD87F8AA" authorityName="Freze" authorityYear="1963" box="[329,515,1793,1815]" pageId="8" pageNumber="516" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
based on the maximum likelihood analysis of the single gene 
<emphasis id="3CF0AD04FFBDFFE00AC8F894FB28F8AA" box="[1168,1196,1793,1815]" italics="true" pageId="8" pageNumber="517">lsr</emphasis>
DNA (above) and 
<emphasis id="3CF0AD04FFBDFFE00B3FF894FA16F8AA" box="[1383,1426,1793,1815]" italics="true" pageId="8" pageNumber="517">COI</emphasis>
(below) datasets. Nodal values depict SH-like approximated likelihood ratio test values (SH-aLRT) (10 000 replicates) followed by standard bootstrap values (1 000 replicates). Only values above 50 are shown. The subfamilies 
<taxonomicName id="C9840A95FFBDFFE00A81F8A9FA17F8EF" authorityName="Freze" authorityYear="1963" box="[1241,1427,1852,1874]" pageId="8" pageNumber="516" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
(green) and 
<taxonomicName id="C9840A95FFBDFFE00F49F8CCFE17F8D2" authorityName="Mola" authorityYear="1929" box="[273,403,1881,1903]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Onchoproteocephalidea" pageId="8" pageNumber="524" phylum="Platyhelminthes" rank="subFamily" subFamily="Gangesiinae">Gangesiinae</taxonomicName>
(blue), along with the outgroups (grey), are set in colour.
</paragraph>
</caption>
<caption id="5AFB219EFFBCFFE10EFBFC73FD22FBCF" ID-DOI="http://doi.org/10.5281/zenodo.7981602" ID-Zenodo-Dep="7981602" httpUri="https://zenodo.org/record/7981602/files/figure.png" pageId="9" pageNumber="518" startId="9.[163,245,998,1020]" targetBox="[163,1442,197,968]" targetPageId="9" targetType="figure">
<paragraph id="0E3B7116FFBCFFE10EFBFC73FD22FBCF" blockId="9.[163,1444,998,1138]" pageId="9" pageNumber="518">
<emphasis id="3CF0AD04FFBCFFE10EFBFC73FE95FC46" bold="true" box="[163,273,998,1020]" pageId="9" pageNumber="518">Figure 2.</emphasis>
Phylogenetic interrelationships of basal groups of 
<taxonomicName id="C9840A95FFBCFFE10D0AFC72FBD7FC40" authorityName="Caira, Jensen, Waeschenbach, Olson &amp; Littlewood" authorityYear="2014" box="[850,1107,999,1021]" class="Cestoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="9" pageNumber="518" phylum="Platyhelminthes" rank="order">Onchoproteocephalidea</taxonomicName>
with focus on the subfamilies 
<taxonomicName id="C9840A95FFBCFFE10EFBFB91FEA2FBA7" authorityName="Mola" authorityYear="1929" box="[163,294,1028,1050]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Onchoproteocephalidea" pageId="9" pageNumber="524" phylum="Platyhelminthes" rank="subFamily" subFamily="Gangesiinae">Gangesiinae</taxonomicName>
and 
<taxonomicName id="C9840A95FFBCFFE10F02FB91FD90FBA7" authorityName="Freze" authorityYear="1963" box="[346,532,1028,1050]" pageId="9" pageNumber="516" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
based on the maximum likelihood analysis of concatenated 
<emphasis id="3CF0AD04FFBCFFE10AD6FB91FB2EFBA7" box="[1166,1194,1028,1050]" italics="true" pageId="9" pageNumber="518">lsr</emphasis>
DNA + 
<emphasis id="3CF0AD04FFBCFFE10AAFFB91FAA6FBA7" box="[1271,1314,1028,1050]" italics="true" pageId="9" pageNumber="518">COI</emphasis>
data. Nodal values depict SH-like approximated likelihood ratio test values (SH-aLRT) (10 000 replicates) followed by standard bootstrap values (1000 replicates). Only values above 50 are shown. The subfamilies 
<taxonomicName id="C9840A95FFBCFFE10DF8FBAAFBDEFBE8" authorityName="Freze" authorityYear="1963" box="[928,1114,1087,1109]" pageId="9" pageNumber="516" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
(green) and 
<taxonomicName id="C9840A95FFBCFFE10A82FBAAFAD8FBE8" authorityName="Mola" authorityYear="1929" box="[1242,1372,1087,1109]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Onchoproteocephalidea" pageId="9" pageNumber="524" phylum="Platyhelminthes" rank="subFamily" subFamily="Gangesiinae">Gangesiinae</taxonomicName>
(blue), along with the outgroups (grey), are set in colour.
</paragraph>
</caption>
<caption id="5AFB219EFFBCFFE10EFBF890FE18F8C9" ID-DOI="http://doi.org/10.5281/zenodo.7981606" ID-Zenodo-Dep="7981606" httpUri="https://zenodo.org/record/7981606/files/figure.png" pageId="9" pageNumber="518" startId="9.[163,241,1797,1819]" targetBox="[323,1283,1194,1765]" targetPageId="9" targetType="figure">
<paragraph id="0E3B7116FFBCFFE10EFBF890FE18F8C9" blockId="9.[163,1443,1797,1908]" pageId="9" pageNumber="518">
<emphasis id="3CF0AD04FFBCFFE10EFBF890FE8EF8A7" bold="true" box="[163,266,1797,1819]" pageId="9" pageNumber="518">Figure 3.</emphasis>
Simplified relationships among the genera of the subfamilies 
<taxonomicName id="C9840A95FFBCFFE10DC8F893FB96F8A1" authorityName="Mola" authorityYear="1929" box="[912,1042,1798,1820]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Onchoproteocephalidea" pageId="9" pageNumber="524" phylum="Platyhelminthes" rank="subFamily" subFamily="Gangesiinae">Gangesiinae</taxonomicName>
and 
<taxonomicName id="C9840A95FFBCFFE10A1BF893FB79F8A1" authorityName="Freze" authorityYear="1963" box="[1091,1277,1798,1820]" pageId="9" pageNumber="516" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
after collapsing nodes with SH-aLRT ≤ 80% and BS ≤ 70–80% on the phylogenetic tree in Figure 2. This simplified tree, having essentially similar topology once the poorly supported nodes on the 
<emphasis id="3CF0AD04FFBCFFE10D58F8D5FC98F8EB" box="[768,796,1856,1878]" italics="true" pageId="9" pageNumber="518">lsr</emphasis>
DNA tree are collapsed, provides the basis for discussions of historical biogeography.
</paragraph>
</caption>
<caption id="5AFB219EFFBFFFE20EC9FA91FB2FFA32" ID-DOI="http://doi.org/10.5281/zenodo.7981608" ID-Zenodo-Dep="7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="10" pageNumber="519" startId="10.[145,223,1284,1306]" targetBox="[147,1420,197,1249]" targetPageId="10" targetType="figure">
<paragraph id="0E3B7116FFBFFFE20EC9FA91FB2FFA32" blockId="10.[145,1425,1284,1423]" pageId="10" pageNumber="519">
<emphasis id="3CF0AD04FFBFFFE20EC9FA91FF7DFAA4" bold="true" box="[145,249,1284,1306]" pageId="10" pageNumber="519">Figure 4.</emphasis>
Line drawings of 
<taxonomicName id="C9840A95FFBFFFE20F9AFA91FD7AFAA7" authority="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash, 2023" authorityName="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash" authorityYear="2023" box="[450,766,1284,1306]" class="Cestoda" family="Proteocephalidae" genus="Gangesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="10" pageNumber="519" phylum="Platyhelminthes" rank="species" species="mukutmanipurensis" status="sp. nov.">
<emphasis id="3CF0AD04FFBFFFE20F9AFA91FD7AFAA7" box="[450,766,1284,1306]" italics="true" pageId="10" pageNumber="519">Gangesia mukutmanipurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C3107FFFBFFFE20D5CFA91FCD4FAA7" box="[772,848,1284,1306]" pageId="10" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="C9840A95FFBFFFE20DD5FA91FBE5FAA7" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[909,1121,1284,1306]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="519" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="3CF0AD04FFBFFFE20DD5FA91FBE5FAA7" box="[909,1121,1284,1306]" italics="true" pageId="10" pageNumber="519">Ompok bimaculatus</emphasis>
</taxonomicName>
(specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal.
</paragraph>
</caption>
<paragraph id="0E3B7116FFBFFFE20EF1F988FE9AF9CD" blockId="10.[145,762,1473,1893]" pageId="10" pageNumber="519">
Proliferative zone 4.7–7.2 mm long and 333–416 wide (
<emphasis id="3CF0AD04FFBFFFE20EB8F9A9FF70F9EC" box="[224,244,1596,1617]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 2). Neck region covered with gladiate spinitriches.
</paragraph>
<paragraph id="0E3B7116FFBFFFE20EF1F9ECFB1BFA49" blockId="10.[145,762,1473,1893]" lastBlockId="10.[809,1425,1473,1893]" pageId="10" pageNumber="519">
Testes medullary, in single field, spherical to oval, 31–81 (50) in diameter (
<emphasis id="3CF0AD04FFBFFFE20FF4F90DFE44F910" box="[428,448,1688,1709]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 115), numbering 82–144 (109) (
<emphasis id="3CF0AD04FFBFFFE20E81F922FF69F971" box="[217,237,1719,1740]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 28), mostly forming single layer (
<figureCitation id="96BF6D93FFBFFFE20C27F923FD77F971" box="[639,755,1718,1740]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="10" pageNumber="519">Figs 4A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFBFFFE20EC9F940FF35F956" box="[145,177,1749,1771]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="10" pageNumber="519">5E</figureCitation>
), occupying 65–80% of length of proglottids. Cirrus-sac elongate, thick-walled, 163–269 (199) long by 63–88 (74) wide [
<emphasis id="3CF0AD04FFBFFFE20F54F886FEA4F895" box="[268,288,1811,1832]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 17; L/W = 2.0–3.7 (2.7)]; C/P = 17–26% (20%), i.e. 1/6–1/4 of width of proglottids. Genital pore irregularly alternating, pre-equatorial, situated at 23–40% (32%) (
<emphasis id="3CF0AD04FFBFFFE20DB8FA54FC70FA6B" box="[992,1012,1473,1494]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 13) of length of proglottids from anterior margin (
<figureCitation id="96BF6D93FFBFFFE20DB6FA4AFBE7FA48" box="[1006,1123,1503,1525]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="10" pageNumber="519">Figs 4A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFBFFFE20A28FA4AFB14FA48" box="[1136,1168,1503,1525]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="10" pageNumber="519">5E</figureCitation>
).
</paragraph>
<paragraph id="0E3B7116FFBFFFE20D19FA6BFA99F89A" blockId="10.[809,1425,1473,1893]" pageId="10" pageNumber="519">
Ovary medullary, 175–338 (228) long, 400–913 (640) wide (
<emphasis id="3CF0AD04FFBFFFE20D2AF988FC02F98F" box="[882,902,1565,1586]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 18), occupying 26–38% (31%) of length of proglottid, bilobed, each lobe spherical to rectangular (
<figureCitation id="96BF6D93FFBFFFE20D69F9CFFC22F9D2" box="[817,934,1626,1648]" captionStart="Figure 3" captionStartId="9.[163,241,1797,1819]" captionTargetBox="[323,1283,1194,1765]" captionTargetId="figure-24@9.[320,1286,1188,1768]" captionTargetPageId="9" captionText="Figure 3. Simplified relationships among the genera of the subfamilies Gangesiinae and Acanthotaeniinae after collapsing nodes with SH-aLRT ≤ 80% and BS ≤ 70–80% on the phylogenetic tree in Figure 2. This simplified tree, having essentially similar topology once the poorly supported nodes on the lsrDNA tree are collapsed, provides the basis for discussions of historical biogeography." figureDoi="http://doi.org/10.5281/zenodo.7981606" httpUri="https://zenodo.org/record/7981606/files/figure.png" pageId="10" pageNumber="519">Figs 3A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFBFFFE20DEBF9CFFC57F9D2" box="[947,979,1626,1647]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="10" pageNumber="519">4E</figureCitation>
); O/P = 55–71% (65%) (
<emphasis id="3CF0AD04FFBFFFE20A86F9CEFB76F9CD" box="[1246,1266,1627,1648]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 18). Mehlis’ gland 81–113 (96) in diameter (
<emphasis id="3CF0AD04FFBFFFE20AFBF9ECFB33F933" box="[1187,1207,1657,1678]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 8), representing 8–11% of width of proglottid. Vagina thick-walled, posterior (58%) or anterior (42%, 
<emphasis id="3CF0AD04FFBFFFE20AD5F922FB25F971" box="[1165,1185,1719,1740]" italics="true" pageId="10" pageNumber="519">N</emphasis>
= 100) to cirrus-sac, with higher concentration of chromophilic cells in its distal (terminal) part; ring-like vaginal sphincter present near genital atrium (
<figureCitation id="96BF6D93FFBFFFE20A17F887FB40F89A" box="[1103,1220,1810,1832]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="10" pageNumber="519">Figs 4A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFBFFFE20A89F887FA89F89A" box="[1233,1293,1810,1832]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="10" pageNumber="519">5E, F</figureCitation>
).
</paragraph>
<paragraph id="0E3B7116FFBFFFE40D19F8A4FACBF8F3" blockId="10.[809,1425,1473,1893]" lastBlockId="12.[809,1426,1634,1870]" lastPageId="12" lastPageNumber="521" pageId="10" pageNumber="519">
Vitelline follicles oval, medullary, some follicles paramuscular (between muscle fibres of inner longitudinal musculature to cortex) (
<figureCitation id="96BF6D93FFB9FFE40C18F9F7FD37F9CA" box="[576,691,1634,1656]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="12" pageNumber="521">Fig. 4C, D</figureCitation>
), present in two longitudinal bands on lateral margin of proglottid, occupying 77–91% (86%) of length of proglottid on poral side, 85–96% (92%) on aporal side in mature proglottids (
<emphasis id="3CF0AD04FFB9FFE40FC6F948FE36F94F" box="[414,434,1757,1778]" italics="true" pageId="12" pageNumber="521">N</emphasis>
= 17); bands interrupted at level of terminal genitalia on ventral side, with few follicles on dorsal side in more than half of mature and pregravid proglottids bands mostly interrupted (92%), both dorsally and ventrally in gravid proglottids (
<figureCitation id="96BF6D93FFB9FFE40B38F9F7FCECF92B" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="12" pageNumber="521">Figs 4A, B</figureCitation>
, 
<figureCitation id="96BF6D93FFB9FFE40D2EF915FC37F92B" box="[886,947,1664,1686]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="12" pageNumber="521">5E, F</figureCitation>
). Uterus medullary, with type 1 development (
<emphasis id="3CF0AD04FFB9FFE40D2DF935FC31F908" box="[885,949,1696,1717]" italics="true" pageId="12" pageNumber="521">sensu</emphasis>
<bibRefCitation id="6A150CE7FFB9FFE40DE6F90AFB6CF909" author="de Chambrier" box="[958,1256,1695,1717]" etAl="et al." firstAuthor="de Chambrier" pageId="12" pageNumber="521" pagination="159 - 171" refId="ref13406" refString="de Chambrier A, Zehnder MP, Vaucher C, Mariaux J. 2004. The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology 57: 159 - 171." type="journal article" year="2004">
de Chambrier 
<emphasis id="3CF0AD04FFB9FFE40A3FF935FB25F909" box="[1127,1185,1695,1717]" italics="true" pageId="12" pageNumber="521">et al.</emphasis>
, 2004
</bibRefCitation>
). In pregravid proglottids, lateral diverticula thin-walled, 17–23 (20) in number on each side (
<emphasis id="3CF0AD04FFB9FFE40A1CF948FBDCF94F" box="[1092,1112,1757,1778]" italics="true" pageId="12" pageNumber="521">N</emphasis>
= 18), occupying up to 58% of proglottid width, may partly overlap ovary (
<figureCitation id="96BF6D93FFB9FFE40B62F96EFCBFF892" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="12" pageNumber="521">Fig. 4A, B</figureCitation>
). Uterus with uterine pores; uterine pores four to seven in number (
<emphasis id="3CF0AD04FFB9FFE40DAEF8ACFB8EF8F3" box="[1014,1034,1849,1870]" italics="true" pageId="12" pageNumber="521">N</emphasis>
= 18), slit-like (
<figureCitation id="96BF6D93FFB9FFE40AE4F8ADFA97F8F3" box="[1212,1299,1848,1870]" captionStart="Figure 4" captionStartId="10.[145,223,1284,1306]" captionTargetBox="[147,1420,197,1249]" captionTargetId="figure-280@10.[143,1426,195,1254]" captionTargetPageId="10" captionText="Figure 4. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, gravid proglottid, ventral view (ZSI/W11083/1/2). B, gravid proglottid, dorsal view (ZSI/W11083/1/2). C, D, cross-sections at level of testicular field and ovary, respectively (ZSI/W11088/1 and ZSI/W11089/1). Abbreviations: doc, dorsal osmoregulatory canal; ilm, internal longitudinal muscles; lud, lateral uterine diverticula; Mg, Mehlis’ gland; ov, ovary; sl, subtegumental layer; te, testes; ut, uterus; vf, vitelline follicles; voc, ventral osmoregulatory canal." figureDoi="http://doi.org/10.5281/zenodo.7981608" httpUri="https://zenodo.org/record/7981608/files/figure.png" pageId="12" pageNumber="521">Figs 4A</figureCitation>
, 
<figureCitation id="96BF6D93FFB9FFE40B78F8ADFAC4F8F3" box="[1312,1344,1848,1870]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="12" pageNumber="521">5E</figureCitation>
).
</paragraph>
<caption id="5AFB219EFFBEFFE30EFBF931FB41F8D7" ID-DOI="http://doi.org/10.5281/zenodo.7981610" ID-Zenodo-Dep="7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="11" pageNumber="520" startId="11.[163,245,1700,1722]" targetBox="[232,1377,199,1669]" targetPageId="11" targetType="figure">
<paragraph id="0E3B7116FFBEFFE30EFBF931FB41F8D7" blockId="11.[163,1444,1700,1898]" pageId="11" pageNumber="520">
<emphasis id="3CF0AD04FFBEFFE30EFBF931FE96F904" bold="true" box="[163,274,1700,1722]" pageId="11" pageNumber="520">Figure 5.</emphasis>
Line drawings of 
<taxonomicName id="C9840A95FFBEFFE30FB5F931FCBFF907" authority="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash, 2023" authorityName="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash" authorityYear="2023" box="[493,827,1700,1722]" class="Cestoda" family="Proteocephalidae" genus="Gangesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="11" pageNumber="520" phylum="Platyhelminthes" rank="species" species="mukutmanipurensis" status="sp. nov.">
<emphasis id="3CF0AD04FFBEFFE30FB5F931FCBFF907" box="[493,827,1700,1722]" italics="true" pageId="11" pageNumber="520">Gangesia mukutmanipurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C3107FFFBEFFE30D1CF931FC13F907" box="[836,919,1700,1722]" pageId="11" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="C9840A95FFBEFFE30D83F931FB3EF907" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[987,1210,1700,1722]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="520" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="3CF0AD04FFBEFFE30D83F931FB3EF907" box="[987,1210,1700,1722]" italics="true" pageId="11" pageNumber="520">Ompok bimaculatus</emphasis>
</taxonomicName>
(specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and 
<emphasis id="3CF0AD04FFBEFFE30EFBF94AFEE1F948" box="[163,357,1759,1781]" italics="true" pageId="11" pageNumber="520">ZSI/W11082/1/1</emphasis>
). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter.
</paragraph>
</caption>
<caption id="5AFB219EFFB9FFE40ECBFA57FA10F98D" ID-DOI="http://doi.org/10.5281/zenodo.7981612" ID-Zenodo-Dep="7981612" httpUri="https://zenodo.org/record/7981612/files/figure.png" pageId="12" pageNumber="521" startId="12.[147,227,1474,1496]" targetBox="[148,1425,196,1444]" targetPageId="12" targetType="figure">
<paragraph id="0E3B7116FFB9FFE40ECBFA57FA10F98D" blockId="12.[147,1428,1474,1584]" pageId="12" pageNumber="521">
<emphasis id="3CF0AD04FFB9FFE40ECBFA57FF79FA6A" bold="true" box="[147,253,1474,1496]" pageId="12" pageNumber="521">Figure 6.</emphasis>
Scanning electron micrographs of 
<taxonomicName id="C9840A95FFB9FFE40C2FFA57FC3CFA65" authority="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash, 2023" authorityName="Marick &amp; Brabec &amp; Choudhury &amp; Scholz &amp; Ash" authorityYear="2023" box="[631,952,1474,1496]" class="Cestoda" family="Proteocephalidae" genus="Gangesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="12" pageNumber="521" phylum="Platyhelminthes" rank="species" species="mukutmanipurensis" status="sp. nov.">
<emphasis id="3CF0AD04FFB9FFE40C2FFA57FC3CFA65" box="[631,952,1474,1496]" italics="true" pageId="12" pageNumber="521">Gangesia mukutmanipurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C3107FFFB9FFE40D98FA57FB8AFA65" box="[960,1038,1474,1496]" pageId="12" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
from 
<emphasis id="3CF0AD04FFB9FFE40A15FA57FAAEFA65" box="[1101,1322,1474,1496]" italics="true" pageId="12" pageNumber="521">
<taxonomicName id="C9840A95FFB9FFE40A15FA57FAA2FA65" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[1101,1318,1474,1496]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="521" phylum="Chordata" rank="species" species="bimaculatus">Ompok bimaculatus</taxonomicName>
.
</emphasis>
A, scolex, subapical view. B, scolex, dorsoventral view. C, detail of rostellum-like organ. D, sucker with coniform spinitriches on the outer rim. E, F, detail of outer rim of the sucker with coniform spinitriches. G, detail of capilliform filitriches on the rostellumlike organ. H, detail of coniform spinitriches on the rostellum-like organ. I, detail of gladiate spinitriches on the neck region.
</paragraph>
</caption>
<caption id="5AFB219EFFB8FFE50EFBFF51FC53FF67" ID-Table-UUID="5AFB219EFFB8FFE50EFBFF51FC53FF67" box="[163,983,196,218]" httpUri="http://table.plazi.org/id/5AFB219EFFB8FFE50EFBFF51FC53FF67" pageId="13" pageNumber="522" startId="13.[163,227,196,217]" targetBox="[163,1437,254,541]" targetIsTable="true" targetPageId="13" targetType="table">
<paragraph id="0E3B7116FFB8FFE50EFBFF51FC53FF67" blockId="13.[163,983,196,218]" box="[163,983,196,218]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFF51FF79FF64" bold="true" box="[163,253,196,217]" pageId="13" pageNumber="522">Table 2.</emphasis>
Prevalence and intensity of infection in Bankura, West Bengal, India
</paragraph>
</caption>
<paragraph id="0E3B7116FFB8FFE50EFBFF6BFA15FDA0" pageId="13" pageNumber="522">
<table id="7C8483B6FFB800170EFBFF6BFA19FDA0" box="[163,1437,254,541]" gridcols="6" gridrows="8" pageId="13" pageNumber="522">
<tr id="B0B47354FFB800170EFBFF6BFA19FE8C" box="[163,1437,254,305]" gridrow="0" isHeader="true" pageId="13" pageNumber="522">
<th id="F3651A28FFB800170EFBFF6BFEF9FE8C" box="[163,381,254,305]" gridcol="0" gridrow="0" pageId="13" pageNumber="522">Locality</th>
<th id="F3651A28FFB800170FB1FF6BFDE2FE8C" box="[489,614,254,305]" gridcol="1" gridrow="0" pageId="13" pageNumber="522">Month/ Year</th>
<th id="F3651A28FFB800170C8CFF6BFCB8FE8C" box="[724,828,254,305]" gridcol="2" gridrow="0" pageId="13" pageNumber="522">No. of fish examined</th>
<th id="F3651A28FFB800170DF0FF6BFB94FE8C" box="[936,1040,254,305]" gridcol="3" gridrow="0" pageId="13" pageNumber="522">No. of fish infected</th>
<th id="F3651A28FFB800170A24FF6BFB6BFE8C" box="[1148,1263,254,305]" gridcol="4" gridrow="0" pageId="13" pageNumber="522">Prevalence</th>
<th id="F3651A28FFB800170B76FF6BFA19FE8C" box="[1326,1437,254,305]" gridcol="5" gridrow="0" pageId="13" pageNumber="522">Intensity (mean)</th>
</tr>
<tr id="B0B47354FFB800170EFBFED8FA19FED5" box="[163,1437,333,360]" gridrow="1" pageId="13" pageNumber="522">
<th id="F3651A28FFB800170EFBFED8FEF9FDA0" box="[163,381,333,541]" gridcol="0" gridrow="1" pageId="13" pageNumber="522" rowspan="7" rowspanBelow="6">Mukutmanipur Dam Lake, Bankura</th>
<td id="F3651A28FFB800170FB1FED8FDE2FED5" box="[489,614,333,360]" gridcol="1" gridrow="1" pageId="13" pageNumber="522">March/2009</td>
<td id="F3651A28FFB800170C8CFED8FCB8FED5" box="[724,828,333,360]" gridcol="2" gridrow="1" pageId="13" pageNumber="522">12</td>
<td id="F3651A28FFB800170DF0FED8FB94FED5" box="[936,1040,333,360]" gridcol="3" gridrow="1" pageId="13" pageNumber="522">1</td>
<td id="F3651A28FFB800170A24FED8FB6BFED5" box="[1148,1263,333,360]" gridcol="4" gridrow="1" pageId="13" pageNumber="522">8%</td>
<td id="F3651A28FFB800170B76FED8FA19FED5" box="[1326,1437,333,360]" gridcol="5" gridrow="1" pageId="13" pageNumber="522">1 (1)</td>
</tr>
<tr id="B0B47354FFB800170EFBFEE5FA19FE3B" box="[163,1437,368,390]" gridrow="2" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FEE5FDE2FE3B" box="[489,614,368,390]" gridcol="1" gridrow="2" pageId="13" pageNumber="522">Jan/2019</td>
<td id="F3651A28FFB800170C8CFEE5FCB8FE3B" box="[724,828,368,390]" gridcol="2" gridrow="2" pageId="13" pageNumber="522">37</td>
<td id="F3651A28FFB800170DF0FEE5FB94FE3B" box="[936,1040,368,390]" gridcol="3" gridrow="2" pageId="13" pageNumber="522">2</td>
<td id="F3651A28FFB800170A24FEE5FB6BFE3B" box="[1148,1263,368,390]" gridcol="4" gridrow="2" pageId="13" pageNumber="522">5%</td>
<td id="F3651A28FFB800170B76FEE5FA19FE3B" box="[1326,1437,368,390]" gridcol="5" gridrow="2" pageId="13" pageNumber="522">2–4 (3)</td>
</tr>
<tr id="B0B47354FFB800170EFBFE1BFA19FE19" box="[163,1437,398,420]" gridrow="3" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FE1BFDE2FE19" box="[489,614,398,420]" gridcol="1" gridrow="3" pageId="13" pageNumber="522">Feb/2019</td>
<td id="F3651A28FFB800170C8CFE1BFCB8FE19" box="[724,828,398,420]" gridcol="2" gridrow="3" pageId="13" pageNumber="522">56</td>
<td id="F3651A28FFB800170DF0FE1BFB94FE19" box="[936,1040,398,420]" gridcol="3" gridrow="3" pageId="13" pageNumber="522">9</td>
<td id="F3651A28FFB800170A24FE1BFB6BFE19" box="[1148,1263,398,420]" gridcol="4" gridrow="3" pageId="13" pageNumber="522">16%</td>
<td id="F3651A28FFB800170B76FE1BFA19FE19" box="[1326,1437,398,420]" gridcol="5" gridrow="3" pageId="13" pageNumber="522">1–4 (1.44)</td>
</tr>
<tr id="B0B47354FFB800170EFBFE38FA19FE7E" box="[163,1437,429,451]" gridrow="4" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FE38FDE2FE7E" box="[489,614,429,451]" gridcol="1" gridrow="4" pageId="13" pageNumber="522">March/2019</td>
<td id="F3651A28FFB800170C8CFE38FCB8FE7E" box="[724,828,429,451]" gridcol="2" gridrow="4" pageId="13" pageNumber="522">37</td>
<td id="F3651A28FFB800170DF0FE38FB94FE7E" box="[936,1040,429,451]" gridcol="3" gridrow="4" pageId="13" pageNumber="522">3</td>
<td id="F3651A28FFB800170A24FE38FB6BFE7E" box="[1148,1263,429,451]" gridcol="4" gridrow="4" pageId="13" pageNumber="522">8%</td>
<td id="F3651A28FFB800170B76FE38FA19FE7E" box="[1326,1437,429,451]" gridcol="5" gridrow="4" pageId="13" pageNumber="522">2–4 (3)</td>
</tr>
<tr id="B0B47354FFB800170EFBFE5EFA19FE5C" box="[163,1437,459,481]" gridrow="5" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FE5EFDE2FE5C" box="[489,614,459,481]" gridcol="1" gridrow="5" pageId="13" pageNumber="522">April/2019</td>
<td id="F3651A28FFB800170C8CFE5EFCB8FE5C" box="[724,828,459,481]" gridcol="2" gridrow="5" pageId="13" pageNumber="522">15</td>
<td id="F3651A28FFB800170DF0FE5EFB94FE5C" box="[936,1040,459,481]" gridcol="3" gridrow="5" pageId="13" pageNumber="522">0</td>
<td id="F3651A28FFB800170A24FE5EFB6BFE5C" box="[1148,1263,459,481]" gridcol="4" gridrow="5" pageId="13" pageNumber="522">0%</td>
<td id="F3651A28FFB800170B76FE5EFA19FE5C" box="[1326,1437,459,481]" gridcol="5" gridrow="5" pageId="13" pageNumber="522">0 (0)</td>
</tr>
<tr id="B0B47354FFB800170EFBFE7CFA19FE42" box="[163,1437,489,511]" gridrow="6" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FE7CFDE2FE42" box="[489,614,489,511]" gridcol="1" gridrow="6" pageId="13" pageNumber="522">Oct/2019</td>
<td id="F3651A28FFB800170C8CFE7CFCB8FE42" box="[724,828,489,511]" gridcol="2" gridrow="6" pageId="13" pageNumber="522">34</td>
<td id="F3651A28FFB800170DF0FE7CFB94FE42" box="[936,1040,489,511]" gridcol="3" gridrow="6" pageId="13" pageNumber="522">0</td>
<td id="F3651A28FFB800170A24FE7CFB6BFE42" box="[1148,1263,489,511]" gridcol="4" gridrow="6" pageId="13" pageNumber="522">0%</td>
<td id="F3651A28FFB800170B76FE7CFA19FE42" box="[1326,1437,489,511]" gridcol="5" gridrow="6" pageId="13" pageNumber="522">0 (0)</td>
</tr>
<tr id="B0B47354FFB800170EFBFD92FA19FDA0" box="[163,1437,519,541]" gridrow="7" pageId="13" pageNumber="522">
<td id="F3651A28FFB800170FB1FD92FDE2FDA0" box="[489,614,519,541]" gridcol="1" gridrow="7" pageId="13" pageNumber="522">Jan/2020</td>
<td id="F3651A28FFB800170C8CFD92FCB8FDA0" box="[724,828,519,541]" gridcol="2" gridrow="7" pageId="13" pageNumber="522">20</td>
<td id="F3651A28FFB800170DF0FD92FB94FDA0" box="[936,1040,519,541]" gridcol="3" gridrow="7" pageId="13" pageNumber="522">5</td>
<td id="F3651A28FFB800170A24FD92FB6BFDA0" box="[1148,1263,519,541]" gridcol="4" gridrow="7" pageId="13" pageNumber="522">25%</td>
<td id="F3651A28FFB800170B76FD92FA19FDA0" box="[1326,1437,519,541]" gridcol="5" gridrow="7" pageId="13" pageNumber="522">1–2 (1.4)</td>
</tr>
</table>
</paragraph>
<paragraph id="0E3B7116FFB8FFE50EE3FDF0FE68FCEC" blockId="13.[163,779,613,849]" pageId="13" pageNumber="522">
Eggs with hyaline, spherical outer envelope, 41–61 (50) in diameter (measured in eggs liberated from uterus in distilled water; 
<emphasis id="3CF0AD04FFB8FFE50FE6FD36FE56FD05" box="[446,466,675,696]" italics="true" pageId="13" pageNumber="522">N</emphasis>
= 15); oncosphere spherical, 13–19 (16) in diameter, with three pairs of embryonic hooks, eight to 8–11 (9) long (
<emphasis id="3CF0AD04FFB8FFE50C59FD75FD91FD48" box="[513,533,736,757]" italics="true" pageId="13" pageNumber="522">N</emphasis>
= 30), diameter with embryophore 20–30 (26); embryophore 2–5 (3) thick, consisting of two layers, outer layer thinner than nucleated inner envelope (
<figureCitation id="96BF6D93FFB8FFE50FD5FCAEFE59FCEC" box="[397,477,827,849]" captionStart="Figure 5" captionStartId="11.[163,245,1700,1722]" captionTargetBox="[232,1377,199,1669]" captionTargetId="figure-24@11.[227,1378,196,1671]" captionTargetPageId="11" captionText="Figure 5. Line drawings of Gangesia mukutmanipurensis sp. nov. from Ompok bimaculatus (specimen accession number in parentheses). A, scolex, subapical view (ZSI/W11086/1). B, C, scolex, dorsoventral view (ZSI/W11085/1 and ZSI/W11082/1/1). D, frontal section of the scolex (ZSI/W11087/1). E, mature proglottid, ventral view (ZSI/W11084/1). F, terminal genitalia (ZSI/W11083/1/2). G, egg drawn in distilled water. Abbreviations: cc, chromophil cell; cs, cirrus-sac; doc, dorsal osmoregulatory canal; eh, embryonic hook; em, embryophore; ga, genital atrium; mi, microtriches; oc, osmoregulatory canal; oe, outer envelope; on, oncosphere; ov, ovary; re, retractor muscles; ro, rostellum-like organ; su, sucker; te, testes; up, uterine pore; va, vagina; vf, vitelline follicles; voc, ventral osmoregulatory canal; vs, vaginal sphincter." figureDoi="http://doi.org/10.5281/zenodo.7981610" httpUri="https://zenodo.org/record/7981610/files/figure.png" pageId="13" pageNumber="522">Fig. 5G</figureCitation>
).
</paragraph>
<paragraph id="0E3B7116FFB8FFE50EFBFC17FE26FC24" blockId="13.[163,418,898,921]" box="[163,418,898,921]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFC17FE26FC24" box="[163,418,898,921]" italics="true" pageId="13" pageNumber="522">Taxonomic summary</emphasis>
</paragraph>
<paragraph id="0E3B7116FFB8FFE50EFBFC53FE3AFC44" blockId="13.[163,777,965,1017]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFC53FDCDFC66" box="[163,585,965,987]" italics="true" pageId="13" pageNumber="522">Ty p e h o s t: O m p o k b i m a c u l a t u s</emphasis>
(B l o c h, 1 7 9 4) (
<taxonomicName id="C9840A95FFB8FFE50EF2FC76FEBFFC44" box="[170,315,995,1017]" class="Actinopterygii" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="522" phylum="Chordata" rank="order">Siluriformes</taxonomicName>
: 
<taxonomicName id="C9840A95FFB8FFE50F1EFC76FE37FC44" authorityName="Cuvier" authorityYear="1816" box="[326,435,995,1017]" class="Actinopterygii" family="Siluridae" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="522" phylum="Chordata" rank="family">Siluridae</taxonomicName>
).
</paragraph>
<paragraph id="0E3B7116FFB8FFE50EFBFBB6FDFBFB8A" blockId="13.[163,639,1058,1080]" box="[163,639,1058,1080]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFBB6FEDEFB8A" box="[163,346,1058,1080]" italics="true" pageId="13" pageNumber="522">Site of infection:</emphasis>
Intestine (anterior part).
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB8FFE50EFBFBC9FD25FB2D" pageId="13" pageNumber="522" type="materials_examined">
<paragraph id="0E3B7116FFB8FFE50EFBFBC9FD25FB2D" blockId="13.[163,777,1115,1168]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFBC9FEC7FBCD" box="[163,323,1115,1137]" italics="true" pageId="13" pageNumber="522">Type locality:</emphasis>
<materialsCitation id="BEEC7B4BFFB8FFE50F17FBCEFD25FB2D" ID-GBIF-Occurrence="4118152305" country="India" latitude="22.963335" location="Mukutmanipur Dam Lake" longLatPrecision="21" longitude="86.78833" pageId="13" pageNumber="522" specimenCount="1" stateProvince="West Bengal">
<location id="0B5B27CDFFB8FFE50F17FBCEFD0AFBCC" LSID="urn:lsid:plazi:treatment:862DC000FFB2FFE50D2EFEE6FB30FB5D:0B5B27CDFFB8FFE50F17FBCEFD0AFBCC" box="[335,654,1115,1137]" country="India" latitude="22.963335" longLatPrecision="21" longitude="86.78833" name="Mukutmanipur Dam Lake" pageId="13" pageNumber="522" stateProvince="West Bengal">Mukutmanipur Dam Lake</location>
, 
<location id="0B5B27CDFFB8FFE50CC3FBCEFC81FBCC" LSID="urn:lsid:plazi:treatment:862DC000FFB2FFE50D2EFEE6FB30FB5D:0B5B27CDFFB8FFE50CC3FBCEFC81FBCC" box="[667,773,1115,1137]" country="India" latitude="22.963335" longLatPrecision="21" longitude="86.78833" name="Bankura" pageId="13" pageNumber="522" stateProvince="West Bengal">Bankura</location>
, 
<collectingRegion id="CC40BFF4FFB8FFE50EFBFBEEFEB0FB2D" box="[163,308,1146,1168]" country="India" name="West Bengal" pageId="13" pageNumber="522">West Bengal</collectingRegion>
, 
<collectingCountry id="76933186FFB8FFE50F66FBEFFEFFFB2D" box="[318,379,1146,1168]" name="India" pageId="13" pageNumber="522">India</collectingCountry>
(
<geoCoordinate id="6BB017D1FFB8FFE50FD2FBEFFD8DFB2D" box="[394,521,1146,1168]" degrees="22" direction="north" minutes="57" orientation="latitude" pageId="13" pageNumber="522" precision="15" seconds="48" value="22.963335">22°57’48&quot;N</geoCoordinate>
, 
<geoCoordinate id="6BB017D1FFB8FFE50C4CFBEFFD10FB2D" box="[532,660,1146,1168]" degrees="86" direction="east" minutes="47" orientation="longitude" pageId="13" pageNumber="522" precision="15" seconds="18" value="86.78833">86°47’18&quot;E</geoCoordinate>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB8FFE50EFBFB2CFD6DFAB6" pageId="13" pageNumber="522" type="description">
<paragraph id="0E3B7116FFB8FFE50EFBFB2CFD6DFAB6" blockId="13.[163,779,1209,1292]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFB2CFDF2FB73" box="[163,630,1209,1230]" italics="true" pageId="13" pageNumber="522">PreƲalence and intensity of infection:</emphasis>
Prevalence 9.5% (20 of 211 
<taxonomicName id="C9840A95FFB8FFE50F01FB42FDC4FB51" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[345,576,1239,1260]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="522" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="3CF0AD04FFB8FFE50F01FB42FDC4FB51" box="[345,576,1239,1260]" italics="true" pageId="13" pageNumber="522">Ompok bimaculatus</emphasis>
</taxonomicName>
examined); mean intensity 1.8 (one to four worms/host) (see 
<tableCitation id="430644ADFFB8FFE50CDEFB63FD5DFAB6" box="[646,729,1270,1292]" captionStart="Table 2" captionStartId="13.[163,227,196,217]" captionTargetBox="[163,1437,254,541]" captionTargetPageId="13" captionText="Table 2. Prevalence and intensity of infection in Bankura, West Bengal, India" httpUri="http://table.plazi.org/id/5AFB219EFFB8FFE50EFBFF51FC53FF67" pageId="13" pageNumber="522" tableUuid="5AFB219EFFB8FFE50EFBFF51FC53FF67">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB8FFE50EFBFAA0FD9DF925" pageId="13" pageNumber="522" type="materials_examined">
<paragraph id="0E3B7116FFB8FFE50EFBFAA0FD22FA78" blockId="13.[163,779,1333,1477]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFAA0FE26FAF7" box="[163,418,1333,1354]" italics="true" pageId="13" pageNumber="522">Specimens deposited:</emphasis>
<materialsCitation id="BEEC7B4BFFB8FFE50FE8FAA0FDFEFAD4" ID-GBIF-Occurrence="4118152303" collectionCode="ZSI, HWML, IPCAS" pageId="13" pageNumber="522" specimenCode="ZSI/W11082/1/1-ZSI/W11083/1/2, HWML-216818, IPCAS C-913/1, MHNG-PLAT-0067053, ZSI/W11084/1, ZSI/W11085/1, ZSI/W11086/1, ZSI/W11087/1, ZSI/W11088/1, ZSI/W11089/1." specimenCount="2" typeStatus="holotype">
<typeStatus id="D13FCFB4FFB8FFE50FE8FAA0FD98FAF6" box="[432,540,1333,1355]" pageId="13" pageNumber="522" type="holotype">Holotype</typeStatus>
and hologenophore 
<emphasis id="3CF0AD04FFB8FFE50EFBFAC1FDF6FAD4" box="[163,626,1364,1385]" italics="true" pageId="13" pageNumber="522">
<specimenCode id="5E22D96DFFB8FFE50EFBFAC1FDF6FAD4" box="[163,626,1364,1385]" pageId="13" pageNumber="522">ZSI/W11082/1/1–ZSI/W11083/1/2</specimenCode>
</emphasis>
.
</materialsCitation>
<materialsCitation id="BEEC7B4BFFB8FFE50CDDFAC1FD22FA78" ID-GBIF-Occurrence="4118152301" collectionCode="ZSI, HWML, IPCAS" pageId="13" pageNumber="522" specimenCode="ZSI/W11082/1/1-ZSI/W11083/1/2, HWML-216818, IPCAS C-913/1, MHNG-PLAT-0067053, ZSI/W11084/1, ZSI/W11085/1, ZSI/W11086/1, ZSI/W11087/1, ZSI/W11088/1, ZSI/W11089/1." specimenCount="2" typeStatus="paratype">
<typeStatus id="D13FCFB4FFB8FFE50CDDFAC1FC81FAD4" box="[645,773,1364,1385]" pageId="13" pageNumber="522" type="paratype">Paratypes</typeStatus>
: 
<specimenCode id="5E22D96DFFB8FFE50EFBFAE7FEECFA35" box="[163,360,1394,1416]" pageId="13" pageNumber="522">HWML-216818</specimenCode>
; 
<specimenCode id="5E22D96DFFB8FFE50F22FAE7FDC7FA3A" box="[378,579,1394,1415]" pageId="13" pageNumber="522">IPCAS C-913/1</specimenCode>
; 
<specimenCode id="5E22D96DFFB8FFE50C0EFAE7FE81FA1A" pageId="13" pageNumber="522">MHNG-PLAT-0067053</specimenCode>
; 
<specimenCode id="5E22D96DFFB8FFE50F55FA04FE34FA1B" box="[269,432,1425,1446]" pageId="13" pageNumber="522">ZSI/W11084/1</specimenCode>
, 
<specimenCode id="5E22D96DFFB8FFE50FE1FA04FDDFFA1A" box="[441,603,1425,1447]" pageId="13" pageNumber="522">ZSI/W11085/1</specimenCode>
, 
<specimenCode id="5E22D96DFFB8FFE50C3CFA04FC82FA1A" box="[612,774,1425,1447]" pageId="13" pageNumber="522">ZSI/W11086/1</specimenCode>
, 
<specimenCode id="5E22D96DFFB8FFE50EFBFA25FEC1FA78" box="[163,325,1456,1477]" pageId="13" pageNumber="522">ZSI/W11087/1</specimenCode>
, 
<specimenCode id="5E22D96DFFB8FFE50F08FA25FE77FA78" box="[336,499,1456,1477]" pageId="13" pageNumber="522">ZSI/W11088/1</specimenCode>
, 
<specimenCode id="5E22D96DFFB8FFE50FA6FA25FD22FA78" box="[510,678,1456,1477]" pageId="13" pageNumber="522">ZSI/W11089/1.</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="0E3B7116FFB8FFE50EFBFA7CFD9DF925" blockId="13.[163,779,1513,1688]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBFA7CFED7FA43" box="[163,339,1513,1534]" italics="true" pageId="13" pageNumber="522">Genetic data:</emphasis>
<materialsCitation id="BEEC7B4BFFB8FFE50F3AFA7CFD9DF925" ID-GBIF-Occurrence="4418409301" pageId="13" pageNumber="522" specimenCount="1">
<emphasis id="3CF0AD04FFB8FFE50F3AFA7CFE00FA43" box="[354,388,1513,1534]" italics="true" pageId="13" pageNumber="522">lsr</emphasis>
DNA gene sequence (partial) (GenBank accession numbers) 
<accessionNumber id="11D7ECF5FFB8FFE50C5CF992FDFBF9A0" box="[516,639,1543,1565]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/ON181446" pageId="13" pageNumber="522" type="GenBank">ON181446</accessionNumber>
, 
<accessionNumber id="11D7ECF5FFB8FFE50CD3F992FC82F9A0" box="[651,774,1543,1565]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/ON181456" pageId="13" pageNumber="522" type="GenBank">ON181456</accessionNumber>
, 
<accessionNumber id="11D7ECF5FFB8FFE50EFBF9B3FE98F981" box="[163,284,1574,1596]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/ON181457" pageId="13" pageNumber="522" type="GenBank">ON181457</accessionNumber>
and 
<emphasis id="3CF0AD04FFB8FFE50F0CF9B3FE07F986" box="[340,387,1574,1595]" italics="true" pageId="13" pageNumber="522">COI</emphasis>
gene sequence (partial) (GenBank accession numbers) 
<accessionNumber id="11D7ECF5FFB8FFE50FFDF9D0FDADF9E6" box="[421,553,1605,1627]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/ON176195" pageId="13" pageNumber="522" type="GenBank">ON176195</accessionNumber>
. Hologenophores: MHNG-PLAT- 0067053; ZSI / W 11087 /1, 
<emphasis id="3CF0AD04FFB8FFE50C90F9F1FD96F92A" italics="true" pageId="13" pageNumber="522">ZSI / W11082/1/1–ZSI/W11083/1/2</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB8FFE50EFBF92EFE83F8B2" pageId="13" pageNumber="522" type="etymology">
<paragraph id="0E3B7116FFB8FFE50EFBF92EFE83F8B2" blockId="13.[163,778,1723,1807]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBF92EFEACF96D" box="[163,296,1723,1744]" italics="true" pageId="13" pageNumber="522">Etymology:</emphasis>
The species name is derived from the Mukutmanipur Dam Lake, the type locality of this parasite.
</paragraph>
</subSubSection>
<subSubSection id="469E229DFFB8FFE50EFBF8A2FB30FB5D" pageId="13" pageNumber="522" type="discussion">
<paragraph id="0E3B7116FFB8FFE50EFBF8A2FB30FB5D" blockId="13.[163,779,1847,1900]" lastBlockId="13.[827,1443,613,1248]" pageId="13" pageNumber="522">
<emphasis id="3CF0AD04FFB8FFE50EFBF8A2FE95F8F1" box="[163,273,1847,1868]" italics="true" pageId="13" pageNumber="522">Remarks:</emphasis>
The new species is a member of 
<taxonomicName id="C9840A95FFB8FFE50CC6F8ADFC8FF8F0" authorityName="Woodland" authorityYear="1924" box="[670,779,1848,1869]" class="Cestoda" family="Proteocephalidae" genus="Gangesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Onchoproteocephalidea" pageId="13" pageNumber="522" phylum="Platyhelminthes" rank="genus">
<emphasis id="3CF0AD04FFB8FFE50CC6F8ADFC8FF8F0" box="[670,779,1848,1869]" italics="true" pageId="13" pageNumber="522">Gangesia</emphasis>
</taxonomicName>
because it possesses characteristics that include: (1) presence of prominent retractor muscles that form a wide band; (2) rostellum-like organ longer than, or as long, as the size of suckers; (3) ventral osmoregulatory canals median (internal) to vitelline follicles; (4) testes in single field; (5) vagina posterior or anterior to the cirrus-sac; (6) lateral band of vitelline follicles occupying almost the total length of the proglottid; and (7) type 1 uterine development (
<emphasis id="3CF0AD04FFB8FFE50A6DFCA9FBF0FCEC" box="[1077,1140,828,849]" italics="true" pageId="13" pageNumber="522">sensu</emphasis>
<bibRefCitation id="6A150CE7FFB8FFE50A23FCAEFA1FFCEC" author="de Chambrier" box="[1147,1435,827,849]" etAl="et al." firstAuthor="de Chambrier" pageId="13" pageNumber="522" pagination="159 - 171" refId="ref13406" refString="de Chambrier A, Zehnder MP, Vaucher C, Mariaux J. 2004. The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology 57: 159 - 171." type="journal article" year="2004">
de Chambrier 
<emphasis id="3CF0AD04FFB8FFE50B47FCA9FAD3FCED" box="[1311,1367,827,849]" italics="true" pageId="13" pageNumber="522">et al.</emphasis>
, 2004
</bibRefCitation>
) (see ‘Generic composition and interrelationship in the Gangesiinae’ in the Discussion for details). The new species differs from all eight congeners by the absence of hooks on the rostellum-like organ and hooklets on the anterior rim of suckers, previously considered to be the most typical feature of the genus. Instead of hooks and hooklets, several rows of coniform spinitriches are present on the rostellum-like organ and the anterior rim of the suckers. Along with this unique scolex morphology, the new species is also typified by a narrow rostellum-like organ (less than 90 µm wide) and indistinguishable dorsal osmoregulatory canals in pregravid and gravid proglottids.
</paragraph>
</subSubSection>
</treatment>
</document>