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<mods:title id="09CE5CC633210ADAB42B255EF4E87F3B">The true identity of Rhopalurus pintoi Mello-Leitão, 1932, with notes on the status and distribution of Rhopalurus crassicauda Caporiacco, 1947 (Scorpiones: Buthidae)</mods:title>
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<subSubSection id="AF826572FFE7FFCCFEB29F6DFD0DF990" box="[268,725,1576,1602]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="E72736F9FFE7FFCCFEB29F6DFD0DF990" blockId="3.[268,725,1576,1636]" box="[268,725,1576,1602]" pageId="3" pageNumber="2">
<heading id="BC6F8195FFE7FFCCFEB29F6DFD0DF990" box="[268,725,1576,1602]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2" reason="4">
<taxonomicName id="20984D7AFFE7FFCCFEB29F6DFD0DF990" authority="Mello-Leitao, 1932" authorityName="Mello-Leitao" authorityYear="1932" box="[268,725,1576,1602]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE7FFCCFEB29F6DFE31F990" bold="true" box="[268,489,1576,1602]" italics="true" pageId="3" pageNumber="2">Rhopalurus pintoi</emphasis>
Mello-Leitão, 1932
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<subSubSection id="AF826572FFE7FFCCFEF89F0FFD44F9B6" box="[326,668,1610,1636]" pageId="3" pageNumber="2" type="description">
<paragraph id="E72736F9FFE7FFCCFEF89F0FFD44F9B6" blockId="3.[268,725,1576,1636]" box="[326,668,1610,1636]" pageId="3" pageNumber="2">
<figureCitation id="7FA32A7CFFE7FFCCFEF89F0FFE0BF9B6" box="[326,467,1610,1636]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="4.[199,275,1665,1688]" captionStartId-1="5.[192,268,1886,1909]" captionStartId-2="7.[192,268,1334,1357]" captionStartId-3="8.[192,268,1926,1949]" captionTargetBox-0="[191,1439,191,1633]" captionTargetBox-1="[228,1392,168,1856]" captionTargetBox-2="[192,1452,192,1304]" captionTargetBox-3="[191,1408,167,1897]" captionTargetId-1="figure-8@5.[228,1392,168,1856]" captionTargetId-2="figure-7@7.[192,1452,192,1304]" captionTargetId-3="figure-8@8.[192,1407,168,1896]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." captionText-1="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." captionText-2="Figure 3: Adult female Rhopalurus pintoi from Rupununi: a) pedipalp; b) closeup of pedipalp movable finger; c) chelicerae and carapace; d) sternopectinal region; e) same specimen, still alive in captivity." captionText-3="Figure 4: Juvenile female Rhopalurus pintoi from Rupununi, still alive in captivity." figureDoi-0="http://doi.org/10.5281/zenodo.4673305" figureDoi-1="http://doi.org/10.5281/zenodo.4673307" figureDoi-2="http://doi.org/10.5281/zenodo.4673311" figureDoi-3="http://doi.org/10.5281/zenodo.4673315" httpUri-0="https://zenodo.org/record/4673305/files/figure.png" httpUri-1="https://zenodo.org/record/4673307/files/figure.png" httpUri-2="https://zenodo.org/record/4673311/files/figure.png" httpUri-3="https://zenodo.org/record/4673315/files/figure.png" pageId="3" pageNumber="2">Figures 14</figureCitation>
,
<figureCitation id="7FA32A7CFFE7FFCCFE5E9F0FFDD6F9B6" box="[480,526,1610,1636]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="12.[192,268,1158,1181]" captionStartId-1="13.[192,268,1926,1949]" captionStartId-2="14.[192,268,1149,1172]" captionTargetBox-0="[191,1440,191,1128]" captionTargetBox-1="[216,1417,178,1897]" captionTargetBox-2="[191,1429,191,1120]" captionTargetId-0="figure-8@12.[192,1439,192,1127]" captionTargetId-1="figure-7@13.[217,1416,179,1896]" captionTargetId-2="figure-9@14.[192,1428,192,1119]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="Figure 7: Geographical distribution of Rhopalurus pintoi (15) and Rhopalurus crassicauda (57): Tepequén [1], Surumu [2], Rio Surumu [3], Rio Tacutú [4], Rupununi [5], 12 km north of Boa Vista [6], Rio Branco [7]." captionText-1="Figure 8: Two views of the habitat where both Rhopalurus pintoi and Rhopalurus crassicauda occur at Rupununi (copyright by Hans-Werner Auer)." captionText-2="Figure 9: View of the microhabitat where both Rhopalurus pintoi and Rhopalurus crassicauda occur at Rupununi (copyright by Hans-Werner Auer)." figureDoi-0="http://doi.org/10.5281/zenodo.4673325" figureDoi-1="http://doi.org/10.5281/zenodo.4673329" figureDoi-2="http://doi.org/10.5281/zenodo.4673331" httpUri-0="https://zenodo.org/record/4673325/files/figure.png" httpUri-1="https://zenodo.org/record/4673329/files/figure.png" httpUri-2="https://zenodo.org/record/4673331/files/figure.png" pageId="3" pageNumber="2">79</figureCitation>
,
<tableCitation id="AA1A0342FFE7FFCCFDA49F0FFD44F9B6" box="[538,668,1610,1636]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="3.[192,257,1110,1133]" captionStartId-1="6.[192,257,434,457]" captionText-0="Table 1: Measurements of four adult Rhopalurus pintoi from Rupununi. Abbreviations: length (L), width (W), posterior width (Wp), depth (H)." captionText-1="Table 2: Variation of pectinal tooth count in Rhopalurus pintoi, including data from MelloLeitão (1932, holotype) and Lourenço &amp; PintodaRocha (1997, as R. piceus). Abbreviations: number of pectines (N), standard deviation (SD)." pageId="3" pageNumber="2">Tables 12</tableCitation>
</paragraph>
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<taxonomicName id="20984D7AFFE7FFCCFF7E9FCCFD7CF972" authority="Mello-Leitao, 1932: 11 - 12" authorityName="Mello-Leitao" authorityPageNumber="11 - 12" authorityYear="1932" box="[192,676,1673,1697]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE7FFCCFF7E9FCCFE58F972" box="[192,384,1673,1697]" italics="true" pageId="3" pageNumber="2">Rhopalurus pintoi</emphasis>
Mello-Leitão, 1932: 1112
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, 1415, 31, 38, 46, figs. 2ac;
<bibRefCitation id="83094B08FFE7FFCCFE0C9FEDFD82F96D" author="PRADO" box="[434,602,1704,1727]" pageId="3" pageNumber="2" refId="ref6005" refString="PRADO, A. 1939. Contribucao ao conhecimento dos escorpioes sul-americanos. Sinopse das especies de" type="book" year="1939">Prado, 1939: 27</bibRefCitation>
, 36; Mello-Leitão, 1945: 266, 284285, fig. 115 (in part);
<bibRefCitation id="83094B08FFE7FFCCFD0B9F83FE4CF92E" author="LOURENCO" pageId="3" pageNumber="2" pagination="107 - 141" refId="ref5851" refString="LOURENCO, W. R. 1982. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones: Buthidae). Revue Arachnologique, 4: 107 - 141." type="journal article" year="1982">Lourenço, 1982: 107108</bibRefCitation>
, 115, 117, fig. 78 (in part);
<bibRefCitation id="83094B08FFE7FFCCFD689FA0FEAFF8C9" author="TERUEL" pageId="3" pageNumber="2" pagination="43 - 54" refId="ref6092" refString="TERUEL, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa, 38: 43 - 54." type="journal article" year="2006">Teruel, 2006: 5152</bibRefCitation>
.
</paragraph>
<paragraph id="E72736F9FFE7FFCCFF7E9E67FBCAFAD7" blockId="3.[192,802,1673,1911]" lastBlockId="3.[830,1441,1232,1469]" pageId="3" pageNumber="2">
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<treatmentCitation id="663910E8FFE7FFCCFF7E9E67FCC3F8EB" author="LOURENCO" box="[192,795,1826,1850]" page="108" pageId="3" pageNumber="2" year="1982">
<taxonomicName id="20984D7AFFE7FFCCFF7E9E67FCC3F8EB" authority=": Lourenco, 1982: 107 - 108" authorityName=": Lourenco" authorityPageNumber="107 - 108" authorityYear="1982" box="[192,795,1826,1850]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="laticauda" subSpecies="pintoi">
<emphasis id="D5ECEAEBFFE7FFCCFF7E9E67FE29F8E8" box="[192,497,1826,1850]" italics="true" pageId="3" pageNumber="2">Rhopalurus laticauda pintoi</emphasis>
:
<bibRefCitation id="83094B08FFE7FFCCFDBC9E67FCC3F8EB" author="LOURENCO" box="[514,795,1826,1849]" pageId="3" pageNumber="2" pagination="107 - 141" refId="ref5851" refString="LOURENCO, W. R. 1982. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones: Buthidae). Revue Arachnologique, 4: 107 - 141." type="journal article" year="1982">Lourenço, 1982: 107108</bibRefCitation>
</taxonomicName>
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, 115, 117119, 121, 134138, figs. 3946, 78; table I (in part, references to the
<typeStatus id="3823885BFFE7FFCCFD9B9E25FD5AF8A5" box="[549,642,1888,1911]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
only); Fet &amp; Lowe, 2000: 220221 (in part, references to the
<typeStatus id="3823885BFFE7FFCCFCD09DABFC13FAD7" box="[878,971,1262,1285]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
only).
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</paragraph>
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<taxonomicName id="20984D7AFFE7FFCCFC809C49FBA5FAF6" authority="Lourenco" authorityName="Lourenco" box="[830,1149,1292,1316]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="D5ECEAEBFFE7FFCCFC809C49FBD3FAF6" box="[830,1035,1292,1316]" italics="true" pageId="3" pageNumber="2">Rhopalurus piceus</emphasis>
Lourenço
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&amp; Pinto-da-Rocha, 1997: 181, 183185, 187191, figs. 4, 6, 8, 10, 1213, 1521, table I; Fet &amp; Lowe, 2000: 221;
<bibRefCitation id="83094B08FFE7FFCCFA8D9C0FFBCCFA52" author="LOURENCO" pageId="3" pageNumber="2" refId="ref5881" refString="LOURENCO, W. R. 2002. Scorpions of Brazil. Les Edit. de l'If, Paris, 307 pp." type="book" year="2002">Lourenço, 2002: 104105</bibRefCitation>
, 110111, 304305, figs. 232238; Lenarducci et al., 2005: 7, table II;
<bibRefCitation id="83094B08FFE7FFCCFB569CCDFA42FA4D" author="TERUEL" box="[1256,1434,1416,1439]" pageId="3" pageNumber="2" pagination="43 - 54" refId="ref6092" refString="TERUEL, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa, 38: 43 - 54." type="journal article" year="2006">Teruel, 2006: 52</bibRefCitation>
.
<emphasis id="D5ECEAEBFFE7FFCCFCD19CE3FBD6FA6F" bold="true" box="[879,1038,1446,1469]" pageId="3" pageNumber="2">New synonym</emphasis>
.
</paragraph>
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<paragraph id="E72736F9FFE7FFCCFC809CAFFB40F948" blockId="3.[830,1441,1513,1690]" pageId="3" pageNumber="2">
<emphasis id="D5ECEAEBFFE7FFCCFC809CAFFC68F9D3" bold="true" box="[830,944,1514,1537]" italics="true" pageId="3" pageNumber="2">Type data</emphasis>
:
<taxonomicName id="20984D7AFFE7FFCCFC789CACFB56F9D3" authorityName="Mello-Leitao" authorityYear="1932" box="[966,1166,1513,1537]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE7FFCCFC789CACFB56F9D3" box="[966,1166,1513,1537]" italics="true" pageId="3" pageNumber="2">Rhopalurus pintoi</emphasis>
</taxonomicName>
:
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<specimenCount id="F19EFD70FFE7FFCCFB1D9CAFFB6FF9D3" box="[1187,1207,1514,1537]" pageId="3" pageNumber="2" type="female"></specimenCount>
<typeStatus id="3823885BFFE7FFCCFB7A9CAFFAF9F9D3" box="[1220,1313,1514,1537]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
(
<collectionCode id="8189AE3CFFE7FFCCFA889CAFFABEF9D3" box="[1334,1382,1514,1537]" country="Brazil" httpUri="http://grbio.org/cool/5rz8-qnyy" name="Colecao de Culturas de Fungos do Instituto Oswaldo Cruz" pageId="3" pageNumber="2">IOC</collectionCode>
, lost
<emphasis id="D5ECEAEBFFE7FFCCFC809F4CFCA1F9F2" box="[830,889,1545,1568]" italics="true" pageId="3" pageNumber="2">sensu</emphasis>
<bibRefCitation id="83094B08FFE7FFCCFC3B9F4DFBE9F9CD" author="LOURENCO" box="[901,1073,1544,1567]" pageId="3" pageNumber="2" pagination="107 - 141" refId="ref5851" refString="LOURENCO, W. R. 1982. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones: Buthidae). Revue Arachnologique, 4: 107 - 141." type="journal article" year="1982">Lourenço, 1982</bibRefCitation>
; see Remarks):
<collectingCountry id="9F8F7669FFE7FFCCFB559F4DFAF7F9CD" box="[1259,1327,1544,1567]" name="Brazil" pageId="3" pageNumber="2">Brazil</collectingCountry>
,
<collectingRegion id="255CF81BFFE7FFCCFA809F4DFA44F9CD" box="[1342,1436,1544,1567]" country="Brazil" name="Roraima" pageId="3" pageNumber="2">Roraima</collectingRegion>
,
<collectingCounty id="0E464E75FFE7FFCCFC809F62FC64F9EC" box="[830,956,1575,1598]" pageId="3" pageNumber="2">Rio Tacutú</collectingCounty>
,
<collectingMunicipality id="0743AC83FFE7FFCCFC749F62FBF4F9EC" box="[970,1068,1575,1598]" pageId="3" pageNumber="2">Brazilian</collectingMunicipality>
border with
<collectingCountry id="9F8F7669FFE7FFCCFB7E9F62FACBF9EC" box="[1216,1299,1575,1598]" name="Guyana" pageId="3" pageNumber="2">Guyana</collectingCountry>
.
<taxonomicName id="20984D7AFFE7FFCCFA9A9F63FC5AF98F" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="D5ECEAEBFFE7FFCCFA9A9F63FC5AF98F" italics="true" pageId="3" pageNumber="2">Rhopalurus piceus</emphasis>
</taxonomicName>
:
<location id="E2476022FFE7FFCCFC259F03FC2CF98F" LSID="urn:lsid:plazi:treatment:6F3187EFFFE7FFC8FEB29F6DFB6BF926:E2476022FFE7FFCCFC259F03FC2CF98F" box="[923,1012,1606,1629]" country="Brazil" county="Rio Tacutu" municipality="Brazilian" name="Juvenile" pageId="3" pageNumber="2" stateProvince="Roraima">Juvenile</location>
</materialsCitation>
<materialsCitation id="57F03CA4FFE7FFCCFBBB9F03FB4CF948" collectingDate="1993-06-15" collectingDateMax="1993-06-22" collectingDateMin="1993-06-15" collectorName="M. Vanzolini" country="Brazil" location="Tepequen" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Roraima" typeStatus="holotype">
<specimenCount id="F19EFD70FFE7FFCCFBBB9F03FBC1F98F" box="[1029,1049,1606,1629]" pageId="3" pageNumber="2" type="female"></specimenCount>
<typeStatus id="3823885BFFE7FFCCFB949F03FB5FF98F" box="[1066,1159,1606,1629]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
(MZSP-15173, not examined; see Remarks):
<collectingCountry id="9F8F7669FFE7FFCCFBE19F21FB7BF9A9" box="[1119,1187,1636,1659]" name="Brazil" pageId="3" pageNumber="2">Brazil</collectingCountry>
,
<collectingRegion id="255CF81BFFE7FFCCFB049F21FAC1F9A9" box="[1210,1305,1636,1659]" country="Brazil" name="Roraima" pageId="3" pageNumber="2">Roraima</collectingRegion>
,
<location id="E2476022FFE7FFCCFA8E9F21FA44F9A9" LSID="urn:lsid:plazi:treatment:6F3187EFFFE7FFC8FEB29F6DFB6BF926:E2476022FFE7FFCCFA8E9F21FA44F9A9" box="[1328,1436,1636,1659]" country="Brazil" name="Tepequen" pageId="3" pageNumber="2" stateProvince="Roraima">Tepequén</location>
,
<date id="93261039FFE7FFCCFC809FC6FC2FF948" box="[830,1015,1667,1690]" pageId="3" pageNumber="2" value="1993-06-15" valueMax="1993-06-22" valueMin="1993-06-15">
<collectingDate id="8362E9D1FFE7FFCCFC809FC6FC2FF948" box="[830,1015,1667,1690]" pageId="3" pageNumber="2" value="1993-06-15" valueMax="1993-06-22" valueMin="1993-06-15">1522 June 1993</collectingDate>
</date>
,
<collectorName id="4A6D532FFFE7FFCCFBBC9FC6FB4CF948" box="[1026,1172,1667,1690]" pageId="3" pageNumber="2">M. Vanzolini</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AF826572FFE7FFC9FC809F83FAAFFE59" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="E72736F9FFE7FFCBFC809F83FC41F8BC" blockId="3.[829,1441,1734,1911]" lastBlockId="4.[830,1441,1725,1902]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<emphasis id="D5ECEAEBFFE7FFCCFC809F83FC74F90F" bold="true" box="[830,940,1734,1757]" italics="true" pageId="3" pageNumber="2">Diagnosis</emphasis>
: species of large size (males
<quantity id="20609B1CFFE7FFCCFAA39F83FA44F90F" box="[1309,1436,1734,1757]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="8.8" metricValueMin="7.7" pageId="3" pageNumber="2" unit="mm" value="82.5" valueMax="88.0" valueMin="77.0">7788 mm</quantity>
, females
<quantity id="20609B1CFFE7FFCCFC239FA0FBCCF92E" box="[925,1044,1765,1788]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="9.1" metricValueMin="8.4" pageId="3" pageNumber="2" unit="mm" value="87.5" valueMax="91.0" valueMin="84.0">8491 mm</quantity>
) for the genus. Coloration entirely blackish, except only for white pectines and yellowish tips of legs and pedipalp fingers. Pedipalps densely hirsute and very slender in both sexes, with hands slightly more incrassate in males; fingers without basal lobe/ notch combination, but with weak scallop in adult males; fingers with 910 principal rows of granules, flanked by a many supernumerary granules. Carapace and tergites very strongly granulose. Sternite III and pectines with stridulatory apparatus well developed; sternite V with a vestigial smooth patch on males. Metasoma distally incrassate on both sexes, much more conspicuously in males; telson vesicle large, subaculear tubercle vestigial and close to the base of aculeus. Pectines with 2730 (mode 28) teeth in males, and 2125 (mode 24) in females; basal plate with a very large and deep pit in females.
</paragraph>
<caption id="B3E76671FFE0FFCBFF799FC4FB7BF94A" ID-DOI="http://doi.org/10.5281/zenodo.4673305" ID-Zenodo-Dep="4673305" box="[199,1187,1665,1688]" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="4" pageNumber="3" startId="4.[199,275,1665,1688]" targetBox="[191,1439,191,1633]" targetPageId="4">
<paragraph id="E72736F9FFE0FFCBFF799FC4FB7BF94A" blockId="4.[199,1187,1665,1688]" box="[199,1187,1665,1688]" pageId="4" pageNumber="3">
<emphasis id="D5ECEAEBFFE0FFCBFF799FC4FEF7F94A" bold="true" box="[199,303,1665,1688]" pageId="4" pageNumber="3">Figure 1:</emphasis>
Adult male
<taxonomicName id="20984D7AFFE0FFCBFE189FC7FD8CF94A" authorityName="Mello-Leitao" authorityYear="1932" box="[422,596,1666,1688]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE0FFCBFE189FC7FD8CF94A" box="[422,596,1666,1688]" italics="true" pageId="4" pageNumber="3">Rhopalurus pintoi</emphasis>
</taxonomicName>
from Rupununi:
<emphasis id="D5ECEAEBFFE0FFCBFD449FC6FCD6F94A" bold="true" box="[762,782,1667,1688]" pageId="4" pageNumber="3">a)</emphasis>
entire dorsal view;
<emphasis id="D5ECEAEBFFE0FFCBFC739FC6FC3AF94A" bold="true" box="[973,994,1667,1688]" pageId="4" pageNumber="3">b)</emphasis>
entire ventral view.
</paragraph>
</caption>
<caption id="B3E76671FFE1FFCAFF7E9E1BFD77F874" ID-DOI="http://doi.org/10.5281/zenodo.4673307" ID-Zenodo-Dep="4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,268,1886,1909]" targetBox="[228,1392,168,1856]" targetPageId="5">
<paragraph id="E72736F9FFE1FFCAFF7E9E1BFD77F874" blockId="5.[192,1442,1886,1958]" pageId="5" pageNumber="4">
<emphasis id="D5ECEAEBFFE1FFCAFF7E9E1BFEF4F8A7" bold="true" box="[192,300,1886,1909]" pageId="5" pageNumber="4">Figure 2:</emphasis>
Adult male
<taxonomicName id="20984D7AFFE1FFCAFE159E1AFD81F8A6" authorityName="Mello-Leitao" authorityYear="1932" box="[427,601,1887,1908]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE1FFCAFE159E1AFD81F8A6" box="[427,601,1887,1908]" italics="true" pageId="5" pageNumber="4">Rhopalurus pintoi</emphasis>
</taxonomicName>
from Rupununi:
<emphasis id="D5ECEAEBFFE1FFCAFCB99E1AFCC2F8A6" bold="true" box="[775,794,1887,1908]" pageId="5" pageNumber="4">a)</emphasis>
pedipalp;
<emphasis id="D5ECEAEBFFE1FFCAFC3A9E1AFC41F8A6" bold="true" box="[900,921,1887,1908]" pageId="5" pageNumber="4">b)</emphasis>
chelicerae and carapace;
<emphasis id="D5ECEAEBFFE1FFCAFB299E1AFB72F8A6" bold="true" box="[1175,1194,1887,1908]" pageId="5" pageNumber="4">c)</emphasis>
sternopectinal region;
<emphasis id="D5ECEAEBFFE1FFCAFA339E1AFA7AF8A6" bold="true" box="[1421,1442,1887,1908]" pageId="5" pageNumber="4">d)</emphasis>
tergite VII, metasoma and telson, dorsal view;
<emphasis id="D5ECEAEBFFE1FFCAFD319E3DFD79F85F" bold="true" box="[655,673,1912,1933]" pageId="5" pageNumber="4">e)</emphasis>
sternite VII, metasoma and telson, ventral view;
<emphasis id="D5ECEAEBFFE1FFCAFB399E3DFB4FF85F" bold="true" box="[1159,1175,1912,1933]" pageId="5" pageNumber="4">f)</emphasis>
metasoma, lateral view;
<emphasis id="D5ECEAEBFFE1FFCAFA309E3DFA7AF85F" bold="true" box="[1422,1442,1912,1933]" pageId="5" pageNumber="4">g)</emphasis>
metasomal segments IVV and telson, lateral view.
</paragraph>
</caption>
<paragraph id="E72736F9FFE2FFC9FD1799A9FABFFECF" blockId="6.[233,1399,236,398]" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FD1799A9FABFFECF" bold="true" pageId="6" pageNumber="5">Pectinal teeth Sex N Mean SD</emphasis>
</paragraph>
<paragraph id="E72736F9FFE2FFC9FE25985FFAAFFE59" blockId="6.[233,1399,236,398]" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FE25985FFBB9FEE6" bold="true" box="[411,1121,282,308]" pageId="6" pageNumber="5">21222324252627282930</emphasis>
<specimenCount id="F19EFD70FFE2FFC9FF579803FECDFEB2" box="[233,277,326,352]" pageId="6" pageNumber="5" type="male">♂♂</specimenCount>
12 1 7 3 1 28.33 ± 0.78
<specimenCount id="F19EFD70FFE2FFC9FF579831FECDFE5C" box="[233,277,372,398]" pageId="6" pageNumber="5" type="female">♀♀</specimenCount>
26 2 5 5 10 4 23.35 ± 1.20
</paragraph>
</subSubSection>
<caption id="B3E76671FFE2FFC9FF7E98F7FB6BFE30" pageId="6" pageNumber="5" startId="6.[192,257,434,457]">
<paragraph id="E72736F9FFE2FFC9FF7E98F7FB6BFE30" blockId="6.[192,1440,434,482]" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FF7E98F7FEC6FE1B" bold="true" box="[192,286,434,457]" pageId="6" pageNumber="5">Table 2:</emphasis>
Variation of pectinal tooth count in
<taxonomicName id="20984D7AFFE2FFC9FDC898F6FCFBFE1B" authorityName="Mello-Leitao" authorityYear="1932" box="[630,803,435,457]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE2FFC9FDC898F6FCFBFE1B" box="[630,803,435,457]" italics="true" pageId="6" pageNumber="5">Rhopalurus pintoi</emphasis>
</taxonomicName>
, including data from MelloLeitão (1932, holotype) and Lourenço &amp; PintodaRocha (1997, as
<taxonomicName id="20984D7AFFE2FFC9FE659888FDEBFE30" box="[475,563,461,482]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="D5ECEAEBFFE2FFC9FE659888FDEBFE30" box="[475,563,461,482]" italics="true" pageId="6" pageNumber="5">R. piceus</emphasis>
</taxonomicName>
). Abbreviations: number of pectines (N), standard deviation (SD).
</paragraph>
</caption>
<subSubSection id="AF826572FFE2FFC9FF7E9B48FE2AFD0E" pageId="6" pageNumber="5" type="distribution">
<paragraph id="E72736F9FFE2FFC9FF7E9B48FE2AFD0E" blockId="6.[192,802,525,732]" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FF7E9B48FE9DFDF6" bold="true" box="[192,325,525,548]" italics="true" pageId="6" pageNumber="5">Distribution</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FEEB9B48FE40FDF6" box="[341,408,525,548]" captionStart="Figure 7" captionStartId="12.[192,268,1158,1181]" captionTargetBox="[191,1440,191,1128]" captionTargetId="figure-8@12.[192,1439,192,1127]" captionTargetPageId="12" captionText="Figure 7: Geographical distribution of Rhopalurus pintoi (15) and Rhopalurus crassicauda (57): Tepequén [1], Surumu [2], Rio Surumu [3], Rio Tacutú [4], Rupununi [5], 12 km north of Boa Vista [6], Rio Branco [7]." figureDoi="http://doi.org/10.5281/zenodo.4673325" httpUri="https://zenodo.org/record/4673325/files/figure.png" pageId="6" pageNumber="5">Fig. 7</figureCitation>
): this species appears to be endemic the Rupununi region comprising the border region of
<collectingCountry id="9F8F7669FFE2FFC9FF7E9B0EFEDCFDB0" box="[192,260,587,610]" name="Brazil" pageId="6" pageNumber="5">Brazil</collectingCountry>
,
<collectingCountry id="9F8F7669FFE2FFC9FEAB9B0EFEB0FDB0" box="[277,360,587,610]" name="Guyana" pageId="6" pageNumber="5">Guyana</collectingCountry>
and
<collectingCountry id="9F8F7669FFE2FFC9FE149B0EFDC6FDB0" box="[426,542,587,610]" name="Venezuela" pageId="6" pageNumber="5">Venezuela</collectingCountry>
. Specimens have been collected only from four localities in
<collectingCountry id="9F8F7669FFE2FFC9FDDF9B2CFD7AFD52" box="[609,674,617,640]" name="Brazil" pageId="6" pageNumber="5">Brazil</collectingCountry>
and one in
<collectingCountry id="9F8F7669FFE2FFC9FF7E9BCDFECEFD4D" box="[192,278,648,671]" name="Guyana" pageId="6" pageNumber="5">Guyana</collectingCountry>
, but it is probably present also in neighboring
<collectingCountry id="9F8F7669FFE2FFC9FF7E9BE2FEE9FD6C" box="[192,305,679,702]" name="Venezuela" pageId="6" pageNumber="5">Venezuela</collectingCountry>
(southern
<collectingRegion id="255CF81BFFE2FFC9FE0F9BE2FD9FFD6C" box="[433,583,679,702]" country="Venezuela" name="Bolivar" pageId="6" pageNumber="5">Bolivar State</collectingRegion>
), where potentially suitable habitat also reaches.
</paragraph>
</subSubSection>
<subSubSection id="AF826572FFE2FFC8FF7E9A46FB6BF926" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="description">
<paragraph id="E72736F9FFE2FFC9FF7E9A46FC25F941" blockId="6.[192,803,771,1928]" lastBlockId="6.[830,1441,525,1683]" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FF7E9A46FE8FFCC8" bold="true" box="[192,343,771,794]" italics="true" pageId="6" pageNumber="5">Redescription</emphasis>
(adult male from Rupununi):
<emphasis id="D5ECEAEBFFE2FFC9FD109A46FCFAFCC8" bold="true" box="[686,802,771,794]" pageId="6" pageNumber="5">coloration</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FF769A64FED0FCEA" box="[200,264,801,824]" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Fig. 1</figureCitation>
) basically dark yellowish to reddish brown, very densely spotted with blackish brown all over the boy and appendages so the entire scorpion looks black to unaided eye, except for the bright white pectines and yellowish tip of legs and pedipalp fingers.
<emphasis id="D5ECEAEBFFE2FFC9FD889AD9FD7BFC61" bold="true" box="[566,675,924,947]" pageId="6" pageNumber="5">Carapace</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FD019AD9FCC2FC61" box="[703,794,924,947]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">Fig. 2b</figureCitation>
) trapezoidal, slightly longer than wide and with all carinae present and coarsely granulose to denticulate: posterior median carinae almost fused to central lateral carinae (both carinae are aligned in straight row, but separate by a small gap of about onegranule length), central median carinae fused to lateral ocular carinae, superciliary carinae fused to anterior median carinae, posterior lateral carinae long and displaced forward to mid portion of carapace; tegument densely granulose and with many coarse granules scattered, much more densely on interocular triangle; median eyes separate by more than one ocular diameter; three pairs of lateral eyes, all relatively large and about the same size.
<emphasis id="D5ECEAEBFFE2FFC9FD399C6FFD3DFA93" bold="true" box="[647,741,1322,1345]" pageId="6" pageNumber="5">Tergites</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FD489C6FFF01FAB2" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Fig. 1a</figureCitation>
) with the same sculpture as on carapace; median carina very strong and coarsely granulose in all tergites; VII with two pairs of strongly granulose lateral carinae.
<emphasis id="D5ECEAEBFFE2FFC9FF7E9CE0FEE2FA6E" bold="true" box="[192,314,1445,1468]" pageId="6" pageNumber="5">Chelicerae</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FEF69CE0FE4DFA6E" box="[328,405,1445,1468]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">Fig. 2b</figureCitation>
) with dentition typical for the genus; tegument smooth, polished.
<emphasis id="D5ECEAEBFFE2FFC9FE479C81FDB0FA09" bold="true" box="[505,616,1476,1499]" pageId="6" pageNumber="5">Pedipalps</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FDC59C81FD29FA09" box="[635,753,1476,1499]" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Figs. 1ab</figureCitation>
,
<figureCitation id="7FA32A7CFFE2FFC9FCBE9C81FCC1FA09" box="[768,793,1476,1499]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">2a</figureCitation>
) orthobothriotaxic A-α, with all segments very slender (femur, patella, and chela each longer than carapace) and densely covered by long, reddish macrosetae. Femur with all carinae crenulate serrate to denticulate, intercarinal tegument densely granulose. Patella with all carinae crenulate to serrate, intercarinal tegument with the same granulation as on femur, internal surface with many large and spiniform granules. Chela oval and slightly incrassate, conspicuously wider than patella; hand with all carinae weakly to moderately granulose, intercarinal tegument coriaceous to finely granulose, with sparse coarser granulation; fingers densely hirsute and without basal lobe/notch combination, but with a weak scallop instead, extending almost the entire length of the fingers when closed. Both fixed and movable fingers with enlarged, clawlike tip and nine principal rows of granules, of which the basalmost is straight and very long (more than twice longer than the remaining), all rows flanked on each side by many supernumerary granules; tip of movable finger with complex dentition, composed of two subrows of four external and two internal granules just basal to distal tooth.
<emphasis id="D5ECEAEBFFE2FFC9FBD09BA1FB7BFD29" bold="true" box="[1134,1187,740,763]" pageId="6" pageNumber="5">Legs</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FB049BA1FAEAFD29" box="[1210,1330,740,763]" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Figs. 1ab</figureCitation>
) with all carinae granulose, intercarinal tegument coriaceous to finely granulose.
<emphasis id="D5ECEAEBFFE2FFC9FC489A64FB81FCEA" bold="true" box="[1014,1113,801,824]" pageId="6" pageNumber="5">Sternum</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FBD99A64FB69FCEA" box="[1127,1201,801,824]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">Fig. 2c</figureCitation>
)
<typeStatus id="3823885BFFE2FFC9FB019A64FB35FCEA" box="[1215,1261,801,824]" pageId="6" pageNumber="5">type</typeStatus>
1 and triangular, typical for the genus.
<emphasis id="D5ECEAEBFFE2FFC9FB8A9A05FB49FC85" bold="true" box="[1076,1169,832,855]" pageId="6" pageNumber="5">Pectines</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FB1A9A05FB2BFC85" box="[1188,1267,832,855]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">Fig. 2c</figureCitation>
) elongate, with basal portion moderately enlarged; pectinal tooth count 28/28; basal plate wider than long, unmodified and with posterior margin slightly convex.
<emphasis id="D5ECEAEBFFE2FFC9FB169AD9FAD5FC61" bold="true" box="[1192,1293,924,947]" pageId="6" pageNumber="5">Sternites</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FAA59AD9FAAAFC61" box="[1307,1394,924,947]" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Figs. 1a</figureCitation>
,
<figureCitation id="7FA32A7CFFE2FFC9FAC09AD9FA40FC61" box="[1406,1432,924,947]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">2c</figureCitation>
) IIIVI smooth and shiny, with sparse large punctations, spiracles elongate and slit-like; stridulatory apparatus well developed; posterior margin of sternite V with a moderately large smooth patch, which is much wider than long, light yellow and slightly bulky; sternite VII granulose, with two pairs of very long and strongly granulose lateral carinae.
<emphasis id="D5ECEAEBFFE2FFC9FBE69D36FB16FB58" bold="true" box="[1112,1230,1139,1162]" pageId="6" pageNumber="5">Metasoma</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FB619D36FA8CFB58" box="[1247,1364,1139,1162]" captionStart="Figure 1" captionStartId="4.[199,275,1665,1688]" captionTargetBox="[191,1439,191,1633]" captionTargetPageId="4" captionText="Figure 1: Adult male Rhopalurus pintoi from Rupununi: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673305" httpUri="https://zenodo.org/record/4673305/files/figure.png" pageId="6" pageNumber="5">Figs. 1ab</figureCitation>
,
<figureCitation id="7FA32A7CFFE2FFC9FADC9D36FA40FB58" box="[1378,1432,1139,1162]" captionStart="Figure 2" captionStartId="5.[192,268,1886,1909]" captionTargetBox="[228,1392,168,1856]" captionTargetId="figure-8@5.[228,1392,168,1856]" captionTargetPageId="5" captionText="Figure 2: Adult male Rhopalurus pintoi from Rupununi: a) pedipalp; b) chelicerae and carapace; c) sternopectinal region; d) tergite VII, metasoma and telson, dorsal view; e) sternite VII, metasoma and telson, ventral view; f) metasoma, lateral view; g) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4673307" httpUri="https://zenodo.org/record/4673307/files/figure.png" pageId="6" pageNumber="5">2dg</figureCitation>
) strongly incrassate distally, with each segment noticeably wider than the preceding, especially on IVV which are inflate; intercarinal tegument coarsely and densely granulose on ventral and lateral surfaces, smooth to sparsely granulose on dorsal surface; segments III with ten complete carinae (even though lateral inframedian carinae is poorly defined in the basal portion of the latter), IIII V with eight, V with five (even though the ventrosubmedian carinae are present on proximal third), all strongly developed and coarsely serrate to denticulate; dorsomedian furrow shallow and narrow on segments IIII, progressively much deeper and wider on segments IVV, specially on the latter where it becomes a deep subtriangular depression; telson with vesicle large, oval and smooth, subaculear tubercle vestigial and very close to the base of the aculeus, which is very long, sharp and shallowly curved.
</paragraph>
<paragraph id="E72736F9FFE2FFC8FC809FFCFDD9FA32" blockId="6.[830,1441,1721,1928]" lastBlockId="7.[192,804,1419,1504]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
<emphasis id="D5ECEAEBFFE2FFC9FC809FFCFC49F902" bold="true" box="[830,913,1721,1744]" italics="true" pageId="6" pageNumber="5">Female</emphasis>
(
<figureCitation id="7FA32A7CFFE2FFC9FC1D9FFCFC3FF902" box="[931,999,1721,1744]" captionStart="Figure 3" captionStartId="7.[192,268,1334,1357]" captionTargetBox="[192,1452,192,1304]" captionTargetId="figure-7@7.[192,1452,192,1304]" captionTargetPageId="7" captionText="Figure 3: Adult female Rhopalurus pintoi from Rupununi: a) pedipalp; b) closeup of pedipalp movable finger; c) chelicerae and carapace; d) sternopectinal region; e) same specimen, still alive in captivity." figureDoi="http://doi.org/10.5281/zenodo.4673311" httpUri="https://zenodo.org/record/4673311/files/figure.png" pageId="6" pageNumber="5">Fig. 3</figureCitation>
,
<tableCitation id="AA1A0342FFE2FFC9FC4B9FFCFBBDF902" box="[1013,1125,1721,1744]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="3.[192,257,1110,1133]" captionStartId-1="6.[192,257,434,457]" captionText-0="Table 1: Measurements of four adult Rhopalurus pintoi from Rupununi. Abbreviations: length (L), width (W), posterior width (Wp), depth (H)." captionText-1="Table 2: Variation of pectinal tooth count in Rhopalurus pintoi, including data from MelloLeitão (1932, holotype) and Lourenço &amp; PintodaRocha (1997, as R. piceus). Abbreviations: number of pectines (N), standard deviation (SD)." pageId="6" pageNumber="5">Tabs. 12</tableCitation>
): in general is similar to the male, but there is a marked sexual dimorphism evidenced by:
<emphasis id="D5ECEAEBFFE2FFC9FCCD9FB3FC4AF8DF" bold="true" box="[883,914,1782,1805]" pageId="6" pageNumber="5">(1)</emphasis>
mesosoma relatively larger and wider;
<emphasis id="D5ECEAEBFFE2FFC9FA3F9FB3FA78F8DF" bold="true" box="[1409,1440,1782,1805]" pageId="6" pageNumber="5">(2)</emphasis>
metasoma less robust, only moderately incrassate distally;
<emphasis id="D5ECEAEBFFE2FFC9FCC19E71FC46F899" bold="true" box="[895,926,1844,1867]" pageId="6" pageNumber="5">(3)</emphasis>
pedipalp chelae noticeably more slender, with carinae stronger;
<emphasis id="D5ECEAEBFFE2FFC9FBBE9E17FBC7F8BB" bold="true" box="[1024,1055,1874,1897]" pageId="6" pageNumber="5">(4)</emphasis>
pedipalp fingers much longer and straight, without scallop;
<emphasis id="D5ECEAEBFFE2FFC9FBE79E34FBA0F85A" bold="true" box="[1113,1144,1905,1928]" pageId="6" pageNumber="5">(5)</emphasis>
genital papillae absent;
<emphasis id="D5ECEAEBFFE2FFC9FA3F9E34FA78F85A" bold="true" box="[1409,1440,1905,1928]" pageId="6" pageNumber="5">(6)</emphasis>
pectines with markedly lower number of teeth, which are also comparative smaller;
<emphasis id="D5ECEAEBFFE3FFC8FE5E9CEFFE27FA13" bold="true" box="[480,511,1450,1473]" pageId="7" pageNumber="6">(7)</emphasis>
basal pectinal plate with a very large and deep discal pit.
</paragraph>
<caption id="B3E76671FFE3FFC8FF7E9C73FC43FAB4" ID-DOI="http://doi.org/10.5281/zenodo.4673311" ID-Zenodo-Dep="4673311" httpUri="https://zenodo.org/record/4673311/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,268,1334,1357]" targetBox="[192,1452,192,1304]" targetPageId="7">
<paragraph id="E72736F9FFE3FFC8FF7E9C73FC43FAB4" blockId="7.[191,1440,1334,1382]" pageId="7" pageNumber="6">
<emphasis id="D5ECEAEBFFE3FFC8FF7E9C73FEF0FA9F" bold="true" box="[192,296,1334,1357]" pageId="7" pageNumber="6">Figure 3:</emphasis>
Adult female
<taxonomicName id="20984D7AFFE3FFC8FE0E9C72FD85FA9F" authorityName="Mello-Leitao" authorityYear="1932" box="[432,605,1335,1357]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pintoi">
<emphasis id="D5ECEAEBFFE3FFC8FE0E9C72FD85FA9F" box="[432,605,1335,1357]" italics="true" pageId="7" pageNumber="6">Rhopalurus pintoi</emphasis>
</taxonomicName>
from Rupununi:
<emphasis id="D5ECEAEBFFE3FFC8FCBC9C7DFCCDFA9F" bold="true" box="[770,789,1336,1357]" pageId="7" pageNumber="6">a)</emphasis>
pedipalp;
<emphasis id="D5ECEAEBFFE3FFC8FCC49C7DFC57FA9F" bold="true" box="[890,911,1336,1357]" pageId="7" pageNumber="6">b)</emphasis>
closeup of pedipalp movable finger;
<emphasis id="D5ECEAEBFFE3FFC8FB409C7DFAC9FA9F" bold="true" box="[1278,1297,1336,1357]" pageId="7" pageNumber="6">c)</emphasis>
chelicerae and carapace;
<emphasis id="D5ECEAEBFFE3FFC8FE9F9C14FEEEFAB4" bold="true" box="[289,310,1361,1382]" pageId="7" pageNumber="6">d)</emphasis>
sternopectinal region;
<emphasis id="D5ECEAEBFFE3FFC8FDAD9C14FDFDFAB4" bold="true" box="[531,549,1361,1382]" pageId="7" pageNumber="6">e)</emphasis>
same specimen, still alive in captivity.
</paragraph>
</caption>
<paragraph id="E72736F9FFE3FFC8FF7E9F43FA98FA32" blockId="7.[191,803,1542,1903]" lastBlockId="7.[830,1441,1419,1504]" pageId="7" pageNumber="6">
<emphasis id="D5ECEAEBFFE3FFC8FF7E9F43FEF1F9CF" bold="true" box="[192,297,1542,1565]" italics="true" pageId="7" pageNumber="6">Variation</emphasis>
: among the examined adults there are two size classes in each sex but inside the same class, males are slightly smaller than females (
<tableCitation id="AA1A0342FFE3FFC8FDBE9F06FD9CF988" box="[512,580,1603,1626]" captionStart="Table 1" captionStartId="3.[192,257,1110,1133]" captionText="Table 1: Measurements of four adult Rhopalurus pintoi from Rupununi. Abbreviations: length (L), width (W), posterior width (Wp), depth (H)." pageId="7" pageNumber="6">Tab. 1</tableCitation>
). Pedipalp fixed and movable fingers almost always have nine principal rows of granules, but in some specimens (mostly juveniles) the basalmost row is divided, giving a count of ten rows (
<figureCitation id="7FA32A7CFFE3FFC8FD489FDAFF03F907" captionStart="Figure 3" captionStartId="7.[192,268,1334,1357]" captionTargetBox="[192,1452,192,1304]" captionTargetId="figure-7@7.[192,1452,192,1304]" captionTargetPageId="7" captionText="Figure 3: Adult female Rhopalurus pintoi from Rupununi: a) pedipalp; b) closeup of pedipalp movable finger; c) chelicerae and carapace; d) sternopectinal region; e) same specimen, still alive in captivity." figureDoi="http://doi.org/10.5281/zenodo.4673311" httpUri="https://zenodo.org/record/4673311/files/figure.png" pageId="7" pageNumber="6">Fig. 3b</figureCitation>
). Pectinal toot counts varied from 2730 (mode 28) in males, and 2125 (mode 25) in females (
<tableCitation id="AA1A0342FFE3FFC8FD2B9F98FD3AF926" box="[661,738,1757,1780]" captionStart="Table 2" captionStartId="6.[192,257,434,457]" captionText="Table 2: Variation of pectinal tooth count in Rhopalurus pintoi, including data from MelloLeitão (1932, holotype) and Lourenço &amp; PintodaRocha (1997, as R. piceus). Abbreviations: number of pectines (N), standard deviation (SD)." pageId="7" pageNumber="6">Tab. 2</tableCitation>
); this species is unique in the genus by the strong sexual dimorphism in pectinal tooth counts (male and female ranges entirely separated), and male counts represent the highest values amongst all
<taxonomicName id="20984D7AFFE3FFC8FE5F9E12FD86F8BD" authorityName="Thorell" authorityYear="1876" box="[481,606,1879,1903]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="D5ECEAEBFFE3FFC8FE5F9E12FD86F8BD" box="[481,606,1879,1903]" italics="true" pageId="7" pageNumber="6">Rhopalurus</emphasis>
</taxonomicName>
species. Juveniles are very similar to the adults in coloration and general morphology, but comparatively have much more slender pedipalps and less incrassate metasoma (
<figureCitation id="7FA32A7CFFE3FFC8FB4C9C8CFAEBFA32" box="[1266,1331,1481,1504]" captionStart="Figure 4" captionStartId="8.[192,268,1926,1949]" captionTargetBox="[191,1408,167,1897]" captionTargetId="figure-8@8.[192,1407,168,1896]" captionTargetPageId="8" captionText="Figure 4: Juvenile female Rhopalurus pintoi from Rupununi, still alive in captivity." figureDoi="http://doi.org/10.5281/zenodo.4673315" httpUri="https://zenodo.org/record/4673315/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="E72736F9FFE3FFC8FC809F43FB6BF926" blockId="7.[830,1441,1542,1780]" pageId="7" pageNumber="6">
<emphasis id="D5ECEAEBFFE3FFC8FC809F43FC27F9CF" bold="true" box="[830,1023,1542,1565]" italics="true" pageId="7" pageNumber="6">Ecological notes</emphasis>
: according to the collector of the samples available to us, about
<specimenCount id="F19EFD70FFE3FFC8FB369F60FAC0F9EE" box="[1160,1304,1573,1596]" pageId="7" pageNumber="6" type="generic">20 specimens</specimenCount>
were found, all under stones inside a small relict patch (100 x
<quantity id="20609B1CFFE3FFC8FAEA9F06FA4FF988" box="[1364,1431,1603,1626]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="6" unit="m" value="100.0">100 m</quantity>
) of primary forest enclaved inside surrounding hilly grasslands on volcanic soils (
<figureCitation id="7FA32A7CFFE3FFC8FB169FC4FB2CF94A" box="[1192,1268,1665,1688]" captionStart="Figure 8" captionStartId="13.[192,268,1926,1949]" captionTargetBox="[216,1417,178,1897]" captionTargetId="figure-7@13.[217,1416,179,1896]" captionTargetPageId="13" captionText="Figure 8: Two views of the habitat where both Rhopalurus pintoi and Rhopalurus crassicauda occur at Rupununi (copyright by Hans-Werner Auer)." figureDoi="http://doi.org/10.5281/zenodo.4673329" httpUri="https://zenodo.org/record/4673329/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
). Only single specimens were found per stone, at an approximate density of four scorpions per
<quantity id="20609B1CFFE3FFC8FBC49FFBFB77F907" box="[1146,1199,1726,1749]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="6" unit="m" value="10.0">10 m</quantity>
<superScript id="10ED9BB1FFE3FFC8FB119FFDFB60F915" attach="left" box="[1199,1208,1720,1735]" fontSize="6" pageId="7" pageNumber="6">2</superScript>
, sympatrically with a single specimen of
<taxonomicName id="20984D7AFFE3FFC8FBB39F98FB74F926" authorityName="Caporiacco" authorityYear="1947" box="[1037,1196,1756,1780]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="D5ECEAEBFFE3FFC8FBB39F98FB74F926" box="[1037,1196,1756,1780]" italics="true" pageId="7" pageNumber="6">R. crassicauda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>