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<document id="4EE3072B42AD365150BE83D00E01F8C4" ID-DOI="10.5281/zenodo.176738" ID-GBIF-Dataset="2dabade9-cae2-445f-a67b-40807d76c658" ID-ISSN="1175-5326" ID-Zenodo-Dep="176738" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838674184" checkinUser="plazi" docAuthor="Assing, Volker" docDate="2007" docId="0D308E1EFF8FFFBBFF39FDDFFCC3FB0F" docLanguage="en" docName="zt01474p054.pdf" docOrigin="Zootaxa 1474" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Vulda ottomana Cameron 1912" docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="F109F666FF88FFB2FFAEFFA8FFD2FFCD" masterDocTitle="On the Xantholinini of Turkey and adjacent regions (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="8" updateTime="1698224072116" updateUser="plazi">
<mods:mods id="B8E341BDC6ACEB2AEDE62C2CE0FEAEC6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8351A275FDD6A2BEBB85808A3E7EC959">
<mods:title id="10AAA3FFB12F01533B57A541180D543D">On the Xantholinini of Turkey and adjacent regions (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="F0E8FF9A4ECB0849647A2BFAE2D4513C" type="personal">
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</mods:role>
<mods:namePart id="1E8E40991844011F82EA1F80C12FDBF3">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="401EB9D56DE4F7DA6E6F57405B70A2D7">text</mods:typeOfResource>
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<mods:part id="F2D337BA2EB78F9E38D2BA3E3DDD599C">
<mods:date id="323F7C0DFF0AA69287A87BDA2D49F0A6">2007</mods:date>
<mods:detail id="AC4BA70AF2423A3D3BD73DCA5873F3C5" type="volume">
<mods:number id="BFA9FA132FF32DB4F9D2BDA1FA18E8BD">1474</mods:number>
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<mods:classification id="EDA999E821CEF4A52A37FC016763F158">journal article</mods:classification>
<mods:identifier id="27A20FFC115605FBE8E233944DDC3298" type="DOI">10.5281/zenodo.176738</mods:identifier>
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<treatment id="0D308E1EFF8FFFBBFF39FDDFFCC3FB0F" ID-DOI="http://doi.org/10.5281/zenodo.6244402" ID-GBIF-Taxon="119359755" ID-Zenodo-Dep="6244402" LSID="urn:lsid:plazi:treatment:0D308E1EFF8FFFBBFF39FDDFFCC3FB0F" httpUri="http://treatment.plazi.org/id/0D308E1EFF8FFFBBFF39FDDFFCC3FB0F" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="CD836C83FF8FFFB5FF39FDDFFE04FD77" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="85263F08FF8FFFB5FF39FDDFFDE2FD5C" blockId="7.[151,560,631,698]" box="[151,560,631,657]" pageId="7" pageNumber="8">
<heading id="DE6E8864FF8FFFB5FF39FDDFFDE2FD5C" bold="true" box="[151,560,631,657]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4299448BFF8FFFB5FF39FDDFFDE2FD5C" ID-CoL="5BSD2" authority="Cameron, 1912" authorityName="Cameron" authorityYear="1912" box="[151,560,631,657]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF8FFFB5FF39FDDFFDE2FD5C" bold="true" box="[151,560,631,657]" pageId="7" pageNumber="8">
<emphasis id="B7EDE31AFF8FFFB5FF39FDDFFE8AFD5C" bold="true" box="[151,344,631,657]" italics="true" pageId="7" pageNumber="8">Vulda ottomana</emphasis>
(
<collectingCountry id="FD8E7F98FF8FFFB5FEC9FDDFFE0DFD5C" box="[359,479,631,657]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
, 1912)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="85263F08FF8FFFB5FF39FD08FE04FD77" blockId="7.[151,560,631,698]" box="[151,470,672,698]" pageId="7" pageNumber="8">
(
<figureCitation id="1DA2238DFF8FFFB5FF31FD08FEDDFD77" box="[159,271,672,698]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="7" pageNumber="8">Figs. 1–6</figureCitation>
,
<figureCitation id="1DA2238DFF8FFFB5FEB2FD08FEBAFD77" box="[284,360,672,698]" captionStart-0="FIGURE 13" captionStart-1="FIGURES 14 – 21" captionStartId-0="11.[151,255,1223,1247]" captionStartId-1="13.[151,269,1783,1807]" captionTargetBox-0="[230,1356,458,1199]" captionTargetBox-1="[309,1234,474,1756]" captionTargetId-0="figure@11.[230,1356,454,1200]" captionTargetId-1="figure@13.[302,1285,474,1759]" captionTargetPageId-0="11" captionTargetPageId-1="13" captionText-0="FIGURE 13. Distribution of the genus Vulda in Turkey: V. ottomana (Cameron) (filled circles), V. vignai Bordoni (open circle), V. cangalica sp. n. (filled squares), and V. brignolii Bordoni (open square). Since the type locality of V. brignolii (&quot; Boraby gölü &quot;) was not identified (misspelling?), the respective square is placed in the centre of Amasya province." captionText-1="FIGURES 14 – 21. Vulda ottomana (Cameron) (14 – 15; holotype of Xantholinus anatolicus Coiffait) and Xantholinus gridellii Coiffait (16 – 21; 20: paratype): 14, 20 – 21 — aedeagus; 15 — internal structures of aedeagus; 16 — habitus; 17 — forebody; 18 — male tergite VIII; 19 — male sternite VIII. Scale bars: 16 – 17: 1.0 mm; 18 – 21: 0.5 mm; 14: 0.2 mm; 15: 0.1 mm." httpUri-0="https://zenodo.org/record/176741/files/figure.png" httpUri-1="https://zenodo.org/record/176742/files/figure.png" pageId="7" pageNumber="8">13–15</figureCitation>
,
<figureCitation id="1DA2238DFF8FFFB5FEDAFD08FE1CFD77" box="[372,462,672,698]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="7" pageNumber="8">99–100</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CD836C83FF8FFFB5FF39FD58FCA2FCEA" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="85263F08FF8FFFB5FF39FD58FD12FCCA" blockId="7.[151,880,752,807]" box="[151,704,752,775]" pageId="7" pageNumber="8">
<taxonomicName id="4299448BFF8FFFB5FF39FD58FD7BFCCA" authority="Cameron, 1912: 1" authorityName="Cameron" authorityPageNumber="1" authorityYear="1912" box="[151,681,752,775]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomanus" subGenus="Vulda">
<emphasis id="B7EDE31AFF8FFFB5FF39FD58FECAFCCA" box="[151,280,752,775]" italics="true" pageId="7" pageNumber="8">Xantholinus</emphasis>
(
<emphasis id="B7EDE31AFF8FFFB5FE89FD58FEB1FCCA" box="[295,355,752,775]" italics="true" pageId="7" pageNumber="8">Vulda</emphasis>
)
<emphasis id="B7EDE31AFF8FFFB5FEDAFD58FE31FCCA" box="[372,483,752,775]" italics="true" pageId="7" pageNumber="8">ottomanus</emphasis>
<collectingCountry id="FD8E7F98FF8FFFB5FE44FD58FD9DFCCA" box="[490,591,752,775]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
, 1912:
<specimenCount id="939FF481FF8FFFB5FD32FD58FD7BFCCA" box="[668,681,752,775]" pageId="7" pageNumber="8" type="generic">1</specimenCount>
</taxonomicName>
f.
</paragraph>
<paragraph id="85263F08FF8FFFB5FF39FCB8FCA2FCEA" blockId="7.[151,880,752,807]" box="[151,880,783,807]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="A5891826FF8FFFB5FF39FCB8FCA2FCEA" box="[151,880,783,807]" pageId="7" pageNumber="8">
<taxonomicName id="4299448BFF8FFFB5FF39FCB8FCD1FCEA" authority="Coiffait, 1965: 121" authorityName="Coiffait" authorityPageNumber="121" authorityYear="1965" box="[151,771,784,807]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolicus" subGenus="Leptophallus">
<emphasis id="B7EDE31AFF8FFFB5FF39FCB8FECAFCEA" box="[151,280,784,807]" italics="true" pageId="7" pageNumber="8">Xantholinus</emphasis>
(
<emphasis id="B7EDE31AFF8FFFB5FE89FCB8FE60FCEA" box="[295,434,784,807]" italics="true" pageId="7" pageNumber="8">Leptophallus</emphasis>
)
<emphasis id="B7EDE31AFF8FFFB5FE6CFCB8FDE3FCEA" box="[450,561,784,807]" italics="true" pageId="7" pageNumber="8">anatolicus</emphasis>
<treatmentCitation id="04381919FF8FFFB5FD96FCB8FCD1FCEA" author="Coiffait" box="[568,771,784,807]" page="121" pageId="7" pageNumber="8" year="1965">
<bibRefCitation id="E10842F9FF8FFFB5FD96FCB8FD02FCEA" author="Coiffait" box="[568,720,784,807]" pageId="7" pageNumber="52" refString="Coiffait, H. (1965) Sur quelques Xantholininae de l'Asie Mineure (Coleoptera, Staphylinidae). Reichenbachia, 5, 120 - 123." type="journal article" year="1965">Coiffait, 1965</bibRefCitation>
:
<specimenCount id="939FF481FF8FFFB5FD75FCB8FCD1FCEA" box="[731,771,784,807]" pageId="7" pageNumber="8" type="generic">121</specimenCount>
</treatmentCitation>
</taxonomicName>
f.;
<emphasis id="B7EDE31AFF8FFFB5FC89FCA7FCA2FCEA" bold="true" box="[807,880,783,807]" pageId="7" pageNumber="8">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CD836C83FF8FFFB5FF39FCFFFE95FB7F" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="85263F08FF8FFFB5FF39FCFFFE8AFC27" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
<emphasis id="B7EDE31AFF8FFFB5FF39FCFFFCD4FCBC" bold="true" box="[151,774,855,881]" pageId="7" pageNumber="8">
<typeStatus id="5A2281AAFF8FFFB5FF39FCFFFF06FCBC" box="[151,212,855,881]" pageId="7" pageNumber="8">Type</typeStatus>
material examined.
<emphasis id="B7EDE31AFF8FFFB5FE4FFCFFFDA0FCBC" bold="true" box="[481,626,855,881]" italics="true" pageId="7" pageNumber="8">V. o t t o m a n a</emphasis>
:
<typeStatus id="5A2281AAFF8FFFB5FD25FCFFFCD3FCBC" box="[651,769,855,881]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
:
</emphasis>
ɗ, &quot;Beikos, 8.'03 M C. / 8448 /
<typeStatus id="5A2281AAFF8FFFB5FB07FCF0FB31FCBF" box="[1193,1251,856,882]" pageId="7" pageNumber="8">Type</typeStatus>
/ M.
<collectingCountry id="FD8E7F98FF8FFFB5FA9EFCF0FA49FCBF" box="[1328,1435,856,882]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
Bequest, B.M. 1955–147 /
<typeStatus id="5A2281AAFF8FFFB5FE74FC28FDE8FC57" box="[474,570,896,922]" pageId="7" pageNumber="8" type="syntype">Syntype</typeStatus>
<taxonomicName id="4299448BFF8FFFB5FDEDFC28FBABFC57" authority="Cameron, 1912" authorityName="Cameron" authorityYear="1912" box="[579,1145,896,922]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomanus" subGenus="Vulda">
Xantholinus (Vulda) ottomanus
<collectingCountry id="FD8E7F98FF8FFFB5FC6EFC28FBFDFC57" box="[960,1071,896,922]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
, 1912
</taxonomicName>
, det. R. G. Booth 2007 /
<typeStatus id="5A2281AAFF8FFFB5FF39FC00FED6FC0F" box="[151,260,936,962]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
<taxonomicName id="4299448BFF8FFFB5FEA3FC00FD44FC0F" authority="Cameron" authorityName="Cameron" box="[269,662,936,962]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomanus">
Xantholinus ottomanus
<collectingCountry id="FD8E7F98FF8FFFB5FD84FC00FD44FC0F" box="[554,662,936,962]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
</taxonomicName>
rev.
<bibRefCitation id="E10842F9FF8FFFB5FD7DFC00FC59FC0F" author="Assing" box="[723,907,936,962]" pageId="7" pageNumber="50" refString="Assing, V. (in press) New species, a new genus, and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beitrage zur Naturkunde Serie A (Biologie) (2007)." type="book" year="2007">V. Assing 2007</bibRefCitation>
/
<taxonomicName id="4299448BFF8FFFB5FC08FC00FB38FC0F" authority="Cameron" authorityName="Cameron" box="[934,1258,936,962]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomana">
Vulda ottomana (
<collectingCountry id="FD8E7F98FF8FFFB5FBDDFC00FB30FC0F" box="[1139,1250,936,962]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
)
</taxonomicName>
det.
<bibRefCitation id="E10842F9FF8FFFB5FA86FC00FF07FC27" author="Assing" pageId="7" pageNumber="50" refString="Assing, V. (in press) New species, a new genus, and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beitrage zur Naturkunde Serie A (Biologie) (2007)." type="book" year="2007">V. Assing 2007</bibRefCitation>
&quot; (
<collectionCode id="E388A7CDFF8FFFB5FF43FC78FE9EFC27" box="[237,332,976,1002]" pageId="7" pageNumber="8">BMNH</collectionCode>
).
</paragraph>
<paragraph id="85263F08FF8FFFB5FF68FC5FFF3FFBAF" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
<emphasis id="B7EDE31AFF8FFFB5FF68FC5FFE24FBDC" bold="true" box="[198,502,1015,1041]" pageId="7" pageNumber="8">
<taxonomicName id="4299448BFF8FFFB5FF68FC5FFEB5FBDC" box="[198,359,1015,1041]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B7EDE31AFF8FFFB5FF68FC5FFEB5FBDC" bold="true" box="[198,359,1015,1041]" italics="true" pageId="7" pageNumber="8">X. anatolicus</emphasis>
</taxonomicName>
:
<typeStatus id="5A2281AAFF8FFFB5FED5FC5FFE20FBDC" box="[379,498,1015,1041]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
:
</emphasis>
ɗ, &quot;Abant Geb. bei Bolu,
<date id="F12719C8FF8FFFB5FC97FC50FC62FBDF" box="[825,944,1016,1042]" pageId="7" pageNumber="8" value="1964-05-30">30.V.1964</date>
/ leg. H. Korge,
<collectingCountry id="FD8E7F98FF8FFFB5FBD1FC50FB36FBDF" box="[1151,1252,1016,1042]" name="Turkey" pageId="7" pageNumber="8">Anatolia</collectingCountry>
bor. /
<taxonomicName id="4299448BFF8FFFB5FA9BFC50FE59FBF7" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov." subGenus="Leptophallus">X. (Leptophallus) anatolicus</taxonomicName>
<taxonomicNameLabel id="ACDE5E61FF8FFFB5FE3DFB88FE01FBF7" box="[403,467,1056,1082]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
H. Coiffait det. 1964 /
<typeStatus id="5A2281AAFF8FFFB5FD44FB88FC85FBF7" box="[746,855,1056,1082]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4299448BFF8FFFB5FCDFFB88FB4DFBF7" authority="Coiffait" authorityName="Coiffait" box="[881,1183,1056,1082]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica">Vulda anatolica (Coiffait)</taxonomicName>
det. V. Assing 2006&quot; (cKor).
</paragraph>
<paragraph id="85263F08FF8FFFB5FF68FBC7FE95FB7F" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
<emphasis id="B7EDE31AFF8FFFB5FF68FBC7FC82FB44" bold="true" box="[198,848,1135,1161]" pageId="7" pageNumber="8">
Additional material examined.
<collectingCountry id="FD8E7F98FF8FFFB5FDFEFBC7FD1DFB44" box="[592,719,1135,1161]" name="Turkey" pageId="7" pageNumber="8">TURKEY</collectingCountry>
: Istanbul:
</emphasis>
1ɗ, &quot;Belgrader Wald&quot;,
<date id="F12719C8FF8FFFB5FBC0FBD8FAC4FB47" box="[1134,1302,1136,1162]" pageId="7" pageNumber="8" value="1954-07-01" valueMax="1954-07-07" valueMin="1954-07-01">1.–7.VII.1954</date>
, leg. Schubert (
<collectionCode id="E388A7CDFF8FFFB5FF7AFB30FEE9FB7F" box="[212,315,1176,1202]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="8">NHMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="CD836C83FF8FFFB5FF68FB17FA49FAE7" pageId="7" pageNumber="8" type="discussion">
<paragraph id="85263F08FF8FFFB5FF68FB17FA49FAE7" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
<emphasis id="B7EDE31AFF8FFFB5FF68FB17FE9BFB14" bold="true" box="[198,329,1215,1241]" pageId="7" pageNumber="8">Comment.</emphasis>
The original description of
<taxonomicName id="4299448BFF8FFFB5FD36FB68FC7BFB14" box="[664,937,1216,1241]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomanus">
<emphasis id="B7EDE31AFF8FFFB5FD36FB68FC7BFB14" box="[664,937,1216,1241]" italics="true" pageId="7" pageNumber="8">Xantholinus ottomanus</emphasis>
</taxonomicName>
is based on a single &quot;
<typeStatus id="5A2281AAFF8FFFB5FB03FB68FB37FB17" box="[1197,1253,1216,1242]" pageId="7" pageNumber="8">type</typeStatus>
&quot; &quot;taken at Beikos, in August, 1903, under a stone&quot; (
<collectingCountry id="FD8E7F98FF8FFFB5FDE1FB40FD6FFACF" box="[591,701,1256,1282]" name="Cameroon" pageId="7" pageNumber="8">Cameron</collectingCountry>
1912). The male
<typeStatus id="5A2281AAFF8FFFB5FC25FB40FC23FACF" box="[907,1009,1256,1282]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
was studied and its habitus and sexual characters were illustrated by
<bibRefCitation id="E10842F9FF8FFFB5FDB1FAB8FD03FAE7" author="Coiffait" box="[543,721,1296,1322]" pageId="7" pageNumber="52" refString="Coiffait, H. (1971) Xantholinini nouveaux ou mal connus de la region palearctique occidentale. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 106 (1970), 429 - 436." type="journal article" year="1971">Coiffait (1971)</bibRefCitation>
, who assigned it to the nominal subgenus of the genus
<taxonomicName id="4299448BFF8FFFB5FAFFFAB8FA41FAE4" box="[1361,1427,1296,1321]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FAFFFAB8FA41FAE4" box="[1361,1427,1296,1321]" italics="true" pageId="7" pageNumber="8">Vulda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CD836C83FF8FFFB5FF68FA90FA57F867" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="85263F08FF8FFFB5FF68FA90FDF9F9D7" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
<taxonomicName id="4299448BFF8FFFB5FF68FA90FE04FA9C" box="[198,470,1336,1361]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B7EDE31AFF8FFFB5FF68FA90FE04FA9C" box="[198,470,1336,1361]" italics="true" pageId="7" pageNumber="8">Xantholinus anatolicus</emphasis>
</taxonomicName>
was originally described based on a single
<typeStatus id="5A2281AAFF8FFFB5FC76FA90FBEDFA9F" box="[984,1087,1336,1362]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
male from the Abant Daġları near Bolu.
<bibRefCitation id="E10842F9FF8FFFB5FEB0FAC8FE1BFAB7" author="Coiffait" box="[286,457,1376,1402]" pageId="7" pageNumber="52" refString="Coiffait, H. (1965) Sur quelques Xantholininae de l'Asie Mineure (Coleoptera, Staphylinidae). Reichenbachia, 5, 120 - 123." type="journal article" year="1965">Coiffait (1965</bibRefCitation>
,
<bibRefCitation id="E10842F9FF8FFFB5FE76FAC8FDC6FAB7" author="Coiffait" box="[472,532,1376,1402]" pageId="7" pageNumber="52" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, sous-familles: Xantholininae et Leptotyphlinae. Supplement a la Nouvelle Revue d´Entomologie, 2 (2), 1 - 651." type="journal article" year="1972">1972</bibRefCitation>
) assigned the species to the subgenus
<taxonomicName id="4299448BFF8FFFB5FC46FAC8FAE0FAB7" authority="Coiffait, 1956" authorityName="Coiffait" authorityYear="1956" box="[1000,1330,1376,1402]" class="Insecta" family="Staphylinidae" genus="Leptophallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FC46FAC8FB53FAB4" box="[1000,1153,1376,1401]" italics="true" pageId="7" pageNumber="8">Leptophallus</emphasis>
<bibRefCitation id="E10842F9FF8FFFB5FB25FAC8FAE0FAB7" author="Coiffait" box="[1163,1330,1376,1402]" pageId="7" pageNumber="51" refString="Coiffait, H. (1956 a) Les Xantholinitae de France et des regions voisines (Col. Staphylinidae). Revue Francaise d´Entomologie, 23, 31 - 76 + i - xvi." type="journal article" year="1956" yearSuffix="a">Coiffait, 1956</bibRefCitation>
</taxonomicName>
a (now a synonym of
<taxonomicName id="4299448BFF8FFFB5FE82FA20FD63FA6F" ID-CoL="8X7JX" authority="Mulsant &amp; Rey, 1877" authorityName="Mulsant &amp; Rey" authorityYear="1877" box="[300,689,1416,1442]" class="Insecta" family="Staphylinidae" genus="Megalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FE82FA20FE7AFA6C" box="[300,424,1416,1441]" italics="true" pageId="7" pageNumber="8">Megalinus</emphasis>
<bibRefCitation id="E10842F9FF8FFFB5FE1EFA20FD63FA6F" author="Mulsant" box="[432,689,1416,1442]" pageId="7" pageNumber="53" refString="Mulsant, E. &amp; Rey, C. (1877) Tribu des brevipennes. Deuxieme famille. Xantholiniens. Memoires de l'Academie des Sciences, Belles-Lettres &amp; Arts de Lyon, 22, 217 - 344." type="journal article" year="1877">Mulsant &amp; Rey, 1877</bibRefCitation>
</taxonomicName>
) of the genus
<taxonomicName id="4299448BFF8FFFB5FCF5FA20FC3BFA6C" ID-CoL="8VS6L" box="[859,1001,1416,1441]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FCF5FA20FC3BFA6C" box="[859,1001,1416,1441]" italics="true" pageId="7" pageNumber="8">Xantholinus</emphasis>
</taxonomicName>
.
<bibRefCitation id="E10842F9FF8FFFB5FC57FA20FB62FA6F" author="Bordoni" box="[1017,1200,1416,1442]" pageId="7" pageNumber="51" refString="Bordoni, A. (1985) Studi sulla sistematica degli Xantholinus Berthold 1827, XIII. Una nuova specie dell'Iran (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica, 63 (1982), 61 - 63." type="journal article" year="1985">Bordoni (1985)</bibRefCitation>
subsequently transferred it to the subgenus
<taxonomicName id="4299448BFF8FFFB5FE62FA18FC9AFA07" authority="Bordoni, 1985" authorityName="Bordoni" authorityYear="1985" box="[460,840,1456,1482]" class="Insecta" family="Staphylinidae" genus="Neoleptophallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FE62FA18FD5EFA04" box="[460,652,1456,1481]" italics="true" pageId="7" pageNumber="8">Neoleptophallus</emphasis>
<bibRefCitation id="E10842F9FF8FFFB5FD36FA18FC9AFA07" author="Bordoni" box="[664,840,1456,1482]" pageId="7" pageNumber="51" refString="Bordoni, A. (1985) Studi sulla sistematica degli Xantholinus Berthold 1827, XIII. Una nuova specie dell'Iran (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica, 63 (1982), 61 - 63." type="journal article" year="1985">Bordoni, 1985</bibRefCitation>
</taxonomicName>
. An examination of the
<typeStatus id="5A2281AAFF8FFFB5FBDBFA18FB09FA07" box="[1141,1243,1456,1482]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
revealed that it refers to the genus
<taxonomicName id="4299448BFF8FFFB5FED9FA70FDE0FA3F" ID-CoL="8XQP9" authority="Jacquelin" authorityName="Jacquelin" box="[375,562,1496,1522]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF8FFFB5FED9FA70FE6BFA3C" box="[375,441,1496,1521]" italics="true" pageId="7" pageNumber="8">Vulda</emphasis>
Jacquelin
</taxonomicName>
du Val, 1853 and that the illustration of the aedeagus provided by
<bibRefCitation id="E10842F9FF8FFFB5FAEFFA70FF0FF9D7" author="Coiffait" pageId="7" pageNumber="52" refString="Coiffait, H. (1965) Sur quelques Xantholininae de l'Asie Mineure (Coleoptera, Staphylinidae). Reichenbachia, 5, 120 - 123." type="journal article" year="1965">Coiffait (1965</bibRefCitation>
,
<bibRefCitation id="E10842F9FF8FFFB5FF44F9A8FEF4F9D7" author="Coiffait" box="[234,294,1536,1562]" pageId="7" pageNumber="52" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, sous-familles: Xantholininae et Leptotyphlinae. Supplement a la Nouvelle Revue d´Entomologie, 2 (2), 1 - 651." type="journal article" year="1972">1972</bibRefCitation>
) is highly misleading.
</paragraph>
<paragraph id="85263F08FF8FFFB5FF68F980FA57F867" blockId="7.[151,1437,855,1962]" pageId="7" pageNumber="8">
At first glance, the
<typeStatus id="5A2281AAFF8FFFB5FE1AF980FDF4F98F" box="[436,550,1576,1602]" pageId="7" pageNumber="8" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="4299448BFF8FFFB5FDFAF980FD31F98C" ID-CoL="5BSD2" box="[596,739,1576,1601]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF8FFFB5FDFAF980FD31F98C" box="[596,739,1576,1601]" italics="true" pageId="7" pageNumber="8">V. ottomana</emphasis>
</taxonomicName>
and
<taxonomicName id="4299448BFF8FFFB5FC8CF980FC63F98C" box="[802,945,1576,1601]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B7EDE31AFF8FFFB5FC8CF980FC63F98C" box="[802,945,1576,1601]" italics="true" pageId="7" pageNumber="8">V. anatolica</emphasis>
</taxonomicName>
appear to refer to different species. The former has large and bulging eyes, a posteriorly somewhat tapering head, long elytra, and fully developed hind wings (
<figureCitation id="1DA2238DFF8FFFB5FE89F9D0FEACF95F" box="[295,382,1656,1682]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="7" pageNumber="8">Fig. 99</figureCitation>
), whereas the latter has small eyes not distinctly projecting from the lateral contours of the head, a head with subparallel lateral margins (dorsal view), distinctly shorter elytra, and reduced hind wings. The primary and secondary sexual characters, however, are practically identical and, moreover, other external characters (coloration, size, proportion, microsculpture, etc.) are highly similar, suggesting that both
<typeStatus id="5A2281AAFF8FFFB5FAF0F958FF07F8FF" pageId="7" pageNumber="8" type="holotype">holotypes</typeStatus>
belong to the same dimorphic species. There is also zoogeographic evidence supporting this conclusion: the male specimen listed above as additional material resembles the
<typeStatus id="5A2281AAFF8FFFB5FC11F8E8FBF4F897" box="[959,1062,1856,1882]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4299448BFF8FFFB5FBE0F8E8FB09F894" box="[1102,1243,1856,1881]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B7EDE31AFF8FFFB5FBE0F8E8FB09F894" box="[1102,1243,1856,1881]" italics="true" pageId="7" pageNumber="8">V. anatolica</emphasis>
</taxonomicName>
both in external and in sexual characters, but was found in Istanbul close to the
<typeStatus id="5A2281AAFF8FFFB5FCDAF8C0FC75F84F" box="[884,935,1896,1922]" pageId="7" pageNumber="8">type</typeStatus>
locality of
<taxonomicName id="4299448BFF8FFFB5FB85F8C0FB64F84C" ID-CoL="5BSD2" box="[1067,1206,1896,1921]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF8FFFB5FB85F8C0FB64F84C" box="[1067,1206,1896,1921]" italics="true" pageId="7" pageNumber="8">V. ottomana</emphasis>
</taxonomicName>
(
<figureCitation id="1DA2238DFF8FFFB5FB6BF8C0FAC9F84F" box="[1221,1307,1896,1922]" captionStart="FIGURE 13" captionStartId="11.[151,255,1223,1247]" captionTargetBox="[230,1356,458,1199]" captionTargetId="figure@11.[230,1356,454,1200]" captionTargetPageId="11" captionText="FIGURE 13. Distribution of the genus Vulda in Turkey: V. ottomana (Cameron) (filled circles), V. vignai Bordoni (open circle), V. cangalica sp. n. (filled squares), and V. brignolii Bordoni (open square). Since the type locality of V. brignolii (&quot; Boraby gölü &quot;) was not identified (misspelling?), the respective square is placed in the centre of Amasya province." httpUri="https://zenodo.org/record/176741/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
). Since the latter is the senior name, it takes precedence, and
<taxonomicName id="4299448BFF8FFFB5FD72F838FCBAF864" box="[732,872,1936,1961]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B7EDE31AFF8FFFB5FD72F838FCBAF864" box="[732,872,1936,1961]" italics="true" pageId="7" pageNumber="8">V. anatolica</emphasis>
</taxonomicName>
is here placed in synonymy with
<emphasis id="B7EDE31AFF8FFFB5FB5DF838FAA8F864" box="[1267,1402,1936,1961]" italics="true" pageId="7" pageNumber="8">V. o t t o m a n a</emphasis>
.
</paragraph>
</subSubSection>
<caption id="D1E66F80FF80FFBAFF39FBA6FB66FB85" httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9" targetBox="[308,1253,197,1012]" targetPageId="8">
<paragraph id="85263F08FF80FFBAFF39FBA6FB66FB85" blockId="8.[151,1436,1038,1096]" pageId="8" pageNumber="9">
<emphasis id="B7EDE31AFF80FFBAFF39FBA6FE91FBEB" bold="true" box="[151,323,1038,1062]" pageId="8" pageNumber="9">FIGURES 1–6.</emphasis>
<taxonomicName id="4299448BFF80FFBAFEE5FBA7FDA1FBE8" authority="Cameron" authorityName="Cameron" box="[331,627,1038,1062]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF80FFBAFEE5FBA7FE27FBEB" box="[331,501,1039,1062]" italics="true" pageId="8" pageNumber="9">Vulda ottomana</emphasis>
(Cameron)
</taxonomicName>
: 1—habitus; 2—forebody; 3—head in lateral view; 4—antenna; 5—aedeagus; 6—internal structure of aedeagus. Scale bars: 1–3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="CD836C83FF80FFBBFF68FBD3FC02FBEF" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="85263F08FF80FFBAFF68FBD3FE1CFA93" blockId="8.[151,1437,1147,1974]" pageId="8" pageNumber="9">
<emphasis id="B7EDE31AFF80FFBAFF68FBD3FEAEFB58" bold="true" box="[198,380,1147,1173]" pageId="8" pageNumber="9">Redescription.</emphasis>
Measurements (in mm) and ratios (
<typeStatus id="5A2281AAFF80FFBAFC98FBD4FC4CFB5B" box="[822,926,1148,1174]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4299448BFF80FFBAFC63FBD4FB8FFB58" box="[973,1117,1148,1173]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF80FFBAFC63FBD4FB8FFB58" box="[973,1117,1148,1173]" italics="true" pageId="8" pageNumber="9">V. ottomana</emphasis>
</taxonomicName>
,
<typeStatus id="5A2281AAFF80FFBAFBC1FBD4FB04FB5B" box="[1135,1238,1148,1174]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4299448BFF80FFBAFAABFBD4FA47FB58" box="[1285,1429,1148,1173]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B7EDE31AFF80FFBAFAABFBD4FA47FB58" box="[1285,1429,1148,1173]" italics="true" pageId="8" pageNumber="9">V. anatolica</emphasis>
</taxonomicName>
, additional specimen): AL: 2.14, 2.17, 1.98; HL: 1.27, 1.43, 1.34; HW: 0.89, 1.00, 0.95; PW: 0.71, 0.84, 0.80; PL: 1.11, 1.31, 1.22; EL: 1.03, 0.88, 0.76; EW: 1.18, 1.03, 0.92; AW: 0.95, 1.01, 0.94; TiL: 0.76, 0.83, 0.79; TaL: 0.68, 0.65, 0.63; ML: 0.72, 0.74, 0.72; TL: 6.9, 8.3, 7.3; HL/HW: 1.42, 1.44, 1.41; PW/HW: 0.80, 0.84, 0.84; PL/PW: 1.56, 1.57, 1.53; EL/PL: 0.93, 0.67, 0.62; EW/PW: 1.45, 1.23, 1.15; AW/EW: 0.93, 0.99, 1.02; TiL/TaL: 1.0 6, 1.28, 1.24.
</paragraph>
<paragraph id="85263F08FF80FFBAFF68FAC4FB45FA1B" blockId="8.[151,1437,1147,1974]" pageId="8" pageNumber="9">
Habitus of brachypterous morph as in
<figureCitation id="1DA2238DFF80FFBAFD27FAC4FD1DFA4B" box="[649,719,1388,1414]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
. Coloration: head blackish brown to blackish; pronotum reddish to dark brown; elytra yellowish to pale reddish; abdomen yellowish brown to brown; legs yellowish to reddish; antennae dark brown, with the basal 3 antennomeres reddish to reddish brown.
</paragraph>
<paragraph id="85263F08FF80FFBAFF6BFA4CFE66F8DB" blockId="8.[151,1437,1147,1974]" pageId="8" pageNumber="9">
Head distinctly oblong (see measurements and ratio HL/HW) and of subrectangular shape (brachypterous morph,
<figureCitation id="1DA2238DFF80FFBAFF5DF9A4FEE8F9EB" box="[243,314,1548,1574]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
) or tapering posteriad (macropterous morph,
<figureCitation id="1DA2238DFF80FFBAFCF6F9A4FC62F9EB" box="[856,944,1548,1574]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="8" pageNumber="9">Fig. 99</figureCitation>
); eyes small (
<figureCitation id="1DA2238DFF80FFBAFBFAF9A4FB4FF9EB" box="[1108,1181,1548,1574]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
), not projecting from lateral outline of head, approximately 1/4 the length of postocular region in dorsal view (brachypterous morph), or large and distinctly bulging (
<figureCitation id="1DA2238DFF80FFBAFDDEF9F4FD1AF9BB" box="[624,712,1628,1654]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="8" pageNumber="9">Fig. 99</figureCitation>
), approximately half the length of postocular region in dorsal view (macropterous morph); dorsal surface with rather sparse fine punctures and interspersed larger punctures; microsculpture absent. Antenna with antennomere I approximately as long as the combined length of antennomeres II–V; II and III oblong and of subequal length; IV weakly transverse; V–X weakly to moderately transverse (
<figureCitation id="1DA2238DFF80FFBAFEF0F954FE75F8DB" box="[350,423,1788,1814]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="85263F08FF80FFBAFF6BF88CFDE5F87B" blockId="8.[151,1437,1147,1974]" pageId="8" pageNumber="9">
Pronotum slender, distinctly narrower and smaller than head (see ratio PW/HW), weakly tapering posteriad; lateral margins straight or shallowly sinuate in dorsal view; dorsal series composed of 10–11 punctures; whole surface with very fine and rather sparse punctures, lateral areas with interspersed coarse punctures; microsculpture absent (
<figureCitation id="1DA2238DFF80FFBAFE07F834FE2EF87B" box="[425,508,1948,1974]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
,
<figureCitation id="1DA2238DFF80FFBAFDA6F834FDFBF87B" box="[520,553,1948,1974]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="8" pageNumber="9">99</figureCitation>
).
</paragraph>
<paragraph id="85263F08FF81FFBBFF6BFF30FAABFEE7" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">
Elytra distinctly shorter and slightly wider than pronotum (brachypterous morph) or almost as long and distinctly wider than pronotum (macropterous morph) (see ratios EL/PL and EW/PW); puncturation more or less well-defined and moderately dense (
<figureCitation id="1DA2238DFF81FFBBFDD6FF40FD1EFECF" box="[632,716,232,258]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="1DA2238DFF81FFBBFD77FF40FD28FECF" box="[729,762,232,258]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="9" pageNumber="10">99</figureCitation>
).
<collectingCountry id="FD8E7F98FF81FFBBFCA1FF40FC98FECF" box="[783,842,232,258]" name="India" pageId="9" pageNumber="10">Hind</collectingCountry>
wings completely reduced (brachypterous morph) or fully developed (macropterous morph). Legs moderately slender (see measurements and ratio TiL/TaL).
</paragraph>
<paragraph id="85263F08FF81FFBBFF6BFE90FD5FFE07" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">Abdomen approximately as wide as (brachypterous morph) or narrower than elytra (macropterous morph), widest at segments VI/VII; puncturation fine, sparser on posterior than on anterior tergites; surface with shallow, but distinct microsculpture; posterior margin of tergite VII with (macropterous morph) or without palisade fringe (brachypterous morph).</paragraph>
<paragraph id="85263F08FF81FFBBFF6BFE7FFF06FDD7" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">
ɗ: aedeagus slender and relatively small, with moderately sclerotised internal tube (
<figureCitation id="1DA2238DFF81FFBBFB68FE70FAE9FE3F" box="[1222,1339,472,498]" captionStart="FIGURES 1 – 6" captionStartId="8.[151,269,1038,1062]" captionTargetBox="[308,1253,197,1012]" captionTargetId="figure@8.[302,1285,197,1012]" captionTargetPageId="8" captionText="FIGURES 1 – 6. Vulda ottomana (Cameron): 1 — habitus; 2 — forebody; 3 — head in lateral view; 4 — antenna; 5 — aedeagus; 6 — internal structure of aedeagus. Scale bars: 1 – 3: 1.0 mm; 4: 0.5 mm; 5: 0.2 mm; 6: 0.1 mm." httpUri="https://zenodo.org/record/176739/files/figure.png" pageId="9" pageNumber="10">Figs. 5–6</figureCitation>
,
<figureCitation id="1DA2238DFF81FFBBFAE5FE70FA4AFE3F" box="[1355,1432,472,498]" captionStart="FIGURES 14 – 21" captionStartId="13.[151,269,1783,1807]" captionTargetBox="[309,1234,474,1756]" captionTargetId="figure@13.[302,1285,474,1759]" captionTargetPageId="13" captionText="FIGURES 14 – 21. Vulda ottomana (Cameron) (14 – 15; holotype of Xantholinus anatolicus Coiffait) and Xantholinus gridellii Coiffait (16 – 21; 20: paratype): 14, 20 – 21 — aedeagus; 15 — internal structures of aedeagus; 16 — habitus; 17 — forebody; 18 — male tergite VIII; 19 — male sternite VIII. Scale bars: 16 – 17: 1.0 mm; 18 – 21: 0.5 mm; 14: 0.2 mm; 15: 0.1 mm." httpUri="https://zenodo.org/record/176742/files/figure.png" pageId="9" pageNumber="10">14–15</figureCitation>
,
<figureCitation id="1DA2238DFF81FFBBFF39FDA8FF15FDD7" box="[151,199,512,538]" captionStart="FIGURES 94 – 100" captionStartId="39.[151,269,1619,1643]" captionTargetBox="[302,1256,346,1602]" captionTargetId="figure@39.[302,1266,346,1605]" captionTargetPageId="39" captionText="FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm." httpUri="https://zenodo.org/record/176757/files/figure.png" pageId="9" pageNumber="10">100</figureCitation>
).
</paragraph>
<paragraph id="85263F08FF81FFBBFF6BFD8FFCCFFCFF" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">
<emphasis id="B7EDE31AFF81FFBBFF6BFD8FFE6EFD8C" bold="true" box="[197,444,551,577]" pageId="9" pageNumber="10">Comparative notes.</emphasis>
Aside from
<taxonomicName id="4299448BFF81FFBBFDF6FD80FD34FD8C" box="[600,742,552,577]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF81FFBBFDF6FD80FD34FD8C" box="[600,742,552,577]" italics="true" pageId="9" pageNumber="10">V. ottomana</emphasis>
</taxonomicName>
, two species of
<taxonomicName id="4299448BFF81FFBBFC1FFD80FC21FD8C" box="[945,1011,552,577]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF81FFBBFC1FFD80FC21FD8C" box="[945,1011,552,577]" italics="true" pageId="9" pageNumber="10">Vulda</emphasis>
</taxonomicName>
have been described from
<collectingCountry id="FD8E7F98FF81FFBBFAEEFD80FA4BFD8F" box="[1344,1433,552,578]" name="Turkey" pageId="9" pageNumber="10">Turkey</collectingCountry>
:
<taxonomicName id="4299448BFF81FFBBFF39FDF8FD5BFDA7" authority="Bordoni, 1973" authorityName="Bordoni" authorityYear="1973" box="[151,649,592,618]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brignolii" subGenus="Typhlodes">
<emphasis id="B7EDE31AFF81FFBBFF39FDF8FF0BFDA4" box="[151,217,592,617]" italics="true" pageId="9" pageNumber="10">Vulda</emphasis>
(
<emphasis id="B7EDE31AFF81FFBBFF45FDF8FEB3FDA4" box="[235,353,592,617]" italics="true" pageId="9" pageNumber="10">Typhlodes</emphasis>
)
<emphasis id="B7EDE31AFF81FFBBFEDCFDF8FE0BFDA4" box="[370,473,592,617]" italics="true" pageId="9" pageNumber="10">brignolii</emphasis>
<bibRefCitation id="E10842F9FF81FFBBFE71FDF8FD5BFDA7" author="Bordoni" box="[479,649,592,618]" pageId="9" pageNumber="51" refString="Bordoni, A. (1973 a) Nuove specie di Xantholininae della Turchia e del Libano (Coleoptera, Staphylinidae). Fragmenta Entomologica, 9, 179 - 194." type="journal article" year="1973" yearSuffix="a">Bordoni, 1973</bibRefCitation>
</taxonomicName>
a (Amasya), and
<taxonomicName id="4299448BFF81FFBBFCFEFDF8FB47FDA7" authority="Bordoni, 1973" authorityName="Bordoni" authorityYear="1973" box="[848,1173,592,618]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vignai" subGenus="Typhlodes">
<emphasis id="B7EDE31AFF81FFBBFCFEFDF8FC8CFDA4" box="[848,862,592,617]" italics="true" pageId="9" pageNumber="10">V</emphasis>
. (
<emphasis id="B7EDE31AFF81FFBBFCD8FDF8FC56FDA4" box="[886,900,592,617]" italics="true" pageId="9" pageNumber="10">T</emphasis>
.)
<emphasis id="B7EDE31AFF81FFBBFC32FDF8FC34FDA4" box="[924,998,592,617]" italics="true" pageId="9" pageNumber="10">vignai</emphasis>
<bibRefCitation id="E10842F9FF81FFBBFC42FDF8FB47FDA7" author="Bordoni" box="[1004,1173,592,618]" pageId="9" pageNumber="51" refString="Bordoni, A. (1973 a) Nuove specie di Xantholininae della Turchia e del Libano (Coleoptera, Staphylinidae). Fragmenta Entomologica, 9, 179 - 194." type="journal article" year="1973" yearSuffix="a">Bordoni, 1973</bibRefCitation>
</taxonomicName>
a (Bolu). From the latter, whose description is based on a single female,
<taxonomicName id="4299448BFF81FFBBFD5CFDD0FCACFD5C" box="[754,894,632,657]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF81FFBBFD5CFDD0FCACFD5C" box="[754,894,632,657]" italics="true" pageId="9" pageNumber="10">V. ottomana</emphasis>
</taxonomicName>
is readily distinguished by much smaller size (
<taxonomicName id="4299448BFF81FFBBFF0FFD08FED5FD74" box="[161,263,672,697]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vignai">
<emphasis id="B7EDE31AFF81FFBBFF0FFD08FED5FD74" box="[161,263,672,697]" italics="true" pageId="9" pageNumber="10">V. vignai</emphasis>
</taxonomicName>
:
<quantity id="426192EDFF81FFBBFEB8FD08FEBCFD77" box="[278,366,672,698]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="10" unit="mm" value="13.0">13 mm</quantity>
), bicoloured body (in
<taxonomicName id="4299448BFF81FFBBFDDCFD08FD0AFD74" box="[626,728,672,697]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vignai">
<emphasis id="B7EDE31AFF81FFBBFDDCFD08FD0AFD74" box="[626,728,672,697]" italics="true" pageId="9" pageNumber="10">V. vignai</emphasis>
</taxonomicName>
uniformly pale brown), and larger eyes. From
<taxonomicName id="4299448BFF81FFBBFAAEFD08FA56FD74" box="[1280,1412,672,697]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B7EDE31AFF81FFBBFAAEFD08FA56FD74" box="[1280,1412,672,697]" italics="true" pageId="9" pageNumber="10">V. brignolii</emphasis>
</taxonomicName>
it is separated by the bicoloured body (in
<taxonomicName id="4299448BFF81FFBBFDD1FD60FCD5FD2C" box="[639,775,712,737]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B7EDE31AFF81FFBBFDD1FD60FCD5FD2C" box="[639,775,712,737]" italics="true" pageId="9" pageNumber="10">V. brignolii</emphasis>
</taxonomicName>
uniformly brown), a distinctly more slender head and pronotum, and a completely different morphology of the aedeagus. For illustrations of the habitus and male sexual characters of these species see
<bibRefCitation id="E10842F9FF81FFBBFDFDFCB0FCCBFCFF" author="Bordoni" box="[595,793,792,818]" pageId="9" pageNumber="51" refString="Bordoni, A. (1973 a) Nuove specie di Xantholininae della Turchia e del Libano (Coleoptera, Staphylinidae). Fragmenta Entomologica, 9, 179 - 194." type="journal article" year="1973" yearSuffix="a">Bordoni (1973a)</bibRefCitation>
.
</paragraph>
<paragraph id="85263F08FF81FFBBFF6BFC97FC02FBEF" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">
<emphasis id="B7EDE31AFF81FFBBFF6BFC97FE37FC94" bold="true" box="[197,485,831,857]" pageId="9" pageNumber="10">Subgeneric placement.</emphasis>
Based on the reductions of eye size, elytral length, hind wings, and palisade fringe at the posterior margin of the abdominal tergite VII, the brachypterous morph would have to be attributed to the subgenus
<taxonomicName id="4299448BFF81FFBBFE36FC38FD78FC67" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[408,682,912,938]" class="Insecta" family="Staphylinidae" genus="Typhlodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF81FFBBFE36FC38FDDEFC64" box="[408,524,912,937]" italics="true" pageId="9" pageNumber="10">Typhlodes</emphasis>
<bibRefCitation id="E10842F9FF81FFBBFDB6FC38FD78FC67" author="Sharp" box="[536,682,912,938]" lastPageId="9" lastPageNumber="54" pageId="9" pageNumber="53" refString="Sharp, D. S. (1873) Description of a new genus and species of blind Coleoptera from Italy. The Entomologist´s Monthly Magazine, 10, 1 - 2." type="journal article" year="1873">Sharp, 1873</bibRefCitation>
</taxonomicName>
, whereas the macropterous morph is currently in the subgenus
<taxonomicName id="4299448BFF81FFBBFF39FC10FF0BFC1C" box="[151,217,952,977]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF81FFBBFF39FC10FF0BFC1C" box="[151,217,952,977]" italics="true" pageId="9" pageNumber="10">Vulda</emphasis>
</taxonomicName>
. The fact that, based on the current subgeneric concept, one of the two morphs would have to be placed in
<taxonomicName id="4299448BFF81FFBBFF19FC48FF2BFC34" box="[183,249,992,1017]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF81FFBBFF19FC48FF2BFC34" box="[183,249,992,1017]" italics="true" pageId="9" pageNumber="10">Vulda</emphasis>
</taxonomicName>
and the other in
<taxonomicName id="4299448BFF81FFBBFE64FC48FDECFC34" box="[458,574,992,1017]" class="Insecta" family="Staphylinidae" genus="Typhlodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7EDE31AFF81FFBBFE64FC48FDECFC34" box="[458,574,992,1017]" italics="true" pageId="9" pageNumber="10">Typhlodes</emphasis>
</taxonomicName>
, suggests that this subgeneric system is highly artificial. Therefore, this and the followings species are attributed to neither of these subgenera.
</paragraph>
</subSubSection>
<subSubSection id="CD836C83FF81FFBBFF6BFB87FCC3FB0F" pageId="9" pageNumber="10" type="distribution">
<paragraph id="85263F08FF81FFBBFF6BFB87FCC3FB0F" blockId="9.[151,1437,152,1218]" pageId="9" pageNumber="10">
<emphasis id="B7EDE31AFF81FFBBFF6BFB87FDF7FB84" bold="true" box="[197,549,1071,1097]" pageId="9" pageNumber="10">Distribution and bionomics.</emphasis>
The species has become known from three localities in northwestern
<collectingCountry id="FD8E7F98FF81FFBBFAC7FB98FF10FBBF" name="Turkey" pageId="9" pageNumber="10">Turkey</collectingCountry>
, one in Bolu (
<typeStatus id="5A2281AAFF81FFBBFEC0FBF0FE70FBBF" box="[366,418,1112,1138]" pageId="9" pageNumber="10">type</typeStatus>
locality of
<taxonomicName id="4299448BFF81FFBBFD80FBF0FD69FBBC" box="[558,699,1112,1137]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B7EDE31AFF81FFBBFD80FBF0FD69FBBC" box="[558,699,1112,1137]" italics="true" pageId="9" pageNumber="10">V. anatolica</emphasis>
</taxonomicName>
) and two in Istanbul province (
<figureCitation id="1DA2238DFF81FFBBFB99FBF0FB42FBBF" box="[1079,1168,1112,1138]" captionStart="FIGURE 13" captionStartId="11.[151,255,1223,1247]" captionTargetBox="[230,1356,458,1199]" captionTargetId="figure@11.[230,1356,454,1200]" captionTargetPageId="11" captionText="FIGURE 13. Distribution of the genus Vulda in Turkey: V. ottomana (Cameron) (filled circles), V. vignai Bordoni (open circle), V. cangalica sp. n. (filled squares), and V. brignolii Bordoni (open square). Since the type locality of V. brignolii (&quot; Boraby gölü &quot;) was not identified (misspelling?), the respective square is placed in the centre of Amasya province." httpUri="https://zenodo.org/record/176741/files/figure.png" pageId="9" pageNumber="10">Fig. 13</figureCitation>
). The three specimens were collected in May, July, and August. The non-type specimen from Istanbul is apparently slightly teneral. The
<typeStatus id="5A2281AAFF81FFBBFF62FB00FEE1FB0F" box="[204,307,1192,1218]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4299448BFF81FFBBFEF7FB00FE37FB0C" box="[345,485,1192,1217]" class="Insecta" family="Staphylinidae" genus="Vulda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ottomana">
<emphasis id="B7EDE31AFF81FFBBFEF7FB00FE37FB0C" box="[345,485,1192,1217]" italics="true" pageId="9" pageNumber="10">V. ottomana</emphasis>
</taxonomicName>
was found under a stone.
</paragraph>
</subSubSection>
</treatment>
</document>
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