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<document id="C6BEFCB1DDEE82F94258E606DC47AD43" ID-CLB-Dataset="39184" ID-DOI="10.11646/zootaxa.4105.5.2" ID-GBIF-Dataset="60ca4dbd-002b-4ca6-999d-9174f9b02324" ID-ISSN="1175-5326" ID-Zenodo-Dep="255425" ID-ZooBank="52401DF9-2901-4AF6-BA17-696D39280B3C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1461736713249" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2016" docId="E338D6753519216EFF05FCD973576EC1" docLanguage="en" docName="zootaxa.4105.5.2.pdf.imf" docOrigin="Zootaxa 4105 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guyalna chlorogena" docType="treatment" docVersion="20" lastPageNumber="438" masterDocId="1F01AE0D351E2166FF92FF9876486E39" masterDocTitle="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" masterLastPageNumber="454" masterPageNumber="430" pageNumber="437" updateTime="1698625263794" updateUser="plazi">
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<mods:title id="5002FEFB85CD98C08291986A733BE2DF">The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species</mods:title>
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<mods:namePart id="59372A9056A83D202D9310557FDA6BBF">Sanborn, Allen F.</mods:namePart>
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<treatment id="E338D6753519216EFF05FCD973576EC1" ID-DOI="http://doi.org/10.5281/zenodo.4951870" ID-GBIF-Taxon="119651787" ID-Zenodo-Dep="4951870" LSID="urn:lsid:plazi:treatment:E338D6753519216EFF05FCD973576EC1" httpUri="http://treatment.plazi.org/id/E338D6753519216EFF05FCD973576EC1" lastPageId="8" lastPageNumber="438" pageId="7" pageNumber="437">
<subSubSection id="238B34E835192161FF05FCD974056D62" box="[151,589,833,859]" pageId="7" pageNumber="437" type="nomenclature">
<paragraph id="6B2E676335192161FF05FCD974056D62" blockId="7.[151,589,833,893]" box="[151,589,833,859]" pageId="7" pageNumber="437">
<heading id="3066D00F35192161FF05FCD974056D62" bold="true" box="[151,589,833,859]" fontSize="11" level="1" pageId="7" pageNumber="437" reason="1">
<taxonomicName id="AC911CE035192161FF05FCD974056D62" ID-CoL="3HN6R" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[151,589,833,859]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="chlorogena">
<emphasis id="59E5BB7135192161FF05FCD974056D62" bold="true" box="[151,589,833,859]" pageId="7" pageNumber="437">
<emphasis id="59E5BB7135192161FF05FCD977C76D62" bold="true" box="[151,399,833,859]" italics="true" pageId="7" pageNumber="437">Guyalna chlorogena</emphasis>
(
<bibRefCitation id="0F001A9235192161FE0CFCD9740D6D62" author="Walker, F." bookContentInfo="260 pp." box="[414,581,833,859]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" pageId="7" pageNumber="437" refId="ref14974" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum" type="book" year="1850">Walker, 1850</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="238B34E835192161FF05FCFC76A36D45" box="[151,235,868,893]" pageId="7" pageNumber="437" type="description">
<paragraph id="6B2E676335192161FF05FCFC76A36D45" blockId="7.[151,589,833,893]" box="[151,235,868,893]" pageId="7" pageNumber="437">
(
<figureCitation id="F3AA7BE635192161FF0DFCFC76AB6D45" box="[159,227,868,893]" captionStart="FIGURE 7" captionStartId="7.[151,250,1662,1684]" captionTargetBox="[151,1436,1145,1640]" captionTargetId="figure@7.[151,1436,1145,1641]" captionTargetPageId="7" captionText="FIGURE 7. Guyalna chlorogena (Walker, 1850). A, holotype male, unknown collection locality (BMNH). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255432/files/figure.png" pageId="7" pageNumber="437">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238B34E835192161FF05FC3474E26DC6" pageId="7" pageNumber="437" type="reference_group">
<paragraph id="6B2E676335192161FF05FC3474526DFB" blockId="7.[151,682,940,1023]" box="[151,538,940,962]" pageId="7" pageNumber="437">
<treatmentCitationGroup id="4B81404D35192161FF05FC3474526DFB" box="[151,538,940,962]" pageId="7" pageNumber="437">
<taxonomicName id="AC911CE035192161FF05FC34745E6DFB" ID-CoL="6HYYW" authority="Walker, 1850" authorityName="Walker" authorityPageNumber="88" authorityYear="1850" box="[151,534,940,962]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="chlorogena">
<emphasis id="59E5BB7135192161FF05FC34772D6DFB" box="[151,357,940,962]" italics="true" pageId="7" pageNumber="437">Fidicina chlorogena</emphasis>
<treatmentCitation id="EA30417235192161FEFEFC34745E6DFB" author="Walker" box="[364,534,940,962]" page="88" pageId="7" pageNumber="437" year="1850">
<bibRefCitation id="0F001A9235192161FEFEFC34745E6DFB" author="Walker, F." bookContentInfo="260 pp." box="[364,534,940,962]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" page="88" pageId="7" pageNumber="437" refId="ref14974" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum" type="book" year="1850">Walker 1850: 88</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="6B2E676335192161FF05FC5277B66DD9" blockId="7.[151,682,940,1023]" box="[151,510,970,992]" pageId="7" pageNumber="437">
<treatmentCitationGroup id="4B81404D35192161FF05FC5277B66DD9" box="[151,510,970,992]" pageId="7" pageNumber="437">
<taxonomicName id="AC911CE035192161FF05FC5277B26DD9" ID-CoL="6HYZG" authority="Walker 1858" authorityName="Walker" authorityPageNumber="15" authorityYear="1858" box="[151,506,970,992]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="basispes">
<emphasis id="59E5BB7135192161FF05FC5277016DD9" box="[151,329,970,992]" italics="true" pageId="7" pageNumber="437">Fidicina basispes</emphasis>
<treatmentCitation id="EA30417235192161FEDDFC5277B26DD9" author="Walker" box="[335,506,970,992]" page="15" pageId="7" pageNumber="437" year="1858">
<bibRefCitation id="0F001A9235192161FEDDFC5277B26DD9" author="Walker, F." bookContentInfo="307 pp." box="[335,506,970,992]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" page="15" pageId="7" pageNumber="437" refId="ref15005" refString="Walker, F. (1858) Supplement. List of the Specimens of Homopterous Insects in the Collection of the British Museum. British Museum Trustees, London, 307 pp." title="Supplement. List of the Specimens of Homopterous Insects in the Collection of the British Museum" type="book" year="1858">Walker 1858: 15</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B2E676335192161FF05FC7174E26DC6" blockId="7.[151,682,940,1023]" box="[151,682,1001,1023]" pageId="7" pageNumber="437">
<treatmentCitationGroup id="4B81404D35192161FF05FC7174E26DC6" box="[151,682,1001,1023]" pageId="7" pageNumber="437">
<taxonomicName id="AC911CE035192161FF05FC71772F6DC6" ID-CoL="3HN6R" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[151,359,1001,1023]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="chlorogena">
<emphasis id="59E5BB7135192161FF05FC71772F6DC6" box="[151,359,1001,1023]" italics="true" pageId="7" pageNumber="437">Guyalna chlorogena</emphasis>
</taxonomicName>
<treatmentCitation id="EA30417235192161FEFFFC7174EE6DC6" author="Boulard" box="[365,678,1001,1023]" page="14" pageId="7" pageNumber="437" year="1996">
<bibRefCitation id="0F001A9235192161FEFFFC7174EE6DC6" author="Boulard, M. &amp; Martinelli, N. M." box="[365,678,1001,1023]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" page="14" pageId="7" pageNumber="437" pagination="11 - 81" part="9" refId="ref12681" refString="Boulard, M. &amp; Martinelli, N. M. (1996) Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11 - 81." title="Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina" type="journal article" year="1996">Boulard &amp; Martinelli 1996: 14</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="238B34E835192161FF05FBB472116A7C" box="[151,1113,1068,1093]" pageId="7" pageNumber="437" type="materials_examined">
<paragraph id="6B2E676335192161FF05FBB472116A7C" blockId="7.[151,1113,1068,1093]" box="[151,1113,1068,1093]" pageId="7" pageNumber="437">
<emphasis id="59E5BB7135192161FF05FBB4777D6A7C" bold="true" box="[151,309,1068,1093]" pageId="7" pageNumber="437">
<typeStatus id="B42AD9C135192161FF05FBB4769A6A7C" box="[151,210,1068,1093]" pageId="7" pageNumber="437">Type</typeStatus>
locality.
</emphasis>
The
<typeStatus id="B42AD9C135192161FEFCFBB577D66A7C" box="[366,414,1069,1093]" pageId="7" pageNumber="437">type</typeStatus>
specimen was described from an unknown collection locality.
</paragraph>
</subSubSection>
<subSubSection id="238B34E83519216EFF55F94276826EED" lastPageId="8" lastPageNumber="438" pageId="7" pageNumber="437" type="discussion">
<paragraph id="6B2E67633519216EFF55F94276826EED" blockId="7.[151,1436,1754,2031]" lastBlockId="8.[151,1437,151,248]" lastPageId="8" lastPageNumber="438" pageId="7" pageNumber="437">
<emphasis id="59E5BB7135192161FF55F942777368CA" bold="true" box="[199,315,1754,1779]" pageId="7" pageNumber="437">Remarks.</emphasis>
<bibRefCitation id="0F001A9235192161FED0F94274CB68CA" author="Boulard, M. &amp; Martinelli, N. M." box="[322,643,1754,1779]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="7" pageNumber="437" pagination="11 - 81" part="9" refId="ref12681" refString="Boulard, M. &amp; Martinelli, N. M. (1996) Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11 - 81." title="Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina" type="journal article" year="1996">Boulard &amp; Martinelli (1996)</bibRefCitation>
transferred the species along with
<taxonomicName id="AC911CE035192161FB9EF94372F968CA" box="[1036,1201,1754,1779]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="bonaerensis">
<emphasis id="59E5BB7135192161FB9EF94372F968CA" box="[1036,1201,1754,1779]" italics="true" pageId="7" pageNumber="437">G. bonaerensis</emphasis>
</taxonomicName>
to the newly erected genus
<taxonomicName id="AC911CE035192161FF70F966770A692E" box="[226,322,1790,1815]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135192161FF70F966770A692E" box="[226,322,1790,1815]" italics="true" pageId="7" pageNumber="437">Guyalna</emphasis>
</taxonomicName>
representing the only two species of the genus at that time. The species is listed by
<bibRefCitation id="0F001A9235192161FAB0F96677106902" author="Boulard, M. &amp; Martinelli, N. M." firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="7" pageNumber="437" pagination="11 - 81" part="9" refId="ref12681" refString="Boulard, M. &amp; Martinelli, N. M. (1996) Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11 - 81." title="Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina" type="journal article" year="1996">Boulard &amp; Martinelli (1996)</bibRefCitation>
as the
<typeStatus id="B42AD9C135192161FE35F8BB779F6902" box="[423,471,1827,1851]" pageId="7" pageNumber="437">type</typeStatus>
species of the subtribe Guyalnina. However, according to Article 63 (ICZN 1999) the name bearing
<typeStatus id="B42AD9C135192161FEA4F8DF772E6966" box="[310,358,1863,1887]" pageId="7" pageNumber="437">type</typeStatus>
for a family-group taxon is a nominal genus as the family-group taxon is based upon the genus (Article 29) so
<taxonomicName id="AC911CE035192161FED9F8F277E469BA" box="[331,428,1898,1923]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135192161FED9F8F277E469BA" box="[331,428,1898,1923]" italics="true" pageId="7" pageNumber="437">Guyalna</emphasis>
</taxonomicName>
is the
<typeStatus id="B42AD9C135192161FE6DF8F3746769BA" box="[511,559,1899,1923]" pageId="7" pageNumber="437">type</typeStatus>
genus for the subtribe not
<taxonomicName id="AC911CE035192161FCFDF8F3724769BA" box="[879,1039,1898,1923]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="chlorogena">
<emphasis id="59E5BB7135192161FCFDF8F3724769BA" box="[879,1039,1898,1923]" italics="true" pageId="7" pageNumber="437">G. chlorogena</emphasis>
</taxonomicName>
as stated in
<bibRefCitation id="0F001A9235192161FB35F8F276AC699E" author="Boulard, M. &amp; Martinelli, N. M." firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="7" pageNumber="437" pagination="11 - 81" part="9" refId="ref12681" refString="Boulard, M. &amp; Martinelli, N. M. (1996) Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11 - 81." title="Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina" type="journal article" year="1996">Boulard &amp; Martinelli (1996)</bibRefCitation>
. The timbal cover expanding over the lateral portion of the timbal cavity (
<figureCitation id="F3AA7BE635192161FBB1F816722F699E" box="[1059,1127,1934,1959]" captionStart="FIGURE 7" captionStartId="7.[151,250,1662,1684]" captionTargetBox="[151,1436,1145,1640]" captionTargetId="figure@7.[151,1436,1145,1641]" captionTargetPageId="7" captionText="FIGURE 7. Guyalna chlorogena (Walker, 1850). A, holotype male, unknown collection locality (BMNH). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255432/files/figure.png" pageId="7" pageNumber="437">Fig. 7</figureCitation>
) supports the interpretation of timbal cover anatomy provided here as characteristic of the genus. The male opercula are larger than congeneric species with the posterior margin extending to form an approximate equilateral triangle rather than the usual right triangle shape of the opercula seen in other members of the genus. Body length
<quantity id="AC69CA863516216EFB83FF0072C46E96" box="[1041,1164,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="8" pageNumber="438" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">30–32 mm</quantity>
, fore wing length
<quantity id="AC69CA863516216EFAC4FF0F768E6EED" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="8" pageNumber="438" unit="mm" value="44.0" valueMax="45.0" valueMin="43.0">43–45 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF55FF4773576EC1" box="[199,1311,223,248]" pageId="8" pageNumber="438" type="distribution">
<paragraph id="6B2E67633516216EFF55FF4773576EC1" blockId="8.[151,1437,151,248]" box="[199,1311,223,248]" pageId="8" pageNumber="438">
<emphasis id="59E5BB713516216EFF55FF4777286EC1" bold="true" box="[199,352,223,248]" pageId="8" pageNumber="438">Distribution.</emphasis>
The species has been reported previously from
<collectingCountry id="138627F33516216EFCEAFF4775F46EC1" box="[888,956,223,248]" name="Brazil" pageId="8" pageNumber="438">Brazil</collectingCountry>
(
<bibRefCitation id="0F001A923516216EFC59FF47722E6EC1" author="Metcalf, Z. P." box="[971,1126,223,248]" firstAuthor="Metcalf" journalOrPublisher="North Carolina State College Contribution" pageId="8" pageNumber="438" pagination="1 - 585" part="1502" refId="ref13609" refString="Metcalf, Z. P. (1963) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i - vii, 1 - 585." title="General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae" type="journal article" year="1963">Metcalf 1963</bibRefCitation>
;
<bibRefCitation id="0F001A923516216EFBE0FF47735B6EC1" author="Sanborn, A. F." bookContentInfo="1001 pp." box="[1138,1299,223,248]" firstAuthor="Sanborn" journalOrPublisher="Elsevier / Academic Press, San Diego" pageId="8" pageNumber="438" refId="ref14140" refString="Sanborn, A. F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier / Academic Press, San Diego, 1001 pp." title="Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet" type="book" year="2013">Sanborn 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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