treatments-xml/data/85/3C/87/853C87FE3715FF9202BDAF8A01DAFA0C.xml

<document ID-GBIF-Dataset="f11667ef-fc68-45d9-a1a8-111a40389151" ID-GBIF-Taxon="114921287" checkinTime="1447846064264" checkinUser="donat" docAuthor="David Emmanuel M. GENERAL" docDate="2015" docId="853C87FE3715FF9202BDAF8A01DAFA0C" docLanguage="en" docName="29882.pdf" docOrigin="Myrmecological News 21" docSource="http://myrmecologicalnews.org/cms/index.php?option=com_content&amp;view=article&amp;id=1195" docTitle="Aretidris David Emmanuel M. GENERAL 2015, gen.n." docType="treatment" docVersion="7" lastPageId="3" lastPageNumber="132" masterDocId="7905FF863714FF91022AAD3F0321FF90" masterDocTitle="Aretidris, a new genus of ants (Hymenoptera: Formicidae: Myrmicinae) from the mountains of Luzon Island, Philippines" masterLastPageNumber="136" masterPageNumber="131" pageId="1" pageNumber="132" updateTime="1643511151075" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Aretidris, a new genus of ants (Hymenoptera: Formicidae: Myrmicinae) from the mountains of Luzon Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>David Emmanuel M. GENERAL</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Myrmecological News</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>21</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>136</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://myrmecologicalnews.org/cms/index.php?option=com_content&amp;view=article&amp;id=1195</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
</mods:mods>
<treatment ID-GBIF-Taxon="114921287" LSID="urn:lsid:plazi:treatment:853C87FE3715FF9202BDAF8A01DAFA0C" httpUri="http://treatment.plazi.org/id/853C87FE3715FF9202BDAF8A01DAFA0C" lastPageId="3" lastPageNumber="132" pageId="1" pageNumber="132">
<subSubSection pageId="1" pageNumber="132" type="nomenclature">
<paragraph box="[151,322,693,717]" pageId="1" pageNumber="132">
<emphasis bold="true" box="[151,322,693,717]" pageId="1" pageNumber="132">
<heading bold="true" box="[151,322,693,717]" fontSize="10" level="1" pageId="1" pageNumber="132" reason="6">
<emphasis bold="true" box="[151,247,693,717]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[151,247,693,717]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[254,322,693,717]" pageId="1" pageNumber="132" rank="genus">gen.n.</taxonomicNameLabel>
</heading>
</emphasis>
</paragraph>
<paragraph box="[151,597,733,756]" pageId="1" pageNumber="132">
<emphasis bold="true" box="[151,303,733,756]" pageId="1" pageNumber="132">
<typeStatus box="[151,209,733,756]" pageId="1" pageNumber="132">Type</typeStatus>
species:
</emphasis>
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[309,540,733,756]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="buenaventei" status="sp.n.">
<emphasis box="[309,540,733,756]" italics="true" pageId="1" pageNumber="132">Aretidris buenaventei</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[547,597,733,756]" pageId="1" pageNumber="132" rank="species">sp.n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="132" type="diagnosis">
<paragraph pageId="1" pageNumber="132">
<emphasis bold="true" box="[189,420,762,786]" pageId="1" pageNumber="132">Diagnosis of worker:</emphasis>
Palp formula: 
<number box="[574,616,763,786]" pageId="1" pageNumber="132" value="3.2">3, 2</number>
. Antennae 
<number box="[733,760,763,786]" pageId="1" pageNumber="132" value="12.0">12</number>
- segmented (including antennal scape) with distinct 
<number box="[700,712,792,815]" pageId="1" pageNumber="132" value="3.0">3</number>
-segmented club. Spur formula: 0, 0. Antennal scrobe absent. In full face view, gena bounded laterally by a longitudinal ridge that runs from dorsal edge of eye to mandibular insertion and is noticeably thickened anteriorly. In full face view, labrum visible below the anterior clypeal margin, even when mouthparts are fully withdrawn into the head. Mandibles with a long narrow basal stem such that a large gap exists between the basal margin of the mandible and the anterior clypeal margin; masticatory margin with a row of 
<number box="[181,209,1085,1108]" pageId="1" pageNumber="132" value="14.0">14</number>
small teeth or denticles. In lateral view, promesonotum strongly convex. Entire length of anterior margin of katepisternum produced into a thickened flange, obscuring the posterior margin of procoxa. Metanotal groove strongly impressed. Propodeum convex with a denticle at the junction between the propodeal dorsum and declivity. Petiole sessile. Protuberance or tooth present above and anterior to petiolar spiracle. Petiole slightly larger than postpetiole. Sting long and functional.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="132" type="description">
<paragraph pageId="1" pageNumber="132">
<emphasis bold="true" box="[189,449,1349,1372]" pageId="1" pageNumber="132">Description of worker:</emphasis>
Palp formula: 
<number box="[610,654,1349,1372]" pageId="1" pageNumber="132" value="3.2">3, 2</number>
. Antennae 
<number box="[151,175,1379,1402]" pageId="1" pageNumber="132" value="12.0">12</number>
-segmented (including antennal scape) with distinct 
<number box="[744,758,1379,1402]" pageId="1" pageNumber="132" value="3.0">3</number>
- segmented club. Antennal scrobe absent. Antennal scape weakly incrassate, reaching beyond the posterior margin of head by at least the thickness of the distal part of scape. In full face view (
<abbreviationData pageId="1" pageNumber="132">
<abbreviationRange box="[353,475,1496,1519]" pageId="1" pageNumber="132" prefix="Figs." type="Arabic" value="4" valueMax="10" valueMin="4" values="4,10">
<figureCitation box="[353,432,1496,1519]" captionStart="Figs. 1 ­ 6" captionStartId="2.[151,200,1571,1594]" captionText="Figs. 1 ­ 6: Aretidris buenaventei sp. n. (holotype). (1) Lateral view of body; (2) head in full­face view; (3) lateral closeup of propodeum and waist segments; (4) close­up of anterior head and mandibles, (5) dorsal view of body; (6) labels." pageId="1" pageNumber="132">
Figs. 
<number box="[416,475,1496,1519]" pageId="1" pageNumber="132" value="4.1">
4, 
<figureCitation box="[445,475,1496,1519]" captionStart="Figs. 7 ­ 12" captionStartId="4.[151,200,1704,1727]" captionText="Figs. 7 ­ 12: Aretidris clousei sp. n. (holotype). (7) Lateral view of body; (8) head in full­face view; (9) lateral close­up of propodeum and waist segments; (10) close­up of anterior head and mandibles; (11) dorsal view of body; (12) labels." pageId="1" pageNumber="132">10</figureCitation>
</number>
</figureCitation>
</abbreviationRange>
), gena with a longitudinal ridge that runs from dorsal edge of eye to mandibular insertion and is noticeably thickened anteriorly. Gena reticulate. Frontal lobes longitudinally striate. Frontal carina short, ending at about mid-length of eye. Clypeus bicarinate; median part narrowly inserted between frontal lobes; anterior margin simple, almost straight, and without an isolated median seta. Labrum visible below anterior clypeal margin, even when mouthparts are fully withdrawn into the head. Mandible smooth, with a long narrowed base such that a large gap exists between the basal margin of the mandible and the anterior clypeal margin, and with 
<number box="[741,767,1819,1842]" pageId="1" pageNumber="132" value="14.0">14</number>
small teeth or denticles; apical and subapical teeth largest and the rest diminishing in size posteriad. Posterior margin of head strongly convex. Lateral margin of head converging anteriorly. Eyes located laterally on the head just anterior of the mid-length, and convex, breaking the lateral margin of head. In lateral view, mandibles bent downward 
<table box="[820,1437,176,2018]" pageId="1" pageNumber="132">
<tr box="[820,1437,176,228]" pageId="1" pageNumber="132">and</tr>
</table>
<td box="[1096,1214,176,228]" pageId="1" pageNumber="132">convex</td>
, 
<td box="[1226,1329,176,228]" pageId="1" pageNumber="132">separated</td>
<td box="[1332,1383,176,228]" pageId="1" pageNumber="132">by</td>
a 
<tr box="[820,1437,235,316]" pageId="1" pageNumber="132">
<td box="[820,918,235,316]" pageId="1" pageNumber="132">
<td box="[1389,1436,176,228]" pageId="1" pageNumber="132">prominently</td>
<td box="[927,1070,235,316]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
impressed 
<td box="[1029,1246,235,316]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
metanotal groove
<td box="[1226,1344,235,316]" pageId="1" pageNumber="132">
. Entire 
<td box="[1335,1404,235,287]" pageId="1" pageNumber="132">
length 
<td box="[1372,1437,235,316]" pageId="1" pageNumber="132">
of anterior 
<td box="[914,990,264,316]" pageId="1" pageNumber="132">margin of katepisternum produced into a thickened flange,</td>
obscuring the posterior margin of procoxa.
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,323,404]" pageId="1" pageNumber="132">
<td box="[820,955,323,404]" pageId="1" pageNumber="132">
Denticle 
<td box="[921,1000,323,375]" pageId="1" pageNumber="132">
present 
<td box="[1009,1095,323,404]" pageId="1" pageNumber="132">
where 
<td box="[1084,1198,323,404]" pageId="1" pageNumber="132">
propodeal 
<td box="[1204,1355,323,404]" pageId="1" pageNumber="132">
dorsum meets 
<td box="[1363,1437,323,404]" pageId="1" pageNumber="132">
propodeal declivity. Propodeal spiracle 
<td box="[1183,1267,352,375]" pageId="1" pageNumber="132">circular</td>
, located 
<td box="[1336,1393,352,404]" pageId="1" pageNumber="132">
at least one diameter 
<td box="[961,1011,381,404]" pageId="1" pageNumber="132">from</td>
edge of declivity. Metapleural gland
</td>
<tr box="[820,1437,411,492]" pageId="1" pageNumber="132">
<td box="[820,948,411,492]" pageId="1" pageNumber="132">
orifice circular
<td box="[920,974,411,463]" pageId="1" pageNumber="132">
, 
<td box="[959,1043,411,492]" pageId="1" pageNumber="132">
directed 
<td box="[1051,1230,411,492]" pageId="1" pageNumber="132">
posteriorly (best 
<td box="[1211,1380,411,492]" pageId="1" pageNumber="132">
observed 
<td box="[1319,1350,411,463]" pageId="1" pageNumber="132">
in 
<td box="[1358,1436,411,492]" pageId="1" pageNumber="132">
oblique posterior view), without guard hairs, located at the apex of the propodeal lobe; 
<td box="[1038,1098,469,492]" pageId="1" pageNumber="132">gland</td>
bulla small
<td box="[1230,1234,469,492]" pageId="1" pageNumber="132">.</td>
Longitudinal ruga
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,499,580]" pageId="1" pageNumber="132">
<td box="[820,933,499,580]" pageId="1" pageNumber="132">
present 
<td box="[906,972,499,580]" pageId="1" pageNumber="132">
above 
<td box="[965,1147,499,580]" pageId="1" pageNumber="132">
metapleural 
<td box="[1113,1240,499,580]" pageId="1" pageNumber="132">
gland 
<td box="[1186,1270,499,580]" pageId="1" pageNumber="132">
orifice
<td box="[1247,1349,499,580]" pageId="1" pageNumber="132">
. Petiole 
<td box="[1357,1436,499,580]" pageId="1" pageNumber="132">
sessile, slightly larger 
<td box="[975,1021,528,551]" pageId="1" pageNumber="132">than</td>
postpetiole; spiracle below 
<td box="[1319,1407,528,580]" pageId="1" pageNumber="132">base of anterior face; a protuberance or tooth above and anterior</td>
to
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,587,668]" pageId="1" pageNumber="132">
<td box="[820,932,587,668]" pageId="1" pageNumber="132">
petiolar 
<td box="[909,995,587,668]" pageId="1" pageNumber="132">
spiracle
<td box="[995,999,587,610]" pageId="1" pageNumber="132">;</td>
node 
<td box="[1065,1253,587,668]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
rounded, junction 
<td box="[1222,1283,587,639]" pageId="1" pageNumber="132">
of convex 
<td box="[1374,1437,587,668]" pageId="1" pageNumber="132">
petiolar dorsum with anterior and posterior faces 
<td box="[1283,1384,616,668]" pageId="1" pageNumber="132">
indistinct; subpetiolar process absent
<td box="[1068,1072,645,668]" pageId="1" pageNumber="132">.</td>
Postpetiole slightly flattened
</td>
<tr box="[820,1437,675,756]" pageId="1" pageNumber="132">
<td box="[820,918,675,756]" pageId="1" pageNumber="132">
dorsally, 
<td box="[887,984,675,756]" pageId="1" pageNumber="132">
dorsum 
<td box="[970,1064,675,756]" pageId="1" pageNumber="132">
forming 
<td box="[1072,1250,675,756]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
rounded angles 
<td box="[1247,1339,675,756]" pageId="1" pageNumber="132">
with 
<td box="[1295,1389,675,756]" pageId="1" pageNumber="132">
anterior 
<td box="[1386,1436,675,756]" pageId="1" pageNumber="132">
and posterior faces; 
<td box="[996,1163,704,756]" pageId="1" pageNumber="132">subpostpetiolar process present as a low rounded lobe in anterior half of</td>
sternite, its anterior 
<tr box="[820,1437,763,844]" pageId="1" pageNumber="132">
<td box="[820,946,763,844]" pageId="1" pageNumber="132">
margin broadly 
<td box="[953,1074,763,844]" pageId="1" pageNumber="132">
emarginate 
<td box="[1088,1208,763,844]" pageId="1" pageNumber="132">
in ventral 
<td box="[1193,1250,763,815]" pageId="1" pageNumber="132">
view
<td box="[1245,1316,763,844]" pageId="1" pageNumber="132">
. First 
<td box="[1312,1436,763,844]" pageId="1" pageNumber="132">
gastral 
<td box="[1389,1436,763,844]" pageId="1" pageNumber="132">
tergite large
<td box="[923,993,792,815]" pageId="1" pageNumber="132">, about</td>
half 
<td box="[1036,1106,792,844]" pageId="1" pageNumber="132">the total length of gaster. Mesotibial and metatibial spurs absent</td>
. Pilosity present as mostly long
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,851,932]" pageId="1" pageNumber="132">
<td box="[820,903,851,932]" pageId="1" pageNumber="132">
erect or 
<td box="[902,1028,851,932]" pageId="1" pageNumber="132">
suberect hairs
<td box="[1036,1116,851,932]" pageId="1" pageNumber="132">
; 
<td box="[1073,1279,851,932]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
shorter erect hairs 
<td box="[1273,1345,851,932]" pageId="1" pageNumber="132">
on 
<td box="[1307,1382,851,932]" pageId="1" pageNumber="132">
coxae; 
<td box="[1380,1436,851,932]" pageId="1" pageNumber="132">
erect hairs on gaster 
<td box="[979,1047,880,903]" pageId="1" pageNumber="132">sparse</td>
. Color reddish-orange to brown, gaster and appendages lighter; antennal club lighter in color
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,939,1020]" pageId="1" pageNumber="132">
<td box="[820,932,939,991]" pageId="1" pageNumber="132">
than 
<td box="[858,982,939,1020]" pageId="1" pageNumber="132">
rest of 
<td box="[947,1058,939,1020]" pageId="1" pageNumber="132">
flagellum 
<td box="[1050,1120,939,1020]" pageId="1" pageNumber="132">
and 
<td box="[1101,1241,939,1020]" pageId="1" pageNumber="132">
scape.
<paragraph box="[858,1245,968,991]" pageId="1" pageNumber="132">
<emphasis bold="true" box="[858,1125,968,991]" pageId="1" pageNumber="132">Queen and male castes:</emphasis>
Unknown.
</paragraph>
<paragraph pageId="1" pageNumber="132">
<emphasis bold="true" box="[858,986,997,1020]" pageId="1" pageNumber="132">Bionomics:</emphasis>
These ants nest in 
<td box="[1197,1260,997,1020]" pageId="1" pageNumber="132">rotten</td>
<td box="[1268,1313,997,1020]" pageId="1" pageNumber="132">logs</td>
<td box="[1321,1359,997,1020]" pageId="1" pageNumber="132">and</td>
<td box="[1367,1436,997,1020]" pageId="1" pageNumber="132">forage</td>
<tr box="[820,1437,1027,1196]" pageId="1" pageNumber="132">
<td box="[820,948,1027,1196]" pageId="1" pageNumber="132">
<td box="[858,1004,1027,1167]" pageId="1" pageNumber="132">
individually 
<td box="[944,1052,1027,1167]" pageId="1" pageNumber="132">
in the 
<td box="[1021,1101,1027,1196]" pageId="1" pageNumber="132">
leaf 
<td box="[1070,1229,1027,1196]" pageId="1" pageNumber="132">
litter during 
<td box="[1201,1345,1027,1196]" pageId="1" pageNumber="132">
the day. 
<td box="[1311,1436,1027,1167]" pageId="1" pageNumber="132">
They 
<td box="[1356,1436,1027,1167]" pageId="1" pageNumber="132">
occur at elevations greater than 
<quantity box="[1061,1128,1056,1079]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="132" unit="m" value="900.0">900 m</quantity>
a.s.l. on 
<td box="[1221,1253,1056,1079]" pageId="1" pageNumber="132">the</td>
island of Luzon.
<subSubSection pageId="1" pageNumber="132" type="etymology">
<paragraph pageId="1" pageNumber="132">
<emphasis bold="true" box="[858,987,1085,1108]" pageId="1" pageNumber="132">Etymology:</emphasis>
<emphasis box="[996,1052,1085,1108]" italics="true" pageId="1" pageNumber="132">Areté</emphasis>
(ἀρετή, Ancient Greek, excellence) + 
<emphasis box="[844,892,1115,1138]" italics="true" pageId="1" pageNumber="132">idris</emphasis>
(ἴδρισ, Greek, ant) (WHEELER 
<number box="[1233,1291,1115,1138]" pageId="1" pageNumber="132" value="1956.0">1956</number>
). Coincidentally, 
<emphasis box="[874,929,1144,1167]" italics="true" pageId="1" pageNumber="132">arête</emphasis>
(French, ridge) also fits the fact that 
<emphasis box="[1330,1436,1144,1167]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="clousei" status="sp.n.">A. clousei</taxonomicName>
</emphasis>
<taxonomicNameLabel rank="species">sp.n.</taxonomicNameLabel>
was collected on a low ridge bordering a mountain 
<tr box="[820,1437,1203,1284]" pageId="1" pageNumber="132">
<td box="[820,915,1203,1284]" pageId="1" pageNumber="132">
lake.
<paragraph pageId="1" pageNumber="132">
<td box="[858,1001,1232,1284]" pageId="1" pageNumber="132">
<emphasis bold="true" box="[858,1073,1232,1255]" pageId="1" pageNumber="132">
Comparative 
<td box="[1008,1073,1232,1255]" pageId="1" pageNumber="132">notes:</td>
</emphasis>
<td box="[1080,1262,1232,1284]" pageId="1" pageNumber="132">
<emphasis box="[1080,1171,1232,1255]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[1080,1171,1232,1255]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
is 
<td box="[1200,1269,1232,1255]" pageId="1" pageNumber="132">unique</td>
<td box="[1276,1346,1232,1255]" pageId="1" pageNumber="132">among</td>
<td box="[1353,1384,1232,1255]" pageId="1" pageNumber="132">ant</td>
<td box="[1389,1437,1232,1284]" pageId="1" pageNumber="132">
genera, with a 
<td box="[944,1077,1261,1284]" pageId="1" pageNumber="132">combination</td>
of narrow-based mandibles with
</td>
</td>
</td>
<tr box="[820,1437,1291,1372]" pageId="1" pageNumber="132">
<td box="[820,958,1291,1372]" pageId="1" pageNumber="132">
a denticulate masticatory 
<td box="[1091,1191,1291,1372]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
margin, 
<td box="[1197,1319,1291,1372]" pageId="1" pageNumber="132">
gena with 
<td box="[1313,1436,1291,1372]" pageId="1" pageNumber="132">
a ridge 
<td box="[1391,1436,1291,1372]" pageId="1" pageNumber="132">
running from 
<td box="[921,972,1320,1372]" pageId="1" pageNumber="132">
the 
<td box="[972,1052,1320,1372]" pageId="1" pageNumber="132">
dorsal 
<td box="[1058,1108,1320,1372]" pageId="1" pageNumber="132">
edge of the eye to the mandibular insertion, palp formula 
<number box="[1059,1071,1349,1372]" pageId="1" pageNumber="132" value="3.0">3</number>
: 
<number box="[1091,1105,1349,1372]" pageId="1" pageNumber="132" value="2.0">2</number>
,
</td>
</td>
</td>
convex propodeum armed with
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,1379,1460]" pageId="1" pageNumber="132">
<td box="[820,923,1379,1460]" pageId="1" pageNumber="132">
a denticle
<td box="[897,976,1379,1460]" pageId="1" pageNumber="132">
, 
<td box="[934,1057,1379,1460]" pageId="1" pageNumber="132">
metapleural gland 
<td box="[1123,1210,1379,1460]" pageId="1" pageNumber="132">
orifice 
<td box="[1205,1335,1379,1460]" pageId="1" pageNumber="132">
at the apex 
<td box="[1325,1386,1379,1460]" pageId="1" pageNumber="132">
of the 
<td box="[1382,1437,1379,1460]" pageId="1" pageNumber="132">
propodeal lobe, and a 
<td box="[1018,1156,1408,1460]" pageId="1" pageNumber="132">
protuberance over 
<td box="[1216,1248,1408,1431]" pageId="1" pageNumber="132">the</td>
petiolar spiracle.
<subSubSection lastPageId="3" lastPageNumber="134" pageId="1" pageNumber="132" type="diagnosis">
<paragraph pageId="1" pageNumber="132">
This genus shares some similar characters, e.g., 
<tr box="[820,1437,1467,1548]" pageId="1" pageNumber="132">
<td box="[820,931,1467,1548]" pageId="1" pageNumber="132">
bicarinate 
<td box="[890,970,1467,1548]" pageId="1" pageNumber="132">
median 
<td box="[933,1064,1467,1548]" pageId="1" pageNumber="132">
clypeus
<td box="[1045,1273,1467,1548]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
<td box="[1036,1072,1467,1519]" pageId="1" pageNumber="132">
, with 
<taxonomicName authorityName="Emery" authorityYear="1897" box="[1132,1273,1467,1490]" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[1132,1273,1467,1490]" italics="true" pageId="1" pageNumber="132">Lordomyrma</emphasis>
</taxonomicName>
<td box="[1219,1360,1467,1548]" pageId="1" pageNumber="132">
EMERY
<td box="[1338,1407,1467,1548]" pageId="1" pageNumber="132">
, 
<td box="[1375,1437,1467,1548]" pageId="1" pageNumber="132">
<number box="[1375,1433,1467,1490]" pageId="1" pageNumber="132" value="1897.0">1897</number>
. However, 
<emphasis box="[933,1029,1496,1519]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[933,1029,1496,1519]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
has narrow-based mandibles, a genal ridge, and a convex propodeum, all characters absent in
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,1555,1636]" pageId="1" pageNumber="132">
<td box="[820,956,1555,1636]" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1897" box="[820,956,1555,1578]" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[820,956,1555,1578]" italics="true" pageId="1" pageNumber="132">Lordomyrma</emphasis>
</taxonomicName>
as 
<td box="[992,1104,1555,1636]" pageId="1" pageNumber="132">
currently 
<td box="[1085,1267,1555,1636]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
defined. 
<td box="[1183,1324,1555,1607]" pageId="1" pageNumber="132">
<emphasis box="[1183,1318,1555,1578]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1897" box="[1183,1318,1555,1578]" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Lordomyrma</taxonomicName>
</emphasis>
, moreover
<td box="[1415,1436,1555,1636]" pageId="1" pageNumber="132">
, has an 
<td box="[898,990,1584,1607]" pageId="1" pageNumber="132">antennal</td>
scrobe (present but poorly 
<td box="[1276,1429,1584,1636]" pageId="1" pageNumber="132">
developed in 
<taxonomicName authorityName="Emery" authorityYear="1897" box="[820,945,1613,1636]" class="Insecta" family="Formicidae" genus="Lordomyrma" genus.outerRound="1" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="furcifera">
<emphasis box="[820,945,1613,1636]" italics="true" pageId="1" pageNumber="132">L. furcifera</emphasis>
</taxonomicName>
<td box="[954,1032,1613,1636]" pageId="1" pageNumber="132">EMERY</td>
, 
<number box="[1048,1104,1613,1636]" pageId="1" pageNumber="132" value="1897.0">1897</number>
, 
<emphasis box="[1118,1267,1613,1636]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Donisthorpe" authorityYear="1940" box="[1118,1267,1613,1636]" class="Insecta" family="Formicidae" genus="Lordomyrma" genus.outerRound="1" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="infundibuli">L. infundibuli</taxonomicName>
</emphasis>
DONISTHORPE
</td>
,
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,1643,1724]" pageId="1" pageNumber="132">
<td box="[820,956,1643,1724]" pageId="1" pageNumber="132">
<number box="[820,874,1643,1666]" pageId="1" pageNumber="132" value="1940.0">1940</number>
<td box="[869,978,1643,1695]" pageId="1" pageNumber="132">
, and 
<taxonomicName authority="MENOZZI, 1923" authorityName="MENOZZI" authorityYear="1923" box="[930,1047,1643,1666]" class="Insecta" family="Formicidae" genus="Lordomyrma" genus.outerRound="1" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="crawleyi">
<emphasis box="[930,1047,1643,1666]" italics="true" pageId="1" pageNumber="132">L. 
<td box="[958,1058,1643,1724]" pageId="1" pageNumber="132">
crawleyi 
<td box="[1054,1220,1643,1724]" pageId="1" pageNumber="132">
MENOZZI, 
<number box="[1164,1219,1643,1666]" pageId="1" pageNumber="132" value="1923.0">1923</number>
<td box="[1207,1330,1643,1724]" pageId="1" pageNumber="132">
<td box="[1219,1232,1643,1666]" pageId="1" pageNumber="132">),</td>
long frontal 
<td box="[1367,1436,1643,1724]" pageId="1" pageNumber="132">
carina, and propodeal spines
<td box="[1028,1100,1672,1724]" pageId="1" pageNumber="132">
, all characters absent in 
<emphasis box="[1339,1436,1672,1695]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
(see 
<abbreviationData pageId="1" pageNumber="132">
TAYLOR 
<number box="[963,1083,1701,1724]" pageId="1" pageNumber="132" value="2009.2012">2009, 2012</number>
). LUCKY &amp; SARNAT (
<td box="[1308,1369,1701,1724]" pageId="1" pageNumber="132">
<number box="[1308,1362,1701,1724]" pageId="1" pageNumber="132" value="2010.0">2010</number>
)
</td>
found 
<tr box="[820,1437,1731,1812]" pageId="1" pageNumber="132">
<td box="[820,956,1731,1812]" pageId="1" pageNumber="132">
that 
<emphasis box="[867,959,1731,1754]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
<td box="[965,1046,1731,1812]" pageId="1" pageNumber="132">
is sister 
<td box="[1036,1103,1731,1812]" pageId="1" pageNumber="132">
to 
<td box="[1078,1251,1731,1812]" pageId="1" pageNumber="132">
<emphasis box="[1082,1218,1731,1754]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1897" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Lordomyrma</taxonomicName>
</emphasis>
<td box="[1218,1224,1731,1754]" pageId="1" pageNumber="132">,</td>
<td box="[1232,1341,1731,1812]" pageId="1" pageNumber="132">
essentially 
<td box="[1349,1436,1731,1812]" pageId="1" pageNumber="132">
disproving Bolton's hunch about the identity of &quot; 
<td box="[1293,1435,1760,1812]" pageId="1" pageNumber="132">
<emphasis box="[1293,1435,1760,1783]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1897" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Lordomyrma</taxonomicName>
</emphasis>
spPH01&quot;
<td box="[919,1000,1789,1812]" pageId="1" pageNumber="132">. WARD</td>
&amp; al. (
<number box="[1078,1134,1789,1812]" pageId="1" pageNumber="132" value="2015.0">2015</number>
) improved the resolution of
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,1819,1900]" pageId="1" pageNumber="132">
<td box="[820,905,1819,1900]" pageId="1" pageNumber="132">
the 
<td box="[860,1019,1819,1900]" pageId="1" pageNumber="132">
phylogenetic 
<td box="[1003,1132,1819,1900]" pageId="1" pageNumber="132">
relationship 
<td box="[1082,1253,1819,1900]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">
of these 
<td box="[1233,1352,1819,1900]" pageId="1" pageNumber="132">
two genera 
<td box="[1328,1400,1819,1900]" pageId="1" pageNumber="132">
and 
<td box="[1400,1436,1819,1900]" pageId="1" pageNumber="132">
recovered them 
<td box="[974,995,1848,1871]" pageId="1" pageNumber="132">in</td>
separate subclades within the newly expanded myrmicine tribe 
<taxonomicName authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
FOREL, 
<number box="[1344,1400,1877,1900]" pageId="1" pageNumber="132" value="1893.0">1893</number>
.
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</abbreviationData>
</td>
</td>
</td>
</td>
</td>
</emphasis>
</taxonomicName>
</td>
</td>
</tr>
</paragraph>
<paragraph pageId="1" pageNumber="132">
<tr box="[820,1437,1907,1988]" pageId="1" pageNumber="132">
<td box="[820,950,1907,1988]" pageId="1" pageNumber="132">
<emphasis box="[858,950,1907,1930]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[858,950,1907,1930]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
<td box="[956,1060,1907,1988]" pageId="1" pageNumber="132">
would 
<td box="[1044,1238,1907,1988]" pageId="1" pageNumber="132">
<td box="[1031,1080,1907,1930]" pageId="1" pageNumber="132">have</td>
been included 
<td box="[1242,1379,1907,1988]" pageId="1" pageNumber="132">
in the 
<td box="[1309,1406,1907,1959]" pageId="1" pageNumber="132">
old 
<td box="[1378,1436,1907,1988]" pageId="1" pageNumber="132">
sense of the 
<td box="[861,979,1936,1988]" pageId="1" pageNumber="132">
myrmicine tribe 
<taxonomicName authorityName="Ashmead" authorityYear="1905" box="[1044,1176,1936,1959]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Stenammini">Stenammini</taxonomicName>
<td box="[1183,1294,1936,1959]" pageId="1" pageNumber="132">ASHMEAD</td>
, 
<number box="[1309,1364,1936,1959]" pageId="1" pageNumber="132" value="1905.0">1905</number>
(BOL- 
<collectionCode box="[820,865,1969,1987]" pageId="1" pageNumber="132">TON</collectionCode>
<number box="[875,934,1965,1988]" pageId="1" pageNumber="132" value="2003.0">2003</number>
).
</td>
Although BOLTON (
<number box="[1181,1238,1965,1988]" pageId="1" pageNumber="132" value="2003.0">2003</number>
) established well
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<tr box="[820,1437,1995,2018]" pageId="1" pageNumber="132">
<td box="[820,853,1995,2018]" pageId="1" pageNumber="132">the</td>
<td box="[860,1017,1995,2018]" colspan="2" colspanRight="1" pageId="1" pageNumber="132">morphological</td>
<td box="[1024,1126,1995,2018]" pageId="1" pageNumber="132">diagnosis</td>
<td box="[1132,1155,1995,2018]" pageId="1" pageNumber="132">of</td>
<td box="[1162,1294,1995,2018]" pageId="1" pageNumber="132">
<taxonomicName authorityName="Ashmead" authorityYear="1905" box="[1162,1294,1995,2018]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Stenammini">Stenammini</taxonomicName>
</td>
<td box="[1294,1299,1995,2018]" pageId="1" pageNumber="132">,</td>
<td box="[1305,1376,1995,2018]" pageId="1" pageNumber="132">WARD</td>
<td box="[1383,1436,1995,2018]" pageId="1" pageNumber="132">&amp; al.</td>
</tr>
</paragraph>
<caption pageId="1" pageNumber="132">
<paragraph pageId="1" pageNumber="132">
Figs. 1 - 6: 
<emphasis box="[275,506,1571,1594]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="buenaventei" status="sp.n.">Aretidris buenaventei</taxonomicName>
</emphasis>
<taxonomicNameLabel rank="species">sp.n.</taxonomicNameLabel>
(holotype). (1) Lateral view of body; (2) head in full-face view; (3) lateral closeup of propodeum and waist segments; (4) close-up of anterior head and mandibles, (5) dorsal view of body; (6) labels.
</paragraph>
</caption>
<paragraph pageId="1" pageNumber="132">
(
<number box="[159,216,1656,1679]" pageId="1" pageNumber="132" value="2015.0">2015</number>
) found many erstwhile stenammine genera distributed throughout the 
<taxonomicName authorityName="Forel" authorityYear="1893" box="[384,555,1685,1708]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
clade. WARD &amp; al. (
<number box="[159,217,1715,1738]" pageId="1" pageNumber="132" value="2015.0">2015</number>
) thus transferred fifteen extant genera out of the 
<taxonomicName authorityName="Ashmead" authorityYear="1905" box="[151,277,1744,1767]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Stenammini">Stenammini</taxonomicName>
into the 
<taxonomicName authorityName="Forel" authorityYear="1893" box="[371,542,1744,1767]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
. Now included in the 
<taxonomicName authorityName="Forel" authorityYear="1893" box="[151,324,1773,1796]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
are all the genera under consideration in this paper, namely, 
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[374,472,1803,1826]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">
<emphasis box="[374,472,1803,1826]" italics="true" pageId="1" pageNumber="132">Aretidris</emphasis>
</taxonomicName>
(= Myrmicine Genus # 
<number box="[739,767,1803,1826]" pageId="1" pageNumber="132" value="26.0">26</number>
PH02 sensu WARD &amp; al. (
<number box="[439,498,1832,1855]" pageId="1" pageNumber="132" value="2015.0">2015</number>
)), 
<emphasis box="[528,671,1832,1855]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1897" box="[528,671,1832,1855]" class="Insecta" family="Formicidae" genus="Lordomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Lordomyrma</taxonomicName>
</emphasis>
, 
<taxonomicName authorityName="Mayr" authorityYear="1865" class="Insecta" family="Formicidae" genus="Vollenhovia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="1" pageNumber="132">Vollenhovia</emphasis>
</taxonomicName>
MAYR, 
<number box="[300,354,1861,1884]" pageId="1" pageNumber="132" value="1865.0">1865</number>
, 
<emphasis box="[367,501,1861,1884]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Wheeler" authorityYear="1935" box="[367,501,1861,1884]" class="Insecta" family="Formicidae" genus="Romblonella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Romblonella</taxonomicName>
</emphasis>
WHEELER, 
<number box="[629,684,1861,1884]" pageId="1" pageNumber="132" value="1935.0">1935</number>
, 
<emphasis italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Calyptomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Calyptomyrmex</taxonomicName>
</emphasis>
EMERY, 
<number box="[357,412,1891,1914]" pageId="1" pageNumber="132" value="1887.0">1887</number>
, 
<emphasis box="[425,558,1891,1914]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="Terayama &amp; Yamane, 2000" authorityName="Terayama &amp; Yamane" authorityYear="2000" box="[425,558,1891,1914]" class="Insecta" family="Formicidae" genus="Lasiomyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Lasiomyrma</taxonomicName>
</emphasis>
TERAYAMA &amp; YA- MANE, 
<number box="[227,281,1920,1943]" pageId="1" pageNumber="132" value="2000.0">2000</number>
and 
<taxonomicName authorityName="Brown" authorityYear="1986" box="[332,455,1920,1943]" class="Insecta" family="Formicidae" genus="Indomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[332,455,1920,1943]" italics="true" pageId="1" pageNumber="132">Indomyrma</emphasis>
</taxonomicName>
BROWN, 
<number box="[558,615,1920,1943]" pageId="1" pageNumber="132" value="1986.0">1986</number>
.
</paragraph>
<paragraph pageId="1" pageNumber="132">
WARD &amp; al. (
<number box="[341,398,1949,1972]" pageId="1" pageNumber="132" value="2015.0">2015</number>
) were unable to sequence 
<emphasis italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="Terayama &amp; Yamane, 2000" authorityName="Terayama &amp; Yamane" authorityYear="2000" class="Insecta" family="Formicidae" genus="Lasiomyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Lasiomyrma</taxonomicName>
</emphasis>
and 
<taxonomicName authorityName="Brown" authorityYear="1986" box="[279,403,1979,2002]" class="Insecta" family="Formicidae" genus="Indomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[279,403,1979,2002]" italics="true" pageId="1" pageNumber="132">Indomyrma</emphasis>
</taxonomicName>
, two endemic Asian genera. Neither genus is morphologically similar to 
<emphasis box="[584,678,2008,2031]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[584,678,2008,2031]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph pageId="1" pageNumber="132">
<emphasis box="[858,991,1656,1679]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="Terayama &amp; Yamane, 2000" authorityName="Terayama &amp; Yamane" authorityYear="2000" box="[858,991,1656,1679]" class="Insecta" family="Formicidae" genus="Lasiomyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Lasiomyrma</taxonomicName>
</emphasis>
has an anterior median clypeus produced into an obtuse angle, 
<number box="[1050,1072,1685,1708]" pageId="1" pageNumber="132" value="11.0">11</number>
-segmented antenna, elongate-triangular mandibles that usually close tightly with anterior clypeal margin (line drawing of frontal head shows a gap in 
<emphasis box="[847,1001,1773,1796]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Terayama &amp; Yamane" authorityYear="2000" class="Insecta" family="Formicidae" genus="Lasiomyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="gracilinoda">L. gracilinoda</taxonomicName>
</emphasis>
TERAYAMA &amp; YAMANE, 
<number box="[1274,1331,1773,1796]" pageId="1" pageNumber="132" value="2000.0">2000</number>
), flat dorsum of promesonotum and propodeum, propodeal spines present, and a pedunculate petiole (TERAYAMA &amp; YAMANE 
<number box="[820,878,1861,1884]" pageId="1" pageNumber="132" value="2000.0">2000</number>
). In contrast, 
<emphasis box="[1024,1118,1861,1884]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[1024,1118,1861,1884]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
has an entire anterior clypeal margin, 
<number box="[912,936,1891,1914]" pageId="1" pageNumber="132" value="12.0">12</number>
-segmented antenna, narrow-based triangular mandibles that always close with a gap between the mandibular basal margin and the anterior clypeal margin, convex dorsum of promesonotum and propodeum, propodeal armament reduced to denticles, and a sessile petiole.
</paragraph>
<paragraph pageId="1" pageNumber="132">
<emphasis box="[189,313,176,199]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Brown" authorityYear="1986" box="[189,313,176,199]" class="Insecta" family="Formicidae" genus="Indomyrma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Indomyrma</taxonomicName>
</emphasis>
has reduced eyes with a total of less than 
<number box="[151,179,205,228]" pageId="1" pageNumber="132" value="12.0">12</number>
ommatidia, an antennal scrobe, albeit weakly developed, bounded dorsally by a distinct frontal carina, median clypeal margin expanded as a notched convex apron that barely covers basal borders of completely closed mandibles, flat dorsum of promesonotum and propodeum, erect hairs in bilateral positions on head, mesosoma, petiole and pospetiole, propodeal spines present, and a pedunculate petiole (
<collectionCode box="[334,420,411,434]" pageId="1" pageNumber="132">BROWN</collectionCode>
<number box="[427,484,411,434]" pageId="1" pageNumber="132" value="1986.0">1986</number>
). In contrast, 
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[630,724,411,434]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">
<emphasis box="[630,724,411,434]" italics="true" pageId="1" pageNumber="132">Aretidris</emphasis>
</taxonomicName>
has larger eyes with about eight ommatidia in its longest axis, no antennal scrobe, a short frontal carina, entire median clypeal margin without an apron, strongly convex dorsum of promesonotum and propodeum, erect hairs on head, mesosoma, petiole and postpetiole abundant but not in bilateral positions, propodeal armament reduced to denticles, and a sessile petiole.
</paragraph>
<paragraph pageId="1" pageNumber="132">
WARD &amp; al. (
<number box="[346,403,645,668]" pageId="1" pageNumber="132" value="2015.0">2015</number>
) found 
<emphasis box="[495,593,645,668]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[495,593,645,668]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
(as Myrmicine Genus # 
<number box="[238,265,675,698]" pageId="1" pageNumber="132" value="26.0">26</number>
<collectionCode box="[272,307,675,698]" pageId="1" pageNumber="132">PH</collectionCode>
02) to be a sister taxon to the genus 
<taxonomicName authorityName="Mayr" authorityYear="1865" class="Insecta" family="Formicidae" genus="Vollenhovia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="1" pageNumber="132">Vollenhovia</emphasis>
</taxonomicName>
in the 
<taxonomicName authorityName="Forel" authorityYear="1893" box="[292,474,704,727]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
. These two genera are not particularly similar morphologically, sharing only a couple of petiolar features: a protuberance over the petiolar spiracle (also found in 
<emphasis box="[381,535,792,815]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Menozzi" authorityYear="1933" class="Insecta" family="Formicidae" genus="Gauromyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Gauromyrmex</taxonomicName>
</emphasis>
MENOZZI, 
<number box="[664,718,792,815]" pageId="1" pageNumber="132" value="1933.0">1933</number>
and 
<taxonomicName authorityName="Wheeler" authorityYear="1935" box="[151,284,821,844]" class="Insecta" family="Formicidae" genus="Romblonella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[151,284,821,844]" italics="true" pageId="1" pageNumber="132">Romblonella</emphasis>
</taxonomicName>
) and a lack of a petiolar peduncle. That 
<emphasis italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
and 
<taxonomicName authorityName="Mayr" authorityYear="1865" box="[257,388,851,874]" class="Insecta" family="Formicidae" genus="Vollenhovia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[257,388,851,874]" italics="true" pageId="1" pageNumber="132">Vollenhovia</emphasis>
</taxonomicName>
together are sister to 
<emphasis italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Calyptomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Calyptomyrmex</taxonomicName>
</emphasis>
is even more unexpected and problematic. I can find no morphological similarities between 
<emphasis box="[555,646,909,932]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[555,646,909,932]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
and 
<emphasis italics="true" pageId="1" pageNumber="132">
<taxonomicName authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Calyptomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Calyptomyrmex</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</paragraph>
</td>
</tr>
</paragraph>
</subSubSection>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</paragraph>
</td>
</td>
</td>
</td>
</td>
</tr>
</abbreviationData>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="132" type="discussion">
<paragraph pageId="1" pageNumber="132">
<emphasis box="[189,283,968,991]" italics="true" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" box="[189,283,968,991]" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.n.">Aretidris</taxonomicName>
</emphasis>
is already included in the myrmicine generic key of GENERAL &amp; ALPERT (
<number box="[466,524,997,1020]" pageId="1" pageNumber="132" value="2012.0">2012</number>
), as &quot;Unnamed Genus PH03&quot;. Only a simple substitution of the name is necessary to key it out among the ant genera of the Philippines.
</paragraph>
</subSubSection>
<subSection pageId="1" pageNumber="132" type="key">
<subSubSection pageId="1" pageNumber="132" type="key">
<paragraph box="[151,326,1101,1124]" pageId="1" pageNumber="132">
<emphasis bold="true" box="[151,326,1101,1124]" pageId="1" pageNumber="132">Key to workers</emphasis>
</paragraph>
<paragraph pageId="1" pageNumber="132">
<number box="[151,164,1144,1167]" pageId="1" pageNumber="132" value="1.0">1</number>
<table box="[212,763,1144,1436]" pageId="1" pageNumber="132">
<tr box="[212,763,1144,1284]" pageId="1" pageNumber="132">
<td box="[228,531,1144,1284]" pageId="1" pageNumber="132">
<td box="[212,271,1144,1284]" pageId="1" pageNumber="132">
Head 
<td box="[251,317,1144,1255]" pageId="1" pageNumber="132">
and mesosoma 
<td box="[398,554,1144,1255]" pageId="1" pageNumber="132">
heavily 
<td box="[528,703,1144,1284]" pageId="1" pageNumber="132">
sculptured
<td box="[641,763,1144,1284]" pageId="1" pageNumber="132">
; erect hairs moderately abundant on head and mesosoma; 
<number box="[283,296,1203,1226]" pageId="1" pageNumber="132" value="1.0">1</number>
gastral tergite slightly less than half the length of gaster; body larger (
<collectionCode box="[569,615,1232,1255]" pageId="1" pageNumber="132">HW</collectionCode>
&gt; 0.88).. ............................................. 
<emphasis bold="true" box="[539,763,1261,1284]" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="buenaventei" status="sp.n.">
<emphasis bold="true" box="[539,557,1261,1284]" italics="true" pageId="1" pageNumber="132">A</emphasis>
. 
<emphasis bold="true" box="[570,703,1261,1284]" italics="true" pageId="1" pageNumber="132">buenaventei</emphasis>
</taxonomicName>
<taxonomicNameLabel rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
<paragraph pageId="1" pageNumber="132">
– 
<tr box="[212,763,1296,1348]" pageId="1" pageNumber="132">
<td box="[212,296,1296,1348]" pageId="1" pageNumber="132">
Head 
<td box="[273,311,1296,1319]" pageId="1" pageNumber="132">and</td>
<td box="[304,426,1296,1348]" pageId="1" pageNumber="132">
mesosoma 
<td box="[433,505,1296,1348]" pageId="1" pageNumber="132">
mostly 
<td box="[510,671,1296,1348]" pageId="1" pageNumber="132">
smooth; erect 
<td box="[657,707,1296,1348]" pageId="1" pageNumber="132">
hairs st sparsely distributed on head and mesosoma; 
<number box="[682,695,1325,1348]" pageId="1" pageNumber="132" value="1.0">1</number>
<tr box="[212,763,1355,1436]" pageId="1" pageNumber="132">
<td box="[212,284,1355,1436]" pageId="1" pageNumber="132">
gastral 
<td box="[302,585,1355,1436]" pageId="1" pageNumber="132">
<td box="[291,474,1355,1407]" pageId="1" pageNumber="132">
tergite extremely large
<td box="[511,668,1355,1436]" pageId="1" pageNumber="132">
, accounting 
<td box="[627,763,1355,1436]" pageId="1" pageNumber="132">
for most 
<td box="[269,329,1384,1436]" pageId="1" pageNumber="132">
of the length of gaster; body smaller (
<collectionCode box="[661,707,1384,1407]" pageId="1" pageNumber="132">HW</collectionCode>
&lt;0.84). .......................................... 
<emphasis bold="true" box="[596,763,1413,1436]" pageId="1" pageNumber="132">
<taxonomicName authority="David Emmanuel M. GENERAL, 2015" authorityName="David Emmanuel M. GENERAL" authorityYear="2015" class="Insecta" family="Formicidae" genus="Aretidris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="clousei" status="sp.n.">
<emphasis bold="true" box="[596,613,1413,1436]" italics="true" pageId="1" pageNumber="132">A</emphasis>
. 
<emphasis bold="true" box="[627,703,1413,1436]" italics="true" pageId="1" pageNumber="132">clousei</emphasis>
</taxonomicName>
<taxonomicNameLabel rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</td>
</td>
</td>
</td>
</td>
</td>
</tr>
</td>
</td>
</td>
</td>
</td>
</tr>
</paragraph>
</table>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>