treatments-xml/data/67/F1/F0/67F1F03C9B955C15B82C0AEF8CEF81BC.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="67F1F03C9B955C15B82C0AEF8CEF81BC" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana occultata Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="B2881074-880F-467C-BDE5-B526AB9F72C5" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-20</mods:number>
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<mods:number>81</mods:number>
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<mods:start>79</mods:start>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e81961</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:B2881074-880F-467C-BDE5-B526AB9F72C5" httpUri="http://treatment.plazi.org/id/67F1F03C9B955C15B82C0AEF8CEF81BC" lastPageNumber="79" pageId="0" pageNumber="79">
<subSubSection pageId="0" pageNumber="79" type="nomenclature">
<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="https://zoobank.org/B2881074-880F-467C-BDE5-B526AB9F72C5" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana occultata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="occultata" status="sp. nov.">Coarctana occultata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">Figs 22</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31I, J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22G</figureCitation>
) with apex broad and rounded. Subgenital plate (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22H</figureCitation>
) with few filiform setae near outer margin and a longer apical group on dorsal surface. Style (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22K</figureCitation>
) apex tapered, curved dorsally and acute. Aedeagus (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22L, M</figureCitation>
) apodemal process slender at base and enlarged subapically; shaft apex with pair of subapical processes short, about 1/5 length of shaft, curved ventrally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 7.8 mm; paratype, male (n = 1) 8.1 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22B</figureCitation>
), frons 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus very wide, 1.1 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins straight and parallel near base and distally strongly convergent toward apex; apex straight. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22C</figureCitation>
), crown-face transition thin, with 4 transverse carinae; clypeus slightly inflated. Profemur AV row with 4-5 setae; PV row with 1-2 setae. Protibia PD row with 4 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 23-24, 12 and 15-16 macrosetae respectively. Metatarsomere I 4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-9 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22A-D</figureCitation>
, 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31I, J</figureCitation>
) reddish-brown. Face (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22B</figureCitation>
) yellowish; frons reddish-brown below crown face transition; antenal pit black. Proepimeron (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22C</figureCitation>
) black. Forewing (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22D</figureCitation>
) with small black maculae on apex of anal veins, cross veins of discal and apical cells; costal and 5th apical cells dark brown. Metatibia (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22E</figureCitation>
) 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin rounded. Valve (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22F</figureCitation>
) 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin slightly excavated. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22G</figureCitation>
), 2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and long, extending ventrally by 1/3 height of lateral lobe; anteroventral margin rounded; posterodorsal margin straight; posteroventral margin rounded; apex broad, rounded. Subgenital plate, in lateral view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22G</figureCitation>
), almost reaching pygofer apex; in ventral view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22H</figureCitation>
), elongated, 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; dorsal surface with few filiform setae near outer margin and a longer apical group on dorsal surface; lateral margins slightly rounded; apex tapered, rounded. Connective (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22I</figureCitation>
) wider than long; anterior margin moderately excavated; dorsal keel and stem reduced. Style, in dorsal view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22J</figureCitation>
), with outer lobe truncated; in lateral view (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22K</figureCitation>
), blade slender, slightly tapered toward apex; dorsal and ventral margins almost straight; ventral margin with apical half serrated; apex curved dorsally, acute. Aedeagus (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme bent posterad, dorsal margin rounded, lateral margins produced laterally; apodemal process slender at base and enlarged subapically, strongly curved dorsally near base; apex rounded; shaft approximately cylindrical, very long and thin, strongly curved dorsally near base, apical half slightly sinuate; apex with pair of subapical processes thin and short, curved ventrally. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2012-08-23" collectorName="R. R. Cavichioli" country="Peru" elevation="985" latitude="-13.3550005" location="Estrada" longLatPrecision="21" longitude="-70.88944" municipality="Peru" specimenCount="1" stateProvince="Cuzco" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male: 
<collectingCountry name="Peru">Peru</collectingCountry>
</emphasis>
: &quot; 
<collectingMunicipality>Peru</collectingMunicipality>
, 
<collectingRegion country="Peru" name="Cusco">Cuzco</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:67F1F03C9B955C15B82C0AEF8CEF81BC:2E710E85BCB1070DFDF6C8B920ACA107" country="Peru" latitude="-13.3550005" longLatPrecision="21" longitude="-70.88944" municipality="Peru" name="Estrada" stateProvince="Cuzco">Estrada</location>
30 C\ 
<locationDeviation location="Quincemil">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.6" unit="km" value="26.0">26km</quantity>
W Quincemil
</locationDeviation>
, 
<collectingDate value="2012-08-23">23.viii.2012</collectingDate>
, (light)\ 
<geoCoordinate degrees="13" direction="south" minutes="21" orientation="latitude" precision="15" seconds="18" value="-13.3550005">13°21′18″S</geoCoordinate>
<geoCoordinate degrees="70" direction="west" minutes="53" orientation="longitude" precision="15" seconds="22" value="-70.88944">70°53′22″W</geoCoordinate>
,\ 
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">985m</elevation>
</quantity>
<collectorName>R.R. Cavichioli</collectorName>
leg.&quot; (MUSM)
</materialsCitation>
. 
<materialsCitation collectingDate="2009-05-01" collectingDateMax="2009-05-08" collectingDateMin="2009-05-01" collectorName="ME Irwin, G" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Bagua" longLatPrecision="21" longitude="-77.63694" municipality="Aguas Verdes" specimenCount="1" stateProvince="Amazonas" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Paratype</typeStatus>
</emphasis>
: &quot; 
<collectingCounty>Peru</collectingCounty>
, Dept. 
<collectingRegion country="Peru" name="Amazonas">Amazonas</collectingRegion>
, Distr.\ 
<collectingMunicipality>
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
Verdes
</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:67F1F03C9B955C15B82C0AEF8CEF81BC:11741100CE9317A64E7B88A6BE3B2793" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Aguas Verdes" name="Bagua" stateProvince="Amazonas">Bagua</location>
/ 
<location LSID="urn:lsid:plazi:treatment:67F1F03C9B955C15B82C0AEF8CEF81BC:6CF03EEFDFC37AEA5DC819AC4CC7619D" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Aguas Verdes" name="Tarapoto Rd" stateProvince="Amazonas">Tarapoto Rd</location>
\ (5N) at 
<location LSID="urn:lsid:plazi:treatment:67F1F03C9B955C15B82C0AEF8CEF81BC:FE03EA453F35CEDE0BCE4B24C6E7CCAA" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Aguas Verdes" name="Km" stateProvince="Amazonas">Km</location>
403, 
<location LSID="urn:lsid:plazi:treatment:67F1F03C9B955C15B82C0AEF8CEF81BC:BC28CB2730CB629761D4D482791B7616" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Aguas Verdes" name="Malaise" stateProvince="Amazonas">Malaise</location>
, 
<collectingDate value="2009-05-01" valueMax="2009-05-08" valueMin="2009-05-01">1-8.V.2009</collectingDate>
\ 
<collectorName>ME Irwin, G</collectorName>
<normalizedToken originalValue="Antón">Anton</normalizedToken>
Maya\ 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
</quantity>
<geoCoordinate degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°41′23″S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°38′13″W</geoCoordinate>
(DZRJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">
The new species name comes from the Latin word 
<normalizedToken originalValue="“occultatum”">&quot;occultatum&quot;</normalizedToken>
and means hidden. This name was chosen because the studied specimens were miss identified as 
<taxonomicName authorityName="DeLong &amp; Freytag" authorityYear="1976" class="Insecta" family="Cicadellidae" genus="Curtara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtara apena" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="apena">
<emphasis italics="true" pageId="0" pageNumber="79">Curtara apena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
Our analysis recovered 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. occultata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="occultata">
<emphasis italics="true" pageId="0" pageNumber="79">C. occultata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
as sister to 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. apena" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="apena">
<emphasis italics="true" pageId="0" pageNumber="79">C. apena</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
with low branch support (SR &lt;50) (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
). These species have a similar coloration (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31A, B, I, J</figureCitation>
), and the subgenital plates (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22H</figureCitation>
) with long filiform setae forming an apical group, and a similar aedeagus shape (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22L</figureCitation>
). However, the new species is differentiated by having a shorter pygofer and the aedeagus with a single pair of subapical processes.
</paragraph>
</subSubSection>
</treatment>
</document>