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<document id="5AD4B96B4D7A85A190587D58EC12E4A9" ID-CLB-Dataset="53713" ID-DOI="10.5852/ejt.2023.872.2131" ID-GBIF-Dataset="31ad2cf7-cce8-4202-9a46-d81b0073d561" ID-ISSN="2118-9773" ID-Zenodo-Dep="8018303" ID-ZooBank="05098E38-AB11-486E-8F28-8567DE6BC19C" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1686244863328" checkinUser="felipe" docAuthor="Ebejer, Martin J." docDate="2023" docId="505487F2B0CBFF59FDE0FCCC08C83F92" docLanguage="en" docName="ejt-872_1-161.pdf" docOrigin="European Journal of Taxonomy 872" docSource="http://zoobank.org/05098e38-ab11-486e-8f28-8567de6bc19c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aphaniosoma verecundum Ebejer 1998" docType="treatment" docVersion="8" lastPageNumber="144" masterDocId="AC6DFF8AB044FFD6FFC7FFC10B313A78" masterDocTitle="The genus Aphaniosoma Becker, 1903 (Diptera: Chyromyidae) in the Eastern Mediterranean and Middle East, with descriptions of new species" masterLastPageNumber="161" masterPageNumber="1" pageNumber="144" updateTime="1699521912180" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D5E837BAE96C97587D49921AA5384772">The genus Aphaniosoma Becker, 1903 (Diptera: Chyromyidae) in the Eastern Mediterranean and Middle East, with descriptions of new species</mods:title>
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<subSubSection id="90E7656FB0CBFF59FDE0FCCC0F3D3950" box="[551,1036,781,808]" pageId="143" pageNumber="144" type="nomenclature">
<paragraph id="D84236E4B0CBFF59FDE0FCCC0F3D3950" blockId="143.[551,1036,781,843]" box="[551,1036,781,808]" pageId="143" pageNumber="144">
<heading id="830A8188B0CBFF59FDE0FCCC0F3D3950" box="[551,1036,781,808]" centered="true" fontSize="11" level="2" pageId="143" pageNumber="144" reason="2">
<taxonomicName id="1FFD4D67B0CBFF59FDE0FCCC0F3D3950" ID-CoL="67MBC" authority="Ebejer, 1998" authorityName="Ebejer" authorityYear="1998" box="[551,1036,781,808]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="verecundum">
<emphasis id="EA89EAF6B0CBFF59FDE0FCCC085E3950" bold="true" box="[551,879,781,808]" italics="true" pageId="143" pageNumber="144">Aphaniosoma verecundum</emphasis>
<bibRefCitation id="BC6C4B15B0CBFF59FCB0FCCF0F3D3950" author="Ebejer M. J." box="[887,1036,782,808]" pageId="143" pageNumber="144" pagination="191 - 230" refId="ref57257" refString="Ebejer M. J. 1998. A Review of the Palaearctic species of Aphaniosoma Becker (Diptera, Chyromyidae), with descriptions of new species and a key for the identification of adults. Mitteilungen Museum fur Naturkunde Berliner Deutsche entomologische Zeitschrift 45 (2): 191 - 230. https: // doi. org / 10.1002 / mmnd. 4810450208" type="journal article" year="1998">Ebejer, 1998</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="90E7656FB0CBFF59FD1FFCF0086A3933" box="[728,859,817,843]" pageId="143" pageNumber="144" type="description">
<paragraph id="D84236E4B0CBFF59FD1FFCF0086A3933" blockId="143.[551,1036,781,843]" box="[728,859,817,843]" pageId="143" pageNumber="144">
<figureCitation id="40C62A61B0CBFF59FD1FFCF0086A3933" box="[728,859,817,843]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="144.[189,232,1963,1989]" captionStartId-1="145.[189,232,1963,1989]" captionTargetBox-0="[339,1249,265,1924]" captionTargetBox-1="[190,1397,265,1924]" captionTargetId-0="figure-13@144.[339,1249,265,1924]" captionTargetId-1="figure-12@145.[190,1397,265,1924]" captionTargetPageId-0="144" captionTargetPageId-1="145" captionText-0="Fig. 83. Aphaniosoma verecundum Ebejer, 1998, ♂, (MJE). A. Habitus, lateral view. B. Head and thorax dorsal view." captionText-1="Fig. 84. Aphaniosoma verecundum Ebejer, 1998, ♂ (MJE). A. Hind trochanter and femur. BC. Hypopygium in situ. B. Postero-ventral view. C. Postero-lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.8018553" figureDoi-1="http://doi.org/10.5281/zenodo.8018555" httpUri-0="https://zenodo.org/record/8018553/files/figure.png" httpUri-1="https://zenodo.org/record/8018555/files/figure.png" pageId="143" pageNumber="144">Figs 8384</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90E7656FB0CBFF59FF7AFCB30A643E3B" pageId="143" pageNumber="144" type="materials_examined">
<paragraph id="D84236E4B0CBFF59FF7AFCB30A9C39F4" blockId="143.[189,429,882,908]" box="[189,429,882,908]" pageId="143" pageNumber="144">
<heading id="830A8188B0CBFF59FF7AFCB30A9C39F4" bold="true" box="[189,429,882,908]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="EA89EAF6B0CBFF59FF7AFCB30A9C39F4" bold="true" box="[189,429,882,908]" pageId="143" pageNumber="144">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B0CBFF59FF7AFC600AC239A7" blockId="143.[189,1399,929,991]" pageId="143" pageNumber="144">
<materialsCitation id="68953CB9B0CBFF59FF7AFC600ADE39A7" ID-GBIF-Occurrence="4127865510" collectingDate="2022-04-30" collectionCode="MJE" collectorName="M. J. Ebejer" country="Cyprus" location="Akrotiri Peninsula" municipality="Kolossi" pageId="143" pageNumber="144" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A0EA7674B0CBFF59FF7AFC600A0139C3" box="[189,304,929,955]" name="Cyprus" pageId="143" pageNumber="144">CYPRUS</collectingCountry>
<specimenCount id="CEFBFD6DB0CBFF59FE8EFC600A4639C4" box="[329,375,929,956]" pageId="143" pageNumber="144" type="male">1 ♂</specimenCount>
;
<location id="DD22603FB0CBFF59FE45FC60096E39C3" LSID="urn:lsid:plazi:treatment:505487F2B0CBFF59FDE0FCCC08C83F92:DD22603FB0CBFF59FE45FC60096E39C3" box="[386,607,929,955]" country="Cyprus" municipality="Kolossi" name="Akrotiri Peninsula" pageId="143" pageNumber="144">Akrotiri Peninsula</location>
,
<quantity id="1F059B01B0CBFF59FDACFC6009AE39C3" box="[619,671,929,955]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="143" pageNumber="144" unit="km" value="3.0">
<locationDeviation id="C9F02FC6B0CBFF59FDACFC6009AE39C3" box="[619,671,929,955]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="143" pageNumber="144" unit="km" value="3.0">3km</locationDeviation>
</quantity>
southwest of
<collectingMunicipality id="3826AC9EB0CBFF59FC83FC60089339C3" box="[836,930,929,955]" pageId="143" pageNumber="144">Kolossi</collectingMunicipality>
;
<date id="AC431024B0CBFF59FC69FC600F7A39C3" box="[942,1099,929,955]" pageId="143" pageNumber="144" value="2022-04-30">
<collectingDate id="BC07E9CCB0CBFF59FC69FC600F7A39C3" box="[942,1099,929,955]" pageId="143" pageNumber="144" value="2022-04-30">30 Apr. 2022</collectingDate>
</date>
;
<collectorName id="75085332B0CBFF59FB90FC630FD139C3" box="[1111,1248,929,956]" pageId="143" pageNumber="144">M.J. Ebejer</collectorName>
leg.; ruderal meadow and marsh;
<collectionCode id="BEECAE21B0CBFF59FE74FC040ADE39A7" box="[435,495,965,991]" pageId="143" pageNumber="144">MJE</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="D84236E4B0CBFF59FF7AFBC70A643E3B" blockId="143.[189,1399,1030,1091]" pageId="143" pageNumber="144">
<materialsCitation id="68953CB9B0CBFF59FF7AFBC70A603E3B" ID-GBIF-Occurrence="4127865795" collectingDate="1998-05-14" collectionCode="MJE" collectorName="M. J. Ebejer" country="Greece" location="Fthiotida" pageId="143" pageNumber="144" specimenCount="1" specimenCount-male="1" stateProvince="Sterea Elada">
<collectingCountry id="A0EA7674B0CBFF59FF7AFBC70A013E58" box="[189,304,1030,1056]" name="Greece" pageId="143" pageNumber="144">GREECE</collectingCountry>
<specimenCount id="CEFBFD6DB0CBFF59FE8DFBC70A4B3E58" box="[330,378,1030,1056]" pageId="143" pageNumber="144" type="male">1 ♂</specimenCount>
;
<collectingRegion id="1A39F806B0CBFF59FE4FFBC709113E58" box="[392,544,1030,1056]" pageId="143" pageNumber="144">Sterea Elada</collectingRegion>
,
<location id="DD22603FB0CBFF59FDEAFBC709AA3E58" LSID="urn:lsid:plazi:treatment:505487F2B0CBFF59FDE0FCCC08C83F92:DD22603FB0CBFF59FDEAFBC709AA3E58" box="[557,667,1030,1056]" country="Greece" name="Fthiotida" pageId="143" pageNumber="144" stateProvince="Sterea Elada">Fthiotida</location>
,
<location id="DD22603FB0CBFF59FD60FBC708703E58" LSID="urn:lsid:plazi:treatment:505487F2B0CBFF59FDE0FCCC08C83F92:DD22603FB0CBFF59FD60FBC708703E58" box="[679,833,1030,1056]" country="Greece" name="Thermopiles" pageId="143" pageNumber="144" stateProvince="Sterea Elada">Thermopiles</location>
;
<date id="AC431024B0CBFF59FC89FBC708C03E58" box="[846,1009,1030,1056]" pageId="143" pageNumber="144" value="1998-05-14">
<collectingDate id="BC07E9CCB0CBFF59FC89FBC708C03E58" box="[846,1009,1030,1056]" pageId="143" pageNumber="144" value="1998-05-14">14 May 1998</collectingDate>
</date>
;
<collectorName id="75085332B0CBFF59FC3AFBC70FB63E58" box="[1021,1159,1030,1056]" pageId="143" pageNumber="144">M.J. Ebejer</collectorName>
leg.; hot spring and marsh;
<collectionCode id="BEECAE21B0CBFF59FED2FBE80A603E3B" box="[277,337,1065,1091]" pageId="143" pageNumber="144">MJE</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90E7656FB0CBFF59FF7AFBAB0E563F1B" pageId="143" pageNumber="144" type="discussion">
<paragraph id="D84236E4B0CBFF59FF7AFBAB0A013EFC" blockId="143.[189,304,1130,1156]" box="[189,304,1130,1156]" pageId="143" pageNumber="144">
<heading id="830A8188B0CBFF59FF7AFBAB0A013EFC" bold="true" box="[189,304,1130,1156]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="EA89EAF6B0CBFF59FF7AFBAB0A013EFC" bold="true" box="[189,304,1130,1156]" pageId="143" pageNumber="144">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B0CBFF59FF7AFB580E563F1B" blockId="143.[187,1399,1177,1379]" pageId="143" pageNumber="144">
This is a species of the
<taxonomicName id="1FFD4D67B0CBFF59FE05FB58093F3ECB" authorityName="Lyneborg" authorityYear="1973" box="[450,526,1177,1203]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="collini">
<emphasis id="EA89EAF6B0CBFF59FE05FB58093F3ECB" box="[450,526,1177,1203]" italics="true" pageId="143" pageNumber="144">collini</emphasis>
</taxonomicName>
group, having long anterior scutal setae, a reduced number of dorsocentrals to 1+3 (
<figureCitation id="40C62A61B0CBFF59FEDEFB7D0AB43EAE" box="[281,389,1212,1238]" captionStart="Fig" captionStartId="144.[189,232,1963,1989]" captionTargetBox="[339,1249,265,1924]" captionTargetId="figure-13@144.[339,1249,265,1924]" captionTargetPageId="144" captionText="Fig. 83. Aphaniosoma verecundum Ebejer, 1998, ♂, (MJE). A. Habitus, lateral view. B. Head and thorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8018553" httpUri="https://zenodo.org/record/8018553/files/figure.png" pageId="143" pageNumber="144">Fig. 83A</figureCitation>
) and prescutellar setae (
<figureCitation id="40C62A61B0CBFF59FD62FB7D08203EAE" box="[677,785,1212,1238]" captionStart="Fig" captionStartId="144.[189,232,1963,1989]" captionTargetBox="[339,1249,265,1924]" captionTargetId="figure-13@144.[339,1249,265,1924]" captionTargetPageId="144" captionText="Fig. 83. Aphaniosoma verecundum Ebejer, 1998, ♂, (MJE). A. Habitus, lateral view. B. Head and thorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8018553" httpUri="https://zenodo.org/record/8018553/files/figure.png" pageId="143" pageNumber="144">Fig. 83B</figureCitation>
). There is no special development on the male hind trochanter (
<figureCitation id="40C62A61B0CBFF59FE85FB1E0A9D3E81" box="[322,428,1247,1273]" captionStart="Fig" captionStartId="145.[189,232,1963,1989]" captionTargetBox="[190,1397,265,1924]" captionTargetId="figure-12@145.[190,1397,265,1924]" captionTargetPageId="145" captionText="Fig. 84. Aphaniosoma verecundum Ebejer, 1998, ♂ (MJE). A. Hind trochanter and femur. BC. Hypopygium in situ. B. Postero-ventral view. C. Postero-lateral view." figureDoi="http://doi.org/10.5281/zenodo.8018555" httpUri="https://zenodo.org/record/8018555/files/figure.png" pageId="143" pageNumber="144">Fig.84A</figureCitation>
). It is best identified on dissection of the male hypopygium. The surstylus is relatively short compared to that of
<taxonomicName id="1FFD4D67B0CBFF59FE3FFAC209BF3F64" authorityName="Lyneborg" authorityYear="1973" box="[504,654,1282,1309]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="hackmani">
<emphasis id="EA89EAF6B0CBFF59FE3FFAC209BF3F64" box="[504,654,1282,1309]" italics="true" pageId="143" pageNumber="144">A. hackmani</emphasis>
</taxonomicName>
and
<taxonomicName id="1FFD4D67B0CBFF59FD0AFAC208773F64" authorityName="Ebejer" authorityYear="1996" box="[717,838,1282,1309]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="harteni">
<emphasis id="EA89EAF6B0CBFF59FD0AFAC208773F64" box="[717,838,1282,1309]" italics="true" pageId="143" pageNumber="144">A. harteni</emphasis>
</taxonomicName>
. The abdominal sternite 6 is distinctive, being small and transversely rectangular with protruding angles and a deep invagination into the abdomen at its middle (
<bibRefCitation id="BC6C4B15B0CBFF59FE9AFA88091D3F1B" author="Ebejer M. J." box="[349,556,1353,1379]" pageId="143" pageNumber="144" pagination="191 - 230" refId="ref57257" refString="Ebejer M. J. 1998. A Review of the Palaearctic species of Aphaniosoma Becker (Diptera, Chyromyidae), with descriptions of new species and a key for the identification of adults. Mitteilungen Museum fur Naturkunde Berliner Deutsche entomologische Zeitschrift 45 (2): 191 - 230. https: // doi. org / 10.1002 / mmnd. 4810450208" type="journal article" year="1998">Ebejer 1998: 228</bibRefCitation>
, figs 8183). The male hypopygium is illustrated in situ (
<figureCitation id="40C62A61B0CBFF59FB0BFA880E6B3F1B" box="[1228,1370,1353,1379]" captionStart="Fig" captionStartId="145.[189,232,1963,1989]" captionTargetBox="[190,1397,265,1924]" captionTargetId="figure-12@145.[190,1397,265,1924]" captionTargetPageId="145" captionText="Fig. 84. Aphaniosoma verecundum Ebejer, 1998, ♂ (MJE). A. Hind trochanter and femur. BC. Hypopygium in situ. B. Postero-ventral view. C. Postero-lateral view." figureDoi="http://doi.org/10.5281/zenodo.8018555" httpUri="https://zenodo.org/record/8018555/files/figure.png" pageId="143" pageNumber="144">Fig. 84BC</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="D84236E4B0CBFF59FF7AFA4B0A693FDC" blockId="143.[189,344,1418,1444]" box="[189,344,1418,1444]" pageId="143" pageNumber="144">
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<emphasis id="EA89EAF6B0CBFF59FF7AFA4B0A693FDC" bold="true" box="[189,344,1418,1444]" pageId="143" pageNumber="144">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B0CBFF59FF7AFA1108C83F92" blockId="143.[189,1017,1488,1515]" box="[189,1017,1488,1515]" pageId="143" pageNumber="144">
<collectingCountry id="A0EA7674B0CBFF59FF7AFA110A223F92" box="[189,275,1488,1514]" name="Greece" pageId="143" pageNumber="144">Greece</collectingCountry>
,
<collectingCountry id="A0EA7674B0CBFF59FED8FA110A463F92" box="[287,375,1488,1514]" name="Cyprus" pageId="143" pageNumber="144">Cyprus</collectingCountry>
,
<collectingCountry id="A0EA7674B0CBFF59FE45FA110AE73F92" box="[386,470,1488,1514]" name="Turkey" pageId="143" pageNumber="144">Turkey</collectingCountry>
(
<bibRefCitation id="BC6C4B15B0CBFF59FE23FA1109463F92" author="Ebejer M. J." box="[484,631,1488,1514]" pageId="143" pageNumber="144" pagination="191 - 230" refId="ref57257" refString="Ebejer M. J. 1998. A Review of the Palaearctic species of Aphaniosoma Becker (Diptera, Chyromyidae), with descriptions of new species and a key for the identification of adults. Mitteilungen Museum fur Naturkunde Berliner Deutsche entomologische Zeitschrift 45 (2): 191 - 230. https: // doi. org / 10.1002 / mmnd. 4810450208" type="journal article" year="1998">Ebejer 1998</bibRefCitation>
,
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). New record for
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