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<mods:title id="3C80C809907A168907F06A7E9D83ED3B">Pseudolitochira integra (Miers, 1884) (Crustacea: Brachyura: Pilumnidae): redescribed and illustrated from micro-CT scanning the type female</mods:title>
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<mods:namePart id="3281F7BD7059DF7F3AB70EAA49D041A3">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="02E49B5FA845252A35C5075DFD2A61EA">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:affiliation id="A57B7D0A51F3A3837C60E5E0F682B582">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, England.</mods:affiliation>
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<mods:affiliation id="789567BDDB2E60182D97B7CDB7D7ED58">Imaging and Analysis Centre, Core Research Laboratories, The Natural History Museum, London SW 7 5 BD, England. Brett. Clark @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5888 - 9223</mods:affiliation>
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<mods:affiliation id="186EE301530C3301A51022BF7E766D8C">Department of Biology, Faculty of Art and Science, Burdur Mehmet Akif Ersoy University, Burdur 15030, Turkey. alikamanli 06 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9118 - 4591</mods:affiliation>
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<taxonomicName id="6ABB8985FFF2FFE21987FDC6C9235C7A" ID-CoL="4NZTW" authority="Miers, 1884" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,609,582,609]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
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<emphasis id="9FCF2E14FFF2FFE21987FDC6CAF15C7B" bold="true" box="[151,435,582,609]" italics="true" pageId="4" pageNumber="381">Pseudolitochira integra</emphasis>
(
<bibRefCitation id="C92A8FF7FFF2FFE218D3FDC6C91B5C7A" author="Miers, E. J." box="[451,601,582,608]" pageId="4" pageNumber="381" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
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<paragraph id="AD04F206FFF2FFE21987FDEBCA505C9F" blockId="4.[151,609,582,645]" box="[151,274,618,645]" pageId="4" pageNumber="381">
(
<figureCitation id="3580EE83FFF2FFE2198FFDEBCA485C9F" box="[159,266,618,645]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,1324,1348]" captionStartId-1="5.[151,250,1629,1653]" captionStartId-2="6.[151,250,1927,1951]" captionStartId-3="7.[151,250,1772,1796]" captionTargetBox-0="[164,1416,181,1289]" captionTargetBox-1="[151,1436,181,1606]" captionTargetBox-2="[151,1436,381,1904]" captionTargetBox-3="[151,1436,181,1749]" captionTargetId-0="figure-18@3.[151,1436,181,1300]" captionTargetId-1="figure-18@5.[151,1436,181,1606]" captionTargetId-2="figure-93@6.[151,1436,381,1904]" captionTargetId-3="figure-18@7.[151,1436,181,1749]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, overall dorsal view; B, buccal cavity and frontal regions; C, outer view of left chela. After Miers (1884: pl. 8 figs. C, c, c)." captionText-1="FIGURE 3. Pseudolitochira integra (Miers, 1884), holotype female (4.9 × 3.4 mm) (NHM 1882.24), Seychelles. A, overall dorsal view; B, left fourth ambulatory leg; C, left ambulatory propodus and dactylus; D, sternopleonal cavity and vulvae." captionText-2="FIGURE 4. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, overall dorsal view; B, ventral view of cephalothorax." captionText-3="FIGURE 5. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, dorsal view of carapace; B, frontal view of cephalothorax; C, outer view of right chela; D, buccal cavity, third maxillipeds and thoracic sternites" figureDoi-0="http://doi.org/10.5281/zenodo.4749241" figureDoi-1="http://doi.org/10.5281/zenodo.4749245" figureDoi-2="http://doi.org/10.5281/zenodo.4749247" figureDoi-3="http://doi.org/10.5281/zenodo.4749251" httpUri-0="https://zenodo.org/record/4749241/files/figure.png" httpUri-1="https://zenodo.org/record/4749245/files/figure.png" httpUri-2="https://zenodo.org/record/4749247/files/figure.png" httpUri-3="https://zenodo.org/record/4749251/files/figure.png" pageId="4" pageNumber="381">Figs. 25</figureCitation>
)
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<treatmentCitationGroup id="8DABD528FFF2FFE21987FD2FCFDA5CDD" box="[151,1176,687,711]" pageId="4" pageNumber="381">
<treatmentCitation id="2C1AD417FFF2FFE21987FD2FC9595CDD" author="Miers, E. J." box="[151,539,687,711]" page="543" pageId="4" pageNumber="381" year="1884">
<taxonomicName id="6ABB8985FFF2FFE21987FD2FC9595CDD" authority="Miers, 1884: 543" authorityName="Miers" authorityPageNumber="543" authorityYear="1884" box="[151,539,687,711]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE21987FD2FCA265CDD" box="[151,356,687,711]" italics="true" pageId="4" pageNumber="381">Carcinoplax integra</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE2187BFD2FC9595CDD" author="Miers, E. J." box="[363,539,687,711]" pageId="4" pageNumber="381" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884: 543</bibRefCitation>
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, pl. 48, fig. C, c; 1886: 233 (list);
<treatmentCitation id="2C1AD417FFF2FFE21A64FD2FCF5A5CDD" author="Ward, M." box="[884,1048,687,711]" page="101" pageId="4" pageNumber="381" year="1942">
<bibRefCitation id="C92A8FF7FFF2FFE21A64FD2FCF5A5CDD" author="Ward, M." box="[884,1048,687,711]" pageId="4" pageNumber="381" pagination="49 - 113" refId="ref9388" refString="Ward, M. (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2), 49 - 113." type="journal article" year="1942">Ward 1942: 101</bibRefCitation>
</treatmentCitation>
, pl. 6, fig. 5.
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</paragraph>
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<treatmentCitationGroup id="8DABD528FFF2FFE21987FD4EC98A5CFC" box="[151,712,718,742]" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FD4ECA125CFF" box="[151,336,718,742]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE21987FD4ECA125CFF" box="[151,336,718,742]" italics="true" pageId="4" pageNumber="381">Litocheira integra</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE21879FD4EC9535CFC" author="Tesch, J. J." box="[361,529,718,742]" page="165" pageId="4" pageNumber="381" year="1918">
<bibRefCitation id="C92A8FF7FFF2FFE21879FD4EC9535CFC" author="Tesch, J. J." box="[361,529,718,742]" pageId="4" pageNumber="381" pagination="149 - 295" refId="ref9056" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918: 165</bibRefCitation>
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(key, discussion).
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</paragraph>
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<treatmentCitationGroup id="8DABD528FFF2FFE21987FD6CC93D5D1E" box="[151,639,748,773]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987FD6CCADC5D1E" box="[151,414,748,772]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FD6CCAC15D1E" box="[151,387,748,772]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integer">Pseudolitochira integer</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE2188EFD6DC9005D1E" author="Serene, R." box="[414,578,748,773]" page="86" pageId="4" pageNumber="381" year="1968">
<bibRefCitation id="C92A8FF7FFF2FFE2188EFD6DC9005D1E" author="Serene, R." box="[414,578,748,773]" pageId="4" pageNumber="381" pagination="33 - 120" refId="ref8843" refString="Serene, R. (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. No. 1. Singapore National Academy of Science, Singapore, pp. 33 - 120." type="book chapter" year="1968">Serène 1968: 86</bibRefCitation>
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(list).
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</paragraph>
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<treatmentCitationGroup id="8DABD528FFF2FFE21987FC8BC93F5D58" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FC8BCACB5D39" ID-CoL="4NZTW" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,393,779,803]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE21987FC8BCACB5D39" box="[151,393,779,803]" italics="true" pageId="4" pageNumber="381">Pseudolitochira integra</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE218B3FC8BC9195D39" author="Turkay, M." box="[419,603,779,803]" page="128" pageId="4" pageNumber="381" year="1975">
<bibRefCitation id="C92A8FF7FFF2FFE218B3FC8BC9195D39" author="Turkay, M." box="[419,603,779,803]" pageId="4" pageNumber="381" pagination="103 - 132" refId="ref9164" refString="Turkay, M. (1975) Zur Kenntnis der Gattung Euchirograpsus mit Bemerkungen zu Brachygrapsus und Litocheira (Crustacea: Decapoda). Senckenbergiana Biologica, 56 (1 / 3), 103 - 132." type="journal article" year="1975">Türkay 1975: 128</bibRefCitation>
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, 12 (list);
<treatmentCitation id="2C1AD417FFF2FFE21BD4FC8CC8045D39" author="Ng, P. K. L." box="[708,838,779,804]" page="79" pageId="4" pageNumber="381" year="1987">
<bibRefCitation id="C92A8FF7FFF2FFE21BD4FC8CC8045D39" author="Ng, P. K. L." box="[708,838,779,804]" pageId="4" pageNumber="381" pagination="69 - 111" refId="ref7912" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111." type="journal article" year="1987">Ng 1987: 79</bibRefCitation>
</treatmentCitation>
, 97 (list);
<treatmentCitation id="2C1AD417FFF2FFE21ABEFC8CCF365D39" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[942,1140,779,804]" page="18" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21ABEFC8CCF365D39" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[942,1140,779,804]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 18</bibRefCitation>
</treatmentCitation>
(discussion);
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<bibRefCitation id="C92A8FF7FFF2FFE21C13FC8CCBAF5D58" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." pageId="4" pageNumber="381" pagination="1 - 286" refId="ref8031" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="9FCF2E14FFF2FFE21C39FC8CCE1E5D39" box="[1321,1372,779,803]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
2008: 144
</bibRefCitation>
</treatmentCitation>
(list);
<treatmentCitation id="2C1AD417FFF2FFE21821FCAACAB75D58" author="Ng, P. K. L. &amp; Lin, C. - W. &amp; Ho, P. - H." box="[305,501,810,834]" page="144" pageId="4" pageNumber="381" year="2018">
<bibRefCitation id="C92A8FF7FFF2FFE21821FCAACAB75D58" author="Ng, P. K. L. &amp; Lin, C. - W. &amp; Ho, P. - H." box="[305,501,810,834]" pageId="4" pageNumber="381" pagination="1 - 17" refId="ref8261" refString="Ng, P. K. L., Lin, C. - W. &amp; Ho, P. - H. (2018) On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on the genus Heteropilumnus De Man, 1895 (Crustacea: Brachyura). Zoological Studies, 57 (12), 1 - 17. https: // doi. org / 10.6620 / ZS. 2018.57 - 12" type="journal article" year="2018">
Ng
<emphasis id="9FCF2E14FFF2FFE21846FCABCACB5D58" box="[342,393,810,834]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
2018: 144
</bibRefCitation>
</treatmentCitation>
(discussion).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FCC8C9655D65" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FCC8C9655D65" pageId="4" pageNumber="381">
<treatmentCitation id="2C1AD417FFF2FFE21987FCC8C9365D7A" author="Lanchester, W. F." box="[151,628,840,865]" page="750" pageId="4" pageNumber="381" year="1900">
<taxonomicName id="6ABB8985FFF2FFE21987FCC8C9365D7A" authority="Lanchester, 1900: 750" authorityName="Lanchester" authorityPageNumber="750" authorityYear="1900" box="[151,628,840,865]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFF2FFE21987FCC8CACA5D7A" box="[151,392,840,864]" italics="true" pageId="4" pageNumber="381">Carcinoplax subinteger</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE2189FFCC8C9365D7A" author="Lanchester, W. F." box="[399,628,840,865]" pageId="4" pageNumber="381" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester, 1900: 750</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 46, fig. 9a (unnecessary replacement name for
<taxonomicName id="6ABB8985FFF2FFE21D96FCC8CBBF5D65" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="9FCF2E14FFF2FFE21D96FCC8CE115D7A" box="[1158,1363,840,864]" italics="true" pageId="4" pageNumber="381">Carcinoplax integer</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21C4BFCC8CBBF5D65" author="Miers, E. J." pageId="4" pageNumber="381" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE2183EFCE8CAB65D65" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[302,500,871,896]" page="15" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE2183EFCE8CAB65D65" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[302,500,871,896]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 15</bibRefCitation>
</treatmentCitation>
, 17).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FC06CE735D84" blockId="4.[151,1437,687,1601]" box="[151,1329,902,926]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FC06CE735D84" box="[151,1329,902,926]" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FC06CBFF5D87" box="[151,189,902,925]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<taxonomicName id="6ABB8985FFF2FFE219D4FC06CAD25D87" authorityName="Miers" authorityYear="1884" box="[196,400,902,926]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="9FCF2E14FFF2FFE219D4FC06CAD25D87" box="[196,400,902,926]" italics="true" pageId="4" pageNumber="381">Carcinoplax integer</emphasis>
</taxonomicName>
—De
<treatmentCitation id="2C1AD417FFF2FFE218DDFC06C9185D84" author="Man, J. G. De" box="[461,602,902,926]" page="93" pageId="4" pageNumber="381" year="1887">
<bibRefCitation id="C92A8FF7FFF2FFE218DDFC06C9185D84" author="Man, J. G. De" box="[461,602,902,926]" pageId="4" pageNumber="381" pagination="65 - 128" refId="ref7189" refString="Man, J. G. De (1887) Part II. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22 (137), 65 - 128, pls. 4 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01455. x" type="journal article" year="1887">Man 1887: 93</bibRefCitation>
</treatmentCitation>
(=
<taxonomicName id="6ABB8985FFF2FFE21B6EFC06CF6F5D84" ID-CoL="3LC49" authority="Ng &amp; Tan, 1988" authorityName="Ng &amp; Tan" authorityYear="1988" box="[638,1069,902,926]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFF2FFE21B6EFC06C8C25D84" box="[638,896,902,926]" italics="true" pageId="4" pageNumber="381">Heteropilumnus holthuisi</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21A96FC06CF6F5D84" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[902,1069,902,926]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan, 1988</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21D4EFC06CE665D84" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1118,1316,902,926]" page="14" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21D4EFC06CE665D84" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1118,1316,902,926]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 14</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FC25CA725DC1" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FC25CA725DC1" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FC25CBFF5DA6" box="[151,189,933,956]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<emphasis id="9FCF2E14FFF2FFE219D5FC24CAC95DA6" box="[197,395,932,956]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE219D5FC24CA325DA6" ID-CoL="3VM2M" authorityName="Miers" authorityYear="1915" box="[197,368,932,956]" class="Malacostraca" family="Pilumnidae" genus="Litochira" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Litochira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE2189BFC24C9055DA7" author="Alcock, A." box="[395,583,932,957]" page="314" pageId="4" pageNumber="381" year="1900">
<bibRefCitation id="C92A8FF7FFF2FFE2189BFC24C9055DA7" author="Alcock, A." box="[395,583,932,957]" pageId="4" pageNumber="381" pagination="279 - 456" refId="ref5619" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa, or Grapsoidea. Journal on the Asiatic Society of Bengal, 69, 2, 279 - 456." type="journal article" year="1900">Alcock 1900: 314</bibRefCitation>
</treatmentCitation>
; De
<treatmentCitation id="2C1AD417FFF2FFE21B68FC25C9BC5DA7" author="Man, J. G. De" box="[632,766,933,957]" page="9" pageId="4" pageNumber="381" year="1929">
<bibRefCitation id="C92A8FF7FFF2FFE21B68FC25C9BC5DA7" author="Man, J. G. De" box="[632,766,933,957]" pageId="4" pageNumber="381" pagination="1 - 26" refId="ref7363" refString="Man, J. G. De (1929) On a collection of decapod and stomatopod Crustacea from Pulau Berhala, an islet situated in the Straits of Malacca. Bijdragen tot de Dierkunde, 26, 1 - 26, pls. I - III. https: // doi. org / 10.1163 / 26660644 - 02601001" type="journal article" year="1929">Man 1929: 9</bibRefCitation>
</treatmentCitation>
, 13 (=
<taxonomicName id="6ABB8985FFF2FFE21A5AFC24CE435DA7" ID-CoL="3LC49" authority="Ng &amp; Tan, 1988" authorityName="Ng &amp; Tan" authorityYear="1988" box="[842,1281,932,957]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFF2FFE21A5AFC24CF0C5DA6" box="[842,1102,932,956]" italics="true" pageId="4" pageNumber="381">Heteropilumnus holthuisi</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21D45FC25CE435DA7" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1109,1281,933,957]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan, 1988</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21C25FC25CA615DC1" author="Ng, P. K. L. &amp; Tan, L. W. H." page="14" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21C25FC25CA615DC1" author="Ng, P. K. L. &amp; Tan, L. W. H." pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 14</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FC62CA615A03" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FC62CA615A03" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FC62CBFF5DE3" box="[151,189,994,1017]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<emphasis id="9FCF2E14FFF2FFE219D6FC62CADA5DE3" box="[198,408,994,1018]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE219D6FC62CA3F5DE3" box="[198,381,994,1018]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Litocheira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE21887FC62C90E5DE0" author="Laurie, R. D." box="[407,588,994,1018]" page="464" pageId="4" pageNumber="381" year="1915">
<bibRefCitation id="C92A8FF7FFF2FFE21887FC62C90E5DE0" author="Laurie, R. D." box="[407,588,994,1018]" pageId="4" pageNumber="381" pagination="407 - 475" refId="ref6904" refString="Laurie, R. D. (1915) XXI. On the Brachyura. In: Reports of the Marine Biology of the Sudanese Red Sea, from Collections made by Cyril Crossland, M. A., B. Sc., F. Z. S. Communicated, with an Introduction, by Prof. W. A. Herdman, D. Sc., F. R. S., F. L. S. Journal of the Linnean Society, Zoology, London, 31 (209), 407 - 475, figs. 1 - 5, pls. 42 - 45. https: // doi. org / 10.1111 / j. 1096 - 3642.1915. tb 00694. x" type="journal article" year="1915">Laurie 1915: 464</bibRefCitation>
</treatmentCitation>
, pl. 45, fig. 2 (=
<taxonomicName id="6ABB8985FFF2FFE21A12FC62CE515DE0" ID-CoL="3LC4G" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[770,1299,994,1018]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="9FCF2E14FFF2FFE21A12FC62C8B45DE0" box="[770,1014,994,1018]" italics="true" pageId="4" pageNumber="381">Heteropilumnus setosus</emphasis>
(A. Milne-Edwards, 1873)
</taxonomicName>
; see De
<treatmentCitation id="2C1AD417FFF2FFE21C7EFC62CA575A03" author="Man, J. G. De" page="8" pageId="4" pageNumber="381" year="1929">
<bibRefCitation id="C92A8FF7FFF2FFE21C7EFC62CA575A03" author="Man, J. G. De" pageId="4" pageNumber="381" pagination="1 - 26" refId="ref7363" refString="Man, J. G. De (1929) On a collection of decapod and stomatopod Crustacea from Pulau Berhala, an islet situated in the Straits of Malacca. Bijdragen tot de Dierkunde, 26, 1 - 26, pls. I - III. https: // doi. org / 10.1163 / 26660644 - 02601001" type="journal article" year="1929">Man 1929: 8</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FBA0CBB25A4C" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FBA0CBB25A4C" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FBA0CBFF5A2D" box="[151,189,1056,1079]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<taxonomicName id="6ABB8985FFF2FFE219D6FB9FCAE65A2D" box="[198,420,1055,1079]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="subintegra">
<emphasis id="9FCF2E14FFF2FFE219D6FB9FCAE65A2D" box="[198,420,1055,1079]" italics="true" pageId="4" pageNumber="381">Litocheira subintegra</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE218AEFB9FC93F5A2D" author="Gordon, I." box="[446,637,1055,1079]" page="549" pageId="4" pageNumber="381" year="1931">
<bibRefCitation id="C92A8FF7FFF2FFE218AEFB9FC93F5A2D" author="Gordon, I." box="[446,637,1055,1079]" pageId="4" pageNumber="381" pagination="525 - 558" refId="ref6261" refString="Gordon, I. (1931) Brachyura from the coast of China. Journal of the Linnean Society of London, Zoology, 37, 525 - 558. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 02365. x" type="journal article" year="1931">Gordon 1931: 549</bibRefCitation>
</treatmentCitation>
, fig. 25b (=
<taxonomicName id="6ABB8985FFF2FFE21A13FB9FCFFE5A2D" ID-CoL="3LC49" authority="Ng &amp; Tan, 1988" authorityName="Ng &amp; Tan" authorityYear="1988" box="[771,1212,1055,1080]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFF2FFE21A13FB9FCF455A2D" box="[771,1031,1055,1079]" italics="true" pageId="4" pageNumber="381">Heteropilumnus holthuisi</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21D1FFBA0CFFE5A2D" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1039,1212,1055,1080]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan, 1988</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21DE1FBA0CBA15A4C" author="Ng, P. K. L. &amp; Tan, L. W. H." page="14" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21DE1FBA0CBA15A4C" author="Ng, P. K. L. &amp; Tan, L. W. H." pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 14</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FBDDCA725A89" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FBDDCA725A89" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FBDDCBFF5A6E" box="[151,189,1117,1140]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<taxonomicName id="6ABB8985FFF2FFE219D3FBDCCA945A6E" box="[195,470,1116,1140]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFF2FFE219D3FBDCCA945A6E" box="[195,470,1116,1140]" italics="true" pageId="4" pageNumber="381">Heteropilumnus subinteger</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE218E1FBDCC9CA5A6F" author="Balss, H." box="[497,648,1116,1141]" page="44" pageId="4" pageNumber="381" year="1933">
<bibRefCitation id="C92A8FF7FFF2FFE218E1FBDCC9CA5A6F" author="Balss, H." box="[497,648,1116,1141]" pageId="4" pageNumber="381" pagination="1 - 47" refId="ref5712" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus und verwandter Gattungen. Capita Zoologica, 4 (3), 1 - 47, pls. 1 - 7." type="journal article" year="1933">Balss 1933: 44</bibRefCitation>
</treatmentCitation>
, pl. 7, figs. 32, 33 (=
<taxonomicName id="6ABB8985FFF2FFE21A4DFBDCCE4B5A6F" ID-CoL="3LC49" authority="Ng &amp; Tan, 1988" authorityName="Ng &amp; Tan" authorityYear="1988" box="[861,1289,1116,1141]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFF2FFE21A4DFBDCCF1C5A6E" box="[861,1118,1116,1140]" italics="true" pageId="4" pageNumber="381">Heteropilumnus holthuisi</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21D74FBDDCE4B5A6F" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1124,1289,1117,1141]" pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan, 1988</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21C29FBDDCA615A89" author="Ng, P. K. L. &amp; Tan, L. W. H." page="14" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21C29FBDDCA615A89" author="Ng, P. K. L. &amp; Tan, L. W. H." pageId="4" pageNumber="381" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 14</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FB1ACA375ACB" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FB1ACA375ACB" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FB1ACBFF5AAB" box="[151,189,1178,1201]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<emphasis id="9FCF2E14FFF2FFE219D4FB1ACA8F5AAB" box="[196,461,1178,1202]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE219D4FB1ACAF05AAB" ID-CoL="4NZTW" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[196,434,1178,1202]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolitochira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE218DDFB1AC9375AA8" author="Naruse, T." box="[461,629,1178,1202]" page="32" pageId="4" pageNumber="381" year="2010">
<bibRefCitation id="C92A8FF7FFF2FFE218DDFB1AC9375AA8" author="Naruse, T." box="[461,629,1178,1202]" pageId="4" pageNumber="381" pagination="1 - 72" refId="ref7864" refString="Naruse, T. (2010) Catalogue of Crustacean Specimens deposited in the University Museum (Fujukan), University of the Ryukyus. Catalogue of Materials Deposited in the University Museum (Fujukan), University of the Ryukyus, 3, 1 - 72." type="journal article" year="2010">Naruse 2010: 32</bibRefCitation>
</treatmentCitation>
(part: RUMF-ZC-973) (=
<taxonomicName id="6ABB8985FFF2FFE21A97FB1ACFB65AA8" ID-CoL="4HVY3" authority="Rathbun, 1911" authorityName="Rathbun" authorityYear="1911" box="[903,1268,1178,1202]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="turgidulus">
<emphasis id="9FCF2E14FFF2FFE21A97FB1ACF165AA8" box="[903,1108,1178,1202]" italics="true" pageId="4" pageNumber="381">Pilumnus turgidulus</emphasis>
<bibRefCitation id="C92A8FF7FFF2FFE21D4CFB1ACFB65AA8" author="Rathbun, M. J." box="[1116,1268,1178,1202]" pageId="4" pageNumber="381" pagination="191 - 261" refId="ref8631" refString="Rathbun, M. J. (1911) Marine Brachyura. In: The Percy Sladen Trust expedition to the Indian Ocean in 1905, Under the leadership of Mr. J. Stanley Gardiner. Vol. III. No. XI. Transactions of the Linnean Society of London, Zoology, Series 2 (Zoology), 14 (2), 191 - 261, figs. 1 &amp; 2, pls. 15 - 20. https: // doi. org / 10.1111 / j. 1096 - 3642.1911. tb 00084. x" type="journal article" year="1911">Rathbun, 1911</bibRefCitation>
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21C36FB1ACA605ACB" author="Maenosono, T." page="33" pageId="4" pageNumber="381" year="2019">
<bibRefCitation id="C92A8FF7FFF2FFE21C36FB1ACA605ACB" author="Maenosono, T." pageId="4" pageNumber="381" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono 2019: 33</bibRefCitation>
</treatmentCitation>
, tab. 1).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FB58CA725B14" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FB58CA725B14" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FB58CBFF5AF5" box="[151,189,1240,1263]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<emphasis id="9FCF2E14FFF2FFE219D2FB57CA8A5AF5" box="[194,456,1239,1263]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE219D2FB57CAEF5AF5" ID-CoL="4NZTW" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[194,429,1239,1263]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolitochira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE218D8FB57C93C5AF5" author="Machida, Y." box="[456,638,1239,1263]" page="53" pageId="4" pageNumber="381" year="2017">
<bibRefCitation id="C92A8FF7FFF2FFE218D8FB57C93C5AF5" author="Machida, Y." box="[456,638,1239,1263]" pageId="4" pageNumber="381" pagination="52 - 57" refId="ref7109" refString="Machida, Y. (2017) Three rare species of crab obtained from the gravel intertidal zone of Uchinoura Bay. Shikoku Natural History Science Research, 10, 52 - 57." type="journal article" year="2017">Machida 2017: 53</bibRefCitation>
</treatmentCitation>
, figs. 2, 3 (=
<taxonomicName id="6ABB8985FFF2FFE21BEDFB57CFBB5AF5" ID-CoL="5CYS4" authority="(Takeda &amp; Miyake, 1969)" baseAuthorityName="Takeda &amp; Miyake" baseAuthorityYear="1969" box="[765,1273,1239,1263]" class="Malacostraca" family="Pilumnidae" genus="Zehntneriana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="amakusae">
<emphasis id="9FCF2E14FFF2FFE21BEDFB57C8AD5AF5" box="[765,1007,1239,1263]" italics="true" pageId="4" pageNumber="381">Zehntneriana amakusae</emphasis>
(
<bibRefCitation id="C92A8FF7FFF2FFE21AEAFB57CFB25AF5" author="Takeda, M. &amp; Miyake, S." box="[1018,1264,1239,1263]" pageId="4" pageNumber="381" pagination="9 - 15" refId="ref8954" refString="Takeda, M. &amp; Miyake, S. (1969) A new species of the family Goneplacidae (Crustacea, Brachyura) from the Amakusa Islands, Kyushu. OHMU, Zoological Laboratory, Faculty of Agriculture, Kyushu University, 2 (1), 9 - 15." type="journal article" year="1969">Takeda &amp; Miyake, 1969</bibRefCitation>
)
</taxonomicName>
; see
<treatmentCitation id="2C1AD417FFF2FFE21C36FB58CA615B14" author="Maenosono, T." page="33" pageId="4" pageNumber="381" year="2019">
<bibRefCitation id="C92A8FF7FFF2FFE21C36FB58CA615B14" author="Maenosono, T." pageId="4" pageNumber="381" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono 2019: 33</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FA95C84F5B36" blockId="4.[151,1437,687,1601]" box="[151,781,1300,1325]" pageId="4" pageNumber="381">
<collectingRegion id="6F7F3CE4FFF2FFE21987FA95CBFF5B36" box="[151,189,1301,1324]" country="Maldives" name="Uthuru" pageId="4" pageNumber="381">Not</collectingRegion>
<taxonomicName id="6ABB8985FFF2FFE219D4FA94C90A5B37" ID-CoL="4NZTW" authority="(Miers, 1884)" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[196,584,1300,1325]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE219D4FA94CAF75B36" box="[196,437,1300,1324]" italics="true" pageId="4" pageNumber="381">Pseudolitochira integra</emphasis>
(
<bibRefCitation id="C92A8FF7FFF2FFE218D2FA94C9035B37" author="Miers, E. J." box="[450,577,1300,1325]" pageId="4" pageNumber="381" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
</taxonomicName>
, identity uncertain:
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FAB3C9F65B51" blockId="4.[151,1437,687,1601]" box="[151,692,1331,1355]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FAB3C9F65B51" box="[151,692,1331,1355]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987FAB3CA3D5B51" box="[151,383,1331,1355]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FAB3CA265B51" box="[151,356,1331,1355]" class="Malacostraca" family="Pilumnidae" genus="Litochira" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="subintegra">Litochira subintegra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE2186FFAB3C96A5B51" author="Tesch, J. J." box="[383,552,1331,1355]" page="175" pageId="4" pageNumber="381" year="1918">
<bibRefCitation id="C92A8FF7FFF2FFE2186FFAB3C96A5B51" author="Tesch, J. J." box="[383,552,1331,1355]" pageId="4" pageNumber="381" pagination="149 - 295" refId="ref9056" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918: 175</bibRefCitation>
</treatmentCitation>
, pl. 16, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FAD2CA415B93" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FAD2CA415B93" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987FAD2CA2A5B73" box="[151,360,1362,1386]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FAD2CA0E5B73" box="[151,332,1362,1386]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Litocheira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE21877FAD2C9005B70" author="Borradaile, L. A." box="[359,578,1362,1386]" page="430" pageId="4" pageNumber="381" year="1903">
<bibRefCitation id="C92A8FF7FFF2FFE21877FAD2C9005B70" author="Borradaile, L. A." box="[359,578,1362,1386]" pageId="4" pageNumber="381" pagination="424 - 443" refId="ref5747" refString="Borradaile, L. A. (1903) Marine crustaceans. IV. Some remarks on the classification of the crabs. V. The crabs of the catometope families. VI. The sand crabs (Oxystomata). VII. The barnacles (Cirripedia). In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, vol. 1, part 4. Cambridge University Press, Cambridge, pp. 424 - 443, pl. XXII," type="book chapter" year="1903">Borradaile 1903: 430</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2C1AD417FFF2FFE21B5DFAD2C84C5B70" author="Bouvier, E. L." box="[589,782,1362,1386]" page="296" pageId="4" pageNumber="381" year="1915">
<bibRefCitation id="C92A8FF7FFF2FFE21B5DFAD2C84C5B70" author="Bouvier, E. L." box="[589,782,1362,1386]" pageId="4" pageNumber="381" pagination="178 - 318" refId="ref5866" refString="Bouvier, E. L. (1915) Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 7 e Serie, 48 (3), 178 - 318, pls. IV - VII." type="journal article" year="1915">Bouvier 1915: 296</bibRefCitation>
</treatmentCitation>
, text-fig. 34; De
<treatmentCitation id="2C1AD417FFF2FFE21AACFAD2CF0F5B70" author="Man, J. G. De" box="[956,1101,1362,1386]" page="13" pageId="4" pageNumber="381" year="1929">
<bibRefCitation id="C92A8FF7FFF2FFE21AACFAD2CF0F5B70" author="Man, J. G. De" box="[956,1101,1362,1386]" pageId="4" pageNumber="381" pagination="1 - 26" refId="ref7363" refString="Man, J. G. De (1929) On a collection of decapod and stomatopod Crustacea from Pulau Berhala, an islet situated in the Straits of Malacca. Bijdragen tot de Dierkunde, 26, 1 - 26, pls. I - III. https: // doi. org / 10.1163 / 26660644 - 02601001" type="journal article" year="1929">Man 1929: 13</bibRefCitation>
</treatmentCitation>
, pl. 2, fig. 4; T.
<treatmentCitation id="2C1AD417FFF2FFE21DE0FAD2CEDA5B70" author="Sakai, T." box="[1264,1432,1362,1386]" page="108" pageId="4" pageNumber="381" year="1955">
<bibRefCitation id="C92A8FF7FFF2FFE21DE0FAD2CEDA5B70" author="Sakai, T." box="[1264,1432,1362,1386]" pageId="4" pageNumber="381" pagination="106 - 113" refId="ref8754" refString="Sakai, T. (1955) On some rare species of crabs from Japan. Bulletin of the Biogeographical Society of Japan, 16 - 19, 106 - 113." type="journal article" year="1955">Sakai 1955: 108</bibRefCitation>
</treatmentCitation>
, fig. 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FA0FC8B95BDC" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FA0FC8B95BDC" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987FA0FCACA5BBD" box="[151,392,1423,1447]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE21987FA0FCACA5BBD" box="[151,392,1423,1447]" italics="true" pageId="4" pageNumber="381">Heteropilumnus integra</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE218B2FA0FC91A5BBD" author="Holthuis, L. B." box="[418,600,1423,1447]" page="22" pageId="4" pageNumber="381" year="1953">
<bibRefCitation id="C92A8FF7FFF2FFE218B2FA0FC91A5BBD" author="Holthuis, L. B." box="[418,600,1423,1447]" pageId="4" pageNumber="381" pagination="1 - 66" refId="ref6380" refString="Holthuis, L. B. (1953) Enumeration of the decapod and stomatopod Crustacea from Pacific coral islands. Atoll Research Bulletin, 24, 1 - 66, 2 maps. https: // doi. org / 10.5479 / si. 00775630.24.1" type="book chapter" year="1953">Holthuis 1953: 22</bibRefCitation>
</treatmentCitation>
; T.
<treatmentCitation id="2C1AD417FFF2FFE21B6AFA0FC85C5BBD" author="Sakai, T." box="[634,798,1423,1447]" page="493" pageId="4" pageNumber="381" year="1976">
<bibRefCitation id="C92A8FF7FFF2FFE21B6AFA0FC85C5BBD" author="Sakai, T." box="[634,798,1423,1447]" pageId="4" pageNumber="381" refId="ref8786" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1 - 3. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp., 251 pls. (plates volume)." type="book" year="1976">Sakai 1976: 493</bibRefCitation>
</treatmentCitation>
, text-fig. 264a, 304;
<treatmentCitation id="2C1AD417FFF2FFE21AFDFA0FCFE55BBD" author="Miyake, S." box="[1005,1191,1423,1447]" page="233" pageId="4" pageNumber="381" year="1983">
<bibRefCitation id="C92A8FF7FFF2FFE21AFDFA0FCFE55BBD" author="Miyake, S." box="[1005,1191,1423,1447]" pageId="4" pageNumber="381" refId="ref7781" refString="Miyake, S. (1983) Brachyura (Crabs). Japanese crustacean decapods and stomatopods in Color. Vol. 2. 1 st Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered fig." type="book" year="1983">Miyake 1983: 233</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2C1AD417FFF2FFE21DA0FA0FCBAD5BDC" author="Nagai, S. &amp; Nomura, K." page="112" pageId="4" pageNumber="381" year="1988">
<bibRefCitation id="C92A8FF7FFF2FFE21DA0FA0FCBAD5BDC" author="Nagai, S. &amp; Nomura, K." pageId="4" pageNumber="381" refId="ref7826" refString="Nagai, S. &amp; Nomura, K. (1988) Crustacea (Brachyura). The guide book of marine organisms of Okinawa. Vol. 7. Shinsei Tosho, Okinawa, 250 pp." type="book" year="1988">Nagai &amp; Nomura 1988: 112</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2C1AD417FFF2FFE219EAFA2ECA3A5BDC" author="Ito, J." box="[250,376,1454,1478]" page="46" pageId="4" pageNumber="381" year="1992">
<bibRefCitation id="C92A8FF7FFF2FFE219EAFA2ECA3A5BDC" author="Ito, J." box="[250,376,1454,1478]" pageId="4" pageNumber="381" pagination="39 - 52" refId="ref6661" refString="Ito, J. (1992) Local marine animals 18. Fukui City Museum of Natural History Research Report, 39, 39 - 52." type="journal article" year="1992">Ito 1992: 46</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2C1AD417FFF2FFE21893FA2EC9335BDC" author="Nomura, K. &amp; Nagai, S. &amp; Asakura, A. &amp; Komai, T." box="[387,625,1454,1478]" page="17" pageId="4" pageNumber="381" year="1996">
<bibRefCitation id="C92A8FF7FFF2FFE21893FA2EC9335BDC" author="Nomura, K. &amp; Nagai, S. &amp; Asakura, A. &amp; Komai, T." box="[387,625,1454,1478]" pageId="4" pageNumber="381" pagination="7 - 21" refId="ref8406" refString="Nomura, K., Nagai, S., Asakura, A. &amp; Komai, T. (1996) A preliminary list of shallow water decapod Crustacea in the Kerama Group, the Ryukyu Archipelago. Bulletin of the Biogeographical Society of Japan, 51, 7 - 21." type="journal article" year="1996">
Nomura
<emphasis id="9FCF2E14FFF2FFE218CCFA2FC94D5BDC" box="[476,527,1454,1478]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
1996: 17
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2C1AD417FFF2FFE21B6CFA2EC84E5BDC" author="Jeng, M. - S." box="[636,780,1454,1478]" page="16" pageId="4" pageNumber="381" year="1997">
<bibRefCitation id="C92A8FF7FFF2FFE21B6CFA2EC84E5BDC" author="Jeng, M. - S." box="[636,780,1454,1478]" pageId="4" pageNumber="381" refId="ref6688" refString="Jeng, M. - S. (1997) Studies on the land and aquatic decapod crustacean fauna of the Kenting National Park (11) - Communities of decapod crustaceans around the sea. Kenting National Park, Ministry of the Interior, Pingtung, ii + 66 pp. [in Chinese]" type="book" year="1997">Jeng 1997: 16</bibRefCitation>
</treatmentCitation>
, 22; K.
<treatmentCitation id="2C1AD417FFF2FFE21A4CFA2EC8B55BDC" author="Sakai, K." box="[860,1015,1454,1478]" page="20" pageId="4" pageNumber="381" year="2003">
<bibRefCitation id="C92A8FF7FFF2FFE21A4CFA2EC8B55BDC" author="Sakai, K." box="[860,1015,1454,1478]" pageId="4" pageNumber="381" pagination="13 - 30" refId="ref8732" refString="Sakai, K. (2003) Standard Japanese names for Japanese crabs. Taxon, 15, 13 - 30." type="journal article" year="2003">Sakai 2003: 20</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987FA4CC8FA5819" blockId="4.[151,1437,687,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987FA4CC8FA5819" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987FA4CCAE25BFE" box="[151,416,1484,1508]" italics="true" pageId="4" pageNumber="381">
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<treatmentCitation id="2C1AD417FFF2FFE2182AFA6BC95F5819" author="Hosie, A. M. &amp; Sampey, A. &amp; Davie P. J. F. &amp; Jones, D. S." box="[314,541,1515,1539]" page="276" pageId="4" pageNumber="381" year="2015">
<bibRefCitation id="C92A8FF7FFF2FFE2182AFA6BC95F5819" author="Hosie, A. M. &amp; Sampey, A. &amp; Davie P. J. F. &amp; Jones, D. S." box="[314,541,1515,1539]" pageId="4" pageNumber="381" pagination="247 - 285" refId="ref6426" refString="Hosie, A. M., Sampey, A., Davie P. J. F. &amp; Jones, D. S. (2015) Kimberley marine biota. Historical data: crustaceans. In: Marine Biodiversity of the Kimberley 1880 s - 2009. Records of the Western Australian Museum, Supplement 84, 247 - 285. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 84.2015.247 - 285" type="journal article" year="2015">
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;
<treatmentCitation id="2C1AD417FFF2FFE21B37FA6CC9A05819" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[551,738,1515,1540]" page="63" pageId="4" pageNumber="381" year="2017">
<bibRefCitation id="C92A8FF7FFF2FFE21B37FA6CC9A05819" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[551,738,1515,1540]" pageId="4" pageNumber="381" pagination="1 - 185" refId="ref8085" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 &amp; 4), 1 - 185. https: // doi. org / 10.6532 / JNTM. 20171270 (3; 4). 01" type="journal article" year="2017">
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<treatmentCitation id="2C1AD417FFF2FFE21BFDFA6CC8F65819" author="Ng, P. K. L. &amp; Lin, C. - W. &amp; Ho, P. - H." box="[749,948,1515,1540]" page="144" pageId="4" pageNumber="381" year="2018">
<bibRefCitation id="C92A8FF7FFF2FFE21BFDFA6CC8F65819" author="Ng, P. K. L. &amp; Lin, C. - W. &amp; Ho, P. - H." box="[749,948,1515,1540]" pageId="4" pageNumber="381" pagination="1 - 17" refId="ref8261" refString="Ng, P. K. L., Lin, C. - W. &amp; Ho, P. - H. (2018) On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on the genus Heteropilumnus De Man, 1895 (Crustacea: Brachyura). Zoological Studies, 57 (12), 1 - 17. https: // doi. org / 10.6620 / ZS. 2018.57 - 12" type="journal article" year="2018">
Ng
<emphasis id="9FCF2E14FFF2FFE21A02FA6CC8075819" box="[786,837,1515,1539]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
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.
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</paragraph>
<paragraph id="AD04F206FFF2FFE21987F98AC9CD5838" blockId="4.[151,1437,687,1601]" box="[151,655,1546,1570]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F98AC9CD5838" box="[151,655,1546,1570]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F98ACADC583B" box="[151,414,1546,1570]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F98ACAC1583B" box="[151,387,1546,1570]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integer">Pseudolitochira integer</taxonomicName>
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<treatmentCitation id="2C1AD417FFF2FFE2188EF98AC9C95838" author="Paulay, G. &amp; Kropp, R. &amp; Ng, P. K. L. &amp; Eldredge, L. G." box="[414,651,1546,1570]" page="498" pageId="4" pageNumber="381" year="2003">
<bibRefCitation id="C92A8FF7FFF2FFE2188EF98AC9C95838" author="Paulay, G. &amp; Kropp, R. &amp; Ng, P. K. L. &amp; Eldredge, L. G." box="[414,651,1546,1570]" pageId="4" pageNumber="381" pagination="456 - 513" refId="ref8459" refString="Paulay, G., Kropp, R., Ng, P. K. L. &amp; Eldredge, L. G. (2003) The crustaceans and pycnogonids of the Mariana Islands. In: Paulay, G. (Ed.), The Marine Biodiversity of Guam and the Marianas. Micronesica, 35 - 36, 456 - 513." type="journal article" year="2003">
Paulay
<emphasis id="9FCF2E14FFF2FFE218F9F98BC95E5838" box="[489,540,1546,1570]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
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</paragraph>
<paragraph id="AD04F206FFF2FFE21987F9A8C891585A" blockId="4.[151,1437,687,1601]" box="[151,979,1576,1601]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F9A8C891585A" box="[151,979,1576,1601]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F9A8CAE9585A" box="[151,427,1576,1600]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F9A8CAD2585A" box="[151,400,1576,1600]" class="Malacostraca" family="Litocheiridae" genus="Pseudolitocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolitocheira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE218BAF9A9C9C9585B" author="Serene, R. &amp; Romimohtarto, K. &amp; Moosa, M. K." box="[426,651,1576,1601]" page="23" pageId="4" pageNumber="381" year="1974">
<bibRefCitation id="C92A8FF7FFF2FFE218BAF9A9C9C9585B" author="Serene, R. &amp; Romimohtarto, K. &amp; Moosa, M. K." box="[426,651,1576,1601]" pageId="4" pageNumber="381" pagination="17 - 26" refId="ref8893" refString="Serene, R., Romimohtarto, K. &amp; Moosa, M. K. (1974) The Hippidea and Brachyura collected by the Rumphius Expedition I. In: Report on the Rumphius Expedition I. (January 6 - February 1, 1973). Oseanologi di Indonesia, 1, pp. 17 - 26." type="journal article" year="1974">
Serène
<emphasis id="9FCF2E14FFF2FFE218E6F9A9C96B585A" box="[502,553,1576,1600]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
1974: 23
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;
<treatmentCitation id="2C1AD417FFF2FFE21B85F9A9C88D585A" author="Marumura, M. &amp; Kosaka, A." box="[661,975,1576,1601]" page="61" pageId="4" pageNumber="381" year="2003">
<bibRefCitation id="C92A8FF7FFF2FFE21B85F9A9C88D585A" author="Marumura, M. &amp; Kosaka, A." box="[661,975,1576,1601]" pageId="4" pageNumber="381" refId="ref7423" refString="Marumura, M. &amp; Kosaka, A. (2003) Catalogue of brachyuran and anomuran crabs collection donated by the late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, Wakayama, 73 pp., 8 pls." type="book" year="2003">Marumura &amp; Kosaka 2003: 61</bibRefCitation>
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<paragraph id="AD04F206FFF2FFE21987F9E6CA5C5864" blockId="4.[151,1436,1638,1846]" box="[151,286,1638,1662]" pageId="4" pageNumber="381">Incerta sedis:</paragraph>
<paragraph id="AD04F206FFF2FFE21987F904CA0A58A1" blockId="4.[151,1436,1638,1846]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F904CA0A58A1" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F904CADD5886" box="[151,415,1668,1692]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F904CAC65886" ID-CoL="4NZTW" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,388,1668,1692]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolitochira integra</taxonomicName>
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<treatmentCitation id="2C1AD417FFF2FFE2188FF905C91E5886" author="Dev Roy, M. K." box="[415,604,1668,1693]" page="60" pageId="4" pageNumber="381" year="2008">
<bibRefCitation id="C92A8FF7FFF2FFE2188FF905C91E5886" author="Dev Roy, M. K." box="[415,604,1668,1693]" pageId="4" pageNumber="381" pagination="1 - 212" refId="ref6108" refString="Dev Roy, M. K. (2008) An annotated checklist of mangrove and coral reef inhabiting brachyuran crabs of India, India. Records of the Zoological Survey of India, Occasional Paper, No. 289, 1 - 212." type="book chapter" year="2008">Dev Roy 2008: 60</bibRefCitation>
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, 168;
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<bibRefCitation id="C92A8FF7FFF2FFE21B89F905C8885887" author="Dev Roy, M. K. &amp; Bhadra, S." box="[665,970,1668,1693]" pageId="4" pageNumber="381" pagination="109 - 269" refId="ref6153" refString="Dev Roy, M. K. &amp; Bhadra, S. (2011) Brachyuran crabs (Crustacea: Decapoda: Brachyura). In: State Fauna Series. 17. Part- 2. Fauna of Tamil Nadu. Zoological Survey of India, Kolkata. Pp. 109 - 269," type="book chapter" year="2011">Dev Roy &amp; Bhadra 2011: 113</bibRefCitation>
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, 186;
<treatmentCitation id="2C1AD417FFF2FFE21D18F905CE725887" author="Dev Roy, M. K. &amp; Nandi, N. C." box="[1032,1328,1668,1693]" page="203" pageId="4" pageNumber="381" year="2012">
<bibRefCitation id="C92A8FF7FFF2FFE21D18F905CE725887" author="Dev Roy, M. K. &amp; Nandi, N. C." box="[1032,1328,1668,1693]" pageId="4" pageNumber="381" pagination="185 - 236" refId="ref6207" refString="Dev Roy, M. K. &amp; Nandi, N. C. (2012) Brachyuran crabs (Crustacea). In: State Fauna Series. 19. Part- 1. Fauna of Andaman and Nicobar Islands. Zoological Survey of India, Kolkata, pp. 185 - 236," type="book chapter" year="2012">Dev Roy &amp; Nandi 2012: 203</bibRefCitation>
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<bibRefCitation id="C92A8FF7FFF2FFE21C2AF904CA0658A1" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." pageId="4" pageNumber="381" pagination="1 - 83" refId="ref9091" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. &amp; Ng, P. K. L. (2018) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi
<emphasis id="9FCF2E14FFF2FFE21C9AF905CBA058A1" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
2018: 60
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.
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</paragraph>
<paragraph id="AD04F206FFF2FFE21987F942C80558C0" blockId="4.[151,1436,1638,1846]" box="[151,839,1730,1754]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F942C80558C0" box="[151,839,1730,1754]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F942CAE958C3" box="[151,427,1730,1754]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F942CAD258C3" box="[151,400,1730,1754]" class="Malacostraca" family="Litocheiridae" genus="Pseudolitocheira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolitocheira integra</taxonomicName>
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<treatmentCitation id="2C1AD417FFF2FFE218BAF942C99E58C0" author="Dev Roy, M. K. &amp; Bhadra, S." box="[426,732,1730,1754]" page="221" pageId="4" pageNumber="381" year="2011">
<bibRefCitation id="C92A8FF7FFF2FFE218BAF942C99E58C0" author="Dev Roy, M. K. &amp; Bhadra, S." box="[426,732,1730,1754]" pageId="4" pageNumber="381" pagination="109 - 269" refId="ref6153" refString="Dev Roy, M. K. &amp; Bhadra, S. (2011) Brachyuran crabs (Crustacea: Decapoda: Brachyura). In: State Fauna Series. 17. Part- 2. Fauna of Tamil Nadu. Zoological Survey of India, Kolkata. Pp. 109 - 269," type="book chapter" year="2011">Dev Roy &amp; Bhadra 2011: 221</bibRefCitation>
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, 231, 256.
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<paragraph id="AD04F206FFF2FFE21987F960C92A58E3" blockId="4.[151,1436,1638,1846]" box="[151,616,1760,1785]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F960C92A58E3" box="[151,616,1760,1785]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F960CAE258E2" box="[151,416,1760,1784]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F960CAC758E2" box="[151,389,1760,1784]" class="Malacostraca" family="Litocheiridae" genus="Heteropilumnas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Heteropilumnas integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE218B0F961C92658E3" author="Rao, G. S. &amp; Suseelan, C. &amp; Kathirvel, M." box="[416,612,1760,1785]" page="74" pageId="4" pageNumber="381" year="1989">
<bibRefCitation id="C92A8FF7FFF2FFE218B0F961C92658E3" author="Rao, G. S. &amp; Suseelan, C. &amp; Kathirvel, M." box="[416,612,1760,1785]" pageId="4" pageNumber="381" pagination="72 - 76" refId="ref8558" refString="Rao, G. S., Suseelan, C. &amp; Kathirvel, M. (1989) Crustacean resources of the Lakshadweep Islands. In: Suseelan, C. (Ed.), CMFRI Bulletin 43. Marine Living Resources of the Union Territory of Lakshadweep. An Indicative Survey with Suggestions for Development. Central Marine Fisheries Research Institute, Cochin, pp. 72 - 76," type="book chapter" year="1989">
Rao
<emphasis id="9FCF2E14FFF2FFE218DFF961C94058E2" box="[463,514,1760,1784]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
1989: 74
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</paragraph>
<paragraph id="AD04F206FFF2FFE21987F97FCF33590D" blockId="4.[151,1436,1638,1846]" box="[151,1137,1791,1815]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F97FCF33590D" box="[151,1137,1791,1815]" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F97FCACB590D" box="[151,393,1791,1815]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF2FFE21987F97FCACB590D" box="[151,393,1791,1815]" italics="true" pageId="4" pageNumber="381">Heteropilumnus integra</emphasis>
</taxonomicName>
<treatmentCitation id="2C1AD417FFF2FFE218B2F97FC99E590D" author="Venkataraman, K. &amp; Jeyabaskaran, R. &amp; Raghuram, K. P. &amp; Alfred, J. R. B." box="[418,732,1791,1815]" page="309" pageId="4" pageNumber="381" year="2004">
<bibRefCitation id="C92A8FF7FFF2FFE218B2F97FC99E590D" author="Venkataraman, K. &amp; Jeyabaskaran, R. &amp; Raghuram, K. P. &amp; Alfred, J. R. B." box="[418,732,1791,1815]" pageId="4" pageNumber="381" pagination="1 - 468" refId="ref9328" refString="Venkataraman, K., Jeyabaskaran, R., Raghuram, K. P. &amp; Alfred, J. R. B. (2004) Bibliography and checklist of corals and coral reef associated organisms of India. Records of the Zoological Survey of India, Occasional Paper, No. 226, 1 - 468." type="book chapter" year="2004">
Venkataraman
<emphasis id="9FCF2E14FFF2FFE21B2AF880C92F590D" box="[570,621,1791,1815]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
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;
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;
<treatmentCitation id="2C1AD417FFF2FFE21A98F97FCF2F590D" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[904,1133,1791,1815]" page="59" pageId="4" pageNumber="381" year="2018">
<bibRefCitation id="C92A8FF7FFF2FFE21A98F97FCF2F590D" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[904,1133,1791,1815]" pageId="4" pageNumber="381" pagination="1 - 83" refId="ref9091" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. &amp; Ng, P. K. L. (2018) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi
<emphasis id="9FCF2E14FFF2FFE21AC8F880CF49590D" box="[984,1035,1791,1815]" italics="true" pageId="4" pageNumber="381">et al.</emphasis>
2018: 59
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD04F206FFF2FFE21987F89EC9AF592C" blockId="4.[151,1436,1638,1846]" box="[151,749,1822,1846]" pageId="4" pageNumber="381">
<treatmentCitationGroup id="8DABD528FFF2FFE21987F89EC9AF592C" box="[151,749,1822,1846]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F89ECAEE592F" box="[151,428,1822,1846]" italics="true" pageId="4" pageNumber="381">
<taxonomicName id="6ABB8985FFF2FFE21987F89ECAD3592F" box="[151,401,1822,1846]" class="Malacostraca" family="Litocheiridae" genus="Pseudolithochira" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="integra">Pseudolithochira integra</taxonomicName>
</emphasis>
<treatmentCitation id="2C1AD417FFF2FFE218BCF89EC9AA592C" author="Innocenti, G. &amp; Manzoni, R." box="[428,744,1822,1846]" page="47" pageId="4" pageNumber="381" year="2017">
<bibRefCitation id="C92A8FF7FFF2FFE218BCF89EC9AA592C" author="Innocenti, G. &amp; Manzoni, R." box="[428,744,1822,1846]" pageId="4" pageNumber="381" pagination="29 - 48" refId="ref6571" refString="Innocenti, G. &amp; Manzoni, R. (2017) Collections of the Natural History Museum Zoological Section &quot; La Specola &quot; of the University of Florence. Crustacea, Class Malacostraca, Order Decapoda. Superfamilies Gecarcinucoidea, Goneplacoidea, Hexapodoidea, Leucosioidea, Majoidea, Orithyioidea, Palicoidea, Parthenopoidea, Pilumnoidea, Atti della Societa toscana di scienze naturali, Memorie, Serie B, 124, 29 - 48. https: // doi. org / 10.2424 / ASTSN. M. 2017.04" type="journal article" year="2017">Innocenti &amp; Manzoni 2017: 47</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5A1A18DFFF2FFE21987F8E4C97559B9" pageId="4" pageNumber="381" type="materials_examined">
<paragraph id="AD04F206FFF2FFE21987F8E4C97559B9" blockId="4.[151,1372,1892,1955]" pageId="4" pageNumber="381">
<emphasis id="9FCF2E14FFF2FFE21987F8E4CA3B5964" bold="true" box="[151,377,1892,1918]" pageId="4" pageNumber="381">Material examined</emphasis>
.
<materialsCitation id="1DD3F85BFFF2FFE21897F8E4C97559B9" ID-GBIF-Occurrence="3330072301" collectorName="H. M. S. Alert. Species &amp; As" country="Seychelles" elevation="15" location="Seychelles" pageId="4" pageNumber="381" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="72004CA4FFF2FFE21897F8E4CAB65964" box="[391,500,1892,1918]" pageId="4" pageNumber="381" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBBD398FFFF2FFE218EEF8E5C9515965" box="[510,531,1893,1919]" pageId="4" pageNumber="381" type="female"></specimenCount>
(4.9 ×
<quantity id="6A435FE3FFF2FFE21B75F8E4C9FE5965" box="[613,700,1892,1919]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="381" unit="mm" value="3.4">3.4 mm</quantity>
) (NHM 1882.24),
<collectingCountry id="D5ACB296FFF2FFE21A9EF8E4CF4B5964" box="[910,1033,1892,1918]" name="Seychelles" pageId="4" pageNumber="381">Seychelles</collectingCountry>
,
<quantity id="6A435FE3FFF2FFE21D04F8E4CFC25965" box="[1044,1152,1892,1919]" metricMagnitude="1" metricUnit="m" metricValue="1.465" metricValueMax="2.2" metricValueMin="0.73" pageId="4" pageNumber="381" unit="m" value="14.65" valueMax="22.0" valueMin="7.3">
<elevation id="26961535FFF2FFE21D04F8E4CFC25965" box="[1044,1152,1892,1919]" metricMagnitude="1" metricUnit="m" metricValue="1.465" metricValueMax="2.2" metricValueMin="0.73" pageId="4" pageNumber="381" unit="m" value="14.65" valueMax="22.0" valueMin="7.3">7.322 m</elevation>
</quantity>
, coll.
<collectorName id="004E97D0FFF2FFE21DD4F8E4CA5C59B8" pageId="4" pageNumber="381">
H.M.S.
<emphasis id="9FCF2E14FFF2FFE21C0DF8E4CE175964" box="[1309,1365,1892,1918]" italics="true" pageId="4" pageNumber="381">Alert</emphasis>
.
<emphasis id="9FCF2E14FFF2FFE219D7F808CA5C59B8" bold="true" box="[199,286,1928,1954]" pageId="4" pageNumber="381">Species</emphasis>
</collectorName>
diagnosis.
<collectorName id="004E97D0FFF2FFE218B0F808CAFD59B8" box="[416,447,1928,1954]" pageId="4" pageNumber="381">As</collectorName>
for genus.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="F9C4A28EFFF3FFE31987F9DDCE205880" ID-DOI="http://doi.org/10.5281/zenodo.4749245" ID-Zenodo-Dep="4749245" httpUri="https://zenodo.org/record/4749245/files/figure.png" pageId="5" pageNumber="382" startId="5.[151,250,1629,1653]" targetBox="[151,1436,181,1606]" targetPageId="5">
<paragraph id="AD04F206FFF3FFE31987F9DDCE205880" blockId="5.[151,1436,1629,1690]" pageId="5" pageNumber="382">
<emphasis id="9FCF2E14FFF3FFE31987F9DDCA4D586C" bold="true" box="[151,271,1629,1654]" pageId="5" pageNumber="382">FIGURE 3</emphasis>
.
<taxonomicName id="6ABB8985FFF3FFE3180DF9DEC9E5586C" authority="(Miers, 1884)" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[285,679,1630,1654]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF3FFE3180DF9DEC952586F" box="[285,528,1630,1654]" italics="true" pageId="5" pageNumber="382">Pseudolitochira integra</emphasis>
(
<bibRefCitation id="C92A8FF7FFF3FFE31B0FF9DEC9E2586C" author="Miers, E. J." box="[543,672,1630,1654]" pageId="5" pageNumber="382" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
</taxonomicName>
, holotype female (4.9 × 3.4 mm) (NHM 1882.24), Seychelles. A, overall dorsal view; B, left fourth ambulatory leg; C, left ambulatory propodus and dactylus; D, sternopleonal cavity and vulvae.
</paragraph>
</caption>
<subSubSection id="E5A1A18DFFF3FFEF19D7F948C8C75D23" lastPageId="9" lastPageNumber="386" pageId="5" pageNumber="382" type="description">
<paragraph id="AD04F206FFF3FFE019D7F948C8745F7F" blockId="5.[151,1436,1736,2014]" lastBlockId="6.[151,1437,151,357]" lastPageId="6" lastPageNumber="383" pageId="5" pageNumber="382">
<emphasis id="9FCF2E14FFF3FFE319D7F948C97158F8" bold="true" box="[199,563,1736,1762]" pageId="5" pageNumber="382">
Description of female
<typeStatus id="72004CA4FFF3FFE318DCF948C97158F8" box="[460,563,1736,1762]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace transversely subovate, width 1.6 times length; surface covered with short pubescence that partially obscures surface, never with long setae along margins; dorsal carapace surface mostly smooth except for small group of low granules behind anterolateral margin, regions almost undiscernible, with only shallow H-shaped gastric grooves barely visible; postfrontal lobes low, just visible. Frontal margin extremely wide, bilobed, margin gently convex, with a low postfrontal ridge; no lateral lobule present on frontal margin, smoothly confluent with supraorbital margin. Supraorbital margin finely granulated, with small submedian indentation, no fissure; infraorbital margin distinctly sinuous, without tooth or spine. Orbits wide; eyes large, ocular peduncle stout, cornea large, extending beyond external orbital tooth when retracted. External orbital tooth low; anterolateral margin relatively short, appears entire, with small granules, no clear indication of lobes, low concavities in parts; joining posterolateral margin as gently curve; posterolateral margin sharply converging towards gently sinuous posterior carapace margin. Suborbital, subhepatic and pterygostomial regions smooth. Antennules folding almost transversely; basal antennal article subrectangular; flagellum entering orbital hiatus. Epistome relatively narrow longitudinally; posterior margin entire, median lobe triangular with shallow median cleft, confluent with gently concave, entire lateral margins. Endostomial ridges distinct.
</paragraph>
<caption id="F9C4A28EFFF0FFE01987F807C96859D9" ID-DOI="http://doi.org/10.5281/zenodo.4749247" ID-Zenodo-Dep="4749247" httpUri="https://zenodo.org/record/4749247/files/figure.png" pageId="6" pageNumber="383" startId="6.[151,250,1927,1951]" targetBox="[151,1436,381,1904]" targetPageId="6">
<paragraph id="AD04F206FFF0FFE01987F807C96859D9" blockId="6.[151,1436,1927,1987]" pageId="6" pageNumber="383">
<emphasis id="9FCF2E14FFF0FFE01987F807CA4C5985" bold="true" box="[151,270,1927,1951]" pageId="6" pageNumber="383">FIGURE 4</emphasis>
.
<taxonomicName id="6ABB8985FFF0FFE0180CF807C9E659BA" authority="(Miers, 1884)" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[284,676,1927,1952]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF0FFE0180CF807C94C5985" box="[284,526,1927,1951]" italics="true" pageId="6" pageNumber="383">Pseudolitochira integra</emphasis>
(
<bibRefCitation id="C92A8FF7FFF0FFE01B0DF807C9DF59BA" author="Miers, E. J." box="[541,669,1927,1952]" pageId="6" pageNumber="383" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
</taxonomicName>
, holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, overall dorsal view; B, ventral view of cephalothorax.
</paragraph>
</caption>
<caption id="F9C4A28EFFF1FFE11987F96CCEDE5939" ID-DOI="http://doi.org/10.5281/zenodo.4749251" ID-Zenodo-Dep="4749251" httpUri="https://zenodo.org/record/4749251/files/figure.png" pageId="7" pageNumber="384" startId="7.[151,250,1772,1796]" targetBox="[151,1436,181,1749]" targetPageId="7">
<paragraph id="AD04F206FFF1FFE11987F96CCEDE5939" blockId="7.[151,1436,1772,1863]" pageId="7" pageNumber="384">
<emphasis id="9FCF2E14FFF1FFE11987F96CCA4C591F" bold="true" box="[151,270,1772,1797]" pageId="7" pageNumber="384">FIGURE 5</emphasis>
.
<taxonomicName id="6ABB8985FFF1FFE1180BF96DC9E0591F" authority="(Miers, 1884)" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[283,674,1773,1797]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFF1FFE1180BF96DC94F591E" box="[283,525,1773,1797]" italics="true" pageId="7" pageNumber="384">Pseudolitochira integra</emphasis>
(
<bibRefCitation id="C92A8FF7FFF1FFE11B0BF96DC9D9591F" author="Miers, E. J." box="[539,667,1773,1797]" pageId="7" pageNumber="384" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
</taxonomicName>
, holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, dorsal view of carapace; B, frontal view of cephalothorax; C, outer view of right chela; D, buccal cavity, third maxillipeds and thoracic sternites
</paragraph>
</caption>
<paragraph id="AD04F206FFF1FFE11987F8AFC9C4595D" blockId="7.[151,1436,1772,1863]" box="[151,646,1839,1863]" pageId="7" pageNumber="384">1 and 2; E, posterior thoracic sternum and pleon.</paragraph>
<caption id="F9C4A28EFFFEFFEE1987F8E5CE0B59DF" ID-DOI="http://doi.org/10.5281/zenodo.4749253" ID-Zenodo-Dep="4749253" httpUri="https://zenodo.org/record/4749253/files/figure.png" pageId="8" pageNumber="385" startId="8.[151,250,1893,1917]" targetBox="[151,1436,181,1869]" targetPageId="8">
<paragraph id="AD04F206FFFEFFEE1987F8E5CE0B59DF" blockId="8.[151,1437,1893,1989]" pageId="8" pageNumber="385">
<emphasis id="9FCF2E14FFFEFFEE1987F8E5CA4E5967" bold="true" box="[151,268,1893,1917]" pageId="8" pageNumber="385">FIGURE 6</emphasis>
.
<taxonomicName id="6ABB8985FFFEFFEE1807F8E5C9825967" authority="Ng &amp; Tan, 1988" authorityName="Ng &amp; Tan" authorityYear="1988" box="[279,704,1893,1918]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFEFFEE1807F8E5C9555967" box="[279,535,1893,1917]" italics="true" pageId="8" pageNumber="385">Heteropilumnus holthuisi</emphasis>
<bibRefCitation id="C92A8FF7FFFEFFEE1B0CF8E6C9825967" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[540,704,1893,1918]" pageId="8" pageNumber="385" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan, 1988</bibRefCitation>
</taxonomicName>
. ♂ (10.1 × 7.8 mm) (NHM 1886.52), Mergui; A, dorsal view of carapace; B, frontal view of maxilliped; C, frontal view of cephalothorax; D, lateral view of cephalothorax; E, anterior thoracic sternum and pleon; F, posterior thoracic sternum and pleon; ♂ (7.5 × 6.3 mm) (NHM 1886.52), Mergui; G, overall dorsal view.
</paragraph>
</caption>
<paragraph id="AD04F206FFFFFFEF19D7FF17C8635EE3" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">Third maxilliped completely covering buccal cavity when closed; ischium short, subrectangular with shallow submedian sulcus; merus quadrate, anteroexternal angle rounded, not auriculiform; exopod relatively stout, reaching to just before distal margin of merus, with long flagellum.</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7FE83CAE25FCB" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">Chelipeds subequal, outer surfaces covered with long and short setae which obscures surfaces. Basis-ischium short, anterior margin granulated. Merus trigonal in cross-section, margins almost smooth or with low granules, not armed. Carpus relatively elongate, surface mostly smooth except for weakly granulose distal surfaces; inner distal angle formed by low, rounded obtuse tooth. Outer surface of chela covered with numerous small granules, reaching to half-length of fingers; fingers shorter than palm, with 2 rows of low ridges, proximal part with granules; cutting margins with low teeth.</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7FE5BC8A95C7B" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">Ambulatory legs proportionately slender, relatively long; second leg longest, fourth leg shortest; surfaces covered with short and long setae which obscures margins. Merus smooth with rounded margins, unarmed. Carpus with shallow subdorsal groove, unarmed. Propodus smooth. Dactylus relatively long, almost straight along proximal corneous part with tip gently curved; dactylo-propodal lock well developed.</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7FDEBCE215CEB" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">Thoracic sternites 1 and 2 completely fused to form triangular plate; deep sinuous suture between sternite 2 and sternite 3; sternites 3 and 4 fused, with shallow broad median grooves demarcating them. Sutures between sternites 4 and 5, 5 and 6 medially interrupted; other sutures complete. Sternopleonal cavity shallow. Vulvae large, subovate, flat, without sternal vulvar covers; occupying anterior two-thirds of sternite 6, touching suture with sternite 5.</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7FD7BC8C75D23" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">Pleon ovate, covering about half of thoracic sternum; somite 1 longitudinally narrow, reaching to coxae of last ambulatory legs, lateral edges gradually tapering; telson triangular.</paragraph>
</subSubSection>
<subSubSection id="E5A1A18DFFFFFFEC19D7FCC3CAF158FB" lastPageId="10" lastPageNumber="387" pageId="9" pageNumber="386" type="discussion">
<paragraph id="AD04F206FFFFFFEF19D7FCC3C8775A0B" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">
<emphasis id="9FCF2E14FFFFFFEF19D7FCC3CA765D47" bold="true" box="[199,308,835,861]" pageId="9" pageNumber="386">Remarks</emphasis>
. The description and figures (reproduced here as
<figureCitation id="3580EE83FFFFFFEF1A73FCC3C8E45D47" box="[867,934,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1324,1348]" captionTargetBox="[164,1416,181,1289]" captionTargetId="figure-18@3.[151,1436,181,1300]" captionTargetPageId="3" captionText="FIGURE 2. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, overall dorsal view; B, buccal cavity and frontal regions; C, outer view of left chela. After Miers (1884: pl. 8 figs. C, c, c)." figureDoi="http://doi.org/10.5281/zenodo.4749241" httpUri="https://zenodo.org/record/4749241/files/figure.png" pageId="9" pageNumber="386">Fig. 2</figureCitation>
) of
<taxonomicName id="6ABB8985FFFFFFEF1AC5FCC3CF005D46" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[981,1090,835,860]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF1AC5FCC3CF005D46" box="[981,1090,835,860]" italics="true" pageId="9" pageNumber="386">P. integra</emphasis>
</taxonomicName>
by
<bibRefCitation id="C92A8FF7FFFFFFEF1D7CFCC3CE425D47" author="Miers, E. J." box="[1132,1280,835,861]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers (1884)</bibRefCitation>
contain some inaccuracies. The anterolateral margin was described and figured as entire (
<bibRefCitation id="C92A8FF7FFFFFFEF1AC3FCE7CF155D9B" author="Miers, E. J." box="[979,1111,871,897]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers 1884</bibRefCitation>
: pl. 8, fig. C) but the structure is lined with small granules (
<figureCitation id="3580EE83FFFFFFEF18CAFC0BC9765DBF" box="[474,564,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1749]" captionTargetId="figure-18@7.[151,1436,181,1749]" captionTargetPageId="7" captionText="FIGURE 5. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, dorsal view of carapace; B, frontal view of cephalothorax; C, outer view of right chela; D, buccal cavity, third maxillipeds and thoracic sternites" figureDoi="http://doi.org/10.5281/zenodo.4749251" httpUri="https://zenodo.org/record/4749251/files/figure.png" pageId="9" pageNumber="386">Fig. 5A</figureCitation>
). The exopod of the third maxilliped was figured as only reaching to just after the edge of the ischium (
<bibRefCitation id="C92A8FF7FFFFFFEF18B6FC2FC9695DD3" author="Miers, E. J." box="[422,555,943,969]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers 1884</bibRefCitation>
: pl. 8, fig. c), but it is actually distinctly longer, reaching to just before the distal edge of the merus (
<figureCitation id="3580EE83FFFFFFEF187DFC53CA855DF7" box="[365,455,978,1005]" captionStart="FIGURE 5" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1749]" captionTargetId="figure-18@7.[151,1436,181,1749]" captionTargetPageId="7" captionText="FIGURE 5. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, dorsal view of carapace; B, frontal view of cephalothorax; C, outer view of right chela; D, buccal cavity, third maxillipeds and thoracic sternites" figureDoi="http://doi.org/10.5281/zenodo.4749251" httpUri="https://zenodo.org/record/4749251/files/figure.png" pageId="9" pageNumber="386">Fig. 5D</figureCitation>
). The outer surface of the chela was figured as smooth (
<bibRefCitation id="C92A8FF7FFFFFFEF1D26FC53CFFE5DF7" author="Miers, E. J." box="[1078,1212,979,1005]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers 1884</bibRefCitation>
: pl. 8, fig. c), but is covered with numerous distinct rounded granules (
<figureCitation id="3580EE83FFFFFFEF1BDFFC77C86A5A0B" box="[719,808,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1749]" captionTargetId="figure-18@7.[151,1436,181,1749]" captionTargetPageId="7" captionText="FIGURE 5. Pseudolitochira integra (Miers, 1884), holotype ♀ (4.9 × 3.4 mm), NHM 1882.24, Seychelles. A, dorsal view of carapace; B, frontal view of cephalothorax; C, outer view of right chela; D, buccal cavity, third maxillipeds and thoracic sternites" figureDoi="http://doi.org/10.5281/zenodo.4749251" httpUri="https://zenodo.org/record/4749251/files/figure.png" pageId="9" pageNumber="386">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7FB9BCABF58A4" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">
The nomenclature of
<taxonomicName id="6ABB8985FFFFFFEF18ADFB9BC9E05A2E" authorityName="Miers" authorityYear="1884" box="[445,674,1051,1077]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF18ADFB9BC9E05A2E" box="[445,674,1051,1077]" italics="true" pageId="9" pageNumber="386">Carcinoplax integra</emphasis>
</taxonomicName>
has also presented a number of taxonomic problems. Shortly after
<bibRefCitation id="C92A8FF7FFFFFFEF1987FBBFCA6C5A43" author="Miers, E. J." box="[151,302,1087,1113]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers (1884)</bibRefCitation>
described his species, the name
<taxonomicName id="6ABB8985FFFFFFEF1BA5FBBFC8695A42" authorityName="Miers" authorityYear="1884" box="[693,811,1087,1113]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF1BA5FBBFC8695A42" box="[693,811,1087,1113]" italics="true" pageId="9" pageNumber="386">C. integra</emphasis>
</taxonomicName>
became confused with another allied taxon. De
<bibRefCitation id="C92A8FF7FFFFFFEF1C79FBBFCA4C5A67" author="Man, J. G. De" pageId="9" pageNumber="386" pagination="65 - 128" refId="ref7189" refString="Man, J. G. De (1887) Part II. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22 (137), 65 - 128, pls. 4 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01455. x" type="journal article" year="1887">Man (1887: 93)</bibRefCitation>
identified
<specimenCount id="BBBD398FFFFFFFEF189CFBE3C9425A67" box="[396,512,1123,1149]" pageId="9" pageNumber="386" type="male">two males</specimenCount>
and a female from Elphinstone Island, Mergui Archipelago as
<taxonomicName id="6ABB8985FFFFFFEF1DC6FBE3CE055A66" box="[1238,1351,1123,1149]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="9FCF2E14FFFFFFEF1DC6FBE3CE055A66" box="[1238,1351,1123,1149]" italics="true" pageId="9" pageNumber="386">C. integer</emphasis>
</taxonomicName>
, which he later referred to “
<taxonomicName id="6ABB8985FFFFFFEF186FFB07C87C5ABB" authority="Lanchester, 1900" authorityName="Lanchester" authorityYear="1900" box="[383,830,1159,1185]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFFFFFEF186FFB07C9305ABB" box="[383,626,1159,1185]" italics="true" pageId="9" pageNumber="386">Litocheira subinteger</emphasis>
<bibRefCitation id="C92A8FF7FFFFFFEF1B6BFB07C87C5ABB" author="Lanchester, W. F." box="[635,830,1159,1185]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester, 1900</bibRefCitation>
</taxonomicName>
” (De
<bibRefCitation id="C92A8FF7FFFFFFEF1A93FB07CF555ABB" author="Man, J. G. De" box="[899,1047,1159,1185]" pageId="9" pageNumber="386" pagination="1 - 26" refId="ref7363" refString="Man, J. G. De (1929) On a collection of decapod and stomatopod Crustacea from Pulau Berhala, an islet situated in the Straits of Malacca. Bijdragen tot de Dierkunde, 26, 1 - 26, pls. I - III. https: // doi. org / 10.1163 / 26660644 - 02601001" type="journal article" year="1929">Man 1929: 9</bibRefCitation>
). The latter name is problematical because De
<bibRefCitation id="C92A8FF7FFFFFFEF180DFB2BCAE35ADF" author="Man, J. G. De" box="[285,417,1195,1221]" pageId="9" pageNumber="386" pagination="1 - 26" refId="ref7363" refString="Man, J. G. De (1929) On a collection of decapod and stomatopod Crustacea from Pulau Berhala, an islet situated in the Straits of Malacca. Bijdragen tot de Dierkunde, 26, 1 - 26, pls. I - III. https: // doi. org / 10.1163 / 26660644 - 02601001" type="journal article" year="1929">Man (1929)</bibRefCitation>
did not realise that
<bibRefCitation id="C92A8FF7FFFFFFEF1B6BFB2BC8055ADF" author="Lanchester, W. F." box="[635,839,1195,1221]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester (1900)</bibRefCitation>
had merely changed the spelling of the species name, from “
<taxonomicName id="6ABB8985FFFFFFEF19F5FB4FCA745AF2" authorityName="Miers" authorityYear="1884" box="[229,310,1231,1256]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF19F5FB4FCA745AF2" box="[229,310,1231,1256]" italics="true" pageId="9" pageNumber="386">integra</emphasis>
</taxonomicName>
” to “
<taxonomicName id="6ABB8985FFFFFFEF186AFB4FCAB25AF3" authorityName="Lanchester" authorityYear="1900" box="[378,496,1231,1257]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFFFFFEF186AFB4FCAB25AF3" box="[378,496,1231,1257]" italics="true" pageId="9" pageNumber="386">subinteger</emphasis>
</taxonomicName>
” because he argued that original
<bibRefCitation id="C92A8FF7FFFFFFEF1A6DFB4FCF565AF3" author="Miers, E. J." box="[893,1044,1231,1257]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers (1884)</bibRefCitation>
name was inaccurate.
<bibRefCitation id="C92A8FF7FFFFFFEF1C31FB4FCBA05B17" author="Lanchester, W. F." pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester (1900)</bibRefCitation>
had specimens from
<collectingCountry id="D5ACB296FFFFFFEF18C6FB73C90A5B17" box="[470,584,1267,1293]" name="Singapore" pageId="9" pageNumber="386">Singapore</collectingCountry>
or
<collectingCountry id="D5ACB296FFFFFFEF1B7FFB73C98C5B17" box="[623,718,1267,1293]" name="Malaysia" pageId="9" pageNumber="386">Malacca</collectingCountry>
(exact location not stated) and believed them to be conspecific with
<taxonomicName id="6ABB8985FFFFFFEF19C3FA97C9125B2B" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[211,592,1303,1329]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="9FCF2E14FFFFFFEF19C3FA97CAFA5B2A" box="[211,440,1303,1329]" italics="true" pageId="9" pageNumber="386">Carcinoplax integer</emphasis>
<bibRefCitation id="C92A8FF7FFFFFFEF18D2FA97C9125B2B" author="Miers, E. J." box="[450,592,1303,1329]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="C92A8FF7FFFFFFEF1B4EFA97C86C5B2B" author="Lanchester, W. F." box="[606,814,1303,1329]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester (1900)</bibRefCitation>
commented that since the anterolateral margin of his specimens were not entire and had low teeth, the name chosen by Miers was inappropriate and he therefore renamed the species “
<taxonomicName id="6ABB8985FFFFFFEF1837FADFC9725B63" authorityName="Lanchester" authorityYear="1900" box="[295,560,1375,1401]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFFFFFEF1837FADFC9725B63" box="[295,560,1375,1401]" italics="true" pageId="9" pageNumber="386">Carcinoplax subinteger</emphasis>
</taxonomicName>
”. This change of name, however, is not permitted under the zoological Code (
<bibRefCitation id="C92A8FF7FFFFFFEF198FFA03CA675B87" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[159,293,1411,1437]" pageId="9" pageNumber="386" refId="ref6509" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. International Commission of Zoological Nomenclature. Fourth Edition. Adopted by the XXI General Assembly of the International Union of Biological Sciences. International Trust for Zoological Nomenclature, in association with the British Museum (Natural History), London, 338 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 72.7) especially since both names share the same
<typeStatus id="72004CA4FFFFFFEF1ABEFA03C89C5B87" box="[942,990,1411,1437]" pageId="9" pageNumber="386">type</typeStatus>
specimen (see
<bibRefCitation id="C92A8FF7FFFFFFEF1D99FA03CE265B87" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[1161,1380,1410,1437]" pageId="9" pageNumber="386" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan 1988: 15</bibRefCitation>
, 17). Consequently, the
<taxonomicName id="6ABB8985FFFFFFEF1875FA27CAB85BDB" authorityName="Lanchester" authorityYear="1900" box="[357,506,1447,1473]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFFFFFEF1875FA27CAB85BDB" box="[357,506,1447,1473]" italics="true" pageId="9" pageNumber="386">C. subinteger</emphasis>
</taxonomicName>
of
<bibRefCitation id="C92A8FF7FFFFFFEF1B0EFA27C9A25BDB" author="Lanchester, W. F." box="[542,736,1447,1473]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester, 1900</bibRefCitation>
, must therefore be treated as an unnecessary replacement name for
<taxonomicName id="6ABB8985FFFFFFEF19D0FA4BC9755BFF" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[192,567,1483,1509]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="9FCF2E14FFFFFFEF19D0FA4BCAE15BFE" box="[192,419,1483,1509]" italics="true" pageId="9" pageNumber="386">Carcinoplax integer</emphasis>
<bibRefCitation id="C92A8FF7FFFFFFEF18BBFA4BC9755BFF" author="Miers, E. J." box="[427,567,1483,1509]" pageId="9" pageNumber="386" pagination="513 - 575" refId="ref7477" refString="Miers, E. J. (1884) Crustacea. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert' 1881 - 2. Part II. Collections from the Western Indian Ocean. British Museum (Natural History), London, pp. 513 - 575, pls. XLVI - LII." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="C92A8FF7FFFFFFEF1B54FA4BC84A5BFF" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[580,776,1483,1509]" pageId="9" pageNumber="386" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan (1988)</bibRefCitation>
also indicated that the
<bibRefCitation id="C92A8FF7FFFFFFEF1D02FA4BCFA25BFF" author="Lanchester, W. F." box="[1042,1248,1483,1509]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester (1900)</bibRefCitation>
specimens were an undescribed species and named it
<taxonomicName id="6ABB8985FFFFFFEF1B23FA6FC80C5813" authorityName="Ng &amp; Tan" authorityYear="1988" box="[563,846,1519,1545]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFFFFEF1B23FA6FC80C5813" box="[563,846,1519,1545]" italics="true" pageId="9" pageNumber="386">Heteropilumnus holthuisi</emphasis>
</taxonomicName>
. Nevertheless,
<taxonomicName id="6ABB8985FFFFFFEF1AEAFA6FCF9E5812" authorityName="Miers" authorityYear="1884" box="[1018,1244,1519,1545]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF1AEAFA6FCF9E5812" box="[1018,1244,1519,1545]" italics="true" pageId="9" pageNumber="386">Carcinoplax integra</emphasis>
</taxonomicName>
and
<taxonomicName id="6ABB8985FFFFFFEF1C01FA6FCA4D5837" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subintegra">
<emphasis id="9FCF2E14FFFFFFEF1C01FA6FCA4D5837" italics="true" pageId="9" pageNumber="386">Carcinoplax subintegra</emphasis>
</taxonomicName>
have sometimes been treated as two separate taxa in the literature (e.g.,
<bibRefCitation id="C92A8FF7FFFFFFEF1D43F993CE565837" author="Tesch, J. J." box="[1107,1300,1554,1581]" pageId="9" pageNumber="386" pagination="149 - 295" refId="ref9056" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918: 165</bibRefCitation>
).
<bibRefCitation id="C92A8FF7FFFFFFEF1C39F993CBA0584B" author="Ng, P. K. L. &amp; Tan, L. W. H." pageId="9" pageNumber="386" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan (1988)</bibRefCitation>
also assigned the specimens of De
<bibRefCitation id="C92A8FF7FFFFFFEF1B6AF9B7C842584B" author="Man, J. G. De" box="[634,768,1591,1617]" pageId="9" pageNumber="386" pagination="65 - 128" refId="ref7189" refString="Man, J. G. De (1887) Part II. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22 (137), 65 - 128, pls. 4 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01455. x" type="journal article" year="1887">Man (1887)</bibRefCitation>
from Mergui to
<taxonomicName id="6ABB8985FFFFFFEF1AD0F9B7CF04584B" authorityName="Ng &amp; Tan" authorityYear="1988" box="[960,1094,1591,1617]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFFFFEF1AD0F9B7CF04584B" box="[960,1094,1591,1617]" italics="true" pageId="9" pageNumber="386">H. holthuisi</emphasis>
</taxonomicName>
, but they did not examine the material. The present study re-examined specimens of De
<bibRefCitation id="C92A8FF7FFFFFFEF1A30F9DBC8E6586F" author="Man, J. G. De" box="[800,932,1627,1653]" pageId="9" pageNumber="386" pagination="65 - 128" refId="ref7189" refString="Man, J. G. De (1887) Part II. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22 (137), 65 - 128, pls. 4 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01455. x" type="journal article" year="1887">Man (1887)</bibRefCitation>
in NHM (
<specimenCount id="BBBD398FFFFFFFEF1D08F9DBCF06586F" box="[1048,1092,1627,1653]" pageId="9" pageNumber="386" type="male">2 ♂</specimenCount>
, 10.1 ×
<quantity id="6A435FE3FFFFFFEF1D8DF9DBCFB7586F" box="[1181,1269,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="386" unit="mm" value="7.8">7.8 mm</quantity>
, 7.5 ×
<quantity id="6A435FE3FFFFFFEF1C50F9DBCEDA586F" box="[1344,1432,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="386" unit="mm" value="6.3">6.3 mm</quantity>
, NHM 1886.52, Mergui Archipelago, coll. Anderson) and confirm here that they are both
<taxonomicName id="6ABB8985FFFFFFEF1D99F9FFCE4C5883" authorityName="Ng &amp; Tan" authorityYear="1988" box="[1161,1294,1663,1689]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFFFFEF1D99F9FFCE4C5883" box="[1161,1294,1663,1689]" italics="true" pageId="9" pageNumber="386">H. holthuisi</emphasis>
</taxonomicName>
as proposed by
<bibRefCitation id="C92A8FF7FFFFFFEF19AAF923CA3758A7" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[186,373,1699,1725]" pageId="9" pageNumber="386" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan (1988</bibRefCitation>
; see
<figureCitation id="3580EE83FFFFFFEF18BBF923CAAC58A4" box="[427,494,1699,1726]" captionStart="FIGURE 6" captionStartId="8.[151,250,1893,1917]" captionTargetBox="[151,1436,181,1869]" captionTargetId="figure-18@8.[151,1436,181,1869]" captionTargetPageId="8" captionText="FIGURE 6. Heteropilumnus holthuisi Ng &amp; Tan, 1988. ♂ (10.1 × 7.8 mm) (NHM 1886.52), Mergui;A, dorsal view of carapace; B, frontal view of maxilliped; C, frontal view of cephalothorax; D, lateral view of cephalothorax; E, anterior thoracic sternum and pleon; F, posterior thoracic sternum and pleon; ♂ (7.5 × 6.3 mm) (NHM 1886.52), Mergui; G, overall dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4749253" httpUri="https://zenodo.org/record/4749253/files/figure.png" pageId="9" pageNumber="386">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="AD04F206FFFFFFEF19D7F947CFEB59A3" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="386">
<bibRefCitation id="C92A8FF7FFFFFFEF19D7F947CA8858FB" author="Borradaile, L. A." box="[199,458,1735,1761]" pageId="9" pageNumber="386" pagination="424 - 443" refId="ref5747" refString="Borradaile, L. A. (1903) Marine crustaceans. IV. Some remarks on the classification of the crabs. V. The crabs of the catometope families. VI. The sand crabs (Oxystomata). VII. The barnacles (Cirripedia). In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, vol. 1, part 4. Cambridge University Press, Cambridge, pp. 424 - 443, pl. XXII," type="book chapter" year="1903">Borradaile (1903: 430)</bibRefCitation>
reported a specimen from
<collectingRegion id="6F7F3CE4FFFFFFEF1BEDF947C87558FB" box="[765,823,1735,1761]" country="Maldives" name="Maale" pageId="9" pageNumber="386">Male</collectingRegion>
Atoll in the
<collectingCountry id="D5ACB296FFFFFFEF1AD7F947CF7058FB" box="[967,1074,1735,1761]" name="Maldives" pageId="9" pageNumber="386">Maldives</collectingCountry>
, but he noted his specimen (sex not specified) differed from the material of
<bibRefCitation id="C92A8FF7FFFFFFEF1B6BF96BC808591F" author="Lanchester, W. F." box="[635,842,1771,1797]" pageId="9" pageNumber="386" pagination="719 - 770" refId="ref6855" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3), 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester (1900)</bibRefCitation>
. He (
<bibRefCitation id="C92A8FF7FFFFFFEF1A93F96BCF78591F" author="Borradaile, L. A." box="[899,1082,1771,1797]" pageId="9" pageNumber="386" pagination="424 - 443" refId="ref5747" refString="Borradaile, L. A. (1903) Marine crustaceans. IV. Some remarks on the classification of the crabs. V. The crabs of the catometope families. VI. The sand crabs (Oxystomata). VII. The barnacles (Cirripedia). In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, vol. 1, part 4. Cambridge University Press, Cambridge, pp. 424 - 443, pl. XXII," type="book chapter" year="1903">Borradaile 1903</bibRefCitation>
) considered it the same as what De
<bibRefCitation id="C92A8FF7FFFFFFEF19D1F88FCA0A5933" author="Man, J. G. De" box="[193,328,1807,1833]" pageId="9" pageNumber="386" pagination="65 - 128" refId="ref7189" refString="Man, J. G. De (1887) Part II. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22 (137), 65 - 128, pls. 4 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01455. x" type="journal article" year="1887">Man (1887)</bibRefCitation>
reported as
<taxonomicName id="6ABB8985FFFFFFEF18C7F88FC92F5933" authorityName="Lanchester" authorityYear="1900" box="[471,621,1807,1833]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="subinteger">
<emphasis id="9FCF2E14FFFFFFEF18C7F88FC92F5933" box="[471,621,1807,1833]" italics="true" pageId="9" pageNumber="386">L. subinteger</emphasis>
</taxonomicName>
as his crab had three, and not two notches, on the anterolateral margin and possessed a relatively narrower front.
<bibRefCitation id="C92A8FF7FFFFFFEF1B9CF8B3C8185957" author="Borradaile, L. A." box="[652,858,1843,1869]" pageId="9" pageNumber="386" pagination="424 - 443" refId="ref5747" refString="Borradaile, L. A. (1903) Marine crustaceans. IV. Some remarks on the classification of the crabs. V. The crabs of the catometope families. VI. The sand crabs (Oxystomata). VII. The barnacles (Cirripedia). In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, vol. 1, part 4. Cambridge University Press, Cambridge, pp. 424 - 443, pl. XXII," type="book chapter" year="1903">Borradaile (1903)</bibRefCitation>
, however, did not elaborate on these differences. Later,
<bibRefCitation id="C92A8FF7FFFFFFEF19F0F8D7CAFD596B" author="Bouvier, E. L." box="[224,447,1878,1905]" pageId="9" pageNumber="386" pagination="178 - 318" refId="ref5866" refString="Bouvier, E. L. (1915) Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 7 e Serie, 48 (3), 178 - 318, pls. IV - VII." type="journal article" year="1915">Bouvier (1915: 296</bibRefCitation>
, fig. 34) recorded and partly figured
<taxonomicName id="6ABB8985FFFFFFEF1A4FF8D7CF6E596A" box="[863,1068,1879,1905]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFFFFEF1A4FF8D7CF6E596A" box="[863,1068,1879,1905]" italics="true" pageId="9" pageNumber="386">Litocheira integra</emphasis>
</taxonomicName>
based on a male 5 ×
<quantity id="6A435FE3FFFFFFEF1C0DF8D7CE1D596B" box="[1309,1375,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="386" unit="mm" value="4.0">4 mm</quantity>
from
<collectingCountry id="D5ACB296FFFFFFEF1987F8FBCA45598F" box="[151,263,1915,1941]" name="Mauritius" pageId="9" pageNumber="386">Mauritius</collectingCountry>
, but compared to the
<typeStatus id="72004CA4FFFFFFEF18E8F8FBC969598F" box="[504,555,1915,1941]" pageId="9" pageNumber="386">type</typeStatus>
, the anterolateral margin he figured was very spinose and the ocular peduncles were too short. This specimen of
<bibRefCitation id="C92A8FF7FFFFFFEF1B1DF81FC9FA59A3" author="Bouvier, E. L." box="[525,696,1950,1977]" pageId="9" pageNumber="386" pagination="178 - 318" refId="ref5866" refString="Bouvier, E. L. (1915) Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 7 e Serie, 48 (3), 178 - 318, pls. IV - VII." type="journal article" year="1915">Bouvier (1915)</bibRefCitation>
almost certainly belongs to another species.
</paragraph>
<paragraph id="AD04F206FFFFFFEC19D7F843CA2C5F07" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,150,1761]" lastPageId="10" lastPageNumber="387" pageId="9" pageNumber="386">
<bibRefCitation id="C92A8FF7FFFFFFEF19D7F843CA1859C7" author="Ward, M." box="[199,346,1987,2013]" pageId="9" pageNumber="386" pagination="49 - 113" refId="ref9388" refString="Ward, M. (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2), 49 - 113." type="journal article" year="1942">Ward (1942)</bibRefCitation>
reported on the brachyuran crabs from
<collectingCountry id="D5ACB296FFFFFFEF1A2AF843C8E559C7" box="[826,935,1987,2013]" name="Mauritius" pageId="9" pageNumber="386">Mauritius</collectingCountry>
and adjacent islands. He listed
<taxonomicName id="6ABB8985FFFFFFEC1C38F843CA855EAB" authority="Bouvier, 1915" authorityName="Bouvier" authorityYear="1915" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" lastPageId="10" lastPageNumber="387" order="Decapoda" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="9FCF2E14FFFFFFEC1C38F843CA595EAB" italics="true" lastPageId="10" lastPageNumber="387" pageId="9" pageNumber="386">Litocheira decharmoyi</emphasis>
<bibRefCitation id="C92A8FF7FFFCFFEC1834FF17CA855EAB" author="Bouvier, E. L." box="[292,455,150,177]" pageId="10" pageNumber="387" pagination="178 - 318" refId="ref5866" refString="Bouvier, E. L. (1915) Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 7 e Serie, 48 (3), 178 - 318, pls. IV - VII." type="journal article" year="1915">Bouvier, 1915</bibRefCitation>
</taxonomicName>
, as belonging to
<taxonomicName id="6ABB8985FFFCFFEC1B99FF17C87E5EAB" authorityName="Ward" authorityYear="1942" box="[649,828,151,177]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1B99FF17C87E5EAB" box="[649,828,151,177]" italics="true" pageId="10" pageNumber="387">Pseudolitochira</emphasis>
</taxonomicName>
, but did not provide an explanation (
<bibRefCitation id="C92A8FF7FFFCFFEC1DF2FF17CED25EAB" author="Ward, M." box="[1250,1424,150,177]" pageId="10" pageNumber="387" pagination="49 - 113" refId="ref9388" refString="Ward, M. (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2), 49 - 113." type="journal article" year="1942">Ward 1942: 52</bibRefCitation>
). He (
<bibRefCitation id="C92A8FF7FFFCFFEC19D9FF3BCAC45ECF" author="Ward, M." box="[201,390,187,213]" pageId="10" pageNumber="387" pagination="49 - 113" refId="ref9388" refString="Ward, M. (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2), 49 - 113." type="journal article" year="1942">Ward 1942: 101</bibRefCitation>
, pl. 6, fig. 5) also stated that
<specimenCount id="BBBD398FFFFCFFEC1BF1FF3BC8075ECF" box="[737,837,187,213]" pageId="10" pageNumber="387" type="female">2 female</specimenCount>
specimens (15 and
<quantity id="6A435FE3FFFCFFEC1D3EFF3BCF3D5ECF" box="[1070,1151,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="387" unit="mm" value="11.0">11 mm</quantity>
in carapace width) of
<taxonomicName id="6ABB8985FFFCFFEC1C98FF3BCBAA5EE2" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1C98FF3BCBAA5EE2" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
were examined from Chagos Islands. From his figure, this material is probably conspecific with
<taxonomicName id="6ABB8985FFFCFFEC1C3FFF5FCEDE5EE2" box="[1327,1436,223,248]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1C3FFF5FCEDE5EE2" box="[1327,1436,223,248]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
as recognised here.
</paragraph>
<paragraph id="AD04F206FFFCFFEC19D7FEA7C9475C27" blockId="10.[151,1437,150,1761]" pageId="10" pageNumber="387">
Further clarification is required for the identity of a number of records assigned to
<taxonomicName id="6ABB8985FFFCFFEC1D7BFEA7CF9A5F5A" box="[1131,1240,295,320]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1D7BFEA7CF9A5F5A" box="[1131,1240,295,320]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
. From the Indian Ocean, Innocenti &amp; Maz (2017: 47) listed
<specimenCount id="BBBD398FFFFCFFEC1B9AFECBC8715F7F" box="[650,819,331,357]" pageId="10" pageNumber="387" type="generic">two specimens</specimenCount>
from
<collectingCountry id="D5ACB296FFFCFFEC1A6DFECBC89E5F7F" box="[893,988,331,357]" name="Somalia" pageId="10" pageNumber="387">Somalia</collectingCountry>
, but they did not figure their material. Workers from
<collectingCountry id="D5ACB296FFFCFFEC182DFEEFCA355F93" box="[317,375,367,393]" name="India" pageId="10" pageNumber="387">India</collectingCountry>
and its nearby islands have recorded and/or listed
<taxonomicName id="6ABB8985FFFCFFEC1AABFEEFCF6B5F92" box="[955,1065,367,392]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1AABFEEFCF6B5F92" box="[955,1065,367,392]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
on several occasions (
<bibRefCitation id="C92A8FF7FFFCFFEC1C3BFEEFCB905FB7" author="Rao, G. S. &amp; Suseelan, C. &amp; Kathirvel, M." pageId="10" pageNumber="387" pagination="72 - 76" refId="ref8558" refString="Rao, G. S., Suseelan, C. &amp; Kathirvel, M. (1989) Crustacean resources of the Lakshadweep Islands. In: Suseelan, C. (Ed.), CMFRI Bulletin 43. Marine Living Resources of the Union Territory of Lakshadweep. An Indicative Survey with Suggestions for Development. Central Marine Fisheries Research Institute, Cochin, pp. 72 - 76," type="book chapter" year="1989">
Rao
<emphasis id="9FCF2E14FFFCFFEC1C73FEEFCEDF5F93" box="[1379,1437,367,393]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
1989
</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC19CFFE13C9455FB7" author="Venkataraman, K. &amp; Jeyabaskaran, R. &amp; Raghuram, K. P. &amp; Alfred, J. R. B." box="[223,519,403,429]" pageId="10" pageNumber="387" pagination="1 - 468" refId="ref9328" refString="Venkataraman, K., Jeyabaskaran, R., Raghuram, K. P. &amp; Alfred, J. R. B. (2004) Bibliography and checklist of corals and coral reef associated organisms of India. Records of the Zoological Survey of India, Occasional Paper, No. 226, 1 - 468." type="book chapter" year="2004">
Venkataraman
<emphasis id="9FCF2E14FFFCFFEC1898FE13CA805FB7" box="[392,450,403,429]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1B04FE13C9FC5FB7" author="Dev Roy, M. K." box="[532,702,403,429]" pageId="10" pageNumber="387" pagination="1 - 212" refId="ref6108" refString="Dev Roy, M. K. (2008) An annotated checklist of mangrove and coral reef inhabiting brachyuran crabs of India, India. Records of the Zoological Survey of India, Occasional Paper, No. 289, 1 - 212." type="book chapter" year="2008">Dev Roy 2008</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1BDBFE13C87F5FB7" author="Rao, D. V." box="[715,829,403,429]" pageId="10" pageNumber="387" refId="ref8526" refString="Rao, D. V. (2010) Field Guide to Corals and Coral Associates of Andaman and Nicobar Islands. Zoological Survey of India, Kolkata, 283 pp." type="book" year="2010">Rao 2010</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1A5AFE13CF775FB7" author="Dev Roy, M. K. &amp; Bhadra, S." box="[842,1077,403,429]" pageId="10" pageNumber="387" pagination="109 - 269" refId="ref6153" refString="Dev Roy, M. K. &amp; Bhadra, S. (2011) Brachyuran crabs (Crustacea: Decapoda: Brachyura). In: State Fauna Series. 17. Part- 2. Fauna of Tamil Nadu. Zoological Survey of India, Kolkata. Pp. 109 - 269," type="book chapter" year="2011">
Dev Roy
<emphasis id="9FCF2E14FFFCFFEC1AA9FE13C8B15FB7" box="[953,1011,403,429]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
2011
</bibRefCitation>
,
<bibRefCitation id="C92A8FF7FFFCFFEC1D52FE13CF3C5FB7" author="Dev Roy, M. K. &amp; Nandi, N. C." box="[1090,1150,403,429]" pageId="10" pageNumber="387" pagination="185 - 236" refId="ref6207" refString="Dev Roy, M. K. &amp; Nandi, N. C. (2012) Brachyuran crabs (Crustacea). In: State Fauna Series. 19. Part- 1. Fauna of Andaman and Nicobar Islands. Zoological Survey of India, Kolkata, pp. 185 - 236," type="book chapter" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1D9AFE13CE215FB7" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[1162,1379,403,429]" pageId="10" pageNumber="387" pagination="1 - 83" refId="ref9091" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. &amp; Ng, P. K. L. (2018) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi
<emphasis id="9FCF2E14FFFCFFEC1DF5FE13CE5D5FB7" box="[1253,1311,403,429]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
2018
</bibRefCitation>
), but their specimens will need to be re-examined to see if they are really this species. The specimens of
<taxonomicName id="6ABB8985FFFCFFEC1DF9FE37CEDE5FCB" authorityName="De Man" authorityYear="1895" box="[1257,1436,439,465]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1DF9FE37CEDE5FCB" box="[1257,1436,439,465]" italics="true" pageId="10" pageNumber="387">Heteropilumnus</emphasis>
</taxonomicName>
or
<taxonomicName id="6ABB8985FFFCFFEC19A7FE5BCA865FEE" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[183,452,475,501]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC19A7FE5BCA865FEE" box="[183,452,475,501]" italics="true" pageId="10" pageNumber="387">Pseudolitochira integra</emphasis>
</taxonomicName>
reported from the central Pacific and
<collectingCountry id="D5ACB296FFFCFFEC1A69FE5BCF765FEF" box="[889,1076,475,501]" name="Marshall Islands" pageId="10" pageNumber="387">Marshall Islands</collectingCountry>
by
<bibRefCitation id="C92A8FF7FFFCFFEC1D72FE5BCE545FEF" author="Holthuis, L. B." box="[1122,1302,474,501]" pageId="10" pageNumber="387" pagination="1 - 66" refId="ref6380" refString="Holthuis, L. B. (1953) Enumeration of the decapod and stomatopod Crustacea from Pacific coral islands. Atoll Research Bulletin, 24, 1 - 66, 2 maps. https: // doi. org / 10.5479 / si. 00775630.24.1" type="book chapter" year="1953">Holthuis (1953)</bibRefCitation>
and
<bibRefCitation id="C92A8FF7FFFCFFEC1C40FE5BCA615C03" author="Paulay, G. &amp; Kropp, R. &amp; Ng, P. K. L. &amp; Eldredge, L. G." pageId="10" pageNumber="387" pagination="456 - 513" refId="ref8459" refString="Paulay, G., Kropp, R., Ng, P. K. L. &amp; Eldredge, L. G. (2003) The crustaceans and pycnogonids of the Mariana Islands. In: Paulay, G. (Ed.), The Marine Biodiversity of Guam and the Marianas. Micronesica, 35 - 36, 456 - 513." type="journal article" year="2003">
Paulay
<emphasis id="9FCF2E14FFFCFFEC1987FE7FCB925C03" box="[151,208,511,537]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
(2003)
</bibRefCitation>
will all need to re-examined. Considering the distribution, it seems unlikely that this material will be conspecific with
<taxonomicName id="6ABB8985FFFCFFEC1846FDA3CA815C26" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[342,451,547,572]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" sensu="stricto" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1846FDA3CA815C26" box="[342,451,547,572]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84FC936FFFFCFFEC18DAFDA3C9475C27" box="[458,517,547,573]" pageId="10" pageNumber="387" sensu="stricto">s. str.</taxonomicNameLabel>
</paragraph>
<paragraph id="AD04F206FFFCFFEC19D7FDC7C8025B2B" blockId="10.[151,1437,150,1761]" pageId="10" pageNumber="387">
The first record of
<taxonomicName id="6ABB8985FFFCFFEC18B9FDC7C95B5C7A" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[425,537,583,608]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC18B9FDC7C95B5C7A" box="[425,537,583,608]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
from the western Pacific was by T.
<bibRefCitation id="C92A8FF7FFFCFFEC1AD7FDC7CFCD5C7B" author="Sakai, T." box="[967,1167,582,609]" pageId="10" pageNumber="387" pagination="106 - 113" refId="ref8754" refString="Sakai, T. (1955) On some rare species of crabs from Japan. Bulletin of the Biogeographical Society of Japan, 16 - 19, 106 - 113." type="journal article" year="1955">Sakai (1955: 108</bibRefCitation>
, text-fig. 3) who listed the species (as
<taxonomicName id="6ABB8985FFFCFFEC1855FDEBC9515C9E" box="[325,531,619,645]" class="Malacostraca" family="Litocheiridae" genus="Litocheira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1855FDEBC9515C9E" box="[325,531,619,645]" italics="true" pageId="10" pageNumber="387">Litocheira integra</emphasis>
</taxonomicName>
) from the Bonin Islands (present day Ogasawara Islands) in
<collectingCountry id="D5ACB296FFFCFFEC1DDEFDEBCE535C9F" box="[1230,1297,619,645]" name="Japan" pageId="10" pageNumber="387">Japan</collectingCountry>
. Tune
<bibRefCitation id="C92A8FF7FFFCFFEC1C4EFDEBCA555CB3" author="Sakai, T." pageId="10" pageNumber="387" refId="ref8786" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1 - 3. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp., 251 pls. (plates volume)." type="book" year="1976">Sakai (1976: 493</bibRefCitation>
, text-fig. 264a) placed the species in
<taxonomicName id="6ABB8985FFFCFFEC1BDCFD0FC83D5CB3" authorityName="De Man" authorityYear="1895" box="[716,895,655,681]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1BDCFD0FC83D5CB3" box="[716,895,655,681]" italics="true" pageId="10" pageNumber="387">Heteropilumnus</emphasis>
</taxonomicName>
(as
<taxonomicName id="6ABB8985FFFCFFEC1AA4FD0FCF805CB2" box="[948,1218,655,681]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1AA4FD0FCF805CB2" box="[948,1218,655,681]" italics="true" pageId="10" pageNumber="387">Heteropilumnus integra</emphasis>
</taxonomicName>
), commenting that his female specimen (5.5 ×
<quantity id="6A435FE3FFFCFFEC18DDFD32C9665CD7" box="[461,548,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="387" unit="mm" value="3.5">3.5 mm</quantity>
) matched the original descriptions except that the anterolateral margin had two rudimentary indentations whereas in the
<typeStatus id="72004CA4FFFCFFEC1B76FD57C9D45CEB" box="[614,662,727,753]" pageId="10" pageNumber="387">type</typeStatus>
female, it was entire. Since then, the name (as a
<taxonomicName id="6ABB8985FFFCFFEC1DD9FD57CE3E5CEB" authorityName="De Man" authorityYear="1895" box="[1225,1404,727,753]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1DD9FD57CE3E5CEB" box="[1225,1404,727,753]" italics="true" pageId="10" pageNumber="387">Heteropilumnus</emphasis>
</taxonomicName>
or
<taxonomicName id="6ABB8985FFFCFFEC1987FD7BCA085D0F" authorityName="Ward" authorityYear="1942" box="[151,330,763,789]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1987FD7BCA085D0F" box="[151,330,763,789]" italics="true" pageId="10" pageNumber="387">Pseudolitochira</emphasis>
</taxonomicName>
) has been used in various reports and lists from
<collectingCountry id="D5ACB296FFFCFFEC1A4EFD7BC8DC5D0F" box="[862,926,763,789]" name="Japan" pageId="10" pageNumber="387">Japan</collectingCountry>
and
<collectingCountry id="D5ACB296FFFCFFEC1AC2FD7BCF665D0F" box="[978,1060,763,789]" name="Taiwan" pageId="10" pageNumber="387">Taiwan</collectingCountry>
(
<bibRefCitation id="C92A8FF7FFFCFFEC1D21FD7BCE7E5D0F" author="Takeda, M. &amp; Miyake, S." box="[1073,1340,763,789]" pageId="10" pageNumber="387" pagination="101 - 115" refId="ref9006" refString="Takeda, M. &amp; Miyake, S. (1976) Crabs of the Ogasawara Islands. I. List of the known species. Researches on Crustacea, 7, 101 - 115. https: // doi. org / 10.18353 / rcustacea. 7.0 _ 101" type="journal article" year="1976">Takeda &amp; Miyake 1976</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1C57FD7BCB905D23" author="Miyake, S." pageId="10" pageNumber="387" refId="ref7781" refString="Miyake, S. (1983) Brachyura (Crabs). Japanese crustacean decapods and stomatopods in Color. Vol. 2. 1 st Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered fig." type="book" year="1983">Miyake 1983</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC19CFFC9FCA005D23" author="Ito, J." box="[223,322,799,825]" pageId="10" pageNumber="387" pagination="39 - 52" refId="ref6661" refString="Ito, J. (1992) Local marine animals 18. Fukui City Museum of Natural History Research Report, 39, 39 - 52." type="journal article" year="1992">Ito 1992</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC185FFC9FC96D5D23" author="Nomura, K. &amp; Nagai, S. &amp; Asakura, A. &amp; Komai, T." box="[335,559,799,825]" pageId="10" pageNumber="387" pagination="7 - 21" refId="ref8406" refString="Nomura, K., Nagai, S., Asakura, A. &amp; Komai, T. (1996) A preliminary list of shallow water decapod Crustacea in the Kerama Group, the Ryukyu Archipelago. Bulletin of the Biogeographical Society of Japan, 51, 7 - 21." type="journal article" year="1996">
Nomura
<emphasis id="9FCF2E14FFFCFFEC18A3FC9FCAAE5D23" box="[435,492,799,825]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
1996
</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1B2CFC9FC9F15D23" author="Jeng, M. - S." box="[572,691,799,825]" pageId="10" pageNumber="387" refId="ref6688" refString="Jeng, M. - S. (1997) Studies on the land and aquatic decapod crustacean fauna of the Kenting National Park (11) - Communities of decapod crustaceans around the sea. Kenting National Park, Ministry of the Interior, Pingtung, ii + 66 pp. [in Chinese]" type="book" year="1997">Jeng 1997</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1BD0FC9FC8255D23" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[704,871,799,825]" pageId="10" pageNumber="387" pagination="1 - 86" refId="ref8340" refString="Ng, P. K. L., Wang, C. - H., Ho, P. - H. &amp; Shih, H. - T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11, i - iv + 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="9FCF2E14FFFCFFEC1BFAFC9FC8665D23" box="[746,804,799,825]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
2001
</bibRefCitation>
,
<bibRefCitation id="C92A8FF7FFFCFFEC1A63FC9FC8EC5D23" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[883,942,799,825]" pageId="10" pageNumber="387" pagination="1 - 185" refId="ref8085" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 &amp; 4), 1 - 185. https: // doi. org / 10.6532 / JNTM. 20171270 (3; 4). 01" type="journal article" year="2017">2017</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1AABFC9FCF0B5D23" author="Motoh, H." box="[955,1097,799,825]" pageId="10" pageNumber="387" pagination="55 - 68" refId="ref7754" refString="Motoh, H. (2003) I. Seafood crabs. Known species. Noto Marine Center Research Report, 9, 55 - 68." type="journal article" year="2003">Motoh 2003</bibRefCitation>
; K.
<bibRefCitation id="C92A8FF7FFFCFFEC1D6AFC9FCFB95D23" author="Sakai, K." box="[1146,1275,799,825]" pageId="10" pageNumber="387" pagination="13 - 30" refId="ref8732" refString="Sakai, K. (2003) Standard Japanese names for Japanese crabs. Taxon, 15, 13 - 30." type="journal article" year="2003">Sakai 2003</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC1C18FC9FCA6C5D47" author="Marumura, M. &amp; Kosaka, A." pageId="10" pageNumber="387" refId="ref7423" refString="Marumura, M. &amp; Kosaka, A. (2003) Catalogue of brachyuran and anomuran crabs collection donated by the late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, Wakayama, 73 pp., 8 pls." type="book" year="2003">Marumura &amp; Kosaka 2003</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC182CFCC3CAA15D47" author="Komatsu, H." box="[316,483,835,861]" pageId="10" pageNumber="387" pagination="219 - 277" refId="ref6821" refString="Komatsu, H. (2011) Crabs dredged off the Ogasawara Islands (Crustacea, Decapoda, Brachyura). Memoirs of the National Science Museum, 47, 219 - 277." type="journal article" year="2011">Komatsu 2011</bibRefCitation>
;
<bibRefCitation id="C92A8FF7FFFCFFEC18E0FCC3C9C75D47" author="Naruse, T." box="[496,645,835,861]" pageId="10" pageNumber="387" pagination="1 - 72" refId="ref7864" refString="Naruse, T. (2010) Catalogue of Crustacean Specimens deposited in the University Museum (Fujukan), University of the Ryukyus. Catalogue of Materials Deposited in the University Museum (Fujukan), University of the Ryukyus, 3, 1 - 72." type="journal article" year="2010">Naruse 2010</bibRefCitation>
).
<bibRefCitation id="C92A8FF7FFFCFFEC1B88FCC3C82C5D47" author="Maenosono, T." box="[664,878,835,861]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
showed that the material listed as
<bibRefCitation id="C92A8FF7FFFCFFEC1DEAFCC3CEDE5D47" author="Naruse, T." box="[1274,1436,835,861]" pageId="10" pageNumber="387" pagination="1 - 72" refId="ref7864" refString="Naruse, T. (2010) Catalogue of Crustacean Specimens deposited in the University Museum (Fujukan), University of the Ryukyus. Catalogue of Materials Deposited in the University Museum (Fujukan), University of the Ryukyus, 3, 1 - 72." type="journal article" year="2010">Naruse (2010)</bibRefCitation>
from Okinawa was mixed, containing both what he regarded as
<taxonomicName id="6ABB8985FFFCFFEC1A74FCE7C8935D9A" box="[868,977,871,896]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1A74FCE7C8935D9A" box="[868,977,871,896]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
as well as
<taxonomicName id="6ABB8985FFFCFFEC1D5CFCE7CB935DBF" authority="Rathbun, 1911" authorityName="Rathbun" authorityYear="1911" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="turgidulus">
<emphasis id="9FCF2E14FFFCFFEC1D5CFCE7CE6D5D9B" box="[1100,1327,871,897]" italics="true" pageId="10" pageNumber="387">Pilumnus turgidulus</emphasis>
<bibRefCitation id="C92A8FF7FFFCFFEC1C26FCE7CB935DBF" author="Rathbun, M. J." pageId="10" pageNumber="387" pagination="191 - 261" refId="ref8631" refString="Rathbun, M. J. (1911) Marine Brachyura. In: The Percy Sladen Trust expedition to the Indian Ocean in 1905, Under the leadership of Mr. J. Stanley Gardiner. Vol. III. No. XI. Transactions of the Linnean Society of London, Zoology, Series 2 (Zoology), 14 (2), 191 - 261, figs. 1 &amp; 2, pls. 15 - 20. https: // doi. org / 10.1111 / j. 1096 - 3642.1911. tb 00084. x" type="journal article" year="1911">Rathbun, 1911</bibRefCitation>
</taxonomicName>
. The record of
<taxonomicName id="6ABB8985FFFCFFEC1891FC0BCAAC5DBE" box="[385,494,907,932]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1891FC0BCAAC5DBE" box="[385,494,907,932]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
by
<bibRefCitation id="C92A8FF7FFFCFFEC1B0BFC0BC9925DBF" author="Machida, Y." box="[539,720,907,933]" pageId="10" pageNumber="387" pagination="52 - 57" refId="ref7109" refString="Machida, Y. (2017) Three rare species of crab obtained from the gravel intertidal zone of Uchinoura Bay. Shikoku Natural History Science Research, 10, 52 - 57." type="journal article" year="2017">Machida (2017)</bibRefCitation>
is incorrect and
<bibRefCitation id="C92A8FF7FFFCFFEC1A80FC0BCF245DBF" author="Maenosono, T." box="[912,1126,907,933]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
referred it to
<taxonomicName id="6ABB8985FFFCFFEC1C15FC0BC9775DD0" authority="(Takeda &amp; Miyake, 1969)" baseAuthorityName="Takeda &amp; Miyake" baseAuthorityYear="1969" class="Malacostraca" family="Pilumnidae" genus="Zehntneriana" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="amakusae">
<emphasis id="9FCF2E14FFFCFFEC1C15FC0BCA455DD3" italics="true" pageId="10" pageNumber="387">Zehntneriana amakusae</emphasis>
(
<bibRefCitation id="C92A8FF7FFFCFFEC1807FC2FC96F5DD0" author="Takeda, M. &amp; Miyake, S." box="[279,557,943,970]" pageId="10" pageNumber="387" pagination="9 - 15" refId="ref8954" refString="Takeda, M. &amp; Miyake, S. (1969) A new species of the family Goneplacidae (Crustacea, Brachyura) from the Amakusa Islands, Kyushu. OHMU, Zoological Laboratory, Faculty of Agriculture, Kyushu University, 2 (1), 9 - 15." type="journal article" year="1969">Takeda &amp; Miyake, 1969</bibRefCitation>
)
</taxonomicName>
instead.
<bibRefCitation id="C92A8FF7FFFCFFEC1B8CFC2FC82E5DD3" author="Maenosono, T." box="[668,876,943,969]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019</bibRefCitation>
: fig. 11A, B, C) described and figured a series of specimens from Okinawa at length and his material agrees well with that reported by T.
<bibRefCitation id="C92A8FF7FFFCFFEC1D61FC53CFBB5DF7" author="Sakai, T." box="[1137,1273,978,1005]" pageId="10" pageNumber="387" pagination="106 - 113" refId="ref8754" refString="Sakai, T. (1955) On some rare species of crabs from Japan. Bulletin of the Biogeographical Society of Japan, 16 - 19, 106 - 113." type="journal article" year="1955">Sakai (1955</bibRefCitation>
,
<bibRefCitation id="C92A8FF7FFFCFFEC1C15FC53CE7C5DF4" author="Sakai, T." box="[1285,1342,979,1006]" pageId="10" pageNumber="387" refId="ref8786" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1 - 3. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp., 251 pls. (plates volume)." type="book" year="1976">1976</bibRefCitation>
) as well as that figured in
<bibRefCitation id="C92A8FF7FFFCFFEC184DFC77C9345A0B" author="Nagai, S. &amp; Nomura, K." box="[349,630,1015,1041]" pageId="10" pageNumber="387" refId="ref7826" refString="Nagai, S. &amp; Nomura, K. (1988) Crustacea (Brachyura). The guide book of marine organisms of Okinawa. Vol. 7. Shinsei Tosho, Okinawa, 250 pp." type="book" year="1988">Nagai &amp; Nomura (1988)</bibRefCitation>
. His specimens compare well with the
<typeStatus id="72004CA4FFFCFFEC1D3DFC77CF1F5A0B" box="[1069,1117,1015,1041]" pageId="10" pageNumber="387">type</typeStatus>
of
<taxonomicName id="6ABB8985FFFCFFEC1D94FC77CFB35A0A" box="[1156,1265,1015,1040]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1D94FC77CFB35A0A" box="[1156,1265,1015,1040]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
except that the carapace is proportionately much wider (width to length ratio 1.8), the frontal margin is not protruding, being only level with the orbits and the anterolateral margin is differently structured, with the lateral area armed with a low lobiform tooth. Despite these similarities, confirmation is required as to the identity of “
<taxonomicName id="6ABB8985FFFCFFEC1D66FBE3CFA05A66" box="[1142,1250,1123,1148]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1D66FBE3CFA05A66" box="[1142,1250,1123,1148]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
” by
<bibRefCitation id="C92A8FF7FFFCFFEC1C0AFBE3CBA75ABB" author="Maenosono, T." pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
. The structures of the posterior margin of the epistome and third maxillipeds are not clear from his figures (
<bibRefCitation id="C92A8FF7FFFCFFEC198FFB2BCA255ADF" author="Maenosono, T." box="[159,359,1195,1221]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 11C) but appear to agree; and the condition of the male pleon (particularly the transversely narrower somites 1 and 3) agrees more with what is known for
<taxonomicName id="6ABB8985FFFCFFEC1A4CFB4FCF4D5AF3" authorityName="De Man" authorityYear="1895" box="[860,1039,1231,1257]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="9FCF2E14FFFCFFEC1A4CFB4FCF4D5AF3" box="[860,1039,1231,1257]" italics="true" pageId="10" pageNumber="387">Heteropilumnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84FC936FFFFCFFEC1D07FB4FCF105AF3" box="[1047,1106,1231,1257]" pageId="10" pageNumber="387" sensu="stricto">s. str.</taxonomicNameLabel>
(see
<bibRefCitation id="C92A8FF7FFFCFFEC1D9CFB4FCE725AF3" author="Ng, P. K. L. &amp; Lin, C. - W. &amp; Ho, P. - H." box="[1164,1328,1231,1257]" pageId="10" pageNumber="387" pagination="1 - 17" refId="ref8261" refString="Ng, P. K. L., Lin, C. - W. &amp; Ho, P. - H. (2018) On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on the genus Heteropilumnus De Man, 1895 (Crustacea: Brachyura). Zoological Studies, 57 (12), 1 - 17. https: // doi. org / 10.6620 / ZS. 2018.57 - 12" type="journal article" year="2018">
Ng
<emphasis id="9FCF2E14FFFCFFEC1DA5FB4FCFAC5AF3" box="[1205,1262,1231,1257]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
2018
</bibRefCitation>
: figs. 7D, 8D). The male and female specimens of
<bibRefCitation id="C92A8FF7FFFCFFEC1B46FB73C8685B17" author="Maenosono, T." box="[598,810,1267,1293]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
will need to re-examined to see if material is congeneric with
<taxonomicName id="6ABB8985FFFCFFEC19C0FA97CA7E5B2A" box="[208,316,1303,1328]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC19C0FA97CA7E5B2A" box="[208,316,1303,1328]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
, but regardless, it is certainly not conspecific.
</paragraph>
<paragraph id="AD04F206FFFCFFEC19D7FABBC8FC5BFF" blockId="10.[151,1437,150,1761]" pageId="10" pageNumber="387">
The material recorded from the
<collectingCountry id="D5ACB296FFFCFFEC1B41FABBC9FC5B4F" box="[593,702,1339,1365]" name="Indonesia" pageId="10" pageNumber="387">Moluccas</collectingCountry>
will also need re-examination. The material identified as
<taxonomicName id="6ABB8985FFFCFFEC1C96FABBCA4D5B63" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="subintegra">
<emphasis id="9FCF2E14FFFCFFEC1C96FABBCA4D5B63" italics="true" pageId="10" pageNumber="387">L. subintegra</emphasis>
</taxonomicName>
by
<bibRefCitation id="C92A8FF7FFFCFFEC182BFADFC9405B63" author="Tesch, J. J." box="[315,514,1374,1401]" pageId="10" pageNumber="387" pagination="149 - 295" refId="ref9056" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918: 175</bibRefCitation>
, pl. 16, fig. 1) is probably another species, its quadrate carapace, weakly marked anterolateral margin and long ambulatory legs are completely different from the
<typeStatus id="72004CA4FFFCFFEC1D07FA03CF055B87" box="[1047,1095,1411,1437]" pageId="10" pageNumber="387">type</typeStatus>
female of
<taxonomicName id="6ABB8985FFFCFFEC1DAEFA03CE685B86" box="[1214,1322,1411,1436]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1DAEFA03CE685B86" box="[1214,1322,1411,1436]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
as well as
<taxonomicName id="6ABB8985FFFCFFEC1987FA27CAF15BDB" authorityName="Ng &amp; Tan" authorityYear="1988" box="[151,435,1447,1473]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFCFFEC1987FA27CAF15BDB" box="[151,435,1447,1473]" italics="true" pageId="10" pageNumber="387">Heteropilumnus holthuisi</emphasis>
</taxonomicName>
.
<bibRefCitation id="C92A8FF7FFFCFFEC18D0FA27C93D5BDB" author="Ng, P. K. L. &amp; Tan, L. W. H." box="[448,639,1447,1473]" pageId="10" pageNumber="387" pagination="13 - 24" refId="ref8172" refString="Ng, P. K. L. &amp; Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng &amp; Tan (1988)</bibRefCitation>
assigned it to
<taxonomicName id="6ABB8985FFFCFFEC1A0EFA27C8E05BDB" authorityName="Ng &amp; Tan" authorityYear="1988" box="[798,930,1447,1473]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="9FCF2E14FFFCFFEC1A0EFA27C8E05BDB" box="[798,930,1447,1473]" italics="true" pageId="10" pageNumber="387">H. holthuisi</emphasis>
</taxonomicName>
with doubt but this seems unlikely. The same is true for the record of
<taxonomicName id="6ABB8985FFFCFFEC18B1FA4BC94C5BFE" box="[417,526,1483,1508]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC18B1FA4BC94C5BFE" box="[417,526,1483,1508]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
by
<bibRefCitation id="C92A8FF7FFFCFFEC1B27FA4BC8565BFF" author="Serene, R. &amp; Romimohtarto, K. &amp; Moosa, M. K." box="[567,788,1483,1509]" pageId="10" pageNumber="387" pagination="17 - 26" refId="ref8893" refString="Serene, R., Romimohtarto, K. &amp; Moosa, M. K. (1974) The Hippidea and Brachyura collected by the Rumphius Expedition I. In: Report on the Rumphius Expedition I. (January 6 - February 1, 1973). Oseanologi di Indonesia, 1, pp. 17 - 26." type="journal article" year="1974">
Serène
<emphasis id="9FCF2E14FFFCFFEC1B9BFA4BC9815BFF" box="[651,707,1483,1509]" italics="true" pageId="10" pageNumber="387">et al.</emphasis>
(1974)
</bibRefCitation>
from this area.
</paragraph>
<paragraph id="AD04F206FFFCFFEC19D7FA6FCAF158FB" blockId="10.[151,1437,150,1761]" pageId="10" pageNumber="387">
The main issue with the identification of
<taxonomicName id="6ABB8985FFFCFFEC1B9EFA6FC9B85812" box="[654,762,1519,1544]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1B9EFA6FC9B85812" box="[654,762,1519,1544]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
, however, is that to date this species has only been described from female specimens and consequently the morphology of the male pleopod remains unknown.
<bibRefCitation id="C92A8FF7FFFCFFEC1DE9F993CB885848" author="Turkay, M." pageId="10" pageNumber="387" pagination="103 - 132" refId="ref9164" refString="Turkay, M. (1975) Zur Kenntnis der Gattung Euchirograpsus mit Bemerkungen zu Brachygrapsus und Litocheira (Crustacea: Decapoda). Senckenbergiana Biologica, 56 (1 / 3), 103 - 132." type="journal article" year="1975">Türkay (1975: 126)</bibRefCitation>
discussed this problem about the lack of male specimens. That being said, the Japanese male figured by
<bibRefCitation id="C92A8FF7FFFCFFEC1987F9DBCA28586F" author="Maenosono, T." box="[151,362,1627,1653]" pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
is probably shared by congeners and his specimens probably belong to
<taxonomicName id="6ABB8985FFFCFFEC1D6BF9DBCE6C586F" authorityName="Ward" authorityYear="1942" box="[1147,1326,1627,1653]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC1D6BF9DBCE6C586F" box="[1147,1326,1627,1653]" italics="true" pageId="10" pageNumber="387">Pseudolitochira</emphasis>
</taxonomicName>
, although it is probably not
<taxonomicName id="6ABB8985FFFCFFEC1875F9FFCA915882" box="[357,467,1663,1688]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="9FCF2E14FFFCFFEC1875F9FFCA915882" box="[357,467,1663,1688]" italics="true" pageId="10" pageNumber="387">P. integra</emphasis>
</taxonomicName>
as redescribed here. The male pleon and male first gonopod he figured (
<bibRefCitation id="C92A8FF7FFFCFFEC1C08F9FFCB9158A7" author="Maenosono, T." pageId="10" pageNumber="387" pagination="19 - 44" refId="ref7143" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. (In Japanese with English abstract)" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: figs. 13C, 14M, N) are typical for pilumnids, resemble those of
<taxonomicName id="6ABB8985FFFCFFEC1ABBF923CFE558A6" box="[939,1191,1699,1725]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="satrai">
<emphasis id="9FCF2E14FFFCFFEC1ABBF923CFE558A6" box="[939,1191,1699,1725]" italics="true" pageId="10" pageNumber="387">Heteropilumnus satrai</emphasis>
</taxonomicName>
and are essentially of the
<taxonomicName id="6ABB8985FFFCFFEC19D0F947CA3158FB" authorityName="De Man" authorityYear="1895" box="[192,371,1735,1761]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="9FCF2E14FFFCFFEC19D0F947CA3158FB" box="[192,371,1735,1761]" italics="true" pageId="10" pageNumber="387">Heteropilumnus</emphasis>
</taxonomicName>
<typeStatus id="72004CA4FFFCFFEC186BF947CAED58FB" box="[379,431,1735,1761]" pageId="10" pageNumber="387">type</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>