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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.966.51921" ID-GBIF-Dataset="57654c43-1ab2-4bf8-8f9c-6c8fae3b9c37" ID-PMC="PMC7494659" ID-Pensoft-Pub="1313-2970-966-153" ID-Pensoft-UUID="4FE5E4CD17135B5289D5F3DC90922CCD" ID-PubMed="32982547" ID-ZooBank="8E1E57D3BE4F452A9C8677B1C4DB3710" ModsDocID="1313-2970-966-153" checkinTime="1599731020277" checkinUser="pensoft" docAuthor="Hsu, Yu-Feng" docDate="2020" docId="0ABA4E9A99AD5BB19E998D73350DB319" docLanguage="en" docName="ZooKeys 966: 153-162" docOrigin="ZooKeys 966" docSource="http://dx.doi.org/10.3897/zookeys.966.51921" docTitle="Famegana nisa Hsu 2020, comb. nov." docType="treatment" docVersion="4" id="4FE5E4CD17135B5289D5F3DC90922CCD" lastPageNumber="153" masterDocId="4FE5E4CD17135B5289D5F3DC90922CCD" masterDocTitle="The identity of Alfred Wallace's mysterious butterfly taxon Lycaena nisa solved: Famegana nisa comb. nov., a senior synonym of F. alsulus (Lepidoptera, Lycaenidae, Polyommatinae)" masterLastPageNumber="162" masterPageNumber="153" pageNumber="153" updateTime="1668169446674" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The identity of Alfred Wallace's mysterious butterfly taxon Lycaena nisa solved: Famegana nisa comb. nov., a senior synonym of F. alsulus (Lepidoptera, Lycaenidae, Polyommatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hsu, Yu-Feng</mods:namePart>
<mods:affiliation>Department of Life Science, National Taiwan Normal University, Taipei, 116, Taiwan, ROC</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-2091-7775</mods:nameIdentifier>
<mods:nameIdentifier type="email">t43018@ntnu.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>966</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>153</mods:start>
<mods:end>162</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.966.51921</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.966.51921</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-966-153</mods:identifier>
<mods:identifier type="ZooBank">8E1E57D3BE4F452A9C8677B1C4DB3710</mods:identifier>
<mods:identifier type="Pensoft-UUID">4FE5E4CD17135B5289D5F3DC90922CCD</mods:identifier>
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<treatment ID-GBIF-Taxon="167581800" LSID="urn:lsid:plazi:treatment:0ABA4E9A99AD5BB19E998D73350DB319" httpUri="http://treatment.plazi.org/id/0ABA4E9A99AD5BB19E998D73350DB319" lastPageNumber="153" pageId="0" pageNumber="153">
<subSubSection pageId="0" pageNumber="153" type="nomenclature">
<paragraph pageId="0" pageNumber="153">
<taxonomicName LSID="0ABA4E9A-99AD-5BB1-9E99-8D73350DB319" authority="(Wallace, 1866)" authorityName="Hsu" authorityYear="2020" baseAuthorityName="Wallace" baseAuthorityYear="1866" class="Insecta" family="Lycaenidae" genus="Famegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Famegana nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa" status="comb. nov.">Famegana nisa (Wallace, 1866)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="153">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">Figures 1-3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="reference_group">
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">Lycaena nisa</taxonomicName>
Wallace, 1866: 360. Type locality: &quot;Takaw, Formosa&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena alsulus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="alsulus">Lycaena alsulus</taxonomicName>
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
, 1869: 75. Type locality: Rockhampton and Upolu [Australia]. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Boisduval" authorityYear="1852" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena exilis" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="exilis">Lycaena exilis</taxonomicName>
Lucas, 1889: 159, figs 13-15. Type locality: Cooktown to Bowen [Australia] (preoccupied by
<taxonomicName authorityName="Boisduval" authorityYear="1852" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena exilis" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="exilis">Lycaena exilis</taxonomicName>
Boisduval, 1852). syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Mathew" authorityYear="1889" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena lulu" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="lulu">Lycaena lulu</taxonomicName>
Mathew, 1889: 312. Type locality: Tongatabu, [Tonga]. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Miskin" authorityYear="1890" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena gracilis" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="gracilis">Lycaena gracilis</taxonomicName>
Miskin, 1890: 37. Type locality: Brisbane to Cooktown [Australia]. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Lucas" authorityYear="1894" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena exiloides" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="exiloides">Lycaena exiloides</taxonomicName>
Lucas, 1891: 47. Replacement name for
<taxonomicName authorityName="Boisduval" authorityYear="1852" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena exilis" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="exilis">Lycaena exilis</taxonomicName>
Lucas, 1889.syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">Zizera nisa</taxonomicName>
:
<bibRefCitation author="Matsumura, S" journalOrPublisher="Zoological Magazine, Tokyo" pageId="0" pageNumber="153" pagination="479 - 481" refId="B20" refString="Matsumura, S, 1909. A list of Formosan butterflies. Zoological Magazine, Tokyo 253: 479 - 481" title="A list of Formosan butterflies." volume="253" year="1909">Matsumura 1909</bibRefCitation>
: 480.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="alsulus">Zizeeria alsulus</taxonomicName>
:
<bibRefCitation author="Waterhouse, GA" journalOrPublisher="Angus &amp; Robertson Publishers, Sydney" pageId="0" pageNumber="153" refId="B34" refString="Waterhouse, GA, Lyell, G, 1914. The Butterflies of Australia. Angus &amp; Robertson Publishers, Sydney" title="The Butterflies of Australia." year="1914">Waterhouse and Lyell 1914</bibRefCitation>
: 106.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera lulu" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="lulu">Zizera lulu</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.1918" author="Rothschild, FRS" journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="153" pagination="387 - 402" refId="B27" refString="Rothschild, FRS, 1915. On the Lepidoptera in the Tring Museum sent by Mr. A. S. Meek from the Admirality Islands, Dampier, and Vulcan Islands. Novitates Zoologicae 22 (3): 387 - 402, DOI: https://doi.org/10.5962/bhl.part.1918" title="On the Lepidoptera in the Tring Museum sent by Mr. A. S. Meek from the Admirality Islands, Dampier, and Vulcan Islands." url="https://doi.org/10.5962/bhl.part.1918" volume="22" year="1915">Rothschild 1915</bibRefCitation>
: 390.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Ribbe" authorityYear="1926" class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera kalawarus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="kalawarus">Zizera kalawarus</taxonomicName>
<bibRefCitation author="Ribbe, C" journalOrPublisher="Entomologische Mitteilungen" pageId="0" pageNumber="153" pagination="78 - 91" refId="B26" refString="Ribbe, C, 1926. Neue Lycaenenformen, hauptsaechlich von Celebes (Lep. Lycaenidae). Entomologische Mitteilungen 15: 78 - 91" title="Neue Lycaenenformen, hauptsaechlich von Celebes (Lep. Lycaenidae)." volume="15" year="1926">Ribbe 1926</bibRefCitation>
: 91;
<bibRefCitation author="Vane Wright, D" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="0" pageNumber="153" pagination="1 - 267" publicationUrl="https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" refId="B35" refString="Vane Wright, D, de Jong, R, 2003. The Butterflies of Sulawesi: Annotated Checklist for a Critical Island Fauna. Zoologische Verhandelingen Leiden 343: 1 - 267, https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" title="The Butterflies of Sulawesi: Annotated Checklist for a Critical Island Fauna." url="https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" volume="343" year="2003">Vane Wright and de Jong 2003</bibRefCitation>
: 155. Type locality: Celebes. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera alsulus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="alsulus">Zizera alsulus</taxonomicName>
:
<bibRefCitation author="Seitz, A" editor="Zeitz, A" journalOrPublisher="IX. Fritz Lehmann Verlag, Stuttgart" pageId="0" pageNumber="153" pagination="925 - 926" refId="B28" refString="Seitz, A, 1927. Genus Zizera. In: Zeitz, A, Ed., The Macrolepidoptera of the World, Vol. IX. Fritz Lehmann Verlag, Stuttgart: 925 - 926" title="Genus Zizera." volumeTitle="The Macrolepidoptera of the World, Vol." year="1927">Seitz 1927</bibRefCitation>
: 926.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">Zizera taiwana</taxonomicName>
Sonan, 1938: 254. Type locality: &quot;Inrin, Formosa.&quot; syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus subsp. eggletoni" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="eggletoni">Zizeeria alsulus eggletoni</taxonomicName>
<bibRefCitation author="Corbet" journalOrPublisher="Angus &amp; Robertson Publishers, Sydney" pageId="0" pageNumber="153" publicationUrl="https://hkjo.lib.hku.hk/exhibits/show/hkjo/browseArticle?book=b27722454&amp;issue=270036" refId="B7" refString="Corbet, 1941. A list of the butterflies of Hong Kong. Hong Kong Naturalist 10 (3/4): 148-165. https://hkjo.lib.hku.hk/exhibits/show/hkjo/browseArticle?book=b27722454&amp;issue=270036" title="A list of the butterflies of Hong Kong. Hong Kong Naturalist 10 (3 / 4): 148 - 165." url="https://hkjo.lib.hku.hk/exhibits/show/hkjo/browseArticle?book=b27722454&amp;issue=270036" year="1941">Corbet 1941</bibRefCitation>
: 150;
<bibRefCitation author="Ek-Amnuay, P" journalOrPublisher="Baan Lae Suan Amarin Printing and Publishing, Bangkok" pageId="0" pageNumber="153" refId="B10" refString="Ek-Amnuay, P, 2012. Butterflies of Thailand, 2nd edition. Baan Lae Suan Amarin Printing and Publishing, Bangkok" title="Butterflies of Thailand, 2 nd edition." year="2012">Ek-Amnuay 2012</bibRefCitation>
: 589. Type locality: Hong Kong, New territory. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizina alsulus subsp. taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="taiwana">Zizina alsulus taiwana</taxonomicName>
:
<bibRefCitation author="Shirozu, T" journalOrPublisher="The Transactions of the Kansai Entomological Society" pageId="0" pageNumber="153" pagination="30 - 42" refId="B30" refString="Shirozu, T, 1944. Notes on some Formosan Butterflies (1). The Transactions of the Kansai Entomological Society 14 (1): 30 - 42" title="Notes on some Formosan Butterflies (1)." volume="14" year="1944">
<normalizedToken originalValue="Shirôzu">Shirozu</normalizedToken>
1944
</bibRefCitation>
: 37;
<bibRefCitation author="Shirozu, T" journalOrPublisher="Hoikusha, Osaka" pageId="0" pageNumber="153" refId="B31" refString="Shirozu, T, 1960. Butterflies of Formosa in Colour. Hoikusha, Osaka" title="Butterflies of Formosa in Colour." year="1960">
<normalizedToken originalValue="Shirôzu">Shirozu</normalizedToken>
1960
</bibRefCitation>
: 334;
<bibRefCitation author="Hsu, YF" journalOrPublisher="Morning Star Publishing Inc., Taichung" pageId="0" pageNumber="153" refId="B13" refString="Hsu, YF, 2013. The Butterflies of Taiwan. Vol. 2. Lycaenidae. Morning Star Publishing Inc., Taichung" title="The Butterflies of Taiwan. Vol. 2. Lycaenidae." year="2013">Hsu: 2013</bibRefCitation>
: 252.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName baseAuthorityName="Herrich-Schäffer" baseAuthorityYear="1869" class="Insecta" family="Lycaenidae" genus="Famegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Famegana alsulus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="alsulus">Famegana alsulus</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.11171" author="Eliot, JN" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="0" pageNumber="153" pagination="371 - 505" refId="B11" refString="Eliot, JN, 1973. The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement. Bulletin of the British Museum (Natural History) Entomology 28: 371 - 505, DOI: https://doi.org/10.5962/bhl.part.11171" title="The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement." url="https://doi.org/10.5962/bhl.part.11171" volume="28" year="1973">Eliot 1973</bibRefCitation>
: 453;
<bibRefCitation author="D'Abrera, B" journalOrPublisher="Hill House, Victoria" pageId="0" pageNumber="153" refId="B8" refString="D'Abrera, B, 1977. Butterflies of the Australian Region. Hill House, Victoria" title="Butterflies of the Australian Region." year="1977">
<normalizedToken originalValue="D’Abrera">D'Abrera</normalizedToken>
1977
</bibRefCitation>
: 361;
<bibRefCitation author="D'Abrera, B" journalOrPublisher="Hill House, Victoria" pageId="0" pageNumber="153" refId="B9" refString="D'Abrera, B, 1986. Butterflies of the Oriental Region, Part III. Hill House, Victoria, 536-672." title="Butterflies of the Oriental Region, Part III. Hill House, Victoria, 536 - 672." year="1986">
<normalizedToken originalValue="D’Abrera">D'Abrera</normalizedToken>
1986
</bibRefCitation>
: 651;
<bibRefCitation author="Parsons, M" journalOrPublisher="Academic Press, London &amp; San Diego" pageId="0" pageNumber="153" refId="B25" refString="Parsons, M, 1999. The Butterflies of Papua New Guinea, Their Systematics and Biology. Academic Press, London &amp; San Diego" title="The Butterflies of Papua New Guinea, Their Systematics and Biology." year="1999">Parsons 1999</bibRefCitation>
: 460;
<bibRefCitation DOI="https://doi.org/10.1071/9780643100770" author="Braby, MF" journalOrPublisher="CSIRO, Collingwood" pageId="0" pageNumber="153" refId="B4" refString="Braby, MF, 2000. Butterflies of Australia. CSIRO, Collingwood, DOI: https://doi.org/10.1071/9780643100770" title="Butterflies of Australia." url="https://doi.org/10.1071/9780643100770" year="2000">Braby 2000</bibRefCitation>
: 839;
<bibRefCitation DOI="https://doi.org/10.1071/9781486301010" author="Braby, MF" journalOrPublisher="CSIRO, Collingwood" pageId="0" pageNumber="153" refId="B5" refString="Braby, MF, 2016. The Complete Guide to Butterflies of Australia, 2nd edition. CSIRO, Collingwood, DOI: https://doi.org/10.1071/9781486301010" title="The Complete Guide to Butterflies of Australia, 2 nd edition." url="https://doi.org/10.1071/9781486301010" year="2016">Braby 2016</bibRefCitation>
: 324.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authority="alsulus" class="Insecta" family="Lycaenidae" genus="Famegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Famegana alsulus subsp. alsulus" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="alsulus">Famegana alsulus alsulus</taxonomicName>
:
<bibRefCitation author="Common, IFB" journalOrPublisher="Angus &amp; Robertson Publishers, Sydney" pageId="0" pageNumber="153" refId="B6" refString="Common, IFB, Waterhouse, DF, 1981. Butterflies of Australia, Revised edition. Angus &amp; Robertson Publishers, Sydney" title="Butterflies of Australia, Revised edition." year="1981">Common and Waterhouse 1981</bibRefCitation>
: 587.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="type material">
<paragraph pageId="0" pageNumber="153">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">Wallace</emphasis>
: The specimen of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">Lycaena nisa</emphasis>
</taxonomicName>
retrieved in NHMUK,
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" italics="true" pageId="0" pageNumber="153">Lectotype</emphasis>
(here designated) (Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">1-3</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="153">
Taiwan • ♀; &quot;♀. Formosa&quot;; &quot;
<taxonomicName lsidName="L. nisa" pageId="0" pageNumber="153" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">L. nisa</emphasis>
</taxonomicName>
Wallace&quot;; &quot;Compare Otis Fab.&quot;; &quot;Moore Coll. 1908-203. Formosa.&quot;; reg. no. 720422; NHMUK.
</paragraph>
<caption doi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153" start="Figures 1–9" startId="F1">
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">Figures 1-9.</emphasis>
Type specimens of
<taxonomicName authorityName="Hsu" authorityYear="2020" baseAuthorityName="Wallace" baseAuthorityYear="1866" class="Insecta" family="Lycaenidae" genus="Famegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Famegana nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">Famegana nisa</emphasis>
</taxonomicName>
(Wallace, 1866)
<emphasis bold="true" pageId="0" pageNumber="153">1-3</emphasis>
lectotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">Lycaena nisa</emphasis>
</taxonomicName>
Wallace, 1866
<emphasis bold="true" pageId="0" pageNumber="153">4-6</emphasis>
holotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Lycaena taiwana</emphasis>
</taxonomicName>
Sonan, 1938
<emphasis bold="true" pageId="0" pageNumber="153">7-9</emphasis>
holotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus subsp. eggletoni" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="eggletoni">
<emphasis italics="true" pageId="0" pageNumber="153">Zizeeria alsulus eggletoni</emphasis>
</taxonomicName>
Corbet, 1941. Scale bar: 0.5 cm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">Sonan</emphasis>
: Three specimens belonging to the type series of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Zizera taiwana</emphasis>
</taxonomicName>
Sonan, 1938 retrieved in TARI, reg. nos. 37, 40, 45.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" italics="true" pageId="0" pageNumber="153">Holotype</emphasis>
(Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">4-6</figureCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="153">
Taiwan • ♂; &quot;Type [round paper, red characters in red circle]&quot;; &quot;Inrin, 30. X. 1932 Col. J. Sonan&quot;; &quot;
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Zizera taiwana</emphasis>
</taxonomicName>
Sonan DET. J. SONAN&quot;; &quot;No. 37&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" italics="true" pageId="0" pageNumber="153">Paratypes</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Taiwan • 1♀ (Allotype); &quot;Allo Type [round paper, orange characters in orange circle]&quot;; &quot;Inrin, 30. X. 1932 Col. J. Sonan&quot;; &quot;
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Zizera taiwana</emphasis>
</taxonomicName>
Sonan DET. J. SONAN&quot;; &quot;No. 45&quot; • 1 ♀; &quot;Para Type [round paper, green characters in green circle]&quot;; &quot;Inrin, 30. X. 1932 Col. J. Sonan&quot;; &quot;
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizera taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Zizera taiwana</emphasis>
</taxonomicName>
Sonan DET. J. SONAN&quot;; &quot;No. 40&quot;
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">Corbet</emphasis>
: Two specimens belonging to the type series of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus subsp. eggletoni" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="eggletoni">
<emphasis italics="true" pageId="0" pageNumber="153">Zizeeria alsulus eggletoni</emphasis>
</taxonomicName>
Corbet, 1941 retrieved in NHMUK with the reg. no. 720438.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" italics="true" pageId="0" pageNumber="153">Holotype</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Hong Kong • ♂ (Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">7-9</figureCitation>
); &quot;♂ Holotype
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus subsp. eggletoni" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="eggletoni">
<emphasis italics="true" pageId="0" pageNumber="153">Zizeeria alsulus eggletoni</emphasis>
</taxonomicName>
Cbt.&quot;; &quot;Hong Kong District. + New Territory 5. IX. 1914. R. W. Barney&quot;, &quot;Type H T [round label with red edge]&quot;, &quot;Brit. Mus. 1921-312&quot;, &quot;
<taxonomicName authorityName="Chapman" authorityYear="1910" class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">Zizeeria</taxonomicName>
GENITALIA slide No. NSC. l.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" italics="true" pageId="0" pageNumber="153">Paratype</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Hong Kong • ♂; &quot;Hong Kong District. + New Territory 5. IX. 1914 R. W. Barney&quot;, &quot;Type Holo-type [round label with red edge]&quot;, &quot;Brit. Mus. 1921-312.&quot; Note: Although this specimen also bears a
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
label, the former bears a label with
<normalizedToken originalValue="Corbet’s">Corbet's</normalizedToken>
hand-written characters indicating that it is the true holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="materials_examined">
<paragraph pageId="0" pageNumber="153">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="153">Australia • 2 ♂ 2♀; Queensland, Mt. Stuart; 26 March 2017; Y. F. Hsu and M. Braby leg.; 1♂ 1♀; Queensland, Cairns, 30 March 2017; Y. F. Hsu and M. Braby leg. Hong Kong; 1♂; Yuen Long District, Shek Wu Wai; 14 October 2009; Y. F. Lo and W. L. Hui leg; 1♀; Ching Mun CP, Tai Mo Shan; 500m; 20. October. 2009; W. L. Hui leg; 1♀; New Territory, Ngau Tam Mei; 100m; 17 December 2018; Y. F. Hsu leg. Hainan• 3♂; Dongfang, Donghe, Nanran; 18 April 2010; Y. F. Lo leg.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="description">
<paragraph pageId="0" pageNumber="153">Descriptions.</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">
Lectotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">Lycaena nisa</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Female (Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">1</figureCitation>
,
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">2</figureCitation>
). Forewing length 10.9 mm. Head hairy, brown, with medial white band on frons. Antennae dark brown, segmented with white. Proboscis brown. Labial palpus hairy, porrect; third segment slender, pointed at distal end, white but brown dorsally. Compound eyes smooth. Thorax and abdomen dark brown dorsally, white ventrally. Forewing broad, somewhat elongate, termen slightly convex. Hindwing rounded. Wing uppersides uniformly brown. Wing undersides ground color white tinged with gray, spotless except for presence of submarginal bands consisted of faint narrow bands proximally and a series of faint brown spots distally along termen of both wings; dot in cell CuA1 prominent, black. Fringe white.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">
Holotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycaena taiwana" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">Lycaena taiwana</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Male (Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">4</figureCitation>
,
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">5</figureCitation>
). Forewing length 10.9 mm. Morphology conformed to those of
<taxonomicName lsidName="L. nisa" pageId="0" pageNumber="153" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">L. nisa</emphasis>
</taxonomicName>
, except metallic purple patches present on uppersides of both wings proximally and those on hindwings.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<emphasis bold="true" pageId="0" pageNumber="153">
Holotype of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Zizeeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zizeeria alsulus subsp. eggletoni" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="subSpecies" species="alsulus" subSpecies="eggletoni">
<emphasis italics="true" pageId="0" pageNumber="153">Zizeeria alsulus eggletoni</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="153">
Male (Figs
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">7</figureCitation>
,
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Type specimens of Famegana nisa (Wallace, 1866) 1 - 3 lectotype of Lycaena nisa Wallace, 1866 4 - 6 holotype of Lycaena taiwana Sonan, 1938 7 - 9 holotype of Zizeeria alsulus eggletoni Corbet, 1941. Scale bar: 0.5 cm." figureDoi="10.3897/zookeys.966.51921.figures1-9" httpUri="https://binary.pensoft.net/fig/451023" pageId="0" pageNumber="153">8</figureCitation>
). Forewing length 11.4 mm. Morphology conformed to those of
<taxonomicName lsidName="L. nisa" pageId="0" pageNumber="153" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">L. nisa</emphasis>
</taxonomicName>
, except metallic purple patches present on uppersides of both wings proximally, and those on hindwings; submarginal bands on wing undersides slightly paler than those of
<taxonomicName lsidName="L. nisa" pageId="0" pageNumber="153" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">L. nisa</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. taiwana" pageId="0" pageNumber="153" rank="species" species="taiwana">
<emphasis italics="true" pageId="0" pageNumber="153">L. taiwana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="distribution">
<paragraph pageId="0" pageNumber="153">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="153">
This species occurs in the Oriental and Australian regions, and western Pacific, including southern China, Taiwan, the Philippines, Thailand, Sulawesi, Australia, the Torres Strait islands, Vanuatu, Fiji, Samoa, and Tonga (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.11171" author="Eliot, JN" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="0" pageNumber="153" pagination="371 - 505" refId="B11" refString="Eliot, JN, 1973. The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement. Bulletin of the British Museum (Natural History) Entomology 28: 371 - 505, DOI: https://doi.org/10.5962/bhl.part.11171" title="The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement." url="https://doi.org/10.5962/bhl.part.11171" volume="28" year="1973">Eliot 1973</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1071/9780643100770" author="Braby, MF" journalOrPublisher="CSIRO, Collingwood" pageId="0" pageNumber="153" refId="B4" refString="Braby, MF, 2000. Butterflies of Australia. CSIRO, Collingwood, DOI: https://doi.org/10.1071/9780643100770" title="Butterflies of Australia." url="https://doi.org/10.1071/9780643100770" year="2000">Braby 2000</bibRefCitation>
;
<bibRefCitation author="Vane Wright, D" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="0" pageNumber="153" pagination="1 - 267" publicationUrl="https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" refId="B35" refString="Vane Wright, D, de Jong, R, 2003. The Butterflies of Sulawesi: Annotated Checklist for a Critical Island Fauna. Zoologische Verhandelingen Leiden 343: 1 - 267, https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" title="The Butterflies of Sulawesi: Annotated Checklist for a Critical Island Fauna." url="https://www.researchgate.net/publication/254911978_The_butterflies_of_Sulawesi_Annotated_checklist_for_a_critical_island_fauna" volume="343" year="2003">Vane Wright and de Jong 2003</bibRefCitation>
;
<bibRefCitation author="Ek-Amnuay, P" journalOrPublisher="Baan Lae Suan Amarin Printing and Publishing, Bangkok" pageId="0" pageNumber="153" refId="B10" refString="Ek-Amnuay, P, 2012. Butterflies of Thailand, 2nd edition. Baan Lae Suan Amarin Printing and Publishing, Bangkok" title="Butterflies of Thailand, 2 nd edition." year="2012">Ek-Amnuay 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="biology_ecology">
<paragraph pageId="0" pageNumber="153">Biology.</paragraph>
<paragraph pageId="0" pageNumber="153">
Larval hostplants of
<taxonomicName authorityName="Hsu" authorityYear="2020" baseAuthorityName="Wallace" baseAuthorityYear="1866" class="Insecta" family="Lycaenidae" genus="Famegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Famegana nisa" order="Lepidoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="nisa">
<emphasis italics="true" pageId="0" pageNumber="153">Famegana nisa</emphasis>
</taxonomicName>
have been reported to include various legume species, such as
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Cajanus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cajanus acutifolius" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="acutifolius">
<emphasis italics="true" pageId="0" pageNumber="153">Cajanus acutifolius</emphasis>
</taxonomicName>
(F.Muell. ex Benth.) Maesen,
<taxonomicName lsidName="C. pubescens" pageId="0" pageNumber="153" rank="species" species="pubescens">
<emphasis italics="true" pageId="0" pageNumber="153">C. pubescens</emphasis>
</taxonomicName>
(Ewart &amp; Morrison) Maesen,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Desmodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Desmodium elegans" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="153">Desmodium elegans</emphasis>
</taxonomicName>
Candolle,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Flemingia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Flemingia macrophylla" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis italics="true" pageId="0" pageNumber="153">Flemingia macrophylla</emphasis>
</taxonomicName>
(Willd.) Merr.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Indigofera pratensis" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="pratensis">
<emphasis italics="true" pageId="0" pageNumber="153">Indigofera pratensis</emphasis>
</taxonomicName>
F. Muell.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Galactia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Galactia tenuiflora" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="tenuiflora">
<emphasis italics="true" pageId="0" pageNumber="153">Galactia tenuiflora</emphasis>
</taxonomicName>
(Klein ex Willdenow) Wight &amp; Arnott,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Phyllodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phyllodium pulchellum" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="pulchellum">
<emphasis italics="true" pageId="0" pageNumber="153">Phyllodium pulchellum</emphasis>
</taxonomicName>
(Linnaeus) Desvaux.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Tephrosia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tephrosia purpurea" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="purpurea">
<emphasis italics="true" pageId="0" pageNumber="153">Tephrosia purpurea</emphasis>
</taxonomicName>
(L.) Pers.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna lanceolata" order="Fabales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis italics="true" pageId="0" pageNumber="153">Vigna lanceolata</emphasis>
</taxonomicName>
Benth.,
<taxonomicName lsidName="V. radiata" pageId="0" pageNumber="153" rank="species" species="radiata">
<emphasis italics="true" pageId="0" pageNumber="153">V. radiata</emphasis>
</taxonomicName>
(L.) Wilczek and
<taxonomicName lsidName="V. vexillata" pageId="0" pageNumber="153" rank="species" species="vexillata">
<emphasis italics="true" pageId="0" pageNumber="153">V. vexillata</emphasis>
</taxonomicName>
(L.) A. Rich (all
<taxonomicName family="Fabaceae" lsidName="" pageId="0" pageNumber="153" rank="family">Fabaceae</taxonomicName>
) (
<bibRefCitation author="Bascombe, MJ" journalOrPublisher="Department of Entomology, British Museum (Natural History), London" pageId="0" pageNumber="153" refId="B2" refString="Bascombe, MJ, Johnston, G, Bascombe, FS, 1999. The Butterflies of Hong Kong. Academic Press, London, x + 422 pp." title="The Butterflies of Hong Kong. Academic Press, London, x + 422 pp." year="1999">Bascombe et al. 1999</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1071/9780643100770" author="Braby, MF" journalOrPublisher="CSIRO, Collingwood" pageId="0" pageNumber="153" refId="B4" refString="Braby, MF, 2000. Butterflies of Australia. CSIRO, Collingwood, DOI: https://doi.org/10.1071/9780643100770" title="Butterflies of Australia." url="https://doi.org/10.1071/9780643100770" year="2000">Braby 2000</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1071/9781486301010" author="Braby, MF" journalOrPublisher="CSIRO, Collingwood" pageId="0" pageNumber="153" refId="B5" refString="Braby, MF, 2016. The Complete Guide to Butterflies of Australia, 2nd edition. CSIRO, Collingwood, DOI: https://doi.org/10.1071/9781486301010" title="The Complete Guide to Butterflies of Australia, 2 nd edition." url="https://doi.org/10.1071/9781486301010" year="2016">2016</bibRefCitation>
). Life histories are illustrated and described in
<bibRefCitation author="Bascombe, MJ" journalOrPublisher="Department of Entomology, British Museum (Natural History), London" pageId="0" pageNumber="153" refId="B2" refString="Bascombe, MJ, Johnston, G, Bascombe, FS, 1999. The Butterflies of Hong Kong. Academic Press, London, x + 422 pp." title="The Butterflies of Hong Kong. Academic Press, London, x + 422 pp." year="1999">Bascombe et al. (1999)</bibRefCitation>
. Eggs are laid near flowers, upon which the larvae feed. Facultative mymecophily does occur.
</paragraph>
</subSubSection>
</treatment>
</document>
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