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<mods:title id="9CF0CE32A67A5075A9C428007F76D2FA">On the taxonomy of the Prabhasa / Zadadra generic complex with descriptions of two new genera and two new species (Lepidoptera, Erebidae, Arctiinae, Lithosiini)</mods:title>
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<mods:namePart id="84E7C99CAC0B7A9136D391C65468F8D7">Dubatolov, Vladimir V.</mods:namePart>
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<mods:affiliation id="3EA2D44B426C7A53F3441D0723DD7C22">Institute of Systematics and Ecology of Animals, SB RAS, Frunze str. 11, RF- 630091, Novosibirsk, Russia &amp; vvdubat @ mail. ru; https: // orcid. org / 0000 - 0001 - 7687 - 2102</mods:affiliation>
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<mods:namePart id="15A0372E9133577BA29F7DC8068DDCDE">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="C34545FFBC2D4BB2E30CBF33AE9DA6E8">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia &amp; National Research Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia</mods:affiliation>
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<mods:namePart id="2155CAA780BF6EC59822885EC6E01AF7">Singh, Navneet</mods:namePart>
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<mods:affiliation id="5BB6A94F430C3C8DFFE39460154D854E">Zoological Survey of India, M-Block, New Alipore, Kolkata 700053, West Bengal, India &amp; nsgill 007 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6657 - 7983</mods:affiliation>
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<mods:affiliation id="53650639927D1BAC20B11600904EC145">Zoological Survey of India, Gangetic Plains Regional Centre, BH Colony, Patna 800026, Bihar, India &amp; joshiarctiidae @ gmail. com; https: // orcid. org / 0000 - 0001 - 8514 - 1272</mods:affiliation>
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<mods:namePart id="E1E9A5B32E59F2716B33F6A05429238E">Černý, Karel</mods:namePart>
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<subSubSection id="C354653FFFEAB256FF73FECBFE1FFE44" box="[151,505,366,393]" pageId="9" pageNumber="528" type="nomenclature">
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Genus
<taxonomicName id="4C4E4D37FFEAB256FF0AFECBFE1FFE44" ID-CoL="95N5L" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[238,505,366,393]" class="Insecta" family="Erebidae" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAA6FFEAB256FF0AFECBFEB3FE45" bold="true" box="[238,341,366,393]" italics="true" pageId="9" pageNumber="528">Zadadra</emphasis>
<bibRefCitation id="EFDF4B45FFEAB256FEB8FECAFE1FFE44" author="Moore, F." box="[348,505,366,393]" pageId="9" pageNumber="528" pagination="3 - 37" refId="ref10104" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="journal article" year="1878">Moore, 1878</bibRefCitation>
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<taxonomicName id="4C4E4D37FFEAB256FF73FE16FE9EFE07" ID-CoL="95N5L" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[151,376,435,459]" class="Insecta" family="Erebidae" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAA6FFEAB256FF73FE16FF08FE07" box="[151,238,435,459]" italics="true" pageId="9" pageNumber="528">Zadadra</emphasis>
<bibRefCitation id="EFDF4B45FFEAB256FF10FE11FE9EFE07" author="Moore, F." box="[244,376,435,459]" pageId="9" pageNumber="528" pagination="3 - 37" refId="ref10104" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="journal article" year="1878">Moore, 1878</bibRefCitation>
</taxonomicName>
,
<emphasis id="B93AEAA6FFEAB256FE66FE16FC8DFE07" box="[386,875,435,459]" italics="true" pageId="9" pageNumber="528">Proceedings of the Zoological Society of London</emphasis>
<emphasis id="B93AEAA6FFEAB256FC95FE16FC42FE07" bold="true" box="[881,932,435,459]" pageId="9" pageNumber="528">1878</emphasis>
: 25.
<typeStatus id="54F58816FFEAB256FC31FE11FBE1FE00" box="[981,1031,436,460]" pageId="9" pageNumber="528">Type</typeStatus>
species:
<taxonomicName id="4C4E4D37FFEAB256FB87FE16FA7EFE07" ID-CoL="BFXY5" authority="Moore, 1872" authorityName="Moore" authorityYear="1872" box="[1123,1432,435,459]" class="Insecta" family="Erebidae" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="distorta">
<emphasis id="B93AEAA6FFEAB256FB87FE16FAE8FE07" box="[1123,1294,435,459]" italics="true" pageId="9" pageNumber="528">Zadadra distorta</emphasis>
<bibRefCitation id="EFDF4B45FFEAB256FAF0FE11FA7EFE07" author="Moore, F." box="[1300,1432,435,459]" pageId="9" pageNumber="528" pagination="555 - 583" refId="ref10067" refString="Moore, F. (1872) Descriptions of new Indian Lepidoptera. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1872 (2), 555 - 583." type="journal article" year="1872">Moore, 1872</bibRefCitation>
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, by monotypy.
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<paragraph id="8BF136B4FFEAB256FF73FDBDFB5EFBCB" blockId="9.[151,1437,536,1031]" pageId="9" pageNumber="528">
<emphasis id="B93AEAA6FFEAB256FF73FDBDFEECFDFE" bold="true" box="[151,266,536,562]" pageId="9" pageNumber="528">Diagnosis</emphasis>
. Small or medium-sized moths. The sexual dimorphism is more expressed in the generic complex (
<figureCitation id="13752A31FFEAB256FA8EFDBDFF06FD9A" captionStart="FIGURES 1118" captionStartId="6.[151,264,1877,1901]" captionTargetBox="[184,1420,183,1828]" captionTargetId="figure-27@6.[151,1436,181,1854]" captionTargetPageId="6" captionText="FIGURES 1118. Zadadra spp.: adults. Depositories of the specimens: 11 in OUMNH; 12 in NHMUK (© The Trustees of the Natural History Museum); 1318 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.4745239" httpUri="https://zenodo.org/record/4745239/files/figure.png" pageId="9" pageNumber="528">Figs 1118</figureCitation>
). The male forewing is shorter and broader than that of female, its distal part is slightly curved posteriorly, and the anal and outer margins are convex. Additionally, the forewing cell of the male is narrow and bearing a longitudinal row of androconial scales continuing to the basal part of the area between the veins Cu
<subScript id="17CA34F1FFEAB256FAFBFD33FAC1FD68" attach="left" box="[1311,1319,662,676]" fontSize="6" pageId="9" pageNumber="528">1</subScript>
and M
<subScript id="17CA34F1FFEAB256FA99FD33FA73FD68" attach="left" box="[1405,1429,662,676]" fontSize="6" pageId="9" pageNumber="528">23</subScript>
. The male hindwing is conspicuously broader than that of female and bearing a wide dark area of androconial scales at the costa (the character unique among the genera of the generic complex). The female forewing is long and narrow, nearly straight. The male genitalia of the genus (
<figureCitation id="13752A31FFEAB256FCBAFD55FC07FCC6" box="[862,993,752,778]" captionStart-0="FIGURES 2730" captionStart-1="FIGURES 3132" captionStartId-0="12.[151,264,1944,1968]" captionStartId-1="13.[151,264,1944,1968]" captionTargetBox-0="[159,1423,181,1909]" captionTargetBox-1="[153,1408,186,1884]" captionTargetId-0="figure-27@12.[151,1436,181,1920]" captionTargetId-1="figure-19@13.[151,1436,181,1920]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 2730. Zadadra spp.: male genitalia. Depositories of the specimens: 27 in OUMNH; 2830 in MWM/ZSM." captionText-1="FIGURES 3132. Zadadra spp.: male genitalia. The specimens are deposited in MWM/ZSM." figureDoi-0="http://doi.org/10.5281/zenodo.4745253" figureDoi-1="http://doi.org/10.5281/zenodo.4745257" httpUri-0="https://zenodo.org/record/4745253/files/figure.png" httpUri-1="https://zenodo.org/record/4745257/files/figure.png" pageId="9" pageNumber="528">Figs 2732</figureCitation>
) are characterized by the combination of the following diagnostic characters: (1) the juxta is narrow, weakly sclerotized, its distal end is connected to the intervalval commissura by a membrane; (2) the intervalval commissura is thick membranous with a weakly sclerotized medial area, connecting from one third to a half of valvae (an autapomorphic feature); (3) the cucullus is weakly sclerotized, well-separated from the distal saccular process; (4) the sacculus is heavily sclerotized, wide, lobe-like (an autapomorphic feature); (5) the distal saccular process is short but broad, robust, without setae. The female genitalia of
<taxonomicName id="4C4E4D37FFEAB256FE89FC6DFE2BFC2E" authorityName="Moore" authorityYear="1878" box="[365,461,968,994]" class="Insecta" family="Erebidae" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAA6FFEAB256FE89FC6DFE2BFC2E" box="[365,461,968,994]" italics="true" pageId="9" pageNumber="528">Zadadra</emphasis>
</taxonomicName>
(
<figureCitation id="13752A31FFEAB256FE3FFC6DFDBFFC2E" box="[475,601,968,994]" captionStart="FIGURES 3338" captionStartId="14.[151,264,1703,1727]" captionTargetBox="[154,1431,186,1644]" captionTargetId="figure-27@14.[151,1436,181,1679]" captionTargetPageId="14" captionText="FIGURES 3338. Lithosiina spp.: female genitalia. Depositories of the specimens: 33 and 36 in NHMUK (© The Trustees of the Natural History Museum); 34 and 38 in CKC; 35 and 37 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.4745261" httpUri="https://zenodo.org/record/4745261/files/figure.png" pageId="9" pageNumber="528">Figs 37, 38</figureCitation>
) display no remarkable differences from those of
<taxonomicName id="4C4E4D37FFEAB256FB65FC6DFB38FC2E" authorityName="Dubatolov, Volynkin &amp; Kishida" authorityYear="2018" box="[1153,1246,968,994]" class="Insecta" family="Erebidae" genus="Chinasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAA6FFEAB256FB65FC6DFB38FC2E" box="[1153,1246,968,994]" italics="true" pageId="9" pageNumber="528">Chinasa</emphasis>
</taxonomicName>
(
<figureCitation id="13752A31FFEAB256FB09FC6DFADBFC2F" box="[1261,1341,968,995]" captionStart="FIGURES 3338" captionStartId="14.[151,264,1703,1727]" captionTargetBox="[154,1431,186,1644]" captionTargetId="figure-27@14.[151,1436,181,1679]" captionTargetPageId="14" captionText="FIGURES 3338. Lithosiina spp.: female genitalia. Depositories of the specimens: 33 and 36 in NHMUK (© The Trustees of the Natural History Museum); 34 and 38 in CKC; 35 and 37 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.4745261" httpUri="https://zenodo.org/record/4745261/files/figure.png" pageId="9" pageNumber="528">Fig. 35</figureCitation>
) but can be distinguished by the presence of sclerotized pockets on the 7th sternite (absent in
<taxonomicName id="4C4E4D37FFEAB256FBAEFC49FB41FBCA" authorityName="Dubatolov, Volynkin &amp; Kishida" authorityYear="2018" box="[1098,1191,1004,1030]" class="Insecta" family="Erebidae" genus="Chinasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAA6FFEAB256FBAEFC49FB41FBCA" box="[1098,1191,1004,1030]" italics="true" pageId="9" pageNumber="528">Chinasa</emphasis>
</taxonomicName>
).
</paragraph>
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