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<document id="EFFBBA75BF2A5EFEAF7BFB4B01DBA412" ID-CLB-Dataset="46124" ID-DOI="10.11646/zootaxa.3784.2.1" ID-GBIF-Dataset="f4530593-bcbe-4092-a140-30de91a6f699" ID-ISSN="1175-5326" ID-Zenodo-Dep="225955" ID-ZooBank="0075CB1A-38B8-4E3E-ABCC-699E3113AAF9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460402627306" checkinUser="plazi" docAuthor="Kaiser, Stefanie" docDate="2014" docId="83031014413EFFCCFF183B48FD63F81C" docLanguage="en" docName="zt03784p119.pdf" docOrigin="Zootaxa 3784 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebefustis juansenii Kaiser, 2014, sp. n." docType="treatment" docVersion="10" lastPageNumber="110" masterDocId="7F3A686C413DFFC5FF8F3E4DFFC6FF89" masterDocTitle="New species of Hebefustis Siebenaller &amp; Hessler 1977 (Isopoda, Asellota, Nannoniscidae) from the Clarion Clipperton Fracture Zone (equatorial NE Pacific)" masterLastPageNumber="119" masterPageNumber="101" pageNumber="104" updateTime="1698299397550" updateUser="ExternalLinkService">
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<mods:title id="172C31DCA4FC433D72C2157CEECECBB4">New species of Hebefustis Siebenaller &amp; Hessler 1977 (Isopoda, Asellota, Nannoniscidae) from the Clarion Clipperton Fracture Zone (equatorial NE Pacific)</mods:title>
</mods:titleInfo>
<mods:name id="D18581792321B249ED66E78F13C7758C" type="personal">
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<mods:namePart id="CBD1F1D8F0649E3555FD5F9C6AF5261D">Kaiser, Stefanie</mods:namePart>
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<mods:title id="F51ACB2DE81E895DCB60D72A5C1ACE09">Zootaxa</mods:title>
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<mods:part id="ECB5DD9E92C3F4779C8A2FD1231FB0BE">
<mods:date id="042EA27CFED9FCB39674D3AFB3066087">2014</mods:date>
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<mods:number id="71FB19DFCB5956D825D06C7371586C87">3784</mods:number>
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<mods:identifier id="BE884B3EAC59FC1471BAD5EAF113B7C3" type="DOI">10.11646/zootaxa.3784.2.1</mods:identifier>
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<treatment id="83031014413EFFCCFF183B48FD63F81C" ID-DOI="http://doi.org/10.5281/zenodo.5698409" ID-GBIF-Taxon="119420207" ID-Zenodo-Dep="5698409" LSID="urn:lsid:plazi:treatment:83031014413EFFCCFF183B48FD63F81C" httpUri="http://treatment.plazi.org/id/83031014413EFFCCFF183B48FD63F81C" lastPageId="9" lastPageNumber="110" pageId="3" pageNumber="104">
<subSubSection id="43B0F289413EFFC6FF183B48FECDFAC8" pageId="3" pageNumber="104" type="nomenclature">
<paragraph id="0B15A102413EFFC6FF183B48FE1FFA96" blockId="3.[151,473,1285,1345]" box="[151,473,1285,1311]" pageId="3" pageNumber="104">
<heading id="505D166E413EFFC6FF183B48FE1FFA96" bold="true" box="[151,473,1285,1311]" fontSize="11" level="1" pageId="3" pageNumber="104" reason="1">
<emphasis id="39DE7D10413EFFC6FF183B48FE1FFA96" bold="true" box="[151,473,1285,1311]" pageId="3" pageNumber="104">
<taxonomicName id="CCAADA81413EFFC6FF183B48FE49FA96" ID-CoL="3JXH7" box="[151,399,1285,1311]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D10413EFFC6FF183B48FE49FA96" bold="true" box="[151,399,1285,1311]" italics="true" pageId="3" pageNumber="104">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B413EFFC6FE193B48FE1FFA96" box="[406,473,1285,1311]" pageId="3" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B15A102413EFFC6FF183B65FECDFAC8" blockId="3.[151,473,1285,1345]" box="[151,267,1320,1345]" pageId="3" pageNumber="104">
(
<figureCitation id="9391BD87413EFFC6FF103B65FEC4FAC8" box="[159,258,1320,1345]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="5.[151,250,1946,1968]" captionStartId-1="6.[151,250,1963,1985]" captionStartId-2="7.[151,250,1572,1594]" captionStartId-3="10.[151,250,1976,1998]" captionStartId-4="11.[151,250,1937,1959]" captionStartId-5="12.[151,250,1965,1987]" captionTargetBox-0="[184,1406,217,1880]" captionTargetBox-1="[209,1358,205,1962]" captionTargetBox-2="[225,1349,212,1538]" captionTargetBox-3="[212,1386,218,1942]" captionTargetBox-4="[289,1284,220,1900]" captionTargetBox-5="[200,1386,218,1915]" captionTargetId-0="figure@5.[166,1429,193,1897]" captionTargetId-1="figure@6.[208,1378,193,1967]" captionTargetId-2="figure@7.[217,1369,193,1550]" captionTargetId-3="figure@10.[188,1398,195,1954]" captionTargetId-4="figure@11.[275,1312,193,1916]" captionTargetId-5="figure@12.[177,1409,193,1935]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." captionText-1="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." captionText-2="FIGURE 4. Hebefustis juansenii sp. n.; A D paratype male (ZMH K 43952); A, habitus, dorsal view. B, antenna. C, antennula. D, pleotelson, ventral view. Scale bar: A = 500 Μm, B C = 100 Μm; D = 250 Μm." captionText-3="FIGURE 5. Hebefustis juansenii sp. n.; A E paratype male (ZMH K 43952); A, habitus, dorsal view. B, left mandible. C, right mandible. D, maxilliped. E, maxillula. F, maxilla, detail: inner endite. Scale bar: A = 500 Μm, B F = 100 Μm." captionText-4="FIGURE 6. Hebefustis juansenii sp. n.; A C paratype male (ZMH K 43952); A, pereopod I, detail: dorsal and ventral claw. B, pereopod II. C, pereopod III. Scale bar: A C = 100 Μm." captionText-5="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri-0="https://zenodo.org/record/225957/files/figure.png" httpUri-1="https://zenodo.org/record/225958/files/figure.png" httpUri-2="https://zenodo.org/record/225959/files/figure.png" httpUri-3="https://zenodo.org/record/225960/files/figure.png" httpUri-4="https://zenodo.org/record/225961/files/figure.png" httpUri-5="https://zenodo.org/record/225962/files/figure.png" pageId="3" pageNumber="104">Figs 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC6FF183B3DFCA7F90D" pageId="3" pageNumber="104" type="materials_examined">
<paragraph id="0B15A102413EFFC6FF183B3DFC64FA00" blockId="3.[151,1437,1392,2029]" box="[151,930,1392,1417]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF183B3DFEFAFA00" bold="true" box="[151,316,1392,1417]" pageId="3" pageNumber="104">
<typeStatus id="D4111FA0413EFFC6FF183B3DFF14FA00" box="[151,210,1392,1417]" pageId="3" pageNumber="104">Type</typeStatus>
fixation.
</emphasis>
<typeStatus id="D4111FA0413EFFC6FECD3B3DFE6BFA00" box="[322,429,1392,1417]" pageId="3" pageNumber="104" type="holotype">Holotype</typeStatus>
, ovigerous female,
<quantity id="CC520CE7413EFFC6FD073B3CFD26FA01" box="[648,736,1393,1417]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="104" unit="mm" value="3.4">3.4 mm</quantity>
, designated here.
</paragraph>
<paragraph id="0B15A102413EFFC6FF483BD9FAF5F9E8" blockId="3.[151,1437,1392,2029]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF483BD9FE7EFA24" bold="true" box="[199,440,1428,1453]" pageId="3" pageNumber="104">Material examined.</emphasis>
<emphasis id="39DE7D10413EFFC6FE483BD9FDF0FA25" box="[455,566,1428,1452]" italics="true" pageId="3" pageNumber="104">
<typeStatus id="D4111FA0413EFFC6FE483BD9FDF7FA25" box="[455,561,1428,1452]" pageId="3" pageNumber="104" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="1DAC6A8B413EFFC6FDCA3BD8FD68FA24" box="[581,686,1428,1453]" pageId="3" pageNumber="104" type="female">1 female</specimenCount>
<typeStatus id="D4111FA0413EFFC6FD323BD9FCD9FA24" box="[701,799,1428,1453]" pageId="3" pageNumber="104" type="holotype">holotype</typeStatus>
(ovigerous),
<quantity id="CC520CE7413EFFC6FC4A3BD8FBE3FA25" box="[965,1061,1429,1453]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="104" unit="mm" value="3.4">3.4 mm</quantity>
,
<collectionCode id="6DBB39C7413EFFC6FBB73BD8FB47FA24" box="[1080,1153,1429,1453]" pageId="3" pageNumber="104">CCFZ</collectionCode>
, equatorial NE Pacific, BIONOD expedition, RV LAtalante,
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, station
<collectionCode id="6DBB39C7413EFFC6FD7C3BF4FCE0FA58" box="[755,806,1465,1489]" httpUri="http://grbio.org/cool/d4va-b13i" name="University of the Basque Country" pageId="3" pageNumber="104">BIO</collectionCode>
12-33 (start:
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,
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,
<quantity id="CC520CE7413EFFC6FB723BF4FA9DFA59" box="[1277,1371,1465,1489]" metricMagnitude="3" metricUnit="m" metricValue="4.133" pageId="3" pageNumber="104" unit="m" value="4133.0">4133 m</quantity>
; end:
<geoCoordinate id="6E9EC7C5413EFFC6FF183B91FEDAFA7C" box="[151,284,1500,1525]" direction="north" orientation="latitude" pageId="3" pageNumber="104" precision="9" value="11.865167">11°51.91N</geoCoordinate>
,
<geoCoordinate id="6E9EC7C5413EFFC6FEA63B90FE06FA7C" box="[297,448,1501,1525]" direction="west" orientation="longitude" pageId="3" pageNumber="104" precision="9" value="-117.05217">117°03.13W</geoCoordinate>
,
<quantity id="CC520CE7413EFFC6FE423B90FDE1FA7C" box="[461,551,1501,1525]" metricMagnitude="3" metricUnit="m" metricValue="4.127" pageId="3" pageNumber="104" unit="m" value="4127.0">4127 m</quantity>
), date:
<date id="7F1487C2413EFFC6FDF33B90FD39FA7C" box="[636,767,1501,1525]" pageId="3" pageNumber="104" value="2012-04-07">07/04/2012</date>
, ZMH-K-43951.
<emphasis id="39DE7D10413EFFC6FC493B90FB87FA7D" box="[966,1089,1501,1524]" italics="true" pageId="3" pageNumber="104">
<typeStatus id="D4111FA0413EFFC6FC493B90FBFDFA7D" box="[966,1083,1501,1524]" pageId="3" pageNumber="104" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="1DAC6A8B413EFFC6FBC53B90FB5FFA7C" box="[1098,1177,1500,1525]" pageId="3" pageNumber="104" type="adult">1 adult</specimenCount>
male,
<quantity id="CC520CE7413EFFC6FB673B90FA84FA7D" box="[1256,1346,1501,1525]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="104" unit="mm" value="2.2">2.2 mm</quantity>
,
<collectionCode id="6DBB39C7413EFFC6FAC03B90FA5EFA7C" box="[1359,1432,1501,1525]" pageId="3" pageNumber="104">CCFZ</collectionCode>
, equatorial NE Pacific, BIONOD expedition, RV LAtalante,
<collectionCode id="6DBB39C7413EFFC6FCD9384CFC4AF990" box="[854,908,1537,1561]" pageId="3" pageNumber="104">EBS</collectionCode>
, from the same station as
<typeStatus id="D4111FA0413EFFC6FB4B384DFAEFF990" box="[1220,1321,1536,1561]" pageId="3" pageNumber="104" type="holotype">holotype</typeStatus>
, ZMH-K- 43952; 1 Manca 1,
<collectionCode id="6DBB39C7413EFFC6FEF13868FE01F9B4" box="[382,455,1573,1597]" pageId="3" pageNumber="104">CCFZ</collectionCode>
, equatorial NE Pacific, BIONOD expedition, RV LAtalante,
<collectionCode id="6DBB39C7413EFFC6FB1A3868FB0DF9B4" box="[1173,1227,1573,1597]" pageId="3" pageNumber="104">EBS</collectionCode>
,
<collectionCode id="6DBB39C7413EFFC6FB553868FACAF9B4" box="[1242,1292,1573,1597]" httpUri="http://grbio.org/cool/d4va-b13i" name="University of the Basque Country" pageId="3" pageNumber="104">BIO</collectionCode>
12
<emphasis id="39DE7D10413EFFC6FAA93868FAF2F9B5" box="[1318,1332,1573,1596]" italics="true" pageId="3" pageNumber="104"></emphasis>
43 (start:
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, 117°34.33,
<quantity id="CC520CE7413EFFC6FE3D3805FDCCF9E9" box="[434,522,1608,1633]" metricMagnitude="3" metricUnit="m" metricValue="4.358" pageId="3" pageNumber="104" unit="m" value="4358.0">4358 m</quantity>
;
<geoCoordinate id="6E9EC7C5413EFFC6FD993804FD5CF9E8" box="[534,666,1609,1633]" direction="north" orientation="latitude" pageId="3" pageNumber="104" precision="9" value="11.805667">11°48.34N</geoCoordinate>
,
<geoCoordinate id="6E9EC7C5413EFFC6FD293805FCFBF9E8" box="[678,829,1608,1633]" direction="west" orientation="longitude" pageId="3" pageNumber="104" precision="9" value="-117.5325">117°31.95W</geoCoordinate>
,
<quantity id="CC520CE7413EFFC6FCC63805FC66F9E8" box="[841,928,1608,1633]" metricMagnitude="3" metricUnit="m" metricValue="4.36" pageId="3" pageNumber="104" unit="m" value="4360.0">4360 m</quantity>
), date:
<date id="7F1487C2413EFFC6FC7D3804FBB2F9E8" box="[1010,1140,1609,1633]" pageId="3" pageNumber="104" value="2012-04-09">09/04/2012</date>
, ZMH-K-43953.
</paragraph>
<paragraph id="0B15A102413EFFC6FF483821FCA7F90D" blockId="3.[151,1437,1392,2029]" box="[199,865,1644,1669]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF483821FEA3F90C" bold="true" box="[199,357,1644,1669]" pageId="3" pageNumber="104">
<typeStatus id="D4111FA0413EFFC6FF483821FEC4F90C" box="[199,258,1644,1669]" pageId="3" pageNumber="104">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="6DBB39C7413EFFC6FEE33820FE73F90C" box="[364,437,1645,1669]" pageId="3" pageNumber="104">CCFZ</collectionCode>
, equatorial NE Pacific,
<quantity id="CC520CE7413EFFC6FD303820FC9BF90D" box="[703,861,1644,1669]" metricMagnitude="3" metricUnit="m" metricValue="4.2435" metricValueMax="4.36" metricValueMin="4.127" pageId="3" pageNumber="104" unit="m" value="4243.5" valueMax="4360.0" valueMin="4127.0">
4127
<emphasis id="39DE7D10413EFFC6FD783820FCC3F90D" box="[759,773,1645,1668]" italics="true" pageId="3" pageNumber="104"></emphasis>
4360 m
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC6FF4838DDFD0EF920" box="[199,712,1680,1705]" pageId="3" pageNumber="104" type="distribution">
<paragraph id="0B15A102413EFFC6FF4838DDFD0EF920" blockId="3.[151,1437,1392,2029]" box="[199,712,1680,1705]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF4838DDFEA6F920" bold="true" box="[199,352,1680,1705]" pageId="3" pageNumber="104">Distribution.</emphasis>
Only known from
<typeStatus id="D4111FA0413EFFC6FDB838DCFDA1F920" box="[567,615,1681,1705]" pageId="3" pageNumber="104">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC6FF4838F9FCBFF944" box="[199,889,1716,1741]" pageId="3" pageNumber="104" type="etymology">
<paragraph id="0B15A102413EFFC6FF4838F9FCBFF944" blockId="3.[151,1437,1392,2029]" box="[199,889,1716,1741]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF4838F9FE88F944" bold="true" box="[199,334,1716,1741]" pageId="3" pageNumber="104">Etymology.</emphasis>
The name is masculine and dedicated to a friend.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC6FF483895FBBBF808" pageId="3" pageNumber="104" type="diagnosis">
<paragraph id="0B15A102413EFFC6FF483895FBBBF808" blockId="3.[151,1437,1392,2029]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF483895FE87F978" bold="true" box="[199,321,1752,1777]" pageId="3" pageNumber="104">Diagnosis.</emphasis>
Maxilliped epipodite reaching upper third of palpal article 2. Pereonites anterior margins 14 frontally directed, rounded; pereonite 2 anterolateral margin bearing a robust spine both in male and female; pereonites 6 and 7 lateral margins rounded; pereonite 6 anterior margin strongly convex. Pleotelson with two pairs of posterolateral spines; proximal pair inserting 0.6 from anterior margin. Uropods not extending beyond pleotelson posterior margin. Male pleopod 1 lateral lobes rounded, distal margin straight.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC6FF4839C1FD13F840" pageId="3" pageNumber="104" type="discussion">
<paragraph id="0B15A102413EFFC6FF4839C1FD13F840" blockId="3.[151,1437,1392,2029]" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF4839C1FEFDF82C" bold="true" box="[199,315,1932,1957]" pageId="3" pageNumber="104">Remarks.</emphasis>
Due to the low number of individuals in the new species, all appendages of
<typeStatus id="D4111FA0413EFFC6FB2839C1FACFF82C" box="[1191,1289,1932,1957]" pageId="3" pageNumber="104" type="holotype">holotype</typeStatus>
female were drawn
<emphasis id="39DE7D10413EFFC6FF6B39FCFEEFF841" box="[228,297,1969,1992]" italics="true" pageId="3" pageNumber="104">in situ</emphasis>
to minimize damage of the specimen.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413EFFC2FF483999FE30F883" lastPageId="7" lastPageNumber="108" pageId="3" pageNumber="104" type="description">
<paragraph id="0B15A102413EFFC1FF483999FD88FE7C" blockId="3.[151,1437,1392,2029]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
<emphasis id="39DE7D10413EFFC6FF483999FDFAF864" bold="true" box="[199,572,2004,2029]" pageId="3" pageNumber="104">
Description of
<typeStatus id="D4111FA0413EFFC6FEF83999FE18F864" box="[375,478,2004,2029]" pageId="3" pageNumber="104" type="holotype">holotype</typeStatus>
female.
</emphasis>
<emphasis id="39DE7D10413EFFC6FDCC3998FD5AF864" box="[579,668,2005,2029]" italics="true" pageId="3" pageNumber="104">Habitus</emphasis>
(
<figureCitation id="9391BD87413EFFC6FD243999FD34F864" box="[683,754,2004,2029]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="3" pageNumber="104">Figs 2</figureCitation>
A, D, 3A). Body length 3.3 pereonite 3 width. Dorsal cuticle surface with numerous setules. Pereonites 13 of similar length; pereonite 1 width 0.8 pereonite 2 width; pereonites anterior margins 14 frontally directed, rounded. Pereonite 2 with a robust spine tipped with a small setule laterally. Pereonites 2 and 3 of equal width, widest; pereonite 2 width 3.5 length; pereonite 3 width 5.3 length. Pereonite 4 longest, width 2.6 length. Pereonites 57 of similar width; pereonites 6 and 7 medially fused, with rounded lateral margins; pereonite 6 with strongly convex anterior margin. Pleotelson length 0.23 body length, nearly as long as wide; width 0.8 pereonite 2 width, with two pairs of well-developed posterolateral spines; proximal pair inserting 0.6 from anterior margin, distal pair inserting 0.9 from anterior margin; posterior margin strongly convex; anterior margin slightly concave. Anus (
<figureCitation id="9391BD874139FFC1FD8A3FD9FD8CFE25" box="[517,586,404,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="4" pageNumber="105">Fig. 2</figureCitation>
D) covered by anus valves laterally. Uropods inserting closely to the anus valves, short in relation to pleotelson, not projecting beyond posterior margin. Operculum length approximately 0.7 pleotelson length (ventrally measured).
</paragraph>
<paragraph id="0B15A1024139FFC1FF483C4DFCBBFDB4" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF483C4DFEA9FD91" box="[199,367,512,536]" italics="true" pageId="4" pageNumber="105">Cephalothorax</emphasis>
(
<figureCitation id="9391BD874139FFC1FE0C3FB2FE16FD91" box="[387,464,511,536]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="4" pageNumber="105">Figs 2</figureCitation>
A, 3A). Length 0.8 width. Anterior and posterior margin slightly rounded; lateral margins straight. Antennae inserting frontolaterally in a deep fold.
</paragraph>
<paragraph id="0B15A1024139FFC1FF483C05FB95FD44" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<taxonomicName id="CCAADA814139FFC1FF483C05FEFCFDE9" box="[199,314,584,608]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="4" pageNumber="105" phylum="Ascomycota" rank="genus">
<emphasis id="39DE7D104139FFC1FF483C05FEFCFDE9" box="[199,314,584,608]" italics="true" pageId="4" pageNumber="105">Antennula</emphasis>
</taxonomicName>
(
<figureCitation id="9391BD874139FFC1FEC63C0AFE4DFDE9" box="[329,395,583,608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1963,1985]" captionTargetBox="[209,1358,205,1962]" captionTargetId="figure@6.[208,1378,193,1967]" captionTargetPageId="6" captionText="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="4" pageNumber="105">Fig. 3</figureCitation>
C). Length 0.12 body length; with 5 articles. First article circular and broadest; with 1 simple seta distolaterally; second article length 1.75 article 1 length, width 0.3 length; with 2 broom setae distally (one broken off). Article 3 and 4 equal in length, length 0.3 article 2 length; article 4 with 2 simple setae. Article 5 length 4 article 4 length, length 8 width; with 1 long aesthetasc and 1 simple seta terminally.
</paragraph>
<paragraph id="0B15A1024139FFC1FF483C95FCBFFC64" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF483C95FEE2FD66" box="[199,292,728,751]" italics="true" pageId="4" pageNumber="105">Antenna</emphasis>
(
<figureCitation id="9391BD874139FFC1FEBB3C9AFEB0FD79" box="[308,374,727,752]" captionStart="FIGURE 3" captionStartId="6.[151,250,1963,1985]" captionTargetBox="[209,1358,205,1962]" captionTargetId="figure@6.[208,1378,193,1967]" captionTargetPageId="6" captionText="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="4" pageNumber="105">Fig. 3</figureCitation>
F). Length one-third body length; with 6 podomere and 11 flagellar articles. Podomere articles 14 short; article 2 with 2 simple setae distally; article 3 with 1 robust spine tipped with a small setule and 1 simple seta distally. Article 5 longest, length 1.4 articles 14 length, length approximately 5.1 width; with 4 simple setae laterally, with 1 broom seta and 2 simple setae distally. Article 6 length 0.8 article 5 length, length approximately 4.0 width; with 3 short simple setae laterally, with 1 broom seta and 1 simple seta distally. Flagellar article 1 longest, about 0.6 podomere article 6 length, length approximately 3.0 width; with 2 simple setae laterally. Flagellar articles 210 of similar length, width decreasing distally; each article with 12 short simple setae distally. Article 11 somewhat shorter; with 6 long slender setae terminally.
</paragraph>
<paragraph id="0B15A1024139FFC1FF483DB5FD67FA84" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF483DB5FE86FB99" box="[199,320,1016,1040]" italics="true" pageId="4" pageNumber="105">Maxilliped</emphasis>
(
<figureCitation id="9391BD874139FFC1FEDF3DBAFE55FB99" box="[336,403,1015,1040]" captionStart="FIGURE 3" captionStartId="6.[151,250,1963,1985]" captionTargetBox="[209,1358,205,1962]" captionTargetId="figure@6.[208,1378,193,1967]" captionTargetPageId="6" captionText="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="4" pageNumber="105">Fig. 3</figureCitation>
B). Left and right maxilliped connected by 2 coupling hooks. Epipodite smooth, triangular, slender, length 3.1 width, reaching upper third of palpal article 2. Palpal article 1 short, width 2.7 length; with 1 simple seta distally. Article 2 and 3 of equal length, about twice article 1 length. Article 2 width 1.3 length; with 3 simple setae laterally, with 5 simple setae distally. Article 3 almost as long as wide, distal margin saw-toothed; with 6 robust sensillae. Article 4 as long as article 1, length 2.5 width, with a distal projection reaching to mid of article 5; with 2 long, slender setae distally. Article 5 length 0.6 article 1 length, twice as long as wide; with 2 slender setae of varying size terminally. Endite distal margin with some robust, dentate setae and several fine setae laterally. Protopod coxa quadrangular; length 2.5 width.
</paragraph>
<paragraph id="0B15A1024139FFC1FF403B55FD5AF9A4" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF403B55FE8DFAB9" box="[207,331,1304,1328]" italics="true" pageId="4" pageNumber="105">Pereopod I</emphasis>
(
<figureCitation id="9391BD874139FFC1FED33B5AFE58FAB9" box="[348,414,1303,1328]" captionStart="FIGURE 3" captionStartId="6.[151,250,1963,1985]" captionTargetBox="[209,1358,205,1962]" captionTargetId="figure@6.[208,1378,193,1967]" captionTargetPageId="6" captionText="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="4" pageNumber="105">Fig. 3</figureCitation>
D) more robust than pereopod 2. Basis not drawn. Ischium length 2.2 width. Merus length about half ischium length, as long as wide; with 2 long robust setae distodorsally, with 2 simple setae of varying size ventrally. Carpus length 1.9 merus length, length 2.5 width; ventral margin with numerous small setae, membranously embedded, and 3 robust unequally bifid setae in between, with 1 long simple seta distoventrally. Propodus length 0.9 carpus length, 2.9 width; with 2 slender simple setae distodorsally, with numerous small setae, membranously embedded, and 3 robust unequally bifid setae in between ventrally. Dactylus length 0.3 propodus length, length twice width; with 3 slender setae medially. Dorsal claw about as long as dactylus; with 2 long, slender setae between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A1024139FFC1FF483875FCD8F8C4" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF483875FE8BF9D9" box="[199,333,1592,1616]" italics="true" pageId="4" pageNumber="105">Pereopod II</emphasis>
(
<figureCitation id="9391BD874139FFC1FED3387AFE58F9D9" box="[348,414,1591,1616]" captionStart="FIGURE 3" captionStartId="6.[151,250,1963,1985]" captionTargetBox="[209,1358,205,1962]" captionTargetId="figure@6.[208,1378,193,1967]" captionTargetPageId="6" captionText="FIGURE 3. Hebefustis juansenii sp. n.; A F, holotype female (ZMH K 43951); A, habitus, dorsal view. B, maxilliped. C, antennula. D E, pereopods I II, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B F = 250 Μm." httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="4" pageNumber="105">Fig. 3</figureCitation>
E). Basis length 3.8 width; with 2 simple setae dorsally, with 2 simple setae ventrally and 1 long simple seta distally. Ischium about half basis length, length 2.4 width; with 1 simple seta ventrally. Merus length 0.6 ischium length, length 1.4 width; with 2 simple setae distodorsally, with 3 more slender setae ventrally. Carpus length twice merus length, length 3.5 width; with 2 slender simple setae dorsally, with 6 stout unequally bifid setae ventrally. Propodus about as long as carpus, length 6 width; with 2 simple setae dorsally, with numerous small setae, membranously embedded, and with 4 stout unequally bifid setae ventrally. Dactylus length one-third propodus length, length 2.6 width; with 3 simple setae medially. Dorsal claw length 0.6 dactylus length; with 1 slender seta (broken off) between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A1024139FFC1FF483915FE2CF81D" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="105">
<emphasis id="39DE7D104139FFC1FF483915FE82F8F9" box="[199,324,1880,1904]" italics="true" pageId="4" pageNumber="105">Operculum</emphasis>
(female pleopod 2;
<figureCitation id="9391BD874139FFC1FDBB391AFDBDF8F9" box="[564,635,1879,1904]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="4" pageNumber="105">Fig. 2</figureCitation>
B, D). Length 1.1 width. Lateral and posterior margin rounded; with numerous (&gt; 40) simple setae.
</paragraph>
<caption id="5FD5F18A4138FFC0FF1839D7FC1FF846" httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="5" pageNumber="106" targetBox="[184,1406,217,1880]" targetPageId="5">
<paragraph id="0B15A1024138FFC0FF1839D7FC1FF846" blockId="5.[151,1436,1946,1999]" pageId="5" pageNumber="106">
<emphasis id="39DE7D104138FFC0FF1839D7FED1F826" bold="true" box="[151,279,1946,1968]" pageId="5" pageNumber="106">FIGURE 2.</emphasis>
<taxonomicName id="CCAADA814138FFC0FEAE39D7FE35F839" box="[289,499,1946,1968]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D104138FFC0FEAE39D7FE35F839" box="[289,499,1946,1968]" italics="true" pageId="5" pageNumber="106">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4138FFC0FE7139D7FDFDF826" box="[510,571,1946,1967]" pageId="5" pageNumber="106" rank="species">
<emphasis id="39DE7D104138FFC0FE7139D7FDFDF826" bold="true" box="[510,571,1946,1967]" pageId="5" pageNumber="106">sp. n.</emphasis>
</taxonomicNameLabel>
; AD, holotype female (ZMH
<emphasis id="39DE7D104138FFC0FC0E39D6FC48F839" box="[897,910,1947,1968]" italics="true" pageId="5" pageNumber="106"></emphasis>
K
<emphasis id="39DE7D104138FFC0FC2F39D6FC6AF839" box="[928,940,1947,1968]" italics="true" pageId="5" pageNumber="106"></emphasis>
43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A
<emphasis id="39DE7D104138FFC0FD2939F7FD75F846" box="[678,691,1978,1999]" italics="true" pageId="5" pageNumber="106"></emphasis>
B = 500 Μm, C
<emphasis id="39DE7D104138FFC0FCC539F7FC91F846" box="[842,855,1978,1999]" italics="true" pageId="5" pageNumber="106"></emphasis>
D = 250 µm.
</paragraph>
</caption>
<caption id="5FD5F18A413BFFC3FF1839E6FAE7F856" httpUri="https://zenodo.org/record/225958/files/figure.png" pageId="6" pageNumber="107" targetBox="[209,1358,205,1962]" targetPageId="6">
<paragraph id="0B15A102413BFFC3FF1839E6FAE7F856" blockId="6.[151,1435,1963,2015]" pageId="6" pageNumber="107">
<emphasis id="39DE7D10413BFFC3FF1839E6FED3F849" bold="true" box="[151,277,1963,1985]" pageId="6" pageNumber="107">FIGURE 3.</emphasis>
<taxonomicName id="CCAADA81413BFFC3FE9239E6FE28F848" box="[285,494,1963,1985]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D10413BFFC3FE9239E6FE28F848" box="[285,494,1963,1985]" italics="true" pageId="6" pageNumber="107">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B413BFFC3FE7939E6FDF7F849" box="[502,561,1963,1984]" pageId="6" pageNumber="107" rank="species">
<emphasis id="39DE7D10413BFFC3FE7939E6FDF7F849" bold="true" box="[502,561,1963,1984]" pageId="6" pageNumber="107">sp. n.</emphasis>
</taxonomicNameLabel>
; AF, holotype female (ZMH
<emphasis id="39DE7D10413BFFC3FCE639E1FCB0F848" box="[873,886,1964,1985]" italics="true" pageId="6" pageNumber="107"></emphasis>
K
<emphasis id="39DE7D10413BFFC3FC0739E1FC52F848" box="[904,916,1964,1985]" italics="true" pageId="6" pageNumber="107"></emphasis>
43951); A, habitus, dorsal view. B, maxilliped. C,
<taxonomicName id="CCAADA81413BFFC3FF183984FF3BF856" box="[151,253,1993,2015]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="6" pageNumber="114" phylum="Ascomycota" rank="genus">antennula</taxonomicName>
. DE, pereopods III, detail: dorsal and ventral claw. F, antenna. Scale bar: A = 500 µm, B
<emphasis id="39DE7D10413BFFC3FB193987FB65F856" box="[1174,1187,1994,2015]" italics="true" pageId="6" pageNumber="107"></emphasis>
F = 250 Μm.
</paragraph>
</caption>
<caption id="5FD5F18A413AFFC2FF183869FBFDF9D1" httpUri="https://zenodo.org/record/225959/files/figure.png" pageId="7" pageNumber="108" targetBox="[225,1349,212,1538]" targetPageId="7">
<paragraph id="0B15A102413AFFC2FF183869FBFDF9D1" blockId="7.[151,1435,1572,1624]" pageId="7" pageNumber="108">
<emphasis id="39DE7D10413AFFC2FF183869FEDEF9B0" bold="true" box="[151,280,1572,1594]" pageId="7" pageNumber="108">FIGURE 4.</emphasis>
<taxonomicName id="CCAADA81413AFFC2FEAB3869FE31F9B3" box="[292,503,1572,1594]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D10413AFFC2FEAB3869FE31F9B3" box="[292,503,1572,1594]" italics="true" pageId="7" pageNumber="108">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B413AFFC2FD8C3869FD87F9B0" box="[515,577,1572,1593]" pageId="7" pageNumber="108" rank="species">
<emphasis id="39DE7D10413AFFC2FD8C3869FD87F9B0" bold="true" box="[515,577,1572,1593]" pageId="7" pageNumber="108">sp. n.</emphasis>
</taxonomicNameLabel>
; AD paratype male (ZMH
<emphasis id="39DE7D10413AFFC2FCFE3868FCB8F9B3" box="[881,894,1573,1594]" italics="true" pageId="7" pageNumber="108"></emphasis>
K
<emphasis id="39DE7D10413AFFC2FC1F3868FC5AF9B3" box="[912,924,1573,1594]" italics="true" pageId="7" pageNumber="108"></emphasis>
43952); A, habitus, dorsal view. B, antenna. C,
<taxonomicName id="CCAADA81413AFFC2FF18380FFF3BF9D1" box="[151,253,1602,1624]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="7" pageNumber="114" phylum="Ascomycota" rank="genus">antennula</taxonomicName>
. D, pleotelson, ventral view. Scale bar: A = 500 Μm, BC = 100 Μm; D = 250 Μm.
</paragraph>
</caption>
<paragraph id="0B15A102413AFFC2FF4838C8FB55F96C" blockId="7.[151,1436,1668,2018]" pageId="7" pageNumber="108">
<emphasis id="39DE7D10413AFFC2FF4838C8FEEEF914" box="[199,296,1669,1693]" italics="true" pageId="7" pageNumber="108">Uropods</emphasis>
(
<figureCitation id="9391BD87413AFFC2FEB838C9FEBDF914" box="[311,379,1668,1693]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="7" pageNumber="108">Fig. 2</figureCitation>
B, C). Biramous. Protopod trapezoid, length 1.2 width; with 3 simple setae distally. Exopodite about as long as protopod, length 2.8 width; with 1 simple seta terminally. Endopodite length 1.3 exopodite length; with 4 simple setae laterally, with 3 broom setae and 1 long, slender simple seta terminally.
</paragraph>
<paragraph id="0B15A102413AFFC2FF4838BCFE30F883" blockId="7.[151,1436,1668,2018]" box="[199,502,1777,1802]" pageId="7" pageNumber="108">
Description
<typeStatus id="D4111FA0413AFFC2FEDF38BFFE77F883" box="[336,433,1778,1802]" pageId="7" pageNumber="108" type="paratype">paratype</typeStatus>
male.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289413AFFCCFF483958FD63F81C" lastPageId="9" lastPageNumber="110" pageId="7" pageNumber="108" type="discussion">
<paragraph id="0B15A102413AFFC2FF483958FDE2F8DB" blockId="7.[151,1436,1668,2018]" pageId="7" pageNumber="108">
<emphasis id="39DE7D10413AFFC2FF483958FEFDF8A7" bold="true" box="[199,315,1813,1838]" pageId="7" pageNumber="108">Remarks.</emphasis>
Details of the habitus, antenna,
<taxonomicName id="CCAADA81413AFFC2FD5F3958FC86F8A7" box="[720,832,1813,1838]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="7" pageNumber="114" phylum="Ascomycota" rank="genus">antennula</taxonomicName>
, maxilliped, pereopods III and uropods are only described when different to female.
</paragraph>
<paragraph id="0B15A102413AFFC2FF483910FDBEF837" blockId="7.[151,1436,1668,2018]" pageId="7" pageNumber="108">
<emphasis id="39DE7D10413AFFC2FF483910FEE6F8FC" box="[199,288,1885,1909]" italics="true" pageId="7" pageNumber="108">Habitus</emphasis>
(
<figureCitation id="9391BD87413AFFC2FEBE3910FEBDF8FC" box="[305,379,1885,1910]" captionStart="FIGURE 4" captionStartId="7.[151,250,1572,1594]" captionTargetBox="[225,1349,212,1538]" captionTargetId="figure@7.[217,1369,193,1550]" captionTargetPageId="7" captionText="FIGURE 4. Hebefustis juansenii sp. n.; A D paratype male (ZMH K 43952); A, habitus, dorsal view. B, antenna. C, antennula. D, pleotelson, ventral view. Scale bar: A = 500 Μm, B C = 100 Μm; D = 250 Μm." httpUri="https://zenodo.org/record/225959/files/figure.png" pageId="7" pageNumber="108">Figs 4</figureCitation>
A, D, 5A) similar to
<typeStatus id="D4111FA0413AFFC2FDE03910FD17F8FF" box="[623,721,1885,1910]" pageId="7" pageNumber="108" type="holotype">holotype</typeStatus>
female. Body length 4.1 pereonite 2 width. Pereonites 14 of similar width; pereonite 4 anterior margin slightly concave. Pleotelson length 0.25 body length. Pleopod 1 length 0.6 pleotelson length (ventrally measured).
</paragraph>
<paragraph id="0B15A102413AFFC2FF483984FC95F86B" blockId="7.[151,1436,1668,2018]" box="[199,851,1993,2018]" pageId="7" pageNumber="108">
<emphasis id="39DE7D10413AFFC2FF483984FEA9F868" box="[199,367,1993,2017]" italics="true" pageId="7" pageNumber="108">Cephalothorax</emphasis>
(
<figureCitation id="9391BD87413AFFC2FEF13984FE03F868" box="[382,453,1993,2018]" captionStart="FIGURE 4" captionStartId="7.[151,250,1572,1594]" captionTargetBox="[225,1349,212,1538]" captionTargetId="figure@7.[217,1369,193,1550]" captionTargetPageId="7" captionText="FIGURE 4. Hebefustis juansenii sp. n.; A D paratype male (ZMH K 43952); A, habitus, dorsal view. B, antenna. C, antennula. D, pleotelson, ventral view. Scale bar: A = 500 Μm, B C = 100 Μm; D = 250 Μm." httpUri="https://zenodo.org/record/225959/files/figure.png" pageId="7" pageNumber="108">Figs 4</figureCitation>
A, 5A). Similar to
<typeStatus id="D4111FA0413AFFC2FD193984FD3EF86B" box="[662,760,1993,2018]" pageId="7" pageNumber="108" type="holotype">holotype</typeStatus>
female.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483ED5FD50FF71" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<taxonomicName id="CCAADA814135FFCDFF483ED5FEFCFF39" box="[199,314,152,176]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="8" pageNumber="109" phylum="Ascomycota" rank="genus">
<emphasis id="39DE7D104135FFCDFF483ED5FEFCFF39" box="[199,314,152,176]" italics="true" pageId="8" pageNumber="109">Antennula</emphasis>
</taxonomicName>
(
<figureCitation id="9391BD874135FFCDFEC33EDAFE55FF39" box="[332,403,151,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1572,1594]" captionTargetBox="[225,1349,212,1538]" captionTargetId="figure@7.[217,1369,193,1550]" captionTargetPageId="7" captionText="FIGURE 4. Hebefustis juansenii sp. n.; A D paratype male (ZMH K 43952); A, habitus, dorsal view. B, antenna. C, antennula. D, pleotelson, ventral view. Scale bar: A = 500 Μm, B C = 100 Μm; D = 250 Μm." httpUri="https://zenodo.org/record/225959/files/figure.png" pageId="8" pageNumber="109">Fig. 4</figureCitation>
B). Length 0.16 body length. Article 1 with 1 broom seta distally; article 2 with 3 broom setae and 1 simple seta distally; article 3 with 1 simple seta; article 4 with 1 broom seta distally; article 5 with 1 long aesthetasc and 2 simple setae terminally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483F49FA95FD91" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483F49FEE2FE92" box="[199,292,260,283]" italics="true" pageId="8" pageNumber="109">Antenna</emphasis>
(
<figureCitation id="9391BD874135FFCDFEBB3F49FEBFFE95" box="[308,377,260,285]" captionStart="FIGURE 4" captionStartId="7.[151,250,1572,1594]" captionTargetBox="[225,1349,212,1538]" captionTargetId="figure@7.[217,1369,193,1550]" captionTargetPageId="7" captionText="FIGURE 4. Hebefustis juansenii sp. n.; A D paratype male (ZMH K 43952); A, habitus, dorsal view. B, antenna. C, antennula. D, pleotelson, ventral view. Scale bar: A = 500 Μm, B C = 100 Μm; D = 250 Μm." httpUri="https://zenodo.org/record/225959/files/figure.png" pageId="8" pageNumber="109">Fig. 4</figureCitation>
C). Length 0.4 body length; with 6 podomere and 8 flagellar articles. Articles 57 stouter than in female. Podomere articles 14 short; article 2 with 2 simple setae distally; article 3 with 1 robust spine tipped with a small setule and 2 simple setae distally. Article 5 length about 3 width; with 3 simple setae and 1 broom seta laterally, with 3 simple setae distally. Article 6 length about as long as article 5 length, length 2.7 width; with 2 simple setae laterally, with 2 broom setae and 4 small simple setae distally. Unidentified ecto-parasites attached to article 6 dorsal cuticle. Flagellar article 1 longest, length 0.8 podomere article 6 length, length 3.1 width; with 2 simple setae laterally and 3 simple setae distally. Flagellar articles 27 of similar length, width decreasing distally; each article with 14 simple setae distally. Article 8 somewhat shorter; with 4 long, slender setae terminally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483C69FEFAFC09" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483C69FEFBFDB5" box="[199,317,548,572]" italics="true" pageId="8" pageNumber="109">Mandibles</emphasis>
(
<figureCitation id="9391BD874135FFCDFEC03C69FE4CFDB4" box="[335,394,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[212,1386,218,1942]" captionTargetId="figure@10.[188,1398,195,1954]" captionTargetPageId="10" captionText="FIGURE 5. Hebefustis juansenii sp. n.; A E paratype male (ZMH K 43952); A, habitus, dorsal view. B, left mandible. C, right mandible. D, maxilliped. E, maxillula. F, maxilla, detail: inner endite. Scale bar: A = 500 Μm, B F = 100 Μm." httpUri="https://zenodo.org/record/225960/files/figure.png" pageId="8" pageNumber="109">Fig.5</figureCitation>
BC). Left and right mandibular palps well developed, consisting of 3 articles and almost reaching incisor. Right mandible palpal article 1 length 0.8 article 2 length. Article 2 with 2 simple setae laterally. Terminal article length about half article 2 length, tapering distally; with a fringe of small setules ventrally and 3 robust setae terminally. Left mandible palpus similar to right mandible; article 2 with fringe of fine setules and 2 simple setae distally. Right mandible incisor process with 7 teeth, left mandible incisor with 7 teeth and a subdistal tooth proximally. Left mandible lacinia mobilis broken off. Right mandible spine row with 10 robust spines and several slender spines in between; dentation decreasing, seta size increasing proximally. Left mandible spine row with 10 robust spines and several slender spines in between, dentation decreasing, seta size increasing proximally. Molar of right and left mandible triangular; right mandible molar with 8, left mandible molar with 7 long, serrate spines distally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483DC1FC68FC41" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483DC1FEF2FC2D" box="[199,308,908,932]" italics="true" pageId="8" pageNumber="109">Maxillula</emphasis>
(
<figureCitation id="9391BD874135FFCDFECB3DC1FE41FC2C" box="[324,391,908,933]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[212,1386,218,1942]" captionTargetId="figure@10.[188,1398,195,1954]" captionTargetPageId="10" captionText="FIGURE 5. Hebefustis juansenii sp. n.; A E paratype male (ZMH K 43952); A, habitus, dorsal view. B, left mandible. C, right mandible. D, maxilliped. E, maxillula. F, maxilla, detail: inner endite. Scale bar: A = 500 Μm, B F = 100 Μm." httpUri="https://zenodo.org/record/225960/files/figure.png" pageId="8" pageNumber="109">Fig. 5</figureCitation>
E). Left and right maxillula inner endites lost during dissection. Outer endite length 4 width; with 11 strong spine-like setae distally, with few simple setae laterally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483D99FC45FBBC" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483D99FED8FC65" box="[199,286,980,1004]" italics="true" pageId="8" pageNumber="109">Maxilla</emphasis>
(
<figureCitation id="9391BD874135FFCDFEA03D99FEB5FC64" box="[303,371,980,1005]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[212,1386,218,1942]" captionTargetId="figure@10.[188,1398,195,1954]" captionTargetPageId="10" captionText="FIGURE 5. Hebefustis juansenii sp. n.; A E paratype male (ZMH K 43952); A, habitus, dorsal view. B, left mandible. C, right mandible. D, maxilliped. E, maxillula. F, maxilla, detail: inner endite. Scale bar: A = 500 Μm, B F = 100 Μm." httpUri="https://zenodo.org/record/225960/files/figure.png" pageId="8" pageNumber="109">Fig. 5</figureCitation>
F). Inner endite longest and broadest; outer margin with several slender setae, with numerous short, simple setae medially, with 5 somewhat more robust setae distally. Medial and outer endite each with 3 robust, long setae. Outer endite with several slender setae laterally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483A0DFA86FBF4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483A0DFE86FBD1" box="[199,320,1088,1112]" italics="true" pageId="8" pageNumber="109">Maxilliped</emphasis>
(
<figureCitation id="9391BD874135FFCDFEDC3A72FE5FFBD1" box="[339,409,1087,1112]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[212,1386,218,1942]" captionTargetId="figure@10.[188,1398,195,1954]" captionTargetPageId="10" captionText="FIGURE 5. Hebefustis juansenii sp. n.; A E paratype male (ZMH K 43952); A, habitus, dorsal view. B, left mandible. C, right mandible. D, maxilliped. E, maxillula. F, maxilla, detail: inner endite. Scale bar: A = 500 Μm, B F = 100 Μm." httpUri="https://zenodo.org/record/225960/files/figure.png" pageId="8" pageNumber="109">Fig. 5</figureCitation>
D). Similar to female maxilliped. One coupling hook broken off, Palpal article 2 with 2 simple setae laterally. Article 3 with 5 robust sensillae distally. Article 5 with 3 long, slender setae distally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483AC5FA57FA84" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483AC5FE84FB29" box="[199,322,1160,1184]" italics="true" pageId="8" pageNumber="109">Pereopod I</emphasis>
(
<figureCitation id="9391BD874135FFCDFEDE3ACAFE55FB29" box="[337,403,1159,1184]" captionStart="FIGURE 6" captionStartId="11.[151,250,1937,1959]" captionTargetBox="[289,1284,220,1900]" captionTargetId="figure@11.[275,1312,193,1916]" captionTargetPageId="11" captionText="FIGURE 6. Hebefustis juansenii sp. n.; A C paratype male (ZMH K 43952); A, pereopod I, detail: dorsal and ventral claw. B, pereopod II. C, pereopod III. Scale bar: A C = 100 Μm." httpUri="https://zenodo.org/record/225961/files/figure.png" pageId="8" pageNumber="109">Fig. 6</figureCitation>
A). Similar to pereopod II. Basis length 2.9 width; with 1 slender simple seta dorsally, with 2 simple setae ventrally and 1 long slender seta distoventrally. Ischium 0.6 basis length; with 1 simple seta dorsally. Carpus with 4 robust unequally bifid setae ventrally. Propodus with 1 slender seta dorsally and 2 slender simple setae distodorsally, with 2 robust unequally bifid setae ventrally and 2 slender setae of varying size distoventrally.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483B55FD5AFA49" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483B55FE88FAB9" box="[199,334,1304,1328]" italics="true" pageId="8" pageNumber="109">Pereopod II</emphasis>
(
<figureCitation id="9391BD874135FFCDFED13B5AFE64FAB9" box="[350,418,1303,1328]" captionStart="FIGURE 6" captionStartId="11.[151,250,1937,1959]" captionTargetBox="[289,1284,220,1900]" captionTargetId="figure@11.[275,1312,193,1916]" captionTargetPageId="11" captionText="FIGURE 6. Hebefustis juansenii sp. n.; A C paratype male (ZMH K 43952); A, pereopod I, detail: dorsal and ventral claw. B, pereopod II. C, pereopod III. Scale bar: A C = 100 Μm." httpUri="https://zenodo.org/record/225961/files/figure.png" pageId="8" pageNumber="109">Fig. 6</figureCitation>
B). Basis with 1 simple seta dorsally, with 3 simple setae ventrally and 1 long simple seta distally. Ischium with 1 simple seta dorsally, with 2 simple setae ventrally. Carpus 2 slender simple setae and 1 broom seta dorsally, with 5 stout unequally bifid setae ventrally. Propodus length 0.7 carpus length, length 6.5 width; with 2 simple setae and 1 broom seta dorsally, with 3 stout unequally bifid setae ventrally. Dactylus with 2 slender setae between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A1024135FFCDFF483B81FAD6F969" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF483B81FE91FA6D" box="[199,343,1484,1508]" italics="true" pageId="8" pageNumber="109">Pereopod III</emphasis>
(
<figureCitation id="9391BD874135FFCDFEE83B81FE6AFA6C" box="[359,428,1484,1509]" captionStart="FIGURE 6" captionStartId="11.[151,250,1937,1959]" captionTargetBox="[289,1284,220,1900]" captionTargetId="figure@11.[275,1312,193,1916]" captionTargetPageId="11" captionText="FIGURE 6. Hebefustis juansenii sp. n.; A C paratype male (ZMH K 43952); A, pereopod I, detail: dorsal and ventral claw. B, pereopod II. C, pereopod III. Scale bar: A C = 100 Μm." httpUri="https://zenodo.org/record/225961/files/figure.png" pageId="8" pageNumber="109">Fig. 6</figureCitation>
C). Basis length 4 width; with 1 simple seta dorsally, with 3 simple setae ventrally and 1 long simple seta distally. Ischium length 0.6 basis length, length 2.6 width; with 2 simple setae dorsally, with 2 simple setae ventrally. Merus length 0.6 ischium length, length 1.7 width; with 1 simple seta distodorsally, with 2 simple setae ventrally. Carpus length 2.5 merus length, length 5.2 width; with 2 slender simple setae dorsally, with 5 stout unequally bifid setae ventrally. Propodus length 0.8 carpus length, length 5.0 width, with 2 simple setae and 1 long broom seta dorsally, with numerous small setae, membranously embedded, 3 stout unequally bifid setae and 1 simple seta ventrally. Dactylus length one-third of propodus length, length 3.6 width; with 3 simple setae medially. Dorsal claw length 0.8 dactylus length; with 2 slender setae between dorsal and ventral claw
</paragraph>
<paragraph id="0B15A1024135FFCDFF4838A1FA5BF854" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="109">
<emphasis id="39DE7D104135FFCDFF4838A1FE8CF88D" box="[199,330,1772,1796]" italics="true" pageId="8" pageNumber="109">Pereopod V</emphasis>
(
<figureCitation id="9391BD874135FFCDFED638A1FE50F88C" box="[345,406,1772,1797]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="8" pageNumber="109">Fig.7</figureCitation>
A). Pereopods V and VII much longer and more slender than pereopods I
<emphasis id="39DE7D104135FFCDFB4938A0FB12F88D" box="[1222,1236,1773,1796]" italics="true" pageId="8" pageNumber="109"></emphasis>
III. Basis length 4 width; with 1 simple seta and 1 long broom seta dorsally, with 4 simple setae ventrally. Ischium length 0.7 basis length, length 3 width; with 1 simple seta dorsally, 3 simple setae medially and with 2 simple setae dorsally. Merus length 0.5 ischium length, length twice width; with 2 long robust setae distodorsally, with 1 simple seta ventrally. Carpus length 2.8 merus length, length 7.2 width; with row of 5 long, slender natatory setae dorsally, with 3 simple setae ventrally, increasing in length distally. Propodus about as long as carpus, length 6.4 width; with row of 9 long, slender natatory setae dorsally, with 2 somewhat shorter simple setae distally, with 1 long, slender seta ventrally.
</paragraph>
<paragraph id="0B15A1024134FFCCFF183EDAFC5BFF5C" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">Dactylus length 0.25 propodus length, length 3.0 width; with 3 simple setae medially. Dorsal claw length 0.7 dactylus length; with 2 slender setae between dorsal and ventral claw</paragraph>
<paragraph id="0B15A1024134FFCCFF483EADFC65FE7C" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483EADFEA7FF71" box="[199,353,224,248]" italics="true" pageId="9" pageNumber="110">Pereopod VII</emphasis>
(
<figureCitation id="9391BD874134FFCCFEFA3E92FE72FF71" box="[373,436,223,248]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig.7</figureCitation>
B). Basis length 4.0 width; with 1 simple seta dorsally, with 2 simple setae ventrally. Ischium about half basis length, length 2.8 width; with 1 simple seta ventrally. Merus length 0.6 ischium length, length 2.1 width, with 2 simple setae distodorsally, with 2 setae of varying size distoventrally. Carpus length twice merus length, length 4.7 width; with 2 simple setae of varying size distodorsally, with 1 long, slender simple seta distoventrally. Propodus about as long as carpus, length 6.0 width, with 1 long, slender simple seta and 1 broom seta distodorsally, with 1 short simple seta ventrally and 1 stout unequally bifid and 1 simple seta distoventrally. Dactylus length one-third propodus length, length 3.5 width; with 3 simple setae medially. Dorsal claw length 0.7 dactylus length; with 2 slender setae between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483C4DFA4BFDB4" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483C4DFEFDFD91" box="[199,315,512,536]" italics="true" pageId="9" pageNumber="110">Pleopod 1</emphasis>
(
<figureCitation id="9391BD874134FFCCFEC33FB2FE57FD91" box="[332,401,511,536]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
C). Length 1.7 proximal width. Distal projection width 0.8 proximal width; lateral margins concave. Lateral lobes rounded; with 1 long simple seta. Distal margins almost straight, with 6 simple setae each.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483C05FDB0FD44" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483C05FEFAFDE9" box="[199,316,584,608]" italics="true" pageId="9" pageNumber="110">Pleopod 2</emphasis>
(
<figureCitation id="9391BD874134FFCCFEC23C0AFE55FDE9" box="[333,403,583,608]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
D). Sympod length 2.4 width, outer margin rounded; with 6 slender simple setae laterally increasing in size towards distal end; inner margin slightly concave. Endopod inserting 0.5 from distal tip of sympod; stylet length 0.7 sympod length, slightly curved, distal end almost reaching tip of sympod. Exopod short and rounded, with few tiny setule laterally.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483C95FAEAFCB1" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483C95FEFCFD79" box="[199,314,728,752]" italics="true" pageId="9" pageNumber="110">Pleopod 3</emphasis>
(
<figureCitation id="9391BD874134FFCCFEC43C9AFE56FD79" box="[331,400,727,752]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
E). Protopodite length 1.5 width, length about half endopodite length. Exopodite about half endopodite length, length 1.3 width, tapering in width distally; with numerous short simple setae laterally and 1 long, robust seta distally. Endopodite length 1.4 width; with 3 long plumose setae distally (1 broken off).
</paragraph>
<paragraph id="0B15A1024134FFCCFF483D09FC31FC2C" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483D09FEFFFCD5" box="[199,313,836,860]" italics="true" pageId="9" pageNumber="110">Pleopod 4</emphasis>
(
<figureCitation id="9391BD874134FFCCFEC83D09FE4DFCD5" box="[327,395,836,861]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
F). Protopodite rectangular, length 0.5 width, about 0.4 endopodite length. Exopodite slender, about as long as endopodite, length 6.5 width; with several thin setules laterally (one-sided) and 1 long robust plumose seta distally. Endopodite ovoid-shaped, about twice as long as wide.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483DFDFDD2FC64" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483DFDFEFCFC4E" box="[199,314,944,968]" italics="true" pageId="9" pageNumber="110">Pleopod 5</emphasis>
(
<figureCitation id="9391BD874134FFCCFEC63DE2FE4AFC41" box="[329,396,943,968]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
G) small oval lobe, without setation, about as long as pleopod 4. Length 2.4 proximal width; width tapering towards distal end.
</paragraph>
<paragraph id="0B15A1024134FFCCFF5E3DB5FDEAFBD1" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF5E3DB5FEF4FB99" box="[209,306,1016,1040]" italics="true" pageId="9" pageNumber="110">Uropods</emphasis>
(drawn
<emphasis id="39DE7D104134FFCCFE1B3DB5FE1AFB86" box="[404,476,1016,1039]" italics="true" pageId="9" pageNumber="110">in situ</emphasis>
;
<figureCitation id="9391BD874134FFCCFE613DBAFDF4FB99" box="[494,562,1015,1040]" captionStart="FIGURE 7" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[200,1386,218,1915]" captionTargetId="figure@12.[177,1409,193,1935]" captionTargetPageId="12" captionText="FIGURE 7. Hebefustis juansenii sp. n.; A H paratype male (ZMH K 43952); A, pereopod V. B, pereopod VII. C G, pleopods 1 5. H, uropod. Scale bar: A H = 100 Μm." httpUri="https://zenodo.org/record/225962/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
H). Protopod trapezoid with 1 simple seta distally. Exopodite with 3 simple setae (1 broken off) terminally. Endopodite with 2 simple setae laterally, with 3 broom setae (1 broken off) and 1 long, slender simple seta terminally.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483A29FCB9FA14" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<emphasis id="39DE7D104134FFCCFF483A29FEFDFBF4" bold="true" box="[199,315,1124,1149]" pageId="9" pageNumber="110">Remarks.</emphasis>
Species in the genus
<taxonomicName id="CCAADA814134FFCCFDA13A29FD62FBF5" box="[558,676,1124,1148]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="39DE7D104134FFCCFDA13A29FD62FBF5" box="[558,676,1124,1148]" italics="true" pageId="9" pageNumber="110">Hebefustis</emphasis>
</taxonomicName>
have been previously assigned to two distinct clusters based on the shape of the lateral margins of pereonites 6 and 7 (acute vs. rounded;
<bibRefCitation id="6F3BDCF34134FFCCFC383ACAFB32FB29" author="Siebenaller" box="[951,1268,1159,1184]" pageId="9" pageNumber="110" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History. 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler 1977</bibRefCitation>
;
<bibRefCitation id="6F3BDCF34134FFCCFA8D3ACAFA57FB29" author="Kaiser" box="[1282,1425,1159,1184]" pageId="9" pageNumber="110" refString="Kaiser, S. (2008) First record and new species of the genus Hebefustis Siebenaller &amp; Hessler, 1977 (Isopoda: Asellota: Nannoniscidae) from the deep Weddell Sea Basin. Zootaxa, 1866, 467 - 481." type="journal article" year="2008">Kaiser 2008</bibRefCitation>
). Aside from the new species rounded margins are also present in
<emphasis id="39DE7D104134FFCCFCE23AE0FF30FB61" italics="true" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFCE23AE0FC17FB4D" box="[877,977,1196,1220]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="alleni">H. alleni</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFC523AE0FBB8FB4D" box="[989,1150,1196,1220]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="mollicellus">H. mollicellus</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFB063AE0FB15FB4D" box="[1161,1235,1197,1220]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="par">H. par</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFB513AE0FAB0FB4D" box="[1246,1398,1197,1220]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="primitivus">H. primitivus</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFA0E3AE0FF30FB61" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="robustus">H. robustus</taxonomicName>
</emphasis>
and
<emphasis id="39DE7D104134FFCCFEBD3A9CFD42FB6E" box="[306,644,1232,1256]" italics="true" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFEBD3A9CFE57FB61" box="[306,401,1232,1256]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="vafer">H. vafer</taxonomicName>
.
<taxonomicName id="CCAADA814134FFCCFE113A9DFD42FB6E" box="[414,644,1232,1256]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">Hebefustis juansenii</taxonomicName>
</emphasis>
<taxonomicNameLabel id="22EDC06B4134FFCCFD003A82FD17FB61" box="[655,721,1231,1256]" pageId="9" pageNumber="110" rank="species">
<emphasis id="39DE7D104134FFCCFD003A82FD17FB61" bold="true" box="[655,721,1231,1256]" pageId="9" pageNumber="110">sp. n.</emphasis>
</taxonomicNameLabel>
can be identified by possessing a robust spine on pereonite 2 antero-lateral margin in both male and female. The following species also bear a spine on pereonite 2:
<emphasis id="39DE7D104134FFCCFA0E3AB8FE6CFAB9" italics="true" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFA0E3AB8FEDEFAB9" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="clareolithis">H. clareolithis</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFEAA3B54FE6CFAB9" box="[293,426,1305,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cornutus">H. cornutus</taxonomicName>
</emphasis>
,
<emphasis id="39DE7D104134FFCCFE363B54FD60FAB9" box="[441,678,1304,1328]" italics="true" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFE363B54FDF8FAB9" box="[441,574,1304,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="robustus">H. robustus</taxonomicName>
,
<taxonomicName id="CCAADA814134FFCCFDC43B54FD60FAB9" box="[587,678,1304,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="vafer">H. vafer</taxonomicName>
</emphasis>
,
<taxonomicName id="CCAADA814134FFCCFD3A3B54FC92FAB9" box="[693,852,1304,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="mollicellus">
<emphasis id="39DE7D104134FFCCFD3A3B54FC92FAB9" box="[693,852,1304,1328]" italics="true" pageId="9" pageNumber="110">H. mollicellus</emphasis>
</taxonomicName>
,
<emphasis id="39DE7D104134FFCCFCEC3B54FB5BFAA6" box="[867,1181,1304,1328]" italics="true" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFCEC3B54FC08FAB9" box="[867,974,1304,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="dispar">H. dispar</taxonomicName>
and
<taxonomicName id="CCAADA814134FFCCFB883B54FB5BFAA6" box="[1031,1181,1304,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="primitivus">H. primitivus</taxonomicName>
</emphasis>
. However, in all these species this feature seems to be sexually dimorphic, as it is only present in males (
<bibRefCitation id="6F3BDCF34134FFCCFBD93B71FA51FADD" author="Siebenaller" box="[1110,1431,1340,1365]" pageId="9" pageNumber="110" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History. 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler 1977</bibRefCitation>
;
<bibRefCitation id="6F3BDCF34134FFCCFF183B12FEE1FAF1" author="Kaiser" box="[151,295,1375,1400]" pageId="9" pageNumber="110" refString="Kaiser, S. (2008) First record and new species of the genus Hebefustis Siebenaller &amp; Hessler, 1977 (Isopoda: Asellota: Nannoniscidae) from the deep Weddell Sea Basin. Zootaxa, 1866, 467 - 481." type="journal article" year="2008">Kaiser 2008</bibRefCitation>
). In
<taxonomicName id="CCAADA814134FFCCFED13B2CFE33FAFE" box="[350,501,1376,1400]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="primitivus">
<emphasis id="39DE7D104134FFCCFED13B2CFE33FAFE" box="[350,501,1376,1400]" italics="true" pageId="9" pageNumber="110">H. primitivus</emphasis>
</taxonomicName>
only the male is illustrated, though. So, for this for this species it is not known whether the occurrence of a spine on pereonite 2 is gender related.
</paragraph>
<paragraph id="0B15A1024134FFCCFF483BEAFAF7F911" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
Furthermore,
<taxonomicName id="CCAADA814134FFCCFEEC3BE4FE2BFA36" box="[355,493,1448,1472]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="juansenii">
<emphasis id="39DE7D104134FFCCFEEC3BE4FE2BFA36" box="[355,493,1448,1472]" italics="true" pageId="9" pageNumber="110">H. juansenii</emphasis>
</taxonomicName>
can be distinguished from all species in the genus by the presence of two pairs of posterolateral spine on the pleotelson, with the proximal pair inserting in an acute angle (≤45°) 0.6 from the pleotelson anterior margin. Two pairs of posterolateral spines can also be found in
<taxonomicName id="CCAADA814134FFCCFBB23BBCFB14F98E" box="[1085,1234,1520,1544]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="primitivus">
<emphasis id="39DE7D104134FFCCFBB23BBCFB14F98E" box="[1085,1234,1520,1544]" italics="true" pageId="9" pageNumber="110">H. primitivus</emphasis>
</taxonomicName>
, but differs in the angle (90°) and distance (0.3) from the anterior margin. The new species can be further delimited from
<taxonomicName id="CCAADA814134FFCCFA0E3858FECFF9C6" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="primitivus">
<emphasis id="39DE7D104134FFCCFA0E3858FECFF9C6" italics="true" pageId="9" pageNumber="110">H. primitivus</emphasis>
</taxonomicName>
by the following characters: pereonites 14 lateral margins acute (vs. rounded); pleotelson lateral margins straight (vs. rounded); uropods clearly extending beyond posterior margin (vs. not visible in dorsal view); male pleopod 1 distal projection forming 90° angle (vs. straight), lateral lobes with 3 simple setae (vs. 1).
</paragraph>
<paragraph id="0B15A1024134FFCCFF4838E9FD63F81C" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="110">
<taxonomicName id="CCAADA814134FFCCFF4838E9FE6DF935" box="[199,427,1700,1724]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D104134FFCCFF4838E9FE6DF935" box="[199,427,1700,1724]" italics="true" pageId="9" pageNumber="110">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4134FFCCFE3E38E9FE37F934" box="[433,497,1700,1725]" pageId="9" pageNumber="110" rank="species">
<emphasis id="39DE7D104134FFCCFE3E38E9FE37F934" bold="true" box="[433,497,1700,1725]" pageId="9" pageNumber="110">sp. n.</emphasis>
</taxonomicNameLabel>
shares some similarities with
<taxonomicName id="CCAADA814134FFCCFCCD38E8FC19F935" box="[834,991,1700,1724]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="mollicellus">
<emphasis id="39DE7D104134FFCCFCCD38E8FC19F935" box="[834,991,1700,1724]" italics="true" pageId="9" pageNumber="110">H. mollicellus</emphasis>
</taxonomicName>
. The latter differs from the new species as follows: male pleopod 1; operculum lateral and distal margin with &lt;20 simple setae (vs.&gt; 40). The new species can be distinguished from
<taxonomicName id="CCAADA814134FFCCFE4D38A0FD82F88D" box="[450,580,1772,1796]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="39DE7D104134FFCCFE4D38A0FD82F88D" box="[450,580,1772,1796]" italics="true" pageId="9" pageNumber="110">H. robustus</emphasis>
</taxonomicName>
based on the following characters: operculum posterior margin straight (vs. rounded); maxilliped epipodite reaching mid of palpal article 3 (vs. upper-third of palpal article 2); pleopod 2 stylet long, clearly extending beyond tip of sympod (vs. almost reaching the tip); sympod outer margin with 9 simple setae (vs. 6).
<taxonomicName id="CCAADA814134FFCCFEA03915FDD2F8E6" box="[303,532,1880,1904]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D104134FFCCFEA03915FDD2F8E6" box="[303,532,1880,1904]" italics="true" pageId="9" pageNumber="110">Hebefustis juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4134FFCCFD93391AFD9BF8F9" box="[540,605,1879,1904]" pageId="9" pageNumber="110" rank="species">
<emphasis id="39DE7D104134FFCCFD93391AFD9BF8F9" bold="true" box="[540,605,1879,1904]" pageId="9" pageNumber="110">sp. n.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="CCAADA814134FFCCFD7B3914FC97F8F9" box="[756,849,1880,1904]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="vafer">
<emphasis id="39DE7D104134FFCCFD7B3914FC97F8F9" box="[756,849,1880,1904]" italics="true" pageId="9" pageNumber="110">H. vafer</emphasis>
</taxonomicName>
as follows: posterolateral projection in female only slightly pronounced (vs. strongly pronounced).
</paragraph>
</subSubSection>
</treatment>
</document>