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<document id="EE10ACDA41CDF35ECB3B3007B2B0210E" ID-CLB-Dataset="299601" ID-DOI="10.11646/zootaxa.5474.5.7" ID-GBIF-Dataset="0adc191b-8f3d-4270-8c14-c08f43b4619d" ID-ISSN="1175-5326" ID-Zenodo-Dep="12682122" ID-ZooBank="7DAFD532-8FE8-4755-B664-57A7102E7E00" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1720373948580" checkinUser="plazi" docAuthor="Thacker, Dimple, Myers, Alan A. &amp; Trivedi, Jigneshkumar N." docDate="2024" docId="423BD1464073EE7044C5FB8DFEC52D12" docLanguage="en" docName="zootaxa.5474.5.7.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan 2016" docType="treatment" docVersion="5" lastPageNumber="569" masterDocId="BE02A93E4072EE764452FF82FFA9296F" masterDocTitle="On a small collection of Maeridae Krapp-Schickel, 2008 (Crustacea: Amphipoda) from Gujarat, India" masterLastPageNumber="583" masterPageNumber="563" pageNumber="564" updateTime="1720769339063" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:title id="FEBB6213B17D3B3EA64A11AAD64AC514">On a small collection of Maeridae Krapp-Schickel, 2008 (Crustacea: Amphipoda) from Gujarat, India</mods:title>
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<mods:roleTerm id="85207655D180C32617164DF8B76FBEC7">Author</mods:roleTerm>
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<mods:namePart id="C009AEFB5AC485AC016D8E13A77992F9">Thacker, Dimple</mods:namePart>
<mods:nameIdentifier id="6FACB74090E7A14F4133656E43722631" type="ORCID">0000-0003-3101-0028</mods:nameIdentifier>
<mods:affiliation id="413E4343FE9907330CF24C912F99E44F">Department of Life sciences, Hemchandracharya North Gujarat University, Patan- 384265, Gujarat, India.</mods:affiliation>
<mods:nameIdentifier id="8931C11164396E920AD5DDB9B9854C93" type="email">dimplethacker789@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5479C994E690A666FEC59DFDB3370584">Myers, Alan A.</mods:namePart>
<mods:nameIdentifier id="137C904F5A9444245C827F64EED98E02" type="ORCID">0000-0003-3256-2123</mods:nameIdentifier>
<mods:affiliation id="CB0A3B29148EC3DDD6E929555E40BE0A">School of Biological, Earth and Environmental Sciences, University College Cork, Cork Enterprise Centre, Distillery Fields, North Mall, Cork, Ireland.</mods:affiliation>
<mods:nameIdentifier id="2ADB76FEDC194EA247C2F149DD0DC902" type="email">bavayia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="80EBE7C577A839F164460879E6609F62">Trivedi, Jigneshkumar N.</mods:namePart>
<mods:affiliation id="EFE01681872BA18F295F1D98100A62FD">Department of Life sciences, Hemchandracharya North Gujarat University, Patan- 384265, Gujarat, India.</mods:affiliation>
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<mods:title id="580788D88BADEE47AE96F9913DC2E213">Zootaxa</mods:title>
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<mods:date id="6CA95BE06571612D502C76F0A623A870">2024</mods:date>
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<mods:number id="4141E364417FAB40EE1B1FFA3B578801">5474</mods:number>
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<treatment id="423BD1464073EE7044C5FB8DFEC52D12" ID-DOI="http://doi.org/10.5281/zenodo.12682177" ID-GBIF-Taxon="234743427" ID-Zenodo-Dep="12682177" LSID="urn:lsid:plazi:treatment:423BD1464073EE7044C5FB8DFEC52D12" httpUri="http://treatment.plazi.org/id/423BD1464073EE7044C5FB8DFEC52D12" lastPageId="6" lastPageNumber="569" pageId="1" pageNumber="564">
<subSubSection id="828833DB4073EE7744C5FB8DFC142D45" box="[151,957,1039,1066]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="CA2D60504073EE7744C5FB8DFC142D45" blockId="1.[151,957,1039,1102]" box="[151,957,1039,1066]" pageId="1" pageNumber="564">
<heading id="9165D73C4073EE7744C5FB8DFC142D45" bold="true" box="[151,957,1039,1066]" fontSize="11" level="1" pageId="1" pageNumber="564" reason="1">
<taxonomicName id="0D921BD34073EE7744C5FB8DFC142D45" ID-CoL="SG9J" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[151,957,1039,1066]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424073EE7744C5FB8DFC142D45" bold="true" box="[151,957,1039,1066]" pageId="1" pageNumber="564">
<emphasis id="F8E6BC424073EE7744C5FB8DFDEB2D45" bold="true" box="[151,578,1039,1066]" italics="true" pageId="1" pageNumber="564">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14073EE77461BFB92FC142D45" author="Myers, A. A. &amp; Nithyanandan, M." box="[585,957,1040,1066]" pageId="1" pageNumber="564" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="828833DB4073EE7744C5FBB6FEEB2D21" box="[151,322,1076,1102]" pageId="1" pageNumber="564" type="description">
<paragraph id="CA2D60504073EE7744C5FBB6FEEB2D21" blockId="1.[151,957,1039,1102]" box="[151,322,1076,1102]" pageId="1" pageNumber="564">
(
<figureCitation id="52A97CD54073EE7744CDFBB6FEAA2D21" box="[159,259,1076,1102]" captionStart="FIGURE 1" captionStartId="2.[151,250,1880,1904]" captionTargetBox="[331,1257,342,1854]" captionTargetId="figure-93@2.[331,1257,342,1854]" captionTargetPageId="2" captionText="FIGURE 1. (A) Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan, 2016 male, (LFSC.ZRC-214): 10 mm, Shivrajpur; (B) Ceradomaera plumosa Ledoyer, 1973, male (LFSC.ZRC-215): 10 mm, Zanzmer; (C) Maera gujaratensis sp. nov., male paratype, (LFSC.ZRC-210): 7 mm, Gopnath; (D) Quadrimaera okha sp. nov., male paratype, (LFSC.ZRC-213): 4.5 mm, Shivrajpur, Gujarat, India." figureDoi="http://doi.org/10.5281/zenodo.12682132" httpUri="https://zenodo.org/record/12682132/files/figure.png" pageId="1" pageNumber="564">Figs. 1A</figureCitation>
,
<figureCitation id="52A97CD54073EE77455DFBB6FE932D21" box="[271,314,1076,1102]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[152,255,1943,1967]" captionStartId-1="4.[152,255,1943,1967]" captionStartId-2="5.[152,255,1753,1777]" captionTargetBox-0="[182,1408,188,1912]" captionTargetBox-1="[181,1407,188,1913]" captionTargetBox-2="[157,1429,187,1723]" captionTargetId-0="figure-19@3.[175,1412,181,1919]" captionTargetId-1="figure-22@4.[174,1414,181,1920]" captionTargetId-2="figure-19@5.[151,1436,181,1729]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan, 2016, male (LFSC.ZRC-214): 10 mm, Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India." captionText-1="FIGURE 3. Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan, 2016, male (LFSC.ZRC-214): 10 mm; female (LFSC. ZRC-214): 7 mm: Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India." captionText-2="FIGURE 4. Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan, 2016, male (LFSC.ZRC-214): 10 mm, Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India." figureDoi-0="http://doi.org/10.5281/zenodo.12682136" figureDoi-1="http://doi.org/10.5281/zenodo.12682138" figureDoi-2="http://doi.org/10.5281/zenodo.12682142" httpUri-0="https://zenodo.org/record/12682136/files/figure.png" httpUri-1="https://zenodo.org/record/12682138/files/figure.png" httpUri-2="https://zenodo.org/record/12682142/files/figure.png" pageId="1" pageNumber="564">2–4</figureCitation>
)
</paragraph>
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<paragraph id="CA2D60504073EE7744C5FBFAFC462DFF" blockId="1.[151,1007,1144,1169]" box="[151,1007,1144,1169]" pageId="1" pageNumber="564">
<taxonomicName id="0D921BD34073EE7744C5FBFAFC902DFF" ID-CoL="SG9J" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[151,825,1144,1169]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424073EE7744C5FBFAFDAB2DFF" box="[151,514,1144,1168]" italics="true" pageId="1" pageNumber="564">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14073EE77465BFBFBFC902DFF" author="Myers, A. A. &amp; Nithyanandan, M." box="[521,825,1144,1169]" pageId="1" pageNumber="564" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</taxonomicName>
, 421, figs. 16–18.
</paragraph>
</subSubSection>
<subSubSection id="828833DB4073EE7744C5FB3CFD782D92" pageId="1" pageNumber="564" type="materials_examined">
<paragraph id="CA2D60504073EE7744C5FB3CFD782D92" blockId="1.[151,1437,1214,2033]" pageId="1" pageNumber="564">
<emphasis id="F8E6BC424073EE7744C5FB3CFE232DB7" bold="true" box="[151,394,1214,1240]" pageId="1" pageNumber="564">Material Examined.</emphasis>
<specimenCount id="DC94ABD94073EE7745C7FB3DFE5B2DB6" box="[405,498,1215,1241]" count="4" pageId="1" pageNumber="564" type="male">4 males</specimenCount>
,
<quantity id="0D6ACDB54073EE774652FB3DFDDB2DB6" box="[512,626,1215,1241]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="1" pageNumber="564" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
;
<materialsCitation id="7AFA6A0D4073EE7746D3FB3DFD642D92" ID-GBIF-Occurrence="4903570310" collectionCode="LFSC, R" latitude="22.3312" longLatPrecision="7" longitude="68.9496" pageId="1" pageNumber="564" specimenCount="2" specimenCount-female="2">
<specimenCount id="DC94ABD94073EE7746D3FB3DFD5D2DB6" box="[641,756,1215,1241]" count="2" pageId="1" pageNumber="564" type="female">2 females</specimenCount>
,
<quantity id="0D6ACDB54073EE774750FB3DFCEE2DB6" box="[770,839,1215,1241]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="564" unit="mm" value="6.0">6 mm</quantity>
&amp;
<quantity id="0D6ACDB54073EE774723FB3DFC112DB6" box="[881,952,1215,1241]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="564" unit="mm" value="7.0">7 mm</quantity>
, (
<collectionCode id="AC83F8954073EE77479DFB3DFBBC2DB6" box="[975,1045,1215,1241]" pageId="1" pageNumber="564">LFSC</collectionCode>
.ZRC-214) Shivrajpur (
<geoCoordinate id="AFA606974073EE774173FB3DFA352DB6" box="[1313,1436,1215,1241]" degrees="22.3312" direction="north" orientation="latitude" pageId="1" pageNumber="564" precision="5" value="22.3312">22.3312 N</geoCoordinate>
<geoCoordinate id="AFA606974073EE7744C5FB61FEA42D92" box="[151,269,1251,1277]" degrees="68.9496" direction="east" orientation="longitude" pageId="1" pageNumber="564" precision="5" value="68.9496">68.9496 E</geoCoordinate>
), 27 February, 2022, coll. D.
<collectionCode id="AC83F8954073EE77461DFB61FDCC2D92" box="[591,613,1251,1277]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
. Thacker
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="828833DB4073EE704495FA84FB2D2A7A" lastPageId="6" lastPageNumber="569" pageId="1" pageNumber="564" type="description">
<paragraph id="CA2D60504073EE774495FA84FDC02C4E" blockId="1.[151,1437,1214,2033]" box="[199,617,1286,1313]" pageId="1" pageNumber="564">
<emphasis id="F8E6BC424073EE774495FA84FEF02C4F" bold="true" box="[199,345,1286,1312]" pageId="1" pageNumber="564">Description.</emphasis>
Based on male,
<quantity id="0D6ACDB54073EE774640FA85FDCD2C4E" box="[530,612,1287,1313]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="564" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="CA2D60504073EE774495FAA9FC9B2FC2" blockId="1.[151,1437,1214,2033]" pageId="1" pageNumber="564">
<emphasis id="F8E6BC424073EE774495FAA9FEA72C2A" bold="true" box="[199,270,1323,1349]" pageId="1" pageNumber="564">Head.</emphasis>
<emphasis id="F8E6BC424073EE774544FAA9FEE22C2B" box="[278,331,1323,1348]" italics="true" pageId="1" pageNumber="564">Eyes</emphasis>
large, ovate.
<emphasis id="F8E6BC424073EE7745B6FAA9FDFE2C2A" box="[484,599,1323,1349]" italics="true" pageId="1" pageNumber="564">Antenna 1</emphasis>
longer than antenna 2; flagellum longer than peduncle; peduncular article 1 0.92 times as long as article 2, article 3 one fifth of the length of article 2; accessory flagellum 7 articulate (fig. 2, A1).
<emphasis id="F8E6BC424073EE774482FAF1FEEB2CE2" box="[208,322,1395,1421]" italics="true" pageId="1" pageNumber="564">Antenna 2</emphasis>
half as long as antenna 1, peduncular article 4 1.23 times as long as article 5; flagellum 16 articulate (fig. 2, A2).
<emphasis id="F8E6BC424073EE774571FA15FE272CDE" box="[291,398,1431,1457]" italics="true" pageId="1" pageNumber="564">Mandible</emphasis>
accessory setal row well developed with 9 setae; palp 3 articulate, article 1 with acute distal tooth, article 3 0.43 times longer than article 2 (fig. 2, Md).
<emphasis id="F8E6BC424073EE77476DFA39FC3C2CBA" box="[831,917,1467,1493]" italics="true" pageId="1" pageNumber="564">Labium</emphasis>
distal margins of outer and inner lobes setose (fig. 2, L).
<emphasis id="F8E6BC424073EE77454BFA5DFE2E2C96" box="[281,391,1503,1529]" italics="true" pageId="1" pageNumber="564">Maxilla 1</emphasis>
inner plate triangular with inner margin setose; outer plate with 8 apical robust setae; palp article 2 apically setose (fig. 2, Mx1).
<emphasis id="F8E6BC424073EE77466BF981FD0D2F72" box="[569,676,1539,1565]" italics="true" pageId="1" pageNumber="564">Maxilla 2</emphasis>
inner plate broader than outer plate; outer plate apically setose, inner plate with dense setae on apical and inner margin, dorsomedial surface with an oblique row of setae (fig. 2, Mx2).
<emphasis id="F8E6BC424073EE7744C5F9C9FEB92F0A" box="[151,272,1611,1637]" italics="true" pageId="1" pageNumber="564">Maxilliped</emphasis>
inner plate apically truncate with 11 apical and 4 marginal plumose setae; outer plate with 7 long to medium robust setae on distal margin, 8 small robust setae on inner margin; palp 4 articulate, article 2 and 3 densely setose, article 4 with a long apical robust seta (fig. 2, Mxp).
</paragraph>
<paragraph id="CA2D60504073EE744495F935FD62282E" blockId="1.[151,1437,1214,2033]" lastBlockId="2.[151,1437,151,321]" lastPageId="2" lastPageNumber="565" pageId="1" pageNumber="564">
<emphasis id="F8E6BC424073EE774495F935FE8B2FBE" bold="true" box="[199,290,1719,1745]" pageId="1" pageNumber="564">Pereon.</emphasis>
<emphasis id="F8E6BC424073EE77457AF935FE132FBE" box="[296,442,1719,1745]" italics="true" pageId="1" pageNumber="564">Gnathopod 1</emphasis>
coxa anteroventral corner acute with few small setae, few simple setae on ventral margin; basis subequal to carpus, anterior margin bare, posterior margin sparsely setose, posterodistal corner with a tuft of simple setae, ischium smaller than merus, with a tuft of simple setae on posterior margin; merus subrectangular, anterodistal corner acutely produced, posterior margin setose; carpus anterior margin 2.41 times as long as merus, posterior margin densely setose, anterior margin with 3 stout setae and 1 thin simple setae, anterodistal corner with a tuft of simple setae; propodus 0.93 times as long as carpus, ovoidal, anterior margin with 5 small simple setae placed at somewhat equal distance, posterior margin densely setose, palm convex; dactylus subequal to palm (fig. 3, G1).
<emphasis id="F8E6BC424073EE774483F831FECD2EA2" box="[209,356,1971,1997]" italics="true" pageId="1" pageNumber="564">Gnathopod 2</emphasis>
asymmetrical.
<emphasis id="F8E6BC424073EE774642F831FD7C2EA2" box="[528,725,1971,1997]" italics="true" pageId="1" pageNumber="564">Left Gnathopod 2</emphasis>
larger than right gnathopod 2, coxa subquadrate with produced posterodistal corner, basis 0.6 times as long as propodus, anterior margin bare, posterior margin with few setae, anterodistal corner acutely produced; ischium smaller than merus; merus subrectangular with anterodistal corner acutely produced, carpus triangular, posterodistally setose; propodus large, subovoidal, palm transverse with 3 small to large shelves, one small subquadrate shelf near the hinge of dactylus followed by a V-shaped excavation, a large subquadrate shelf, a large U-shaped excavation and a medium sized triangular shelf, posterodistal corner of the palm concave; dactylus subequal to palm (fig. 3, MG2).
</paragraph>
<caption id="9EED30D84070EE7444C5F8DAFE0B2EB3" ID-DOI="http://doi.org/10.5281/zenodo.12682132" ID-Zenodo-Dep="12682132" httpUri="https://zenodo.org/record/12682132/files/figure.png" pageId="2" pageNumber="565" startId="2.[151,250,1880,1904]" targetBox="[331,1257,342,1854]" targetPageId="2" targetType="figure">
<paragraph id="CA2D60504070EE7444C5F8DAFE0B2EB3" blockId="2.[151,1437,1880,2012]" pageId="2" pageNumber="565">
<emphasis id="F8E6BC424070EE7444C5F8DAFEB62E1F" bold="true" box="[151,287,1880,1904]" pageId="2" pageNumber="565">FIGURE 1. (</emphasis>
A)
<taxonomicName id="0D921BD34070EE74456DF8DAFC712E1F" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[319,984,1880,1905]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424070EE74456DF8DAFD0E2E1F" box="[319,679,1880,1904]" italics="true" pageId="2" pageNumber="565">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14070EE7446FFF8DBFC712E1F" author="Myers, A. A. &amp; Nithyanandan, M." box="[685,984,1880,1905]" pageId="2" pageNumber="565" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</taxonomicName>
male, (LFSC.ZRC-214): 10 mm, Shivrajpur; (B)
<taxonomicName id="0D921BD34070EE744493F8FEFDE32EFA" authority="Ledoyer, 1973" authorityName="Ledoyer" authorityYear="1973" box="[193,586,1916,1941]" class="Malacostraca" family="Maeridae" genus="Ceradomaera" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="plumosa">
<emphasis id="F8E6BC424070EE744493F8FEFE022EFB" box="[193,427,1916,1940]" italics="true" pageId="2" pageNumber="565">Ceradomaera plumosa</emphasis>
<bibRefCitation id="AE031DA14070EE7445E1F8FEFDE32EFA" author="Ledoyer, M." box="[435,586,1916,1941]" pageId="2" pageNumber="565" pagination="51 - 94" refId="ref7655" refString="Ledoyer, M. (1973) Etude des Amphipodes Gammariens des biotopes de substrats sableux et sablo-vaseux de la region de Tulear et de Nosy-Be (Madagascar). Tethys Supplement, 5, 51 - 94." type="journal article" year="1973">Ledoyer, 1973</bibRefCitation>
</taxonomicName>
, male (LFSC.ZRC-215): 10 mm, Zanzmer; (C)
<taxonomicName id="0D921BD34070EE744068F8FFFAAB2EFB" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2024" box="[1082,1282,1917,1940]" class="Malacostraca" family="Maeridae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="gujaratensis" status="sp. nov.">
<emphasis id="F8E6BC424070EE744068F8FFFAAB2EFB" box="[1082,1282,1917,1940]" italics="true" pageId="2" pageNumber="565">Maera gujaratensis</emphasis>
</taxonomicName>
<emphasis id="F8E6BC424070EE744158F8FFFACA2EFA" bold="true" box="[1290,1379,1917,1941]" pageId="2" pageNumber="565">
<taxonomicNameLabel id="E3D501394070EE744158F8FFFAF62EFA" box="[1290,1375,1917,1941]" pageId="2" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male paratype, (LFSC.ZRC-210): 7 mm, Gopnath; (D)
<taxonomicName id="0D921BD34070EE7446FAF822FCC42ED7" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2024" box="[680,877,1952,1976]" class="Malacostraca" family="Maeridae" genus="Quadrimaera" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="okha" status="sp. nov.">
<emphasis id="F8E6BC424070EE7446FAF822FCC42ED7" box="[680,877,1952,1976]" italics="true" pageId="2" pageNumber="565">Quadrimaera okha</emphasis>
</taxonomicName>
<emphasis id="F8E6BC424070EE74472AF823FC7A2ED6" bold="true" box="[888,979,1953,1977]" pageId="2" pageNumber="565">
<taxonomicNameLabel id="E3D501394070EE74472AF823FC672ED6" box="[888,974,1953,1977]" pageId="2" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male paratype, (LFSC.ZRC-213): 4.5 mm, Shivrajpur, Gujarat, India.
</paragraph>
</caption>
<caption id="9EED30D84071EE7544CAF815FD8B2EBC" ID-DOI="http://doi.org/10.5281/zenodo.12682136" ID-Zenodo-Dep="12682136" httpUri="https://zenodo.org/record/12682136/files/figure.png" pageId="3" pageNumber="566" startId="3.[152,255,1943,1967]" targetBox="[182,1408,188,1912]" targetPageId="3" targetType="figure">
<paragraph id="CA2D60504071EE7544CAF815FD8B2EBC" blockId="3.[151,1437,1943,2004]" pageId="3" pageNumber="566">
<emphasis id="F8E6BC424071EE7544CAF815FEB32EC0" bold="true" box="[152,282,1943,1967]" pageId="3" pageNumber="566">FIGURE 2.</emphasis>
<taxonomicName id="0D921BD34071EE754570F815FC672EC0" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[290,974,1943,1968]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424071EE754570F815FD392EC0" box="[290,656,1943,1967]" italics="true" pageId="3" pageNumber="566">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14071EE7546CAF81AFC672EC0" author="Myers, A. A. &amp; Nithyanandan, M." box="[664,974,1943,1968]" pageId="3" pageNumber="566" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</taxonomicName>
, male (LFSC.ZRC-214): 10 mm, Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India.
</paragraph>
</caption>
<caption id="9EED30D84076EE7244CAF815FCFF2EBC" ID-DOI="http://doi.org/10.5281/zenodo.12682138" ID-Zenodo-Dep="12682138" httpUri="https://zenodo.org/record/12682138/files/figure.png" pageId="4" pageNumber="567" startId="4.[152,255,1943,1967]" targetBox="[181,1407,188,1913]" targetPageId="4" targetType="figure">
<paragraph id="CA2D60504076EE7244CAF815FCFF2EBC" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="567">
<emphasis id="F8E6BC424076EE7244CAF815FEBE2EC0" bold="true" box="[152,279,1943,1967]" pageId="4" pageNumber="567">FIGURE 3.</emphasis>
<taxonomicName id="0D921BD34076EE72454EF815FC1F2EC0" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[284,950,1943,1968]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424076EE72454EF815FD2D2EC0" box="[284,644,1943,1967]" italics="true" pageId="4" pageNumber="567">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14076EE7246DBF81AFC1F2EC0" author="Myers, A. A. &amp; Nithyanandan, M." box="[649,950,1943,1968]" pageId="4" pageNumber="567" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</taxonomicName>
, male (LFSC.ZRC-214): 10 mm; female (LFSC. ZRC-214): 7 mm: Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India.
</paragraph>
</caption>
<caption id="9EED30D84077EE7344CAF95BFD8B2E7A" ID-DOI="http://doi.org/10.5281/zenodo.12682142" ID-Zenodo-Dep="12682142" httpUri="https://zenodo.org/record/12682142/files/figure.png" pageId="5" pageNumber="568" startId="5.[152,255,1753,1777]" targetBox="[157,1429,187,1723]" targetPageId="5" targetType="figure">
<paragraph id="CA2D60504077EE7344CAF95BFD8B2E7A" blockId="5.[151,1437,1753,1813]" pageId="5" pageNumber="568">
<emphasis id="F8E6BC424077EE7344CAF95BFEB32F9E" bold="true" box="[152,282,1753,1777]" pageId="5" pageNumber="568">FIGURE 4.</emphasis>
<taxonomicName id="0D921BD34077EE734570F95BFC672F9E" authority="Myers &amp; Nithyanandan, 2016" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[290,974,1753,1777]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424077EE734570F95BFD392F9E" box="[290,656,1753,1777]" italics="true" pageId="5" pageNumber="568">Ceradocus (Denticeradocus) alama</emphasis>
<bibRefCitation id="AE031DA14077EE7346CAF95BFC672F9E" author="Myers, A. A. &amp; Nithyanandan, M." box="[664,974,1753,1777]" pageId="5" pageNumber="568" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
</taxonomicName>
, male (LFSC.ZRC-214): 10 mm, Shivrajpur, (22.3312 N 68.9496 E), Gujarat, India.
</paragraph>
</caption>
<paragraph id="CA2D60504077EE704495F8E5FA95280A" blockId="5.[151,1436,1895,2030]" lastBlockId="6.[151,1437,151,1149]" lastPageId="6" lastPageNumber="569" pageId="5" pageNumber="568">
<emphasis id="F8E6BC424077EE734495F8E5FEE52EEE" box="[199,332,1895,1921]" italics="true" pageId="5" pageNumber="568">Pereopod 3</emphasis>
longer than pereopod 4; coxa subquadrate, basis slender, both margins with long simple setae proximally while short robust setae medially and distally; merus 0.62 times as long as basis; carpus and propodus subequal; dactylus 0.42 times as long as propodus (fig. 4, P3).
<emphasis id="F8E6BC424077EE73470AF82DFC702EA6" box="[856,985,1967,1993]" italics="true" pageId="5" pageNumber="568">Pereopod 4</emphasis>
coxa broader than long, basis anteriorly slender, medially broader, merus 1.17 times longer than carpus; propodus 1.1 times longer than carpus; dactylus 0.4 times longer than propodus (fig. 4, P4).
<emphasis id="F8E6BC424074EE7046C5FF15FCB229DE" box="[663,795,151,177]" italics="true" pageId="6" pageNumber="569">Pereopod 5</emphasis>
basis subrectangular, anterior margin lightly undulated, posterior strongly undulated; merus 0.8 times as long as carpus; dactylus 0.33 times longer than propodus (fig. 4, P5).
<emphasis id="F8E6BC424074EE70449FFF5DFEE72997" box="[205,334,222,249]" italics="true" pageId="6" pageNumber="569">Pereopod 6</emphasis>
longer than pereopod 7; basis subrectangular with anterior margin lightly undulated and posterior margin strongly undulated; merus 0.79 times longer than carpus; carpus subequal to propodus; dactylus 0.3 times as long as propodus (fig. 4, P6).
<emphasis id="F8E6BC424074EE70458FFEA5FDF4282E" box="[477,605,295,321]" italics="true" pageId="6" pageNumber="569">Pereopod 7</emphasis>
basis similar as pereopod 6 but less undulated on both margins; merus 0.65 times as long as carpus; carpus subequal to propodus; dactylus 0.36 times as long as propodus (fig. 4, P7).
</paragraph>
<paragraph id="CA2D60504074EE704495FEEDFCA22BEA" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="569">
<emphasis id="F8E6BC424074EE704495FEEDFEB828E6" bold="true" box="[199,273,367,393]" pageId="6" pageNumber="569">Pleon.</emphasis>
<emphasis id="F8E6BC424074EE704544FEEDFE0D28E6" box="[278,420,367,393]" italics="true" pageId="6" pageNumber="569">Epimera 1–3</emphasis>
with dorsal, posteroventral and ventral teeth.
<emphasis id="F8E6BC424074EE7047C7FEEDFB8D28E6" box="[917,1060,367,393]" italics="true" pageId="6" pageNumber="569">Epimera 1–3</emphasis>
with 3, 4 and 5 spines respectively on ventral margin (fig. 2, Ep1–3).
<emphasis id="F8E6BC424074EE704673FE11FD6028C2" box="[545,713,403,429]" italics="true" pageId="6" pageNumber="569">Urosomite 1–2</emphasis>
with dorsal and posteroventral spines.
<emphasis id="F8E6BC424074EE7040DAFE11FB5C28C2" box="[1160,1269,403,429]" italics="true" pageId="6" pageNumber="569">Uropod 1</emphasis>
peduncle 1.17 times as long as outer ramus, with well-developed basofacial seta and long spur at distal margin; outer ramus slightly longer than inner ramus (fig. 4, U1).
<emphasis id="F8E6BC424074EE7046CDFE59FCA5289A" box="[671,780,475,501]" italics="true" pageId="6" pageNumber="569">Uropod 2</emphasis>
peduncle 0.74 times as long as outer ramus; inner ramus slightly shorter than outer ramus (fig. 4, U2).
<emphasis id="F8E6BC424074EE7046F1FE7DFCA62B76" box="[675,783,511,537]" italics="true" pageId="6" pageNumber="569">Uropod 3</emphasis>
peduncle 0.64 times as long as outer ramus; outer ramus slightly longer than inner ramus (fig. 4, U3). Telson deeply cleft, 1.21 times longer than wide, both halves pointed distally, each half with 5 setae on inner margin and 1 long and 1 short robust distal seta, distal long seta reaching between 0.5 to 0.6 times of the telsonic lobes (fig. 4, T).
</paragraph>
<paragraph id="CA2D60504074EE704495FD0DFD582BA2" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="569">
<emphasis id="F8E6BC424074EE704495FD0DFD012BC7" bold="true" box="[199,680,654,681]" pageId="6" pageNumber="569">Female (sexually dimorphic characters).</emphasis>
<emphasis id="F8E6BC424074EE7046FDFD0DFCEB2BC6" box="[687,834,655,681]" italics="true" pageId="6" pageNumber="569">Gnathopod 2</emphasis>
both similar in shape as male minor gnathopod 2, but one larger and more setose than another (fig. 3, FG2).
</paragraph>
<paragraph id="CA2D60504074EE704495FD54FB2D2A7A" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="569">
<emphasis id="F8E6BC424074EE704495FD54FDA12B9F" bold="true" box="[199,520,726,752]" pageId="6" pageNumber="569">Geographical distribution.</emphasis>
<materialsCitation id="7AFA6A0D4074EE704640FD55FE2F2A7A" ID-GBIF-Occurrence="4903570305" country="Kuwait" location="Kuwait" pageId="6" pageNumber="569" specimenCount="1">
The species was so far reported only from its
<typeStatus id="1529DEF24074EE704062FD55FBC92B9E" box="[1072,1120,727,753]" pageId="6" pageNumber="569">type</typeStatus>
locality,
<collectingCountry id="B28520C04074EE70409CFD55FA892B9E" box="[1230,1312,727,753]" name="Kuwait" pageId="6" pageNumber="569">Kuwait</collectingCountry>
(
<bibRefCitation id="AE031DA14074EE704161FD55FED72A7A" author="Myers, A. A. &amp; Nithyanandan, M." pageId="6" pageNumber="569" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan, 2016</bibRefCitation>
)
</materialsCitation>
, and here we have reported this species for the first time from
<collectingCountry id="B28520C04074EE704011FD79FB292A7A" box="[1091,1152,763,789]" name="India" pageId="6" pageNumber="569">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="828833DB4074EE704495FC9DFEC52D12" pageId="6" pageNumber="569" type="discussion">
<paragraph id="CA2D60504074EE704495FC9DFEC52D12" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="569">
<emphasis id="F8E6BC424074EE704495FC9DFE922A56" bold="true" box="[199,315,799,825]" pageId="6" pageNumber="569">Remarks.</emphasis>
The specimen examined here agrees with the description and illustrations of
<taxonomicName id="0D921BD34074EE7040C8FC9DFA9C2A56" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[1178,1333,799,825]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424074EE7040C8FC9DFA9C2A56" box="[1178,1333,799,825]" italics="true" pageId="6" pageNumber="569">C. (D.) alama</emphasis>
</taxonomicName>
given by
<bibRefCitation id="AE031DA14074EE7044C5FCC1FE5C2A32" author="Myers, A. A. &amp; Nithyanandan, M." box="[151,501,835,861]" pageId="6" pageNumber="569" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan (2016)</bibRefCitation>
. However, the specimen examined here shows some variations, such as: gnathopod 1 propodus convex (fig. 3, G1) (vs. concave in
<taxonomicName id="0D921BD34074EE7046FAFCE5FCED2AEE" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[680,836,871,897]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424074EE7046FAFCE5FCED2AEE" box="[680,836,871,897]" italics="true" pageId="6" pageNumber="569">C. (D.) alama</emphasis>
</taxonomicName>
cf.
<bibRefCitation id="AE031DA14074EE70473DFCE5FB132AEE" author="Myers, A. A. &amp; Nithyanandan, M." box="[879,1210,871,897]" pageId="6" pageNumber="569" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan 2016</bibRefCitation>
: fig. 15 MG1), cone gland reaching nearly half of the peduncular article 3 (fig. 2, A2) (vs. reaching more than half of the peduncular article
<quantity id="0D6ACDB54074EE7044B6FC2DFEA62AA6" box="[228,271,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="569" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="0D921BD34074EE704547FC2DFE192AA6" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[277,432,943,969]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424074EE704547FC2DFE192AA6" box="[277,432,943,969]" italics="true" pageId="6" pageNumber="569">C. (D.) alama</emphasis>
</taxonomicName>
cf.
<bibRefCitation id="AE031DA14074EE704588FC2DFC8C2AA6" author="Myers, A. A. &amp; Nithyanandan, M." box="[474,805,943,969]" pageId="6" pageNumber="569" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan 2016</bibRefCitation>
: fig. 16, Hd), pereopod 5 posterodistal corner is rounded (fig. 4, P5) (vs. posterodistal corner with a very strong, acute spine in
<taxonomicName id="0D921BD34074EE704793FC51FBC82A82" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[961,1121,979,1005]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424074EE704793FC51FBC82A82" box="[961,1121,979,1005]" italics="true" pageId="6" pageNumber="569">C. (D.) alama</emphasis>
</taxonomicName>
cf.
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: fig.18, P5) and Uropod 3 rami are having robust setae on both margins (fig. 5, U3) (vs. robust setae on only one margin in
<taxonomicName id="0D921BD34074EE704568FB99FE7E2D5A" authorityName="Myers &amp; Nithyanandan" authorityYear="2016" box="[314,471,1051,1077]" class="Malacostraca" family="Maeridae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="alama" subGenus="Denticeradocus">
<emphasis id="F8E6BC424074EE704568FB99FE7E2D5A" box="[314,471,1051,1077]" italics="true" pageId="6" pageNumber="569">C. (D.) alama</emphasis>
</taxonomicName>
cf.
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: fig. 17, U3). The specimen examined here (
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) is larger than the specimen of
<bibRefCitation id="AE031DA14074EE7045BEFBBDFCE12D36" author="Myers, A. A. &amp; Nithyanandan, M." box="[492,840,1087,1113]" pageId="6" pageNumber="569" pagination="401 - 429" refId="ref7749" refString="Myers, A. A. &amp; Nithyanandan, M. (2016) The Amphipoda of Sea City, Kuwait. The Senticaudata (Crustacea). Zootaxa, 4072 (4), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 4072.4.1" type="journal article" year="2016">Myers &amp; Nithyanandan (2016)</bibRefCitation>
(
<quantity id="0D6ACDB54074EE70470AFBBCFC1D2D36" box="[856,948,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="569" unit="mm" value="6.5">6.5 mm</quantity>
); thus, these variations could be because of the size difference.
</paragraph>
</subSubSection>
</treatment>
</document>
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