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<document ID-DOI="10.11646/zootaxa.5256.6.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7762456" ID-ZooBank="057285E1-92C9-4D76-921A-CE3C85D0D3B1" checkinTime="1679558471249" checkinUser="plazi" docAuthor="Neethling, J. A. &amp; Neethling, C." docDate="2023" docId="EB305F51B50FF66BFF1EF93EFADBFD9C" docLanguage="en" docName="zootaxa.5256.6.1.pdf" docOrigin="Zootaxa 5256 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.6.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnobisium octoflagellatum Beier 1947" docType="treatment" docVersion="3" lastPageNumber="529" masterDocId="17092729B518F677FF89FF81FFD1FF85" masterDocTitle="A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)" masterLastPageNumber="543" masterPageNumber="501" pageNumber="524" updateTime="1679591149964" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neethling, J. A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1702-9566</mods:nameIdentifier>
<mods:affiliation>Terrestrial Invertebrates Department, National Museum, P. O. Box 266, Bloemfontein 9300, South Africa. ja. neethling @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 1702 - 9566</mods:affiliation>
<mods:nameIdentifier type="email">ja.neethling@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neethling, C.</mods:namePart>
<mods:nameIdentifier type="ORCID">0009-0003-6492-4798</mods:nameIdentifier>
<mods:affiliation>Research Quality and Support, Clinglobal Limited, P. O. Box 11186, Bloemfontein 9321, South Africa. carmen. neethling @ clinglobal. com; https: // orcid. org / 0009 - 0003 - 6492 - 4798</mods:affiliation>
<mods:nameIdentifier type="email">carmen.neethling@clinglobal.com</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-23</mods:number>
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<mods:detail type="volume">
<mods:number>5256</mods:number>
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<mods:detail type="issue">
<mods:number>6</mods:number>
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<subSubSection box="[151,662,1727,1754]" pageId="23" pageNumber="524" type="nomenclature">
<paragraph blockId="23.[151,662,1727,1790]" box="[151,662,1727,1754]" pageId="23" pageNumber="524">
<heading bold="true" box="[151,662,1727,1754]" fontSize="11" level="1" pageId="23" pageNumber="524" reason="1">
<taxonomicName ID-CoL="3HQK3" authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[151,662,1727,1754]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis bold="true" box="[151,662,1727,1754]" pageId="23" pageNumber="524">
<emphasis bold="true" box="[151,517,1727,1754]" italics="true" pageId="23" pageNumber="524">Gymnobisium octoflagellatum</emphasis>
<bibRefCitation author="Beier, M." box="[524,662,1727,1754]" pageId="23" pageNumber="524" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,276,1764,1790]" pageId="23" pageNumber="524" type="description">
<paragraph blockId="23.[151,662,1727,1790]" box="[151,276,1764,1790]" pageId="23" pageNumber="524">
<figureCitation box="[151,276,1764,1790]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="24.[151,250,1932,1956]" captionStartId-1="26.[151,250,1005,1029]" captionStartId-2="27.[151,250,1691,1715]" captionTargetBox-0="[151,1435,273,1909]" captionTargetBox-1="[152,1433,383,981]" captionTargetBox-2="[171,1417,327,1649]" captionTargetId-0="figure-53@24.[151,1435,273,1909]" captionTargetId-1="figure-119@26.[151,1435,381,981]" captionTargetId-2="figure-83@27.[151,1436,309,1668]" captionTargetPageId-0="24" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURE 18. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947, Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." captionText-1="FIGURE 19. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947 right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." captionText-2="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7762514" figureDoi-1="http://doi.org/10.5281/zenodo.7762518" figureDoi-2="http://doi.org/10.5281/zenodo.7762522" httpUri-0="https://zenodo.org/record/7762514/files/figure.png" httpUri-1="https://zenodo.org/record/7762518/files/figure.png" httpUri-2="https://zenodo.org/record/7762522/files/figure.png" pageId="23" pageNumber="524">Figs 1820</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="524" type="reference_group">
<paragraph blockId="23.[151,1436,1832,1887]" pageId="23" pageNumber="524">
<treatmentCitationGroup pageId="23" pageNumber="524">
<treatmentCitation author="Beier, M." box="[151,678,1832,1856]" page="292" pageId="23" pageNumber="524" year="1947">
<taxonomicName ID-CoL="3HQK3" authority="Beier, 1947: 291 - 292" authorityName="Beier" authorityPageNumber="291 - 292" authorityYear="1947" box="[151,678,1832,1856]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis box="[151,455,1832,1856]" italics="true" pageId="23" pageNumber="524">Gymnobisium octoflagellatum</emphasis>
<bibRefCitation author="Beier, M." box="[460,678,1832,1856]" pageId="23" pageNumber="524" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947: 291292</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 4; Vitali-di Castri, 1970: 126; Harvey, 1991: 312;
<treatmentCitation author="Dippenaar-Schoeman, A. S. &amp; Harvey, M. S." page="92" pageId="23" pageNumber="524" year="2000">
<bibRefCitation author="Dippenaar-Schoeman, A. S. &amp; Harvey, M. S." pageId="23" pageNumber="524" pagination="89 - 102" refId="ref27783" refString="Dippenaar-Schoeman, A. S. &amp; Harvey, M. S. (2000) A check list of the pseudoscorpions of South Africa (Arachnida: Pseudoscorpiones). Koedoe, 43, 89 - 102. https: // doi. org / 10.4102 / koedoe. v 43 i 2.201" type="journal article" year="2000">Dippenaar-Schoeman &amp; Harvey, 2000: 92</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Harvey, M. S. &amp; Huey, J. &amp; Hillyer, M. &amp; McIntyre, E. &amp; Giribet, G." box="[414,643,1863,1887]" page="76" pageId="23" pageNumber="524" year="2016">
<bibRefCitation author="Harvey, M. S. &amp; Huey, J. &amp; Hillyer, M. &amp; McIntyre, E. &amp; Giribet, G." box="[414,643,1863,1887]" pageId="23" pageNumber="524" pagination="75 - 85" refId="ref28141" refString="Harvey, M. S., Huey, J., Hillyer, M., McIntyre, E. &amp; Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics, 30, 75 - 85. https: // doi. org / 10.1071 / IS 15044" type="journal article" year="2016">
Harvey
<emphasis box="[494,546,1863,1887]" italics="true" pageId="23" pageNumber="524">et al.</emphasis>
2016: 76
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="525" pageId="23" pageNumber="524" type="materials_examined">
<paragraph blockId="23.[151,1436,1928,1991]" pageId="23" pageNumber="524">
<materialsCitation collectionCode="SAMC" collectorName="R. F. Lawrance" country="South Africa" latitude="-34.066666" location="Viljoen's Pass" longLatPrecision="1201" longitude="19.066668" municipality="Grabouw" pageId="23" pageNumber="524" specimenCode="B8509" specimenCount="1" specimenCount-adult="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis bold="true" box="[151,269,1928,1954]" pageId="23" pageNumber="524">
<typeStatus box="[151,264,1928,1954]" pageId="23" pageNumber="524">Holotype</typeStatus>
:
</emphasis>
<specimenCount box="[276,361,1928,1954]" pageId="23" pageNumber="524" type="adult">1 Adult</specimenCount>
,
<collectingCountry box="[371,575,1928,1954]" name="South Africa" pageId="23" pageNumber="524">SOUTH AFRICA</collectingCountry>
,
<collectingRegion box="[584,741,1928,1954]" country="South Africa" name="Western Cape" pageId="23" pageNumber="524">Western Cape</collectingRegion>
,
<collectingMunicipality box="[752,857,1928,1954]" pageId="23" pageNumber="524">Grabouw</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB50FF660FCEAF809FBD1F827" box="[867,1024,1928,1954]" country="South Africa" latitude="-34.066666" longLatPrecision="1201" longitude="19.066668" municipality="Grabouw" name="Viljoen's Pass" pageId="23" pageNumber="524" stateProvince="Western Cape">Viljoens Pass</location>
[
<geoCoordinate box="[1037,1116,1929,1956]" degrees="34" direction="south" minutes="4" orientation="latitude" pageId="23" pageNumber="524" precision="925" value="-34.066666">34°4S</geoCoordinate>
,
<geoCoordinate box="[1127,1209,1929,1956]" degrees="19" direction="east" minutes="4" orientation="longitude" pageId="23" pageNumber="524" precision="925" value="19.066668">19°4E</geoCoordinate>
], leg.
<collectorName box="[1274,1436,1928,1954]" pageId="23" pageNumber="524">R.F. Lawrance</collectorName>
(
<collectionCode box="[159,240,1964,1990]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="23" pageNumber="524" type="Museum">SAMC</collectionCode>
<specimenCode box="[247,325,1964,1991]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="23" pageNumber="524" type="Museum">B8509</specimenCode>
), examined.
</materialsCitation>
</paragraph>
<paragraph blockId="24.[151,1436,150,249]" pageId="24" pageNumber="525">
<emphasis bold="true" box="[199,302,150,176]" pageId="24" pageNumber="525">Epitype:</emphasis>
<materialsCitation collectingDate="2016-04-04" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Grabouw" elevation="587" latitude="-33.066666" location="Hottentots Holland Nature Reserve" longLatPrecision="1207" longitude="19.05" municipality="Viljoen's Pass" pageId="24" pageNumber="525" specimenCode="NMBAP 00338" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
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,
<collectingCountry box="[347,553,151,177]" name="South Africa" pageId="24" pageNumber="525">SOUTH AFRICA</collectingCountry>
,
<collectingRegion box="[565,725,151,177]" country="South Africa" name="Western Cape" pageId="24" pageNumber="525">Western Cape</collectingRegion>
,
<collectingCounty box="[738,843,151,177]" pageId="24" pageNumber="525">Grabouw</collectingCounty>
,
<collectingMunicipality box="[855,1017,151,177]" pageId="24" pageNumber="525">Viljoens Pass</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB500F66FFB8FFF16FA49FF34" box="[1030,1432,151,177]" country="South Africa" county="Grabouw" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Viljoen's Pass" name="Hottentots Holland Nature Reserve" pageId="24" pageNumber="525" stateProvince="Western Cape">Hottentots Holland Nature Reserve</location>
,
<geoCoordinate box="[151,228,188,215]" degrees="33" direction="south" minutes="4" orientation="latitude" pageId="24" pageNumber="525" precision="925" value="-33.066666">33°4S</geoCoordinate>
,
<geoCoordinate box="[243,321,188,215]" degrees="19" direction="east" minutes="3" orientation="longitude" pageId="24" pageNumber="525" precision="925" value="19.05">19°3E</geoCoordinate>
,
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<quantity box="[336,399,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="24" pageNumber="525" unit="m" value="587.0">587m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB500F66FFE57FF3AFD78FF50" box="[478,681,187,213]" country="South Africa" county="Grabouw" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Viljoen's Pass" name="Mountain Fynbos" pageId="24" pageNumber="525" stateProvince="Western Cape">Mountain Fynbos</location>
,
<collectorName box="[695,746,187,213]" pageId="24" pageNumber="525">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[958,1123,187,213]" pageId="24" pageNumber="525">J.A. Neethling</collectorName>
&amp;
<collectorName box="[1164,1312,187,213]" pageId="24" pageNumber="525">C. Neethling</collectorName>
,
<date box="[1326,1436,187,213]" pageId="24" pageNumber="525" value="2016-04-04">
<collectingDate box="[1326,1436,187,213]" pageId="24" pageNumber="525" value="2016-04-04">4.IV.2016</collectingDate>
</date>
(
<specimenCode box="[159,341,223,249]" collectionCode="NMBAP" pageId="24" pageNumber="525">NMBAP 00338</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762514" ID-Zenodo-Dep="7762514" httpUri="https://zenodo.org/record/7762514/files/figure.png" pageId="24" pageNumber="525" startId="24.[151,250,1932,1956]" targetBox="[151,1435,273,1909]" targetPageId="24">
<paragraph blockId="24.[151,1436,1932,1992]" pageId="24" pageNumber="525">
<emphasis bold="true" box="[151,289,1932,1956]" pageId="24" pageNumber="525">FIGURE 18.</emphasis>
Digital microscope photographs of
<taxonomicName authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[659,1093,1932,1956]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis box="[659,966,1932,1956]" italics="true" pageId="24" pageNumber="525">Gymnobisium octoflagellatum</emphasis>
<bibRefCitation author="Beier, M." box="[973,1093,1932,1956]" pageId="24" pageNumber="525" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947</bibRefCitation>
</taxonomicName>
, Female (
<emphasis bold="true" box="[1195,1213,1932,1956]" pageId="24" pageNumber="525">A</emphasis>
,
<emphasis bold="true" box="[1227,1244,1932,1956]" pageId="24" pageNumber="525">B</emphasis>
) and Male (
<emphasis bold="true" box="[1371,1389,1932,1956]" pageId="24" pageNumber="525">C</emphasis>
,
<emphasis bold="true" box="[1403,1421,1932,1956]" pageId="24" pageNumber="525">D</emphasis>
).
<emphasis bold="true" box="[151,169,1968,1992]" pageId="24" pageNumber="525">A</emphasis>
,
<emphasis bold="true" box="[182,207,1968,1992]" pageId="24" pageNumber="525">C.</emphasis>
Dorsal view;
<emphasis bold="true" box="[350,367,1968,1992]" pageId="24" pageNumber="525">B</emphasis>
,
<emphasis bold="true" box="[379,404,1968,1992]" pageId="24" pageNumber="525">D.</emphasis>
Ventral view. Scale bar: 1.00 mm.
</paragraph>
</caption>
<subSubSection pageId="25" pageNumber="526" type="diagnosis">
<paragraph blockId="25.[151,1437,150,969]" pageId="25" pageNumber="526">
<emphasis bold="true" box="[199,322,150,176]" pageId="25" pageNumber="526">Diagnosis.</emphasis>
Moderately sized species (chela length
<specimenCount box="[758,779,151,177]" pageId="25" pageNumber="526" type="female"></specimenCount>
<quantity box="[784,947,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.605" metricValueMax="1.65" metricValueMin="1.56" pageId="25" pageNumber="526" unit="mm" value="1.605" valueMax="1.65" valueMin="1.56">1.561.65 mm</quantity>
,
<geoCoordinate box="[957,1031,151,179]" degrees="1.39" direction="west" orientation="longitude" pageId="25" pageNumber="526" precision="555" value="-1.39">
<quantity box="[983,1031,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.42" metricValueMin="1.39" pageId="25" pageNumber="526" unit="mm" value="1.4049999999999998" valueMax="1.42" valueMin="1.39">1.39</quantity>
</geoCoordinate>
1.42 mm), with chelae significantly more bulbous in females than in males (chela ratio
<specimenCount box="[721,742,187,213]" pageId="25" pageNumber="526" type="female"></specimenCount>
3.253.30 versus
<geoCoordinate box="[945,1022,187,215]" degrees="4.06" direction="west" orientation="longitude" pageId="25" pageNumber="526" precision="555" value="-4.06">♂ 4.06</geoCoordinate>
4.21 times longer than broad); fixed finger multi-toothed structure with eight strongly sclerotized teeth; cheliceral hand with five setae; galea with eight to ten rami; rallum with eight to nine blades. Differs from all congeners by having a minimum of eight rami on the galea versus a maximum of seven in all other species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="527" pageId="25" pageNumber="526" type="description">
<paragraph blockId="25.[151,1437,150,969]" box="[199,337,330,356]" pageId="25" pageNumber="526">
<emphasis bold="true" box="[199,337,330,356]" pageId="25" pageNumber="526">Description</emphasis>
</paragraph>
<paragraph blockId="25.[151,1437,150,969]" pageId="25" pageNumber="526">
<emphasis box="[199,319,367,393]" italics="true" pageId="25" pageNumber="526">Carapace:</emphasis>
Somewhat broader than long, subquadrate, overall smooth and without furrow. Very dark brown in females, appearing almost black, lighter in males. Two pairs of eyes located dorso-laterally at the front (
<figureCitation captionStart="FIGURE 18" captionStartId="24.[151,250,1932,1956]" captionTargetBox="[151,1435,273,1909]" captionTargetId="figure-53@24.[151,1435,273,1909]" captionTargetPageId="24" captionText="FIGURE 18. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947, Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762514" httpUri="https://zenodo.org/record/7762514/files/figure.png" pageId="25" pageNumber="526">Fig. 18A, C</figureCitation>
). Small seta located on the dorso-anterior margin of each of the posterior eyes. Female carapace with 20 setae, arranged 4: 4: 4: 2: 6. Male carapace possess mostly the same chaetotaxy as female, though some specimens varied by having 18 setae, arranged 4: 3: 4: 2: 5.
</paragraph>
<paragraph blockId="25.[151,1437,150,969]" pageId="25" pageNumber="526">
<emphasis box="[199,314,547,573]" italics="true" pageId="25" pageNumber="526">Abdomen:</emphasis>
Tergites and sternites smooth, undivided and uniseriate. Female abdominal tergites same colour as carapace. Male tergites same colour as carapace. Lateral and posterior edges of tergites IXI lighter in colour, more so in males, tergite XII uniformly lighter in colour. The tergal setae are located on the posterior edge of each tergite, within the lighter coloured band. Female sternites mostly same colour as tergites, slightly lighter, and strongly sclerotized. Female genital operculum tan to light brown in colour. Male sternites III tan; IIIVIII light brown, becoming slightly darker to posterior of abdomen; IXXI dark brown. Tergite and sternite XII brown (
<figureCitation box="[1288,1424,727,753]" captionStart="FIGURE 18" captionStartId="24.[151,250,1932,1956]" captionTargetBox="[151,1435,273,1909]" captionTargetId="figure-53@24.[151,1435,273,1909]" captionTargetPageId="24" captionText="FIGURE 18. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947, Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762514" httpUri="https://zenodo.org/record/7762514/files/figure.png" pageId="25" pageNumber="526">Fig. 18B, D</figureCitation>
). Sternites I and II are fused and hardly distinguishable. Female operculum with 11 setae, nine loosely clustered in two groups centrally at the posterior edge of sternite II, single seta situated lateral from each group, at the posterior margin of sternite. Males with 21 setae at the operculum cluster on sternite II, seven along the posterior margin of the sternite at the genital opening, and an additional 14 clustered loosely just anterior of these, centrally on sternite II. Male sternite III with 10 setae situated along the anterior margin, at the genital opening. Pleural membrane cream to light yellow in colour, faintly papillate.
</paragraph>
<paragraph blockId="25.[151,1436,1015,1077]" box="[151,1348,1015,1041]" pageId="25" pageNumber="526">Tergal chaetotaxy: 6(6): 8(8): 10(9): 12(9): 12(11): 12(11): 11(11): 10(10): 10(11): 9(9): 13(9): 2(2).</paragraph>
<paragraph blockId="25.[151,1436,1015,1077]" box="[151,1436,1050,1077]" pageId="25" pageNumber="526">Sternal chaetotaxy: 11(21):?(?): 18(24): 17(15): 15(14): 15(17): 17(15): 15(15): 12(15): 12(11): 13(9): 2(2).</paragraph>
<paragraph blockId="25.[151,1437,1123,2013]" pageId="25" pageNumber="526">
<emphasis box="[199,300,1123,1149]" italics="true" pageId="25" pageNumber="526">Pedipalp</emphasis>
: All segments smooth, orange-brown in females, slightly lighter in males. Trochanter cone-shaped and rounded anteriorly, curved slightly posteriorly. Base narrow, widening distally. Setae present on the anterior surface. Femur slender, 3.713.83 (
<specimenCount box="[462,480,1195,1221]" pageId="25" pageNumber="526" type="female"></specimenCount>
) to 4.054.10 (♂) times longer than wide, somewhat constricted basally, widening just distal of base. Deep disto-prolateral excavation present. Setae absent from base, though distributed evenly over rest of surface. Patella constricted and slightly angled at base, widening markedly distal at around a third segment length, 2.632.71 (
<specimenCount box="[363,381,1303,1329]" pageId="25" pageNumber="526" type="female"></specimenCount>
) to 2.923.00 (♂) times longer than wide. Several small lyriform fissures present on the dorsal surface, just distal of base. Two larger lyriform fissures present latero-dorsally from these, where the segment starts to widen. Shallow disto-prolateral excavation present. Setae sparse at base, more numerous on wide part of patella (
<figureCitation box="[159,262,1411,1437]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="25" pageNumber="526">Fig. 20B</figureCitation>
).
</paragraph>
<paragraph blockId="25.[151,1437,1123,2013]" pageId="25" pageNumber="526">
<emphasis box="[199,266,1447,1473]" italics="true" pageId="25" pageNumber="526">Chela</emphasis>
: In both sexes smooth and uniformly orange-brown, slightly lighter in males. Female hand larger and markedly more bulbous, strongly convex on the prolateral edge, slightly less so on the retrolateral edge (
<figureCitation captionStart="FIGURE 19" captionStartId="26.[151,250,1005,1029]" captionTargetBox="[152,1433,383,981]" captionTargetId="figure-119@26.[151,1435,381,981]" captionTargetPageId="26" captionText="FIGURE 19. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947 right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762518" httpUri="https://zenodo.org/record/7762518/files/figure.png" pageId="25" pageNumber="526">Fig. 19A, B</figureCitation>
). Male hand smaller, markedly less convex prolaterally (
<figureCitation box="[803,939,1519,1545]" captionStart="FIGURE 19" captionStartId="26.[151,250,1005,1029]" captionTargetBox="[152,1433,383,981]" captionTargetId="figure-119@26.[151,1435,381,981]" captionTargetPageId="26" captionText="FIGURE 19. Digital microscope photographs of Gymnobisium octoflagellatum Beier, 1947 right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762518" httpUri="https://zenodo.org/record/7762518/files/figure.png" pageId="25" pageNumber="526">Fig. 19C, D</figureCitation>
). Pedicel as long as wide in females, slightly longer than wide in males. Retrolateral condyle small but distinct and rounded in both sexes. Fingers narrow, same length as hand (without pedicel) to slightly longer than hand (without pedicel) in females. Male chelal fingers narrow and slightly shorter in length than hand (with pedicel). Fingers curved slightly prolaterally in both sexes. Fixed and movable chelal fingers with eight and four trichobothria respectively (
<figureCitation box="[1055,1159,1663,1689]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="25" pageNumber="526">Fig. 20A</figureCitation>
).
</paragraph>
<paragraph blockId="25.[151,1437,1123,2013]" lastBlockId="26.[151,1436,150,357]" lastPageId="26" lastPageNumber="527" pageId="25" pageNumber="526">
Terminal multi-toothed structure extends well beyond venom tooth of movable finger in both sexes, curving downward. Female structure slightly more curved than male. In both sexes the structure consists of a large terminal tooth preceded by a row of six smaller, highly sclerotized teeth that reduce in size away from the terminal tooth. This row is situated slightly prolaterally behind the terminal tooth. Located retrolaterally at the base of the smaller teeth is another larger tooth (
<figureCitation box="[474,579,1843,1869]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="25" pageNumber="526">Fig. 20D</figureCitation>
). Both fingers with rounded, juxtadentate teeth situated on flexible lamellae. Lamella on female fixed finger originating roughly one fifth finger length from base, originating between
<emphasis box="[1362,1387,1879,1905]" italics="true" pageId="25" pageNumber="526">sb</emphasis>
and
<emphasis box="[151,170,1915,1940]" italics="true" pageId="25" pageNumber="526">st</emphasis>
on movable finger. Lamella on male fixed finger originating slightly less than one third finger length from base, originating between
<emphasis box="[384,409,1951,1977]" italics="true" pageId="25" pageNumber="526">sb</emphasis>
and
<emphasis box="[466,485,1951,1976]" italics="true" pageId="25" pageNumber="526">st</emphasis>
on movable finger. Female fixed finger with 5052 lightly sclerotized, juxtadentate teeth, 1011 proximal to lamella base, together with eight strongly sclerotized teeth on distal structure. Male fixed finger with 5052 lightly sclerotized, juxtadentate teeth, 1112 proximal to lamella base, together with eight strongly sclerotized teeth on distal structure. Female movable finger with 4143 lightly sclerotized, juxtadentate teeth, nine proximal to lamella base, 3133 on lamella, and one small, lightly sclerotized tooth at the posterior base of the venom tooth. Male movable finger with 4549 lightly sclerotized, juxtadentate teeth, 1314 proximal to lamella base, 3134 on lamella, and one small, lightly sclerotized tooth at the posterior base of the venom tooth. Two thickened, spine-like seta present on distal end of movable finger in both sexes.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762518" ID-Zenodo-Dep="7762518" httpUri="https://zenodo.org/record/7762518/files/figure.png" pageId="26" pageNumber="527" startId="26.[151,250,1005,1029]" targetBox="[152,1433,383,981]" targetPageId="26">
<paragraph blockId="26.[151,1437,1005,1065]" pageId="26" pageNumber="527">
<emphasis bold="true" box="[151,290,1005,1029]" pageId="26" pageNumber="527">FIGURE 19.</emphasis>
Digital microscope photographs of
<taxonomicName authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[668,1106,1005,1029]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis box="[668,977,1005,1029]" italics="true" pageId="26" pageNumber="527">Gymnobisium octoflagellatum</emphasis>
<bibRefCitation author="Beier, M." box="[985,1106,1005,1029]" pageId="26" pageNumber="527" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947</bibRefCitation>
</taxonomicName>
right chela: Female (
<emphasis bold="true" box="[1332,1350,1005,1029]" pageId="26" pageNumber="527">A</emphasis>
,
<emphasis bold="true" box="[1365,1382,1005,1029]" pageId="26" pageNumber="527">B</emphasis>
) and Male (
<emphasis bold="true" box="[218,236,1041,1065]" pageId="26" pageNumber="527">C</emphasis>
,
<emphasis bold="true" box="[249,267,1041,1065]" pageId="26" pageNumber="527">D</emphasis>
).
<emphasis bold="true" box="[288,306,1041,1065]" pageId="26" pageNumber="527">A</emphasis>
,
<emphasis bold="true" box="[319,344,1041,1065]" pageId="26" pageNumber="527">C.</emphasis>
Retrolateral view;
<emphasis bold="true" box="[539,556,1041,1065]" pageId="26" pageNumber="527">B</emphasis>
,
<emphasis bold="true" box="[568,593,1041,1065]" pageId="26" pageNumber="527">D.</emphasis>
Dorsal view. Scale bar: 1.00 mm.
</paragraph>
</caption>
<paragraph blockId="26.[151,1437,1111,2038]" pageId="26" pageNumber="527">
<emphasis box="[199,309,1111,1137]" italics="true" pageId="26" pageNumber="527">Chelicera</emphasis>
: Hand with five long and acuminate setae,
<emphasis box="[804,827,1112,1137]" italics="true" pageId="26" pageNumber="527">es</emphasis>
shorter (
<figureCitation box="[929,1034,1111,1137]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="26" pageNumber="527">Fig. 20C</figureCitation>
). Fixed finger with 12 teeth in both sexes. Female moveable finger with eight teeth, male with seven. Female galea divided into two major rami, terminating in three smaller rami on one branch and five on the other (
<figureCitation box="[933,1034,1183,1209]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="26" pageNumber="527">Fig. 20E</figureCitation>
). Male galea divided into two major rami, each terminating in five smaller rami (
<figureCitation box="[646,746,1219,1245]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="26" pageNumber="527">Fig. 20F</figureCitation>
). Female rallum with eight long, acuminate blades (
<figureCitation box="[1326,1424,1219,1245]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="26" pageNumber="527">Fig 20G</figureCitation>
), male rallum with nine blades (
<figureCitation box="[492,597,1255,1281]" captionStart="FIGURE 20" captionStartId="27.[151,250,1691,1715]" captionTargetBox="[171,1417,327,1649]" captionTargetId="figure-83@27.[151,1436,309,1668]" captionTargetPageId="27" captionText="FIGURE 20. Gymnobisium octoflagellatum Beier, 1947. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Female chelal fingers, anterior retrolateral view; E. Female galea; F. Male galea; G. Female rallum; H. Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="26" pageNumber="527">Fig. 20H</figureCitation>
). Lamina exterior absent in both sexes.
</paragraph>
<paragraph blockId="26.[151,1437,1111,2038]" pageId="26" pageNumber="527">
<emphasis box="[199,377,1291,1317]" italics="true" pageId="26" pageNumber="527">Coxae and legs</emphasis>
: Pedipalpal coxae same colour, or slightly lighter than rest of pedipalps. Leg coxae yellowbrown in females, tan in males. Legs light brown to tan in both sexes. All legs diplotarsate with subterminal tarsal setae dentate distally; claws simple; arolium shorter than claws.
</paragraph>
<paragraph blockId="26.[151,1437,1111,2038]" pageId="26" pageNumber="527">
<emphasis bold="true" box="[199,455,1399,1425]" pageId="26" pageNumber="527">Measurements (mm):</emphasis>
Body length
<specimenCount box="[607,628,1400,1426]" pageId="26" pageNumber="527" type="female"></specimenCount>
1.88
<geoCoordinate box="[698,775,1399,1427]" degrees="1.99" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-1.99">1.99 ♂</geoCoordinate>
1.781.83; Carapace
<specimenCount box="[1019,1040,1400,1426]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.660.68 x 0.690.74 (0.920.96)
<geoCoordinate box="[151,231,1435,1463]" degrees="0.54" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.54">♂ 0.54</geoCoordinate>
0.57 x 0.600.61 (0.900.93); Chelicera
<specimenCount box="[712,733,1436,1462]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.510.54 x 0.280.29 (1.821.86)
<geoCoordinate box="[1149,1228,1435,1463]" degrees="0.40" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.4">♂ 0.40</geoCoordinate>
0.43 x 0.200.22 (1.952.00), movable finger length
<specimenCount box="[558,579,1472,1498]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.30
<geoCoordinate box="[651,730,1471,1499]" degrees="0.32" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.32">0.32 ♂</geoCoordinate>
0.240.25; Pedipalps: femur
<specimenCount box="[1069,1090,1472,1498]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.880.89 x 0.230.24 (3.71 3.83)
<geoCoordinate box="[217,294,1507,1535]" degrees="0.78" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.78">♂ 0.78</geoCoordinate>
0.81 x 0.190.20 (4.054.10), patella
<specimenCount box="[731,752,1508,1534]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.760.79 x 0.280.30 (2.632.71)
<geoCoordinate box="[1156,1233,1507,1535]" degrees="0.66" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.66">♂ 0.66</geoCoordinate>
0.70 x 0.220.24 (2.923.00), chela
<specimenCount box="[361,382,1544,1570]" pageId="26" pageNumber="527" type="female"></specimenCount>
1.561.65 x 0.480.50 (3.253.30)
<geoCoordinate box="[784,860,1543,1571]" degrees="1.39" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-1.39">♂ 1.39</geoCoordinate>
1.42 x 0.330.35 (4.064.21), hand
<specimenCount box="[1276,1297,1544,1570]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.840.89 x 0.480.50 (1.751.78)
<geoCoordinate box="[407,483,1579,1607]" degrees="0.71" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.71">♂ 0.71</geoCoordinate>
0.73 x 0.330.35 (2.082.15), movable finger length
<specimenCount box="[1092,1113,1580,1606]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.75
<geoCoordinate box="[1182,1259,1579,1607]" degrees="0.80" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.8">0.80 ♂</geoCoordinate>
0.680.69; Leg I: femur
<specimenCount box="[253,274,1616,1642]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.420.44 x 0.110.12 (3.673.82)
<geoCoordinate box="[688,767,1615,1643]" degrees="0.36" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.36">♂ 0.36</geoCoordinate>
0.38 x 0.08 (4.504.75), patella
<specimenCount box="[1151,1172,1616,1642]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.310.33 x 0.100.11 (3.003.10)
<geoCoordinate box="[289,366,1651,1679]" degrees="0.28" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.28">♂ 0.28</geoCoordinate>
x 0.09 (3.11), tibia
<specimenCount box="[587,608,1652,1678]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.380.39 x 0.08 (4.754.87)
<geoCoordinate box="[948,1024,1651,1679]" degrees="0.31" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.31">♂ 0.31</geoCoordinate>
0.34 x 0.07 (4.434.86), metatarsus
<specimenCount box="[151,172,1688,1714]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.190.20 x 0.07 (2.712.86)
<geoCoordinate box="[507,583,1687,1715]" degrees="0.17" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.17">♂ 0.17</geoCoordinate>
0.18 x 0.06 (2.832.57), tarsus
<specimenCount box="[941,962,1688,1714]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.260.27 x 0.06 (4.334.50)
<geoCoordinate box="[1298,1373,1687,1715]" degrees="0.22" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.22">♂ 0.22</geoCoordinate>
0.23 x 0.06 (3.673.83); Leg IV femoropatella
<specimenCount box="[634,655,1724,1750]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.800.84 x 0.180.19 (4.424.44)
<geoCoordinate box="[1069,1148,1723,1751]" degrees="0.69" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.69">♂ 0.69</geoCoordinate>
0.76 x 0.140.17 (4.47 4.93), tibia
<specimenCount box="[285,306,1760,1786]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.710.73 x 0.100.11 (6.647.10)
<geoCoordinate box="[721,800,1759,1787]" degrees="0.57" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.57">♂ 0.57</geoCoordinate>
0.62 x 0.090.11 (5.636.33), metatarsus
<specimenCount box="[1294,1315,1760,1786]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.250.27 x 0.08 (3.123.37)
<geoCoordinate box="[375,455,1795,1823]" degrees="0.22" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.22">♂ 0.22</geoCoordinate>
0.25 x 0.07 (3.143.57), tarsus
<specimenCount box="[834,855,1796,1822]" pageId="26" pageNumber="527" type="female"></specimenCount>
0.330.36 x 0.07 (4.715.14)
<geoCoordinate box="[1210,1290,1795,1823]" degrees="0.29" direction="west" orientation="longitude" pageId="26" pageNumber="527" precision="555" value="-0.29">♂ 0.29</geoCoordinate>
0.32 x 0.07 (4.144.57).
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="529" pageId="26" pageNumber="527" type="discussion">
<paragraph blockId="26.[151,1437,1111,2038]" lastBlockId="27.[151,1436,151,285]" lastPageId="27" lastPageNumber="528" pageId="26" pageNumber="527">
<emphasis bold="true" box="[199,315,1867,1893]" pageId="26" pageNumber="527">Remarks.</emphasis>
Upon examination of the
<typeStatus box="[625,726,1867,1893]" pageId="26" pageNumber="527">holotype</typeStatus>
, we found that the specimen had degraded to such an extent as to be all but unusable. The specimen was broken into parts, the main body separate from the two pedipalps. The trochanter, femur and patella of the left palp were missing. All sclerotized structures had become completely transparent, resulting in the entire body to be light yellow to cream in colour. Delicate structures such as the galea and rallum of the chelicerae were not present and due to the completely clear cuticula, neither the chaetotaxy nor the sex of the specimen could be determined. As such, a new epitype was designated and deposited in the arachnology collection of the National Museum in Bloemfontein. The sex of the
<typeStatus box="[923,1021,187,213]" pageId="27" pageNumber="528">holotype</typeStatus>
was never mentioned in the original description, with the specimen listed merely as “
<specimenCount box="[707,764,223,249]" pageId="27" pageNumber="528" type="generic">1 ad.</specimenCount>
”. By comparing the original description to newly collected material, the
<typeStatus box="[298,396,259,285]" pageId="27" pageNumber="528">holotype</typeStatus>
is most likely a small female specimen.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762522" ID-Zenodo-Dep="7762522" httpUri="https://zenodo.org/record/7762522/files/figure.png" pageId="27" pageNumber="528" startId="27.[151,250,1691,1715]" targetBox="[171,1417,327,1649]" targetPageId="27">
<paragraph blockId="27.[151,1437,1691,1787]" pageId="27" pageNumber="528">
<emphasis bold="true" box="[151,283,1691,1715]" pageId="27" pageNumber="528">FIGURE 20</emphasis>
.
<taxonomicName authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[298,737,1691,1715]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis box="[298,606,1691,1715]" italics="true" pageId="27" pageNumber="528">Gymnobisium octoflagellatum</emphasis>
<bibRefCitation author="Beier, M." box="[614,737,1691,1715]" pageId="27" pageNumber="528" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947</bibRefCitation>
</taxonomicName>
.
<emphasis bold="true" box="[748,773,1691,1715]" pageId="27" pageNumber="528">A.</emphasis>
Female right chela, retrolateral view;
<emphasis bold="true" box="[1172,1195,1691,1715]" pageId="27" pageNumber="528">B.</emphasis>
Female right pedipalp, dorsal view;
<emphasis bold="true" box="[280,305,1727,1751]" pageId="27" pageNumber="528">C.</emphasis>
Female left chelicera, dorsal view;
<emphasis bold="true" box="[663,687,1727,1751]" pageId="27" pageNumber="528">D.</emphasis>
Female chelal fingers, anterior retrolateral view;
<emphasis bold="true" box="[1184,1207,1727,1751]" pageId="27" pageNumber="528">E.</emphasis>
Female galea;
<emphasis bold="true" box="[1359,1378,1727,1751]" pageId="27" pageNumber="528">F.</emphasis>
Male galea;
<emphasis bold="true" box="[218,244,1763,1787]" pageId="27" pageNumber="528">G.</emphasis>
Female rallum;
<emphasis bold="true" box="[411,437,1763,1787]" pageId="27" pageNumber="528">H.</emphasis>
Male rallum. Scale bars: A, B: 0.50 mm; C, D: 0.20 mm; EH: 0.10 mm.
</paragraph>
</caption>
<paragraph blockId="27.[151,1437,1833,2040]" pageId="27" pageNumber="528">
According to the literature, no additional specimens other than the
<typeStatus box="[1001,1099,1833,1859]" pageId="27" pageNumber="528">holotype</typeStatus>
were ever collected, though three nymphal specimens, housed in the Royal Museum of Central Africa and misidentified as
<taxonomicName baseAuthorityName="Tullgren" baseAuthorityYear="1907" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="quadrispinosum">
<emphasis italics="true" pageId="27" pageNumber="528">Gymnobisium quadrispinosum</emphasis>
</taxonomicName>
, were collected from 19601962. No collection date was mentioned for the
<typeStatus box="[1176,1274,1905,1931]" pageId="27" pageNumber="528">holotype</typeStatus>
in the original description, and thus additional adult specimens of the species were then only recollected from the type locality in 2016, at least 69 years after the original
<typeStatus box="[601,703,1977,2003]" pageId="27" pageNumber="528">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="27.[151,1437,1833,2040]" lastBlockId="28.[151,1436,151,537]" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528">
The chela of
<taxonomicName authorityName="Beier" authorityYear="1947" box="[350,690,2013,2039]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="octoflagellatum">
<emphasis box="[350,690,2013,2039]" italics="true" pageId="27" pageNumber="528">Gymnobisium octoflagellatum</emphasis>
</taxonomicName>
are nearly identical to those of
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[1054,1300,2013,2040]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="prionotogladiatum" status="sp. nov.">
<emphasis box="[1054,1300,2013,2040]" italics="true" pageId="27" pageNumber="528">G. prionotogladiatum</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1308,1399,2013,2039]" pageId="27" pageNumber="528">
<taxonomicNameLabel box="[1308,1399,2013,2039]" pageId="27" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in regard to the overall chelal shape, the degree to which female chela differ from those of males, as well as the overall structure, and number of teeth, of the modified terminal tooth on the fixed chelal finger. That said, the species can easily be distinguished by the difference in size, colouration, trichobothrial arrangement, and number of teeth on the chelal fingers.
</paragraph>
<paragraph blockId="28.[151,1436,151,537]" pageId="28" pageNumber="529">
<emphasis bold="true" box="[199,566,294,321]" pageId="28" pageNumber="529">Additional material examined.</emphasis>
<materialsCitation box="[574,1096,294,321]" country="South Africa" location="Western Cape" pageId="28" pageNumber="529" specimenCount="1" stateProvince="Western Cape">
<emphasis bold="true" box="[574,788,294,320]" pageId="28" pageNumber="529">
<collectingCountry box="[574,788,294,320]" name="South Africa" pageId="28" pageNumber="529">SOUTH AFRICA</collectingCountry>
</emphasis>
:
<emphasis box="[803,960,295,321]" italics="true" pageId="28" pageNumber="529">
<collectingRegion box="[803,960,295,321]" country="South Africa" name="Western Cape" pageId="28" pageNumber="529">Western Cape</collectingRegion>
</emphasis>
: 2 Nymphs
</materialsCitation>
,
<materialsCitation collectingDate="1960-12" collectingDateMax="2016-04-04" collectingDateMin="1960-12" collectionCode="RMCA, NMBAP" collectorName="N. Leleup" country="South Africa" county="Town" elevation="587" latitude="-33.066666" location="Bats Cave" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" pageId="28" pageNumber="529" specimenCode="RMCA 127850, RMCA 127852, NMBAP 00322, NMBAP 00323" specimenCount="6" specimenCount-female="6" stateProvince="Western Cape">
<collectingRegion box="[1107,1165,295,321]" country="South Africa" name="Western Cape" pageId="28" pageNumber="529">Cape</collectingRegion>
<collectingCounty box="[1173,1239,295,321]" pageId="28" pageNumber="529">Town</collectingCounty>
,
<collectingMunicipality box="[1250,1432,295,321]" pageId="28" pageNumber="529">Table Mountain</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFF1EFECAFED8FEE0" box="[151,265,331,357]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Bats Cave" pageId="28" pageNumber="529" stateProvince="Western Cape">Bats Cave</location>
[
<geoCoordinate box="[277,370,332,359]" degrees="33" direction="south" minutes="58" orientation="latitude" pageId="28" pageNumber="529" precision="925" value="-33.966667">33°58S</geoCoordinate>
,
<geoCoordinate box="[381,477,332,359]" degrees="18" direction="east" minutes="25" orientation="longitude" pageId="28" pageNumber="529" precision="925" value="18.416666">18°25E</geoCoordinate>
],
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFE67FECAFDF0FEE0" box="[494,545,331,357]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Leaf" pageId="28" pageNumber="529" stateProvince="Western Cape">Leaf</location>
litter of
<taxonomicName box="[639,774,331,357]" class="Magnoliopsida" family="Sapotaceae" genus="Podocarpus" kingdom="Plantae" order="Ericales" pageId="28" pageNumber="529" phylum="Tracheophyta" rank="genus">
<emphasis box="[639,774,331,357]" italics="true" pageId="28" pageNumber="529">Podocarpus</emphasis>
</taxonomicName>
trees, leg.
<collectorName box="[893,1008,331,357]" pageId="28" pageNumber="529">N. Leleup</collectorName>
,
<date box="[1017,1119,331,357]" pageId="28" pageNumber="529" value="1960-12">
<collectingDate box="[1017,1119,331,357]" pageId="28" pageNumber="529" value="1960-12">XII.1960</collectingDate>
</date>
(
<specimenCode box="[1132,1308,330,357]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="28" pageNumber="529" type="Museum">RMCA 127850</specimenCode>
, previously ZA 43);
</materialsCitation>
<materialsCitation collectingDate="1960-12" collectingDateMax="2016-04-04" collectingDateMin="1960-12" collectionCode="RMCA, NMBAP" collectorName="N. Leleup" country="South Africa" county="Town" elevation="587" latitude="-33.066666" location="Bats Cave" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" pageId="28" pageNumber="529" specimenCode="RMCA 127850, RMCA 127852, NMBAP 00322, NMBAP 00323" specimenCount="6" specimenCount-female="6" stateProvince="Western Cape">
1
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFE99FEEEFE4BFE0C" box="[272,410,367,393]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Clanwilliam" pageId="28" pageNumber="529" stateProvince="Western Cape">Clanwilliam</location>
[
<geoCoordinate box="[426,520,368,395]" degrees="32" direction="south" minutes="20" orientation="latitude" pageId="28" pageNumber="529" precision="925" value="-32.333332">32°20S</geoCoordinate>
,
<geoCoordinate box="[533,613,368,395]" degrees="19" direction="east" minutes="5" orientation="longitude" pageId="28" pageNumber="529" precision="925" value="19.083334">19°5E</geoCoordinate>
],
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFDF3FEEEFD7CFE0C" box="[634,685,367,393]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Leaf" pageId="28" pageNumber="529" stateProvince="Western Cape">Leaf</location>
litter under large rock, leg.
<collectorName box="[1007,1124,367,393]" pageId="28" pageNumber="529">N. Leleup</collectorName>
,
<date box="[1136,1237,367,393]" pageId="28" pageNumber="529" value="1962-07">
<collectingDate box="[1136,1237,367,393]" pageId="28" pageNumber="529" value="1962-07">VII.1962</collectingDate>
</date>
(
<specimenCode box="[1254,1432,366,393]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="28" pageNumber="529" type="Museum">RMCA 127852</specimenCode>
, previously ZA 99);
</materialsCitation>
<materialsCitation collectingDate="1960-12" collectingDateMax="2016-04-04" collectingDateMin="1960-12" collectionCode="RMCA, NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Town" elevation="587" latitude="-33.066666" location="Bats Cave" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" pageId="28" pageNumber="529" specimenCode="RMCA 127850, RMCA 127852, NMBAP 00322, NMBAP 00323" specimenCount="6" specimenCount-female="6" stateProvince="Western Cape">
13
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFE1EFE12FDFBFE28" box="[407,554,403,429]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Tritonymphs" pageId="28" pageNumber="529" stateProvince="Western Cape">Tritonymphs</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFDBCFE12FD4FFE28" box="[565,670,403,429]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Grabouw" pageId="28" pageNumber="529" stateProvince="Western Cape">Grabouw</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFD20FE12FC9BFE28" box="[681,842,403,429]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Viljoen's Pass" pageId="28" pageNumber="529" stateProvince="Western Cape">Viljoens Pass</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFCDFFE12FB35FE28" box="[854,1252,403,429]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Hottentots Holland Nature Reserve" pageId="28" pageNumber="529" stateProvince="Western Cape">Hottentots Holland Nature Reserve</location>
,
<geoCoordinate box="[1263,1341,404,431]" degrees="33" direction="south" minutes="4" orientation="latitude" pageId="28" pageNumber="529" precision="925" value="-33.066666">33°4S</geoCoordinate>
,
<geoCoordinate box="[1353,1432,404,431]" degrees="19" direction="east" minutes="3" orientation="longitude" pageId="28" pageNumber="529" precision="925" value="19.05">19°3E</geoCoordinate>
,
<elevation box="[151,274,438,465]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="28" pageNumber="529" unit="m" value="587.0">
<quantity box="[151,215,438,465]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="28" pageNumber="529" unit="m" value="587.0">587m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFE97FE36FE34FE54" box="[286,485,439,465]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Mountain Fynbos" pageId="28" pageNumber="529" stateProvince="Western Cape">Mountain Fynbos</location>
, Leaf litter sifting, leg.
<collectorName box="[743,904,439,465]" pageId="28" pageNumber="529">J.A. Neethling</collectorName>
&amp;
<collectorName box="[938,1082,439,465]" pageId="28" pageNumber="529">C. Neethling</collectorName>
,
<date box="[1093,1230,438,465]" pageId="28" pageNumber="529" value="2015-12-13">
<collectingDate box="[1093,1230,438,465]" pageId="28" pageNumber="529" value="2015-12-13">13.XII.2015</collectingDate>
</date>
(
<specimenCode box="[1243,1424,439,465]" collectionCode="NMBAP" pageId="28" pageNumber="529">NMBAP 00322</specimenCode>
);
</materialsCitation>
<materialsCitation collectingDate="1960-12" collectingDateMax="2016-04-04" collectingDateMin="1960-12" collectionCode="RMCA, NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Town" elevation="587" latitude="-33.066666" location="Bats Cave" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" pageId="28" pageNumber="529" specimenCode="RMCA 127850, RMCA 127852, NMBAP 00322, NMBAP 00323" specimenCount="6" specimenCount-female="6" stateProvince="Western Cape">
<specimenCount box="[151,187,475,501]" pageId="28" pageNumber="529" type="female">6♀</specimenCount>
, 6♂, 1 Tritonymph, Grabouw, Viljoens Pass,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFD4EFE5AFB89FE70" box="[711,1112,475,501]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Hottentots Holland Nature Reserve" pageId="28" pageNumber="529" stateProvince="Western Cape">Hottentots Holland Nature Reserve</location>
,
<geoCoordinate box="[1124,1201,476,503]" degrees="33" direction="south" minutes="4" orientation="latitude" pageId="28" pageNumber="529" precision="925" value="-33.066666">33°4S</geoCoordinate>
,
<geoCoordinate box="[1214,1293,476,503]" degrees="19" direction="east" minutes="3" orientation="longitude" pageId="28" pageNumber="529" precision="925" value="19.05">19°3E</geoCoordinate>
,
<elevation box="[1305,1431,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="28" pageNumber="529" unit="m" value="587.0">
<quantity box="[1305,1369,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="28" pageNumber="529" unit="m" value="587.0">587m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B50FF66BFF1EF93EFADBFD9C:6646B89CB504F66BFF1EFE7EFEB1FD9C" box="[151,352,511,537]" country="South Africa" county="Town" latitude="-33.066666" longLatPrecision="1207" longitude="19.05" municipality="Table Mountain" name="Mountain Fynbos" pageId="28" pageNumber="529" stateProvince="Western Cape">Mountain Fynbos</location>
, Leaf litter sifting, leg.
<collectorName box="[616,778,511,537]" pageId="28" pageNumber="529">J.A. Neethling</collectorName>
&amp;
<collectorName box="[814,960,511,537]" pageId="28" pageNumber="529">C. Neethling</collectorName>
,
<date box="[971,1081,511,537]" pageId="28" pageNumber="529" value="2016-04-04">
<collectingDate box="[971,1081,511,537]" pageId="28" pageNumber="529" value="2016-04-04">4.IV.2016</collectingDate>
</date>
(
<specimenCode box="[1095,1278,511,537]" collectionCode="NMBAP" pageId="28" pageNumber="529">NMBAP 00323</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>