293 lines
37 KiB
XML
293 lines
37 KiB
XML
<document id="9EF79EC884E7CA922A5BFE58E7AAD366" ID-DOI="10.11646/zootaxa.5380.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10254532" ID-ZooBank="76CC750C-19F2-4308-94B2-3DB61BC13E3F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701678877052" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="03A787C8E914420DCF964EAD11AFA4C1" docLanguage="en" docName="zootaxa.5380.5.2.pdf" docOrigin="Zootaxa 5380 (5)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5380.5.2/52399" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omaliopsis obliquesignata Shavrin 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="455" masterDocId="FF9EFFB0E91C4204CF0148111060A319" masterDocTitle="Four new species of the genus Omaliopsis Jeannel, 1940 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China" masterLastPageNumber="460" masterPageNumber="446" pageNumber="454" updateTime="1701680077583" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CF3940A1796D0EBB63504F264A016373">Four new species of the genus Omaliopsis Jeannel, 1940 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China</mods:title>
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<mods:name id="CA1A8317FAEB4AC0C414983A00FC5017" type="personal">
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<mods:namePart id="0ACB2BF0803866F1F13A4457503C57E9">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="B40DE760067EECB0021CDA2FDB44C23C">Zootaxa</mods:title>
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<mods:part id="A213E506B0BAD7C9AC5E31847C56DDDF">
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<mods:date id="DE49898135ACB7BC64F0FBDCED493812">2023</mods:date>
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<mods:number id="E3B145680EE4BF6B7A368643A9C7C6E4">2023-12-04</mods:number>
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<mods:identifier id="EDFEE31A53F0CEC594A305C8C60D62D2" type="DOI">10.11646/zootaxa.5380.5.2</mods:identifier>
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<treatment id="03A787C8E914420DCF964EAD11AFA4C1" ID-DOI="http://doi.org/10.5281/zenodo.10254538" ID-Zenodo-Dep="10254538" LSID="urn:lsid:plazi:treatment:03A787C8E914420DCF964EAD11AFA4C1" httpUri="http://treatment.plazi.org/id/03A787C8E914420DCF964EAD11AFA4C1" lastPageId="9" lastPageNumber="455" pageId="8" pageNumber="454">
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<subSubSection id="C3146555E914420CCF964EAD1241A5CE" box="[151,545,1724,1751]" pageId="8" pageNumber="454" type="nomenclature">
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<paragraph id="8BB136DEE914420CCF964EAD1241A5CE" blockId="8.[151,545,1724,1787]" box="[151,545,1724,1751]" pageId="8" pageNumber="454">
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<heading id="D0F981B2E914420CCF964EAD1241A5CE" bold="true" box="[151,545,1724,1751]" fontSize="11" level="1" pageId="8" pageNumber="454" reason="1">
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<emphasis id="B97AEACCE914420CCF964EAD1241A5CE" bold="true" box="[151,545,1724,1751]" pageId="8" pageNumber="454">
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<taxonomicName id="4C0E4D5DE914420CCF964EAD11B8A5CE" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,472,1724,1751]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="454" phylum="Arthropoda" rank="species" species="obliquesignata" status="sp. nov.">
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<emphasis id="B97AEACCE914420CCF964EAD11B8A5CE" bold="true" box="[151,472,1724,1751]" italics="true" pageId="8" pageNumber="454">Omaliopsis obliquesignata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957B7E914420CCEDE4EAC1241A5CE" box="[479,545,1725,1751]" pageId="8" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3146555E914420CCF964EF1110DA5E2" box="[151,365,1760,1787]" pageId="8" pageNumber="454" type="description">
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<paragraph id="8BB136DEE914420CCF964EF1110DA5E2" blockId="8.[151,545,1724,1787]" box="[151,365,1760,1787]" pageId="8" pageNumber="454">
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(
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<figureCitation id="13352A5BE914420CCF9E4EF11087A5E2" box="[159,231,1760,1787]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[228,1359,192,1905]" captionTargetId="figure-21@3.[216,1371,179,1917]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Omaliopsis: 1—O. fraterna (holotype), 2—O. hlavaci (holotype), 3—O. obliquesignata (holotype), 4—O. smetanai (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10254536" httpUri="https://zenodo.org/record/10254536/files/figure.png" pageId="8" pageNumber="454">Figs 3</figureCitation>
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,
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<figureCitation id="13352A5BE914420CCFF54EF01172A5E2" box="[244,274,1761,1787]" captionStart="FIGURE 17" captionStartId="6.[151,250,1548,1572]" captionTargetBox="[151,1436,453,1525]" captionTargetId="figure-120@6.[151,1436,453,1525]" captionTargetPageId="6" captionText="FIGURE 17. Distribution of Omaliopsis in China: O. fraterna (circles), O. hlavaci (triangles), O. obliquesignata (black star), O. smetanai (diamonds)." figureDoi="http://doi.org/10.5281/zenodo.10254548" httpUri="https://zenodo.org/record/10254548/files/figure.png" pageId="8" pageNumber="454">17</figureCitation>
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,
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<figureCitation id="13352A5BE914420CCE1F4EF11105A5E2" box="[286,357,1760,1787]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="8" pageNumber="454">25–31</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146555E914420DCF964F3812FCA3CD" lastPageId="9" lastPageNumber="455" pageId="8" pageNumber="454" type="materials_examined">
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<paragraph id="8BB136DEE914420CCF964F381223A4CB" blockId="8.[151,1437,1832,2039]" pageId="8" pageNumber="454">
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<emphasis id="B97AEACCE914420CCF964F381128A45B" bold="true" box="[151,328,1832,1859]" pageId="8" pageNumber="454">Type material.</emphasis>
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<typeStatus id="54B5887CE914420CCE504F3911D9A45B" box="[337,441,1832,1858]" pageId="8" pageNumber="454" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D08FD57E914420CCEC24F3811B8A45A" box="[451,472,1833,1859]" count="1" pageId="8" pageNumber="454" type="male">♂</specimenCount>
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(
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<figureCitation id="13352A5BE914420CCEEB4F391251A45A" box="[490,561,1832,1859]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[228,1359,192,1905]" captionTargetId="figure-21@3.[216,1371,179,1917]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Omaliopsis: 1—O. fraterna (holotype), 2—O. hlavaci (holotype), 3—O. obliquesignata (holotype), 4—O. smetanai (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10254536" httpUri="https://zenodo.org/record/10254536/files/figure.png" pageId="8" pageNumber="454">Fig. 3</figureCitation>
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): ‘
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<collectingCountry id="F319764EE914420CCD4F4F3812D0A45A" box="[590,688,1833,1859]" name="China" pageId="8" pageNumber="454">CHINA</collectingCountry>
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: S-Shaanxi (Qinling Shan) | pass on rd. Zhouzhi – Foping, |
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<quantity id="4CF69B3BE914420CCA734F3910DBA47F" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="454" unit="km" value="105.0">105 km</quantity>
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SW Xi`an, N-slope, |
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<quantity id="4CF69B3BE914420CCEBF4F5C1277A47E" box="[446,535,1869,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="8" pageNumber="454" unit="m" value="1990.0">1990 m</quantity>
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,
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<geoCoordinate id="EE3A5019E914420CCD244F5C12EFA47E" box="[549,655,1869,1895]" degrees="33" direction="north" minutes="44" orientation="latitude" pageId="8" pageNumber="454" precision="925" value="33.733334">33°44’ N</geoCoordinate>
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,
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<geoCoordinate id="EE3A5019E914420CCD9D4F5D1368A47E" box="[668,776,1868,1895]" degrees="107" direction="east" minutes="59" orientation="longitude" pageId="8" pageNumber="454" precision="925" value="107.98333">107°59’E</geoCoordinate>
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| leg. M. Schülke [C01-01]’ <printed>, ‘2./
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<date id="FFB0101EE914420CCBF84F5C15EAA47E" box="[1273,1418,1869,1895]" pageId="8" pageNumber="454" value="2001-07-04">4. VII. 2001</date>
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, | small creek valley, mixed | deciduous forest, bamboo, | small meadows, dead wood, | mushrooms (sifted) [C01-01]’ <printed>, ‘Sammlung | M. Schülke | Berlin’ <printed>, ‘
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<typeStatus id="54B5887CE914420CCC0C4F8413C6A4B6" box="[781,934,1941,1967]" pageId="8" pageNumber="454" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C0E4D5DE914420CCCB64F851457A4B7" authorityName="Jeannel" authorityYear="1940" box="[951,1079,1940,1966]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="454" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEACCE914420CCCB64F851457A4B7" box="[951,1079,1940,1966]" italics="true" pageId="8" pageNumber="454">Omaliopsis</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4C0E4D5DE914420CCB484F85148FA4B7" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1097,1263,1940,1966]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="454" phylum="Arthropoda" rank="species" species="obliquesignata" status="sp. nov.">
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<emphasis id="B97AEACCE914420CCB484F85148FA4B7" box="[1097,1263,1940,1966]" italics="true" pageId="8" pageNumber="454">obliquesignata</emphasis>
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</taxonomicName>
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<emphasis id="B97AEACCE914420CCBF44F841554A4B6" bold="true" box="[1269,1332,1941,1967]" pageId="8" pageNumber="454">
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<taxonomicNameLabel id="A24957B7E914420CCBF44F841554A4B6" box="[1269,1332,1941,1967]" pageId="8" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ <red, printed> (cSch).
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</paragraph>
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<paragraph id="8BB136DEE914420DCFC64FCC12FCA3CD" blockId="8.[151,1437,1832,2039]" lastBlockId="9.[151,1437,151,2008]" lastPageId="9" lastPageNumber="455" pageId="8" pageNumber="454">
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<typeStatus id="54B5887CE914420CCFC64FCC1154A4EE" box="[199,308,2013,2039]" pageId="8" pageNumber="454" type="paratype">Paratypes</typeStatus>
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(
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<specimenCount id="9D08FD57E914420CCE464FCC11EDA4EE" box="[327,397,2013,2039]" count="9" pageId="8" pageNumber="454" type="male">9 ♂♂</specimenCount>
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,
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<specimenCount id="9D08FD57E914420CCE9D4FCC11BFA4EE" box="[412,479,2013,2039]" count="7" pageId="8" pageNumber="454" type="female">7 ♀♀</specimenCount>
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):
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<specimenCount id="9D08FD57E914420CCEFB4FCC125DA4EE" box="[506,573,2013,2039]" count="9" pageId="8" pageNumber="454" type="male">9 ♂♂</specimenCount>
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(
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<specimenCount id="9D08FD57E914420CCD514FCC12BEA4EF" box="[592,734,2012,2039]" count="2" pageId="8" pageNumber="454" type="generic">2 specimens</specimenCount>
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dissected),
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<specimenCount id="9D08FD57E914420CCC6B4FCC13CDA4EE" box="[874,941,2013,2039]" count="7" pageId="8" pageNumber="454" type="female">7 ♀♀</specimenCount>
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(
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<specimenCount id="9D08FD57E914420CCCC14FCC142EA4EF" box="[960,1102,2012,2039]" count="2" pageId="8" pageNumber="454" type="generic">2 specimens</specimenCount>
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dissected):
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<materialsCitation id="3B663C83E914420DCBDA4FCC121DA3A8" lastPageId="9" lastPageNumber="455" pageId="8" pageNumber="454" specimenCount="1" typeStatus="paratype">sama data as the holotype, with additional red printed label</materialsCitation>
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: ‘
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<typeStatus id="54B5887CE915420DCD8C4886137DA3A8" box="[653,797,151,177]" pageId="9" pageNumber="455" type="paratype">PARATYPE</typeStatus>
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|
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<taxonomicName id="4C0E4D5DE915420DCC37488613D6A3A8" authorityName="Jeannel" authorityYear="1940" box="[822,950,151,177]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEACCE915420DCC37488613D6A3A8" box="[822,950,151,177]" italics="true" pageId="9" pageNumber="455">Omaliopsis</emphasis>
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</taxonomicName>
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||
|
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||
<taxonomicName id="4C0E4D5DE915420DCCCC48861413A3A8" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[973,1139,151,177]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="obliquesignata" status="sp. nov.">
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||
<emphasis id="B97AEACCE915420DCCCC48861413A3A8" box="[973,1139,151,177]" italics="true" pageId="9" pageNumber="455">obliquesignata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCB7C488614DEA3A8" bold="true" box="[1149,1214,151,177]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCB7C488614DEA3A8" box="[1149,1214,151,177]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.V. des. 2023’ (
|
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<specimenCount id="9D08FD57E915420DCFE748AB1148A3CD" box="[230,296,186,212]" count="2" pageId="9" pageNumber="455" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD57E915420DCE3548AB1115A3CD" box="[308,373,186,212]" count="2" pageId="9" pageNumber="455" type="female">2 ♀♀</specimenCount>
|
||
: cSh;
|
||
<specimenCount id="9D08FD57E915420DCEBB48AB119BA3CD" box="[442,507,186,212]" count="7" pageId="9" pageNumber="455" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD57E915420DCD0648AB1228A3CD" box="[519,584,186,213]" count="5" pageId="9" pageNumber="455" type="female">5 ♀♀</specimenCount>
|
||
: cSch).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146555E915420DCFC648CC1315A41D" pageId="9" pageNumber="455" type="description">
|
||
<paragraph id="8BB136DEE915420DCFC648CC10B9A278" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
<emphasis id="B97AEACCE915420DCFC648CC1139A3EE" bold="true" box="[199,345,221,247]" pageId="9" pageNumber="455">Description.</emphasis>
|
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Measurements (n=17): HW: 0.38–0.42; HL: 0.24–0.30; OL: 0.09–0.10; TL: 0.04–0.05; AL (
|
||
<typeStatus id="54B5887CE915420DCF9E49111168A203" box="[159,264,256,282]" pageId="9" pageNumber="455" type="holotype">holotype</typeStatus>
|
||
): 0.67; PL: 0.31–0.35; PW: 0.54–0.55; ESL: 0.64–0.67; EW: 0.72–0.73; MTbL (
|
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<typeStatus id="54B5887CE915420DCB9D49111565A203" box="[1180,1285,256,282]" pageId="9" pageNumber="455" type="holotype">holotype</typeStatus>
|
||
): 0.32; MTrL (
|
||
<typeStatus id="54B5887CE915420DCF9E49321168A224" box="[159,264,291,317]" pageId="9" pageNumber="455" type="holotype">holotype</typeStatus>
|
||
): 0.18 (MTrL 1–4: 0.06; MTrL 5: 0.12); AW: 0.64–0.67; AedL: 0.36–0.47; BL: 1.92–2.37 (
|
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<typeStatus id="54B5887CE915420DCA30493215F9A224" box="[1329,1433,291,317]" pageId="9" pageNumber="455" type="holotype">holotype</typeStatus>
|
||
: 2.10).
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</paragraph>
|
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<paragraph id="8BB136DEE915420DCFC6497B132EA01E" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
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||
Habitus as in
|
||
<figureCitation id="13352A5BE915420DCE69497B11CEA29D" box="[360,430,362,388]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[228,1359,192,1905]" captionTargetId="figure-21@3.[216,1371,179,1917]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Omaliopsis: 1—O. fraterna (holotype), 2—O. hlavaci (holotype), 3—O. obliquesignata (holotype), 4—O. smetanai (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10254536" httpUri="https://zenodo.org/record/10254536/files/figure.png" pageId="9" pageNumber="455">Fig. 3</figureCitation>
|
||
. Body yellow-brown to reddish brown, with distinctly darker head and abdomen; basal and/or lateral portions of pronotum, middle and lateral portions of elytra, paratergites, apical margins of abdominal tergites and apex of abdomen yellowish; each elytron with narrow brownish spot in mediobasal and with moderately wide obligue spot in apical half, starting from about mediolateral margin toward medioapical part slightly not reaching suture, lateroapical portions of each elytron darkened in some specimens; antennomeres 7–11 brownish; mouthparts, antennomeres 1–6 and legs yellow. Punctation of head irregular and very sparse, denser in middle and sometimes without punctures between ocelli; punctation of pronotum about as that in middle portion of head, but denser, sparser and finer in medioapical and/or mediobasal portions; punctation of elytra about that in protonum, but slightly denser, sometimes denser and coarser around scutellum, finer and sparser along suture. Frontoclypeal part with fine transverse microreticulation, infraorbital portions with longitudinal meshes, middle of head without distinct microsculpture; pronotum and elytra without microreticulation; scutellum without or with fine transverse meshes; abdominal tergites with dense isodiametric sculpture.
|
||
</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64B011303A0E7" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">Head 1.4–1.5 times as broad as long, with narrow frontoclypeal portion and distinctly convex supra-antennal elevations, forming very long linear parallel ridges reaching level of basal third of eyes; anteriomedian depressions moderately deep and elongate; anteocellar foveae short and deep, slightly longer than diameter of ocelli; dorsal margins near eyes with fine indistinct one or two elongate wrinkles. Antennomeres 5–8 progressively broadened apicad; antennomere 3 slightly shorter and distinctly narrower than 2, 4 slightly less than twice shorter than 3, 5 slightly broader than 4, 7 slightly longer and distinctly broader than 6, 7 indistinctly longer and broader than 6, 8 slightly broader than 7, 9–10 slightly longer and broader than 8.</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64C161240A77E" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">Pronotum 1.5–1.7 times as broad as long, 1.3–1.4 times as broad as head, widest in about middle, slightly more narrowed posteriad than anteriad; basal part of surface of disc with two indistinct and shallow or distinct and moderately deep wide depressions.</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64C6113F7A792" blockId="9.[151,1437,151,2008]" box="[199,919,1136,1163]" pageId="9" pageNumber="455">Elytra slightly broader than long and twice as long as pronotum.</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64C82135BA7B4" blockId="9.[151,1437,151,2008]" box="[199,827,1171,1197]" pageId="9" pageNumber="455">Metatarsi slightly less than twice shorter than metatibia.</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64CA612E2A7C8" blockId="9.[151,1437,151,2008]" box="[199,642,1207,1233]" pageId="9" pageNumber="455">Abdomen slightly narrower than elytra.</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64CCB145AA698" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
Male. Posterior margin of abdominal tergite VIII rounded (
|
||
<figureCitation id="13352A5BE915420DCC724CCB13AAA7ED" box="[883,970,1242,1268]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 27</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII widely emarginate (
|
||
<figureCitation id="13352A5BE915420DCEAF4CEC1263A60E" box="[430,515,1277,1303]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 28</figureCitation>
|
||
). Aedeagus with moderately wide basal portion, gradually narrowed toward small rounded apex; parameres narrow, slightly broadened apically, not reaching apex of median lobe, with two moderately long apical setae; internal sac long and wide, with paired elongate and wide structures in basal portion and very short thin flagellum between them (
|
||
<figureCitation id="13352A5BE915420DCEE94D76125CA698" box="[488,572,1382,1409]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 25</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="13352A5BE915420DCCE24D761455A698" box="[995,1077,1383,1409]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 26</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64D9B11BCA6F3" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
Female. Posterior margin of abdominal tergite VIII (
|
||
<figureCitation id="13352A5BE915420DCC334D9B13E8A6BD" box="[818,904,1418,1444]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 29</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="13352A5BE915420DCB734D9B14A8A6BD" box="[1138,1224,1418,1444]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 30</figureCitation>
|
||
) rounded. Female accessoty sclerite leaf-shaped, short, from widest basal portion gradually narrowed toward rounded apex (
|
||
<figureCitation id="13352A5BE915420DCA3A4DBC15EFA6DE" box="[1339,1423,1453,1479]" captionStart="FIGURES 25–31" captionStartId="10.[151,264,1617,1641]" captionTargetBox="[159,1428,209,1586]" captionTargetId="figure-16@10.[151,1436,201,1594]" captionTargetPageId="10" captionText="FIGURES 25–31. Omaliopsis obliquesignata: 25–aedeagus, parameral view, 26–aedeagus, lateral view, 27–male tergite VIII, 28–male sternite VIII, 29–female tergite VIII, 30–female sternite 8, 31–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10254552" httpUri="https://zenodo.org/record/10254552/files/figure.png" pageId="9" pageNumber="455">Fig. 31</figureCitation>
|
||
). Spermatheca not recognized.
|
||
</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64DE21315A41D" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
<emphasis id="B97AEACCE915420DCFC64DE211D3A517" bold="true" box="[199,435,1523,1550]" pageId="9" pageNumber="455">Comparative notes.</emphasis>
|
||
Based on the general shapes of the convex forebody and the aedeagus, and the lack of postocular ridges,
|
||
<taxonomicName id="4C0E4D5DE915420DCE684E061253A528" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[361,563,1559,1585]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="obliquesignata" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCE684E06111DA528" box="[361,381,1559,1585]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCE8C4E061253A528" box="[397,563,1559,1585]" italics="true" pageId="9" pageNumber="455">obliquesignata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCD3D4E06121DA528" bold="true" box="[572,637,1559,1585]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCD3D4E06121DA528" box="[572,637,1559,1585]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C0E4D5DE915420DCC174E0613F6A528" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[790,918,1559,1585]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="fraterna" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCC174E06134AA528" box="[790,810,1559,1585]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCC3B4E0613F6A528" box="[826,918,1559,1585]" italics="true" pageId="9" pageNumber="455">fraterna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCC9F4E061380A528" bold="true" box="[926,992,1559,1585]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCC9F4E061380A528" box="[926,992,1559,1585]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C0E4D5DE915420DCCEE4E061405A528" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1007,1125,1559,1585]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="hlavaci" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCCEE4E061463A528" box="[1007,1027,1559,1585]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCB124E061405A528" box="[1043,1125,1559,1585]" italics="true" pageId="9" pageNumber="455">hlavaci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCB6F4E0614CFA528" bold="true" box="[1134,1199,1559,1585]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCB6F4E0614CFA528" box="[1134,1199,1559,1585]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0E4D5DE915420DCBE84E061512A529" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1257,1394,1559,1585]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="smetanai" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCBE84E06149DA528" box="[1257,1277,1559,1585]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCA0C4E061512A529" box="[1293,1394,1559,1584]" italics="true" pageId="9" pageNumber="455">smetanai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCA7A4E0610CEA54D" bold="true" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCA7A4E0610CEA54D" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
From
|
||
<taxonomicName id="4C0E4D5DE915420DCFFB4E2B1119A54D" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[250,377,1594,1620]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="fraterna" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCFFB4E2B116EA54D" box="[250,270,1594,1620]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCE1C4E2B1119A54D" box="[285,377,1594,1620]" italics="true" pageId="9" pageNumber="455">fraterna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCE814E2B11A0A54D" bold="true" box="[384,448,1594,1620]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCE814E2B11A0A54D" box="[384,448,1594,1620]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it differs by the paler forebody, slightly broader and shorter elytra, and slightly broader median lobe of the aedeagus. From
|
||
<taxonomicName id="4C0E4D5DE915420DCD224E4C12F7A56E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[547,663,1629,1656]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="hlavaci" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCD224E4C1257A56E" box="[547,567,1629,1655]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCD444E4C12F7A56E" box="[581,663,1629,1655]" italics="true" pageId="9" pageNumber="455">hlavaci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCD9F4E4C12BDA56E" bold="true" box="[670,733,1629,1655]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCD9F4E4C12BDA56E" box="[670,733,1629,1655]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it differs by the paler coloration, sparser punctation of the head and the pronotum, and longer elytra. From
|
||
<taxonomicName id="4C0E4D5DE915420DCD764E90129EA583" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[631,766,1665,1691]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="455" phylum="Arthropoda" rank="species" species="smetanai" status="sp. n.">
|
||
<emphasis id="B97AEACCE915420DCD764E9012EBA582" box="[631,651,1665,1691]" italics="true" pageId="9" pageNumber="455">O</emphasis>
|
||
.
|
||
<emphasis id="B97AEACCE915420DCD984E90129EA583" box="[665,766,1665,1690]" italics="true" pageId="9" pageNumber="455">smetanai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCC054E911323A583" bold="true" box="[772,835,1664,1690]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCC054E911323A583" box="[772,835,1664,1690]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it differs by the paler elytra with the presence of wide spots in medioapical portions. From all these species,
|
||
<taxonomicName id="4C0E4D5DE915420DCDF34EB513DAA5A4" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[754,954,1699,1726]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="459" phylum="Arthropoda" rank="species" species="obliquesignata" status="sp. n.">
|
||
O.
|
||
<emphasis id="B97AEACCE915420DCC154EB213DAA5A4" box="[788,954,1699,1725]" italics="true" pageId="9" pageNumber="455">obliquesignata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEACCE915420DCCC04EB51460A5A7" bold="true" box="[961,1024,1700,1726]" pageId="9" pageNumber="455">
|
||
<taxonomicNameLabel id="A24957B7E915420DCCC04EB51460A5A7" box="[961,1024,1700,1726]" pageId="9" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the different coloration of the elytra, slightly shorter parameres, details of the structure of the internal sac, and the shape of the female accessory sclerite. For additional detail see the key below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146555E915420DCFC64F1C1160A453" pageId="9" pageNumber="455" type="distribution">
|
||
<paragraph id="8BB136DEE915420DCFC64F1C1160A453" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
<emphasis id="B97AEACCE915420DCFC64F1C1101A43E" bold="true" box="[199,353,1805,1831]" pageId="9" pageNumber="455">Distribution.</emphasis>
|
||
The new species is known only from the
|
||
<typeStatus id="54B5887CE915420DCC374F1F1306A431" box="[822,870,1806,1832]" pageId="9" pageNumber="455">type</typeStatus>
|
||
locality in Qinling Shan, southern
|
||
<collectingRegion id="49CAF83CE915420DCBF04F1C152EA43E" box="[1265,1358,1805,1831]" country="China" name="Shaanxi" pageId="9" pageNumber="455">Shaanxi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F319764EE915420DCA584F1C15FCA43E" box="[1369,1436,1805,1831]" name="China" pageId="9" pageNumber="455">China</collectingCountry>
|
||
(
|
||
<figureCitation id="13352A5BE915420DCF9E4F211093A453" box="[159,243,1840,1866]" captionStart="FIGURE 17" captionStartId="6.[151,250,1548,1572]" captionTargetBox="[151,1436,453,1525]" captionTargetId="figure-120@6.[151,1436,453,1525]" captionTargetPageId="6" captionText="FIGURE 17. Distribution of Omaliopsis in China: O. fraterna (circles), O. hlavaci (triangles), O. obliquesignata (black star), O. smetanai (diamonds)." figureDoi="http://doi.org/10.5281/zenodo.10254548" httpUri="https://zenodo.org/record/10254548/files/figure.png" pageId="9" pageNumber="455">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146555E915420DCFC64F4511AFA4C1" pageId="9" pageNumber="455" type="etymology">
|
||
<paragraph id="8BB136DEE915420DCFC64F451361A488" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
<emphasis id="B97AEACCE915420DCFC64F45112EA477" bold="true" box="[199,334,1876,1902]" pageId="9" pageNumber="455">Etymology.</emphasis>
|
||
The specific epithet is the Latine adjective
|
||
<emphasis id="B97AEACCE915420DCC344F421386A474" box="[821,998,1875,1901]" italics="true" pageId="9" pageNumber="455">obliquesignatus</emphasis>
|
||
, -
|
||
<emphasis id="B97AEACCE915420DCCFC4F45146BA474" box="[1021,1035,1876,1901]" italics="true" pageId="9" pageNumber="455">a</emphasis>
|
||
, -
|
||
<emphasis id="B97AEACCE915420DCB224F451425A474" box="[1059,1093,1876,1901]" italics="true" pageId="9" pageNumber="455">um</emphasis>
|
||
(with oblique marks). It refers to the presence of oblique darkened spots on the elytra.
|
||
</paragraph>
|
||
<paragraph id="8BB136DEE915420DCFC64F8B11AFA4C1" blockId="9.[151,1437,151,2008]" pageId="9" pageNumber="455">
|
||
<emphasis id="B97AEACCE915420DCFC64F8B112BA4AD" bold="true" box="[199,331,1946,1972]" pageId="9" pageNumber="455">Bionomics.</emphasis>
|
||
Specimens were collected by sifting dead wood and mushrooms in deciduous forest with bamboo at elevation of
|
||
<quantity id="4CF69B3BE915420DCE3F4FAF11F3A4C1" box="[318,403,1982,2008]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="9" pageNumber="455" unit="m" value="1990.0">1990 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |