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<document id="E318B917C9D62DFADDD512A2806392F7" ID-DOI="10.5281/zenodo.4504241" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504241" ID-ZooBank="8DB4D80A-9B4B-443C-8F24-2234E093D347" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465845244" checkinUser="carolina" docAuthor="Tumpeesuwan, Sakboworn &amp; Tumpeesuwan, Chanidaporn" docDate="2015" docId="0385EF45555EFFADFC36B0F379359931" docLanguage="en" docName="RafflesBZool.63.287-292.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Pearsonia lamphunensis Tumpeesuwan &amp; Tumpeesuwan 2015, new species" docType="treatment" docUuid="B38197C4-D533-41F7-9F87-D570233F9915" docUuidSource="ZooBank" docVersion="5" lastPageNumber="291" masterDocId="FFBC973D555FFFA9FFAFB41D7A329F04" masterDocTitle="First record and description of a new species of the land snail genus Pearsonia Kobelt, 1902 (Cyclophoridae: Pterocyclinae) from Thailand, with a note on radula morphology" masterLastPageNumber="292" masterPageNumber="287" pageNumber="288" updateTime="1698866043417" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CC8ABE284D69A55E281051F4449F30DC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="980068BC071BF3937F1D3F4F6F5200C3">First record and description of a new species of the land snail genus Pearsonia Kobelt, 1902 (Cyclophoridae: Pterocyclinae) from Thailand, with a note on radula morphology</mods:title>
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<mods:name id="B6B6C0E5C6E2698E3E1F9317C1170D33" type="personal">
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<mods:namePart id="9A64524E58EAEA7DDA1B6F78FEAF157D">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:namePart id="B77588BCD6B5F9CD410A89C0FDF540FB">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:title id="703554D1F1BFF202D257C4837E6CAC87">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="8917925D7BE51DBA3DD3F73383FFEFFF">2015</mods:date>
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<mods:number id="2A677D8F0CC82C4025E96391D4270F7A">2015-07-01</mods:number>
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<mods:number id="B1D3929E844B934490113938CEF881F2">63</mods:number>
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<mods:classification id="82D1187A1DECA79B7958D1C04CEDFB6D">journal article</mods:classification>
<mods:identifier id="DDF56D3F9052D422FC6FC197C175CC8E" type="DOI">10.5281/zenodo.4504241</mods:identifier>
<mods:identifier id="C23D49C785E7E1112AF4FF638C0D844F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="2994DAEFFD4D2EE53E5BC789992CAC61" type="Zenodo-Dep">4504241</mods:identifier>
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<treatment id="0385EF45555EFFADFC36B0F379359931" LSID="urn:lsid:zoobank.org:act:B38197C4-D533-41F7-9F87-D570233F9915" httpUri="http://treatment.plazi.org/id/0385EF45555EFFADFC36B0F379359931" lastPageId="4" lastPageNumber="291" pageId="1" pageNumber="288">
<subSubSection id="C3360DD8555EFFA8FC36B0F37F729A02" box="[921,1344,1262,1286]" pageId="1" pageNumber="288" type="nomenclature">
<paragraph id="8B935E53555EFFA8FC36B0F37F729A02" blockId="1.[921,1344,1262,1318]" box="[921,1344,1262,1286]" pageId="1" pageNumber="288">
<heading id="D0DBE93F555EFFA8FC36B0F37F729A02" bold="true" box="[921,1344,1262,1286]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="288" reason="2">
<emphasis id="B9588241555EFFA8FC36B0F37F729A02" bold="true" box="[921,1344,1262,1286]" pageId="1" pageNumber="288">
<taxonomicName id="4C2C25D0555EFFA8FC36B0F37E999A02" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[921,1195,1262,1286]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="288" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555EFFA8FC36B0F37E999A02" bold="true" box="[921,1195,1262,1286]" italics="true" pageId="1" pageNumber="288">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555EFFA8FB15B0F37F729A02" box="[1210,1344,1262,1286]" pageId="1" pageNumber="288" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3360DD8555EFFA8FBA7B1137EE29A22" box="[1032,1232,1294,1318]" pageId="1" pageNumber="288" type="description">
<paragraph id="8B935E53555EFFA8FBA7B1137EE29A22" blockId="1.[921,1344,1262,1318]" box="[1032,1232,1294,1318]" pageId="1" pageNumber="288">
<figureCitation id="131742D6555EFFA8FBA7B1137E429A22" box="[1032,1136,1294,1318]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[135,170,1135,1157]" captionStartId-1="3.[113,148,690,712]" captionStartId-2="3.[113,148,2002,2024]" captionTargetBox-0="[135,1473,189,1120]" captionTargetBox-1="[114,1452,189,675]" captionTargetBox-2="[113,1451,854,1988]" captionTargetId-0="figure-8@2.[135,1473,189,1120]" captionTargetId-1="figure-51@3.[102,797,190,687]" captionTargetId-2="figure-129@3.[113,1451,854,1988]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="Fig. 2. Pearsonia lamphunensis, new species. A, Type locality habitat in dolomitic limestone hill, Lamphun Province; B, Aestivating living snail in natural habitat. Paratype (NHMSU-0007); C, Female partially retracted with the operculum (op) showing; D, vaginal groove (vg) near right tentacle visible." captionText-1="Fig. 3. Pearsonia lamphunensis, new species, dolomitic limestone hill Fig. 4. Operculum of Pearsonia lamphunensis, new species. near Tham Erawan, Lamphun Province. Holotype (NHMSU-0006). A, Paratype (NHMSU-0007). A, outer surface; B, inner surface; dorsal view; B, ventral view; C, apertural view; D, side view. Arrow C, side view; D, close up view of outer surface." captionText-2="Fig. 5. Pearsonia lamphunensis, new species, radula of paratype (NHMSU-0007). A, curled position, most central teeth were covered by lateral teeth; B, close up view of left side of radula; C, uncurled position, all cuspidal features are visible; D, close up view of right side of radula (st)." figureDoi-0="http://doi.org/10.5281/zenodo.4504245" figureDoi-1="http://doi.org/10.5281/zenodo.4504247" figureDoi-2="http://doi.org/10.5281/zenodo.4504249" httpUri-0="https://zenodo.org/record/4504245/files/figure.png" httpUri-1="https://zenodo.org/record/4504247/files/figure.png" httpUri-2="https://zenodo.org/record/4504249/files/figure.png" pageId="1" pageNumber="288">Figs. 2–5</figureCitation>
;
<tableCitation id="C6AE6BE8555EFFA8FBD2B1137EE29A22" box="[1149,1232,1294,1318]" captionStart="Table 1" captionStartId="2.[136,191,1270,1292]" captionText="Table 1. Comparison of shell morphology between Pearsonia lamphunensis, new species, and the recently described and two geographically close Pearsonia species." pageId="1" pageNumber="288">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3360DD8555EFFA8FC82B15379D19942" pageId="1" pageNumber="288" type="materials_examined">
<paragraph id="8B935E53555EFFA8FC82B15379D19942" blockId="1.[813,1452,1358,1606]" pageId="1" pageNumber="288">
<emphasis id="B9588241555EFFA8FC82B15379EF9A62" bold="true" box="[813,989,1358,1382]" pageId="1" pageNumber="288">Type material.</emphasis>
<materialsCitation id="3B44540E555EFFA8FC48B1537F9A9AC2" collectingDate="2014-06" country="Thailand" location="Malacological Laboratory Team of Mahasarakham University" pageId="1" pageNumber="288" specimenCount="1" stateProvince="Lamphun" typeStatus="holotype">
<typeStatus id="5497E0F1555EFFA8FC48B1537E609A62" box="[999,1106,1358,1382]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
: NHMSU-0006 (
<figureCitation id="131742D6555EFFA8FABAB1537F699A62" box="[1301,1371,1358,1382]" captionStart="Fig" captionStartId="3.[113,148,690,712]" captionTargetBox="[114,1452,189,675]" captionTargetId="figure-51@3.[102,797,190,687]" captionTargetPageId="3" captionText="Fig. 3. Pearsonia lamphunensis, new species, dolomitic limestone hill Fig. 4. Operculum of Pearsonia lamphunensis, new species. near Tham Erawan, Lamphun Province. Holotype (NHMSU-0006). A, Paratype (NHMSU-0007). A, outer surface; B, inner surface; dorsal view; B, ventral view; C, apertural view; D, side view. Arrow C, side view; D, close up view of outer surface." figureDoi="http://doi.org/10.5281/zenodo.4504247" httpUri="https://zenodo.org/record/4504247/files/figure.png" pageId="1" pageNumber="288">Fig. 3</figureCitation>
) (shell height 9.0 mm, shell width
<quantity id="4CD4F3B6555EFFA8FBD4B1737EDB9A82" box="[1147,1257,1390,1414]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="1" pageNumber="288" unit="mm" value="16.9">16.9 mm</quantity>
), Tham Erawan,
<collectingRegion id="49E890B1555EFFA8FC82B19379CC9AA2" box="[813,1022,1422,1446]" country="Thailand" name="Lamphun" pageId="1" pageNumber="288">Lamphun Province</collectingRegion>
, northern
<collectingCountry id="F33B1EC3555EFFA8FBC2B1937EFD9AA2" box="[1133,1231,1422,1446]" name="Thailand" pageId="1" pageNumber="288">Thailand</collectingCountry>
, coll.
<location id="8EF30888555EFFA8FABCB1937F199AC2" LSID="urn:lsid:plazi:treatment:0385EF45555EFFADFC36B0F379359931:8EF30888555EFFA8FABCB1937F199AC2" country="Thailand" name="Malacological Laboratory Team of Mahasarakham University" pageId="1" pageNumber="288" stateProvince="Lamphun">Malacological Laboratory Team of Mahasarakham University</location>
,
<date id="FF927893555EFFA8FA98B1B27F9A9AC2" box="[1335,1448,1454,1478]" pageId="1" pageNumber="288" value="2014-06">
<collectingDate id="EFD6817B555EFFA8FA98B1B27F9A9AC2" box="[1335,1448,1454,1478]" pageId="1" pageNumber="288" value="2014-06">June 2014</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="5497E0F1555EFFA8FC82B1D279AB9AE3" box="[813,921,1487,1511]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
: NHMSU-0007 (two shells and three specimens in ethanol; shell height 9.1–10.0 mm, shell width 15.7–17.0 mm);
<collectionCode id="ED3DC696555EFFA8FCC2B21379919922" box="[877,931,1550,1574]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="288">ZRC</collectionCode>
.
<collectionCode id="ED3DC696555EFFA8FC09B21379D69922" box="[934,996,1550,1574]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="1" pageNumber="288" type="Herbarium">MOL</collectionCode>
.5792 (one shell; shell height
<quantity id="4CD4F3B6555EFFA8FAB4B2137F5D9922" box="[1307,1391,1550,1574]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="288" unit="mm" value="9.2">9.2 mm</quantity>
, shell width
<quantity id="4CD4F3B6555EFFA8FCDDB23379E59942" box="[882,983,1582,1606]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="1" pageNumber="288" unit="mm" value="16.3">16.3 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3360DD8555EFFA8FC82B2737F4199A2" pageId="1" pageNumber="288" type="etymology">
<paragraph id="8B935E53555EFFA8FC82B2737F4199A2" blockId="1.[813,1451,1646,1703]" pageId="1" pageNumber="288">
<emphasis id="B9588241555EFFA8FC82B273799D9982" bold="true" box="[813,943,1646,1670]" pageId="1" pageNumber="288">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C2C25D0555EFFA8FB0CB2737F059982" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[1187,1335,1646,1670]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="288" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555EFFA8FB0CB2737F059982" box="[1187,1335,1646,1670]" italics="true" pageId="1" pageNumber="288">lamphunensis</emphasis>
</taxonomicName>
, is named after its
<typeStatus id="5497E0F1555EFFA8FC27B292798499A3" box="[904,950,1679,1703]" pageId="1" pageNumber="288">type</typeStatus>
locality
<collectingRegion id="49E890B1555EFFA8FBB8B2937ED799A2" box="[1047,1253,1678,1702]" country="Thailand" name="Lamphun" pageId="1" pageNumber="288">Lamphun Province</collectingRegion>
of
<collectingCountry id="F33B1EC3555EFFA8FAA3B2937F5C99A2" box="[1292,1390,1678,1702]" name="Thailand" pageId="1" pageNumber="288">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3360DD8555EFFA8FC82B2D37E249842" pageId="1" pageNumber="288" type="diagnosis">
<paragraph id="8B935E53555EFFA8FC82B2D37E249842" blockId="1.[813,1452,1742,1862]" pageId="1" pageNumber="288">
<emphasis id="B9588241555EFFA8FC82B2D379AD99E2" bold="true" box="[813,927,1742,1766]" pageId="1" pageNumber="288">Diagnosis.</emphasis>
Shell possesses backward pointing sutural tube (st) and horizontal, somewhat linguiform, peristome projection (pp). Zig-zag brown stripes present on both dorsal and ventral sides of shell.
</paragraph>
</subSubSection>
<subSubSection id="C3360DD8555EFFADFC82B37378589D51" lastPageId="4" lastPageNumber="291" pageId="1" pageNumber="288" type="description">
<paragraph id="8B935E53555EFFABFC82B37379359853" blockId="1.[813,1452,1902,2086]" lastBlockId="2.[136,775,1695,1879]" lastPageId="2" lastPageNumber="289" pageId="1" pageNumber="288">
<emphasis id="B9588241555EFFA8FC82B37379859882" bold="true" box="[813,951,1902,1926]" pageId="1" pageNumber="288">Description.</emphasis>
Shell: Dextral, sub-discoidal depressed, widely umbilicate, height 9.0–10.0 mm, shell width 15.7–17.0 mm, slightly thin, whorls 5¼–5½, rounded, spire a little raised. Background color pale brown, translucent, shiny, dorsal side with broad dark brown v-shaped or zig-zag stripes crossing the whorls; peripheral band broad, dark brownish; ventral side with many fine zig-zag brown stripes. Peristome double, the inner lip with sinus, at the parieto-columella angle, the outer lip expanded and forms a horizontal linguiform wing (peristome projection) near the suture. Sutural tube situated a little distance (ca.
<quantity id="4CD4F3B6555DFFABFEE9B3037BFB9833" box="[326,457,1822,1847]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.3" metricValueMin="3.6" pageId="2" pageNumber="289" unit="mm" value="4.45" valueMax="5.3" valueMin="3.6">3.6–5.3 mm</quantity>
.) from the peristome, in which the distal end is pointed backward (
<figureCitation id="131742D6555DFFABFDA2B32278639853" box="[525,593,1855,1879]" captionStart="Fig" captionStartId="2.[135,170,1135,1157]" captionTargetBox="[135,1473,189,1120]" captionTargetId="figure-8@2.[135,1473,189,1120]" captionTargetPageId="2" captionText="Fig. 2. Pearsonia lamphunensis, new species. A, Type locality habitat in dolomitic limestone hill, Lamphun Province; B, Aestivating living snail in natural habitat. Paratype (NHMSU-0007); C, Female partially retracted with the operculum (op) showing; D, vaginal groove (vg) near right tentacle visible." figureDoi="http://doi.org/10.5281/zenodo.4504245" httpUri="https://zenodo.org/record/4504245/files/figure.png" pageId="2" pageNumber="289">Figs 2</figureCitation>
B–D, 3A, C, D).
</paragraph>
<caption id="DF530EDB555DFFABFF28B0727B859BBB" ID-DOI="http://doi.org/10.5281/zenodo.4504245" ID-Zenodo-Dep="4504245" httpUri="https://zenodo.org/record/4504245/files/figure.png" pageId="2" pageNumber="289" startId="2.[135,170,1135,1157]" targetBox="[135,1473,189,1120]" targetPageId="2">
<paragraph id="8B935E53555DFFABFF28B0727B859BBB" blockId="2.[135,1474,1135,1215]" pageId="2" pageNumber="289">
Fig. 2.
<taxonomicName id="4C2C25D0555DFFABFF7DB06D7BF19B80" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[210,451,1135,1157]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555DFFABFF7DB06D7BF19B80" box="[210,451,1135,1157]" italics="true" pageId="2" pageNumber="289">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555DFFABFE7DB06D787E9B81" box="[466,588,1135,1157]" pageId="2" pageNumber="289" rank="species">new species</taxonomicNameLabel>
. A, Type locality habitat in dolomitic limestone hill, Lamphun Province; B, Aestivating living snail in natural habitat. Paratype (NHMSU-0007); C, Female partially retracted with the operculum (op) showing; D, vaginal groove (vg) near right tentacle visible.
</paragraph>
</caption>
<caption id="DF530EDB555DFFABFF27B0EB7B459A2C" pageId="2" pageNumber="289">
<paragraph id="8B935E53555DFFABFF27B0EB7B459A2C" blockId="2.[136,1475,1270,1321]" pageId="2" pageNumber="289">
Table 1. Comparison of shell morphology between
<taxonomicName id="4C2C25D0555DFFABFDDFB0EA796E9A0F" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[624,860,1270,1292]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555DFFABFDDFB0EA796E9A0F" box="[624,860,1270,1292]" italics="true" pageId="2" pageNumber="289">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555DFFABFCC8B0EB79EE9A08" box="[871,988,1270,1292]" pageId="2" pageNumber="289" rank="species">new species</taxonomicNameLabel>
, and the recently described and two geographically close
<taxonomicName id="4C2C25D0555DFFABFF6EB10E7B169A2C" authorityName="Kobelt" authorityYear="1902" box="[193,292,1299,1320]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="B9588241555DFFABFF6EB10E7B169A2C" box="[193,292,1299,1320]" italics="true" pageId="2" pageNumber="289">Pearsonia</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="8B935E53555DFFABFF6DB1477FA09954" pageId="2" pageNumber="289">
<table id="F92CACF3555D0056FF38B1477F829954" box="[151,1456,1370,1616]" gridcols="7" gridrows="5" pageId="2" pageNumber="289">
<tr id="351C5C11555D0056FF38B1477F829A74" box="[151,1456,1370,1392]" gridrow="0" pageId="2" pageNumber="289">
<th id="76CD356D555D0056FF38B1477B059A74" box="[151,311,1370,1392]" gridcol="0" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFF6DB1477B3F9A74" bold="true" box="[194,269,1370,1392]" pageId="2" pageNumber="289">Species</emphasis>
</th>
<th id="76CD356D555D0056FED4B1477BE09A74" box="[379,466,1370,1392]" gridcol="1" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFED4B1477BE09A74" bold="true" box="[379,466,1370,1392]" pageId="2" pageNumber="289">sh (mm)</emphasis>
</th>
<th id="76CD356D555D0056FD99B14778A49A74" box="[566,662,1370,1392]" gridcol="2" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFD96B14778A19A74" bold="true" box="[569,659,1370,1392]" pageId="2" pageNumber="289">sw (mm)</emphasis>
</th>
<th id="76CD356D555D0056FD4BB14779549A74" box="[740,870,1370,1392]" gridcol="3" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFD4BB14779549A6B" bold="true" box="[740,870,1370,1392]" pageId="2" pageNumber="289">Sutural tube</emphasis>
</th>
<th id="76CD356D555D0056FC1BB1477E279A74" box="[948,1045,1370,1392]" gridcol="4" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFC78B14779C09A6B" bold="true" box="[983,1010,1370,1391]" pageId="2" pageNumber="289">pp</emphasis>
</th>
<th id="76CD356D555D0056FBFCB1477EC69A74" box="[1107,1268,1370,1392]" gridcol="5" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFBFCB1477EC69A6B" bold="true" box="[1107,1268,1370,1392]" pageId="2" pageNumber="289">Zig-zag pattern</emphasis>
</th>
<th id="76CD356D555D0056FAB9B1477F829A74" box="[1302,1456,1370,1392]" gridcol="6" gridrow="0" pageId="2" pageNumber="289">
<emphasis id="B9588241555DFFABFA80B1477FA49A74" bold="true" box="[1327,1430,1370,1392]" pageId="2" pageNumber="289">Reference</emphasis>
</th>
</tr>
<tr id="351C5C11555D0056FF38B18C7F829AA3" box="[151,1456,1425,1447]" gridrow="1" pageId="2" pageNumber="289">
<th id="76CD356D555D0056FF38B18C7B059AA3" box="[151,311,1425,1447]" gridcol="0" gridrow="1" pageId="2" pageNumber="289">
<taxonomicName id="4C2C25D0555DFFABFF38B18C7B169AA2" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[151,292,1425,1446]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555DFFABFF38B18C7B169AA2" box="[151,292,1425,1446]" italics="true" pageId="2" pageNumber="289">P. bhamoensis</emphasis>
</taxonomicName>
</th>
<td id="76CD356D555D0056FED4B18C7BE09AA3" box="[379,466,1425,1447]" gridcol="1" gridrow="1" pageId="2" pageNumber="289">5.5</td>
<td id="76CD356D555D0056FD99B18C78A49AA3" box="[566,662,1425,1447]" gridcol="2" gridrow="1" pageId="2" pageNumber="289">11.0</td>
<td id="76CD356D555D0056FD4BB18C79549AA3" box="[740,870,1425,1447]" gridcol="3" gridrow="1" pageId="2" pageNumber="289">backward</td>
<td id="76CD356D555D0056FC1BB18C7E279AA3" box="[948,1045,1425,1447]" gridcol="4" gridrow="1" pageId="2" pageNumber="289">horizontal</td>
<td id="76CD356D555D0056FBFCB18C7EC69AA3" box="[1107,1268,1425,1447]" gridcol="5" gridrow="1" pageId="2" pageNumber="289">absent</td>
<td id="76CD356D555D0056FAB9B18C7F829AA3" box="[1302,1456,1425,1447]" gridcol="6" gridrow="1" pageId="2" pageNumber="289">
<bibRefCitation id="EFBD23A2555DFFABFA8AB18C7F929AA3" author="Gude GK" box="[1317,1440,1425,1447]" pageId="2" pageNumber="289" refId="ref3341" refString="Gude GK (1921) Land operculates (Cyclophoridae, Truncatellidae, Assimindeidae, Helicinidae). In: Shipley AS &amp; Marshall GAK (eds.). The Fauna of British India including Ceylon and Burma. Mollusca III Taylor &amp; Francis, Bombay, India. 386 pp." type="book" year="1921">Gude (1921)</bibRefCitation>
</td>
</tr>
<tr id="351C5C11555D0056FF38B1DD7F829AD2" box="[151,1456,1472,1494]" gridrow="2" pageId="2" pageNumber="289">
<th id="76CD356D555D0056FF38B1DD7B059AD2" box="[151,311,1472,1494]" gridcol="0" gridrow="2" pageId="2" pageNumber="289">
<taxonomicName id="4C2C25D0555DFFABFF38B1DD7ADC9AD1" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[151,238,1472,1493]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555DFFABFF38B1DD7ADC9AD1" box="[151,238,1472,1493]" italics="true" pageId="2" pageNumber="289">P. avana</emphasis>
</taxonomicName>
</th>
<td id="76CD356D555D0056FED4B1DD7BE09AD2" box="[379,466,1472,1494]" gridcol="1" gridrow="2" pageId="2" pageNumber="289">8.0</td>
<td id="76CD356D555D0056FD99B1DD78A49AD2" box="[566,662,1472,1494]" gridcol="2" gridrow="2" pageId="2" pageNumber="289">17.0</td>
<td id="76CD356D555D0056FD4BB1DD79549AD2" box="[740,870,1472,1494]" gridcol="3" gridrow="2" pageId="2" pageNumber="289">backward</td>
<td id="76CD356D555D0056FC1BB1DD7E279AD2" box="[948,1045,1472,1494]" gridcol="4" gridrow="2" pageId="2" pageNumber="289">vertical</td>
<td id="76CD356D555D0056FBFCB1DD7EC69AD2" box="[1107,1268,1472,1494]" gridcol="5" gridrow="2" pageId="2" pageNumber="289">dorsal side</td>
<td id="76CD356D555D0056FAB9B1DD7F829AD2" box="[1302,1456,1472,1494]" gridcol="6" gridrow="2" pageId="2" pageNumber="289">
<bibRefCitation id="EFBD23A2555DFFABFA8AB1DD7F929AD2" author="Gude GK" box="[1317,1440,1472,1494]" pageId="2" pageNumber="289" refId="ref3341" refString="Gude GK (1921) Land operculates (Cyclophoridae, Truncatellidae, Assimindeidae, Helicinidae). In: Shipley AS &amp; Marshall GAK (eds.). The Fauna of British India including Ceylon and Burma. Mollusca III Taylor &amp; Francis, Bombay, India. 386 pp." type="book" year="1921">Gude (1921)</bibRefCitation>
</td>
</tr>
<tr id="351C5C11555D0056FF38B1F37F829925" box="[151,1456,1518,1569]" gridrow="3" pageId="2" pageNumber="289">
<th id="76CD356D555D0056FF38B1F37B059925" box="[151,311,1518,1569]" gridcol="0" gridrow="3" pageId="2" pageNumber="289">
<taxonomicName id="4C2C25D0555DFFABFF38B1F27B169907" authorityName="Marzaki &amp; Clements" authorityYear="2013" box="[151,292,1518,1540]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="tembatensis">
<emphasis id="B9588241555DFFABFF38B1F27B169907" box="[151,292,1518,1540]" italics="true" pageId="2" pageNumber="289">P. tembatensis</emphasis>
</taxonomicName>
</th>
<td id="76CD356D555D0056FED4B1F37BE09925" box="[379,466,1518,1569]" gridcol="1" gridrow="3" pageId="2" pageNumber="289">8.6–11.8</td>
<td id="76CD356D555D0056FD99B1F378A49925" box="[566,662,1518,1569]" gridcol="2" gridrow="3" pageId="2" pageNumber="289">23.8–27.0</td>
<td id="76CD356D555D0056FD4BB1F379549925" box="[740,870,1518,1569]" gridcol="3" gridrow="3" pageId="2" pageNumber="289">backward</td>
<td id="76CD356D555D0056FC1BB1F37E279925" box="[948,1045,1518,1569]" gridcol="4" gridrow="3" pageId="2" pageNumber="289">horizontal</td>
<td id="76CD356D555D0056FBFCB1F37EC69925" box="[1107,1268,1518,1569]" gridcol="5" gridrow="3" pageId="2" pageNumber="289">absent</td>
<td id="76CD356D555D0056FAB9B1F37F829925" box="[1302,1456,1518,1569]" gridcol="6" gridrow="3" pageId="2" pageNumber="289">Masaki &amp; Clement (2013)</td>
</tr>
<tr id="351C5C11555D0056FF38B2277F829954" box="[151,1456,1594,1616]" gridrow="4" pageId="2" pageNumber="289">
<th id="76CD356D555D0056FF38B2277B059954" box="[151,311,1594,1616]" gridcol="0" gridrow="4" pageId="2" pageNumber="289">
<taxonomicName id="4C2C25D0555DFFABFF38B2267B05994B" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[151,311,1594,1616]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555DFFABFF38B2267B05994B" box="[151,311,1594,1616]" italics="true" pageId="2" pageNumber="289">P. lamphunensis</emphasis>
</taxonomicName>
</th>
<td id="76CD356D555D0056FED4B2277BE09954" box="[379,466,1594,1616]" gridcol="1" gridrow="4" pageId="2" pageNumber="289">9.0–10.0</td>
<td id="76CD356D555D0056FD99B22778A49954" box="[566,662,1594,1616]" gridcol="2" gridrow="4" pageId="2" pageNumber="289">15.7–17.0</td>
<td id="76CD356D555D0056FD4BB22779549954" box="[740,870,1594,1616]" gridcol="3" gridrow="4" pageId="2" pageNumber="289">backward</td>
<td id="76CD356D555D0056FC1BB2277E279954" box="[948,1045,1594,1616]" gridcol="4" gridrow="4" pageId="2" pageNumber="289">horizontal</td>
<td id="76CD356D555D0056FBFCB2277EC69954" box="[1107,1268,1594,1616]" gridcol="5" gridrow="4" pageId="2" pageNumber="289">Both sides</td>
<td id="76CD356D555D0056FAB9B2277F829954" box="[1302,1456,1594,1616]" gridcol="6" gridrow="4" pageId="2" pageNumber="289">this study</td>
</tr>
</table>
</paragraph>
<paragraph id="8B935E53555DFFABFF27B3627BCB9733" blockId="2.[136,776,1919,2103]" pageId="2" pageNumber="289">
Body: Pale brown to dark brown body, foot dark brown. Black eye spot present at the base of short brown tentacles. All three living specimens available for study are female, with a vaginal groove (vg) on the right side of the head (
<figureCitation id="131742D6555DFFABFF3FB3E27AD49713" box="[144,230,2047,2071]" captionStart="Fig" captionStartId="2.[135,170,1135,1157]" captionTargetBox="[135,1473,189,1120]" captionTargetId="figure-8@2.[135,1473,189,1120]" captionTargetPageId="2" captionText="Fig. 2. Pearsonia lamphunensis, new species. A, Type locality habitat in dolomitic limestone hill, Lamphun Province; B, Aestivating living snail in natural habitat. Paratype (NHMSU-0007); C, Female partially retracted with the operculum (op) showing; D, vaginal groove (vg) near right tentacle visible." figureDoi="http://doi.org/10.5281/zenodo.4504245" httpUri="https://zenodo.org/record/4504245/files/figure.png" pageId="2" pageNumber="289">Fig. 2D</figureCitation>
), which runs from the genital opening downwards to the area near the right tentacle.
</paragraph>
<paragraph id="8B935E53555DFFABFCECB2827E9B9713" blockId="2.[835,1475,1695,2071]" pageId="2" pageNumber="289">Operculum: Solid, round, bi-layered; thick calcified layer form outer part of operculum, and thin corneous layer forms inner surface. The outer surface rather convex, multispiral with more than seven counter-clockwise whorls, covered with fine growth lines, nucleus normally abraded. Internal corneous layer concave and wider than calcified layer and outer surface, therefore, bending outwards at edges; opercular disc attachment site large, circular, and can be divided into 3 parts, the middle part forms horseshoe shape area, the outer part and the large circular nucleus depressed. Periphery bearing imbricated semi-circular growth line, which is concave on the left-side (Fig. 4).</paragraph>
<caption id="DF530EDB555CFFAAFFDEB6AF7F189C06" ID-DOI="http://doi.org/10.5281/zenodo.4504247" ID-Zenodo-Dep="4504247" httpUri="https://zenodo.org/record/4504247/files/figure.png" pageId="3" pageNumber="290" startId="3.[113,148,690,712]" targetBox="[114,1452,189,675]" targetPageId="3">
<paragraph id="8B935E53555CFFAAFFDEB6AF7F189C06" blockId="3.[113,1453,690,799]" pageId="3" pageNumber="290">
Fig. 3.
<taxonomicName id="4C2C25D0555CFFAAFF19B6AE7B919DC3" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[182,419,690,712]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555CFFAAFF19B6AE7B919DC3" box="[182,419,690,712]" italics="true" pageId="3" pageNumber="290">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555CFFAAFE00B6AE78179DCC" box="[431,549,690,712]" pageId="3" pageNumber="290" rank="species">new species</taxonomicNameLabel>
, dolomitic limestone hill Fig. 4. Operculum of
<taxonomicName id="4C2C25D0555CFFAAFB81B6AE7F129DC3" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[1070,1312,690,712]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555CFFAAFB81B6AE7F129DC3" box="[1070,1312,690,712]" italics="true" pageId="3" pageNumber="290">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555CFFAAFA81B6AE7F9B9DCC" box="[1326,1449,690,712]" pageId="3" pageNumber="290" rank="species">new species</taxonomicNameLabel>
. near Tham Erawan, Lamphun Province. Holotype (NHMSU-0006). A, Paratype (NHMSU-0007). A, outer surface; B, inner surface; dorsal view; B, ventral view; C, apertural view; D, side view. Arrow C, side view; D, close up view of outer surface.
</paragraph>
</caption>
<paragraph id="8B935E53555CFFAAFFDEB7147B8B9C1B" blockId="3.[113,1453,690,799]" box="[113,441,777,799]" pageId="3" pageNumber="290">indicates position of sutural tube.</paragraph>
<caption id="DF530EDB555CFFAAFFDEB3CF7B149726" ID-DOI="http://doi.org/10.5281/zenodo.4504249" ID-Zenodo-Dep="4504249" httpUri="https://zenodo.org/record/4504249/files/figure.png" pageId="3" pageNumber="290" startId="3.[113,148,2002,2024]" targetBox="[113,1451,854,1988]" targetPageId="3">
<paragraph id="8B935E53555CFFAAFFDEB3CF7B149726" blockId="3.[113,1452,2002,2082]" pageId="3" pageNumber="290">
Fig. 5.
<taxonomicName id="4C2C25D0555CFFAAFF14B3CE7B9998E3" authority="Tumpeesuwan &amp; Tumpeesuwan, 2015" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[187,427,2002,2024]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="lamphunensis" status="sp. nov.">
<emphasis id="B9588241555CFFAAFF14B3CE7B9998E3" box="[187,427,2002,2024]" italics="true" pageId="3" pageNumber="290">Pearsonia lamphunensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B3F3A555CFFAAFE14B3CE780698EC" box="[443,564,2002,2024]" pageId="3" pageNumber="290" rank="species">new species</taxonomicNameLabel>
, radula of paratype (NHMSU-0007). A, curled position, most central teeth were covered by lateral teeth; B, close up view of left side of radula; C, uncurled position, all cuspidal features are visible; D, close up view of right side of radula (st).
</paragraph>
</caption>
<paragraph id="8B935E53555BFFADFF27B4A078589D51" blockId="4.[136,776,189,597]" pageId="4" pageNumber="291">
Radula: Taenioglossate, the sole specimen examined has at least 84 transverse rows and each row comprises seven teeth (1-2-1-2-1), including one left marginal tooth, two left lateral teeth, one central tooth, two right lateral teeth and one right marginal tooth. Central tooth is low, stout mushroom shape, pentacuspid, central cusp very large trapezoid shape, lateral and marginal cusps, small triangle knobs. Inner and outer lateral teeth tetracuspid, the first and second cusps are small robust finger shaped, the third cusp is very large shovel-shaped and the fourth cusp is small finger shaped. Marginal tooth, tricuspid, possesses the first finger shaped cusp, the second cusp, tongue shaped and the third cusp large triangular shape (
<figureCitation id="131742D6555BFFADFE2BB6207BF19D51" box="[388,451,573,597]" captionStart="Fig" captionStartId="3.[113,148,2002,2024]" captionTargetBox="[113,1451,854,1988]" captionTargetId="figure-129@3.[113,1451,854,1988]" captionTargetPageId="3" captionText="Fig. 5. Pearsonia lamphunensis, new species, radula of paratype (NHMSU-0007). A, curled position, most central teeth were covered by lateral teeth; B, close up view of left side of radula; C, uncurled position, all cuspidal features are visible; D, close up view of right side of radula (st)." figureDoi="http://doi.org/10.5281/zenodo.4504249" httpUri="https://zenodo.org/record/4504249/files/figure.png" pageId="4" pageNumber="291">Fig. 5</figureCitation>
A–D, Table 2).
</paragraph>
</subSubSection>
<subSubSection id="C3360DD8555BFFADFF27B66079359931" pageId="4" pageNumber="291" type="discussion">
<paragraph id="8B935E53555BFFADFF27B66078B49CD1" blockId="4.[136,775,637,981]" pageId="4" pageNumber="291">
<emphasis id="B9588241555BFFADFF27B6607B309D91" bold="true" box="[136,258,637,661]" pageId="4" pageNumber="291">Remarks.</emphasis>
The new species differs from
<taxonomicName id="4C2C25D0555BFFADFD23B66378069DB1" authority="Marzaki &amp; Clements, 2013" authorityName="Marzaki &amp; Clements" authorityYear="2013" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="tembatensis">
<emphasis id="B9588241555BFFADFD23B6637B349DB1" italics="true" pageId="4" pageNumber="291">Pearsonia tembatensis</emphasis>
Marzaki &amp; Clements, 2013
</taxonomicName>
, by its higher spire, presence of a broader dark brown peripheral band and zigzag stripes on both dorsal and ventral sides of shell. It is easily distinguished from
<taxonomicName id="4C2C25D0555BFFADFE30B6E07AFB9C31" authority="Sutcharit &amp; Panha" authorityName="Sutcharit &amp; Panha" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="291" phylum="Mollusca" rank="species" species="diluvium">
<emphasis id="B9588241555BFFADFE30B6E078B09C11" box="[415,642,765,789]" italics="true" pageId="4" pageNumber="291">Pterocyclos diluvium</emphasis>
Sutcharit &amp; Panha
</taxonomicName>
in
<bibRefCitation id="EFBD23A2555BFFADFF5DB7007BEF9C31" author="Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[242,477,797,821]" pageId="4" pageNumber="291" pagination="330 - 338" refId="ref3790" refString="Sutcharit C, Tongkerd P &amp; Panha S (2014) The land snails genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with descriptions of two new species. Raffle Bulletin of Zoology, 62: 330 - 338." type="journal article" year="2014">Sutcharit et al., 2014</bibRefCitation>
, by the backward pointing sutural tube and horizontal somewhat linguiform peristome projection.
<taxonomicName id="4C2C25D0555BFFADFEABB74078DE9C71" authority="(Pfeiffer, 1854)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" box="[260,748,861,885]" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="291" phylum="Mollusca" rank="species" species="pterocycloides">
<emphasis id="B9588241555BFFADFEABB74078099C71" box="[260,571,861,885]" italics="true" pageId="4" pageNumber="291">Opisthoporus pterocycloides</emphasis>
(
<bibRefCitation id="EFBD23A2555BFFADFDE5B74078D69C71" author="Pfeiffer L" box="[586,740,861,885]" pageId="4" pageNumber="291" pagination="299 - 303" refId="ref3754" refString="Pfeiffer L (1854) Descriptions of eighteen new species of Cyclostomacea, from Mr. Cuming's collection. Proceedings of the Zoological Society of London, 22: 299 - 303, pl. 35" type="journal article" year="1854">Pfeiffer, 1854</bibRefCitation>
)
</taxonomicName>
is very similarly patterned, but differs from the new species by its flat spire, expanded outer peristome, forward pointing sutural tube, and absence of a peripheral band.
</paragraph>
<paragraph id="8B935E53555BFFADFF27B7E079359931" blockId="4.[136,776,1021,1589]" pageId="4" pageNumber="291">
Shell morphology of
<taxonomicName id="4C2C25D0555BFFADFE23B7E3789C9B11" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[396,686,1021,1045]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555BFFADFE23B7E3789C9B11" box="[396,686,1021,1045]" italics="true" pageId="4" pageNumber="291">Pearsonia lamphunensis</emphasis>
</taxonomicName>
appear superficially similar to
<bibRefCitation id="EFBD23A2555BFFADFE34B00078709B31" author="Gude GK" box="[411,578,1053,1077]" pageId="4" pageNumber="291" refId="ref3341" refString="Gude GK (1921) Land operculates (Cyclophoridae, Truncatellidae, Assimindeidae, Helicinidae). In: Shipley AS &amp; Marshall GAK (eds.). The Fauna of British India including Ceylon and Burma. Mollusca III Taylor &amp; Francis, Bombay, India. 386 pp." type="book" year="1921">Gude’s (1921)</bibRefCitation>
illustrations and descriptions of
<taxonomicName id="4C2C25D0555BFFADFE88B02378FC9B51" authority="(Theobald, 1876)" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[295,718,1085,1109]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555BFFADFE88B02378269B51" box="[295,532,1085,1109]" italics="true" pageId="4" pageNumber="291">Pearsonia bhamoensis</emphasis>
(
<bibRefCitation id="EFBD23A2555BFFADFD8EB02078F59B51" author="Theobald W" box="[545,711,1085,1109]" pageId="4" pageNumber="291" pagination="184 - 189" refId="ref3838" refString="Theobald W (1876) Descriptions of some new land- and freshwater shells from India and Burmah. Journal of the Asiatic Society of Bengal, 45 (2) 3: 184 - 189, pl. 19." type="journal article" year="1876">Theobald, 1876</bibRefCitation>
)
</taxonomicName>
from Bhamo, and
<taxonomicName id="4C2C25D0555BFFADFEBAB04378BE9B71" authority="(Blanford, 1863)" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[277,652,1117,1141]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555BFFADFEBAB0437BFE9B71" box="[277,460,1118,1141]" italics="true" pageId="4" pageNumber="291">Pearsonia avana</emphasis>
(
<bibRefCitation id="EFBD23A2555BFFADFE72B04078B69B71" author="Blanford WT" box="[477,644,1117,1141]" pageId="4" pageNumber="291" pagination="319 - 327" refId="ref3232" refString="Blanford WT (1863) Contribution to Indian malacology, No. IV. Description of new land shells from Ava, and other parts of Burma. Journal of the Asiatic Society of Bengal, 32 (2): 319 - 327." type="journal article" year="1863">Blanford, 1863</bibRefCitation>
)
</taxonomicName>
from
<collectingRegion id="49E890B1555BFFADFD7EB04079349B71" box="[721,774,1117,1141]" country="Myanmar" name="Shan" pageId="4" pageNumber="291">Shan</collectingRegion>
Hill to the east of Ava, northeast
<collectingCountry id="F33B1EC3555BFFADFE56B06378569B92" box="[505,612,1150,1174]" name="Myanmar" pageId="4" pageNumber="291">Myanmar</collectingCountry>
. The shell size of
<taxonomicName id="4C2C25D0555BFFADFF07B0837B809BB1" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[168,434,1181,1205]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555BFFADFF07B0837B809BB1" box="[168,434,1181,1205]" italics="true" pageId="4" pageNumber="291">Pearsonia lamphunensis</emphasis>
</taxonomicName>
is however significantly larger and the spire is higher than
<taxonomicName id="4C2C25D0555BFFADFE00B0A378579BD1" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[431,613,1214,1237]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555BFFADFE00B0A378579BD1" box="[431,613,1214,1237]" italics="true" pageId="4" pageNumber="291">Pearsonia avana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C25D0555BFFADFD36B0A37B3A9BF1" baseAuthorityName="Theobald" baseAuthorityYear="1876" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555BFFADFD36B0A37B3A9BF1" italics="true" pageId="4" pageNumber="291">Pearsonia bhamoensis</emphasis>
</taxonomicName>
(see
<tableCitation id="C6AE6BE8555BFFADFEE7B0C07BAF9BF1" box="[328,413,1245,1269]" captionStart="Table 1" captionStartId="2.[136,191,1270,1292]" captionText="Table 1. Comparison of shell morphology between Pearsonia lamphunensis, new species, and the recently described and two geographically close Pearsonia species." pageId="4" pageNumber="291">Table 1</tableCitation>
). The peristome projection (pp) of
<taxonomicName id="4C2C25D0555BFFADFF06B0E37B8A9A11" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[169,440,1277,1301]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555BFFADFF06B0E37B8A9A11" box="[169,440,1277,1301]" italics="true" pageId="4" pageNumber="291">Pearsonia lamphunensis</emphasis>
</taxonomicName>
is horizontal as in
<taxonomicName id="4C2C25D0555BFFADFD38B0E37B349A31" baseAuthorityName="Theobald" baseAuthorityYear="1876" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555BFFADFD38B0E37B349A31" italics="true" pageId="4" pageNumber="291">Pearsonia bhamoensis</emphasis>
</taxonomicName>
, but unlike
<taxonomicName id="4C2C25D0555BFFADFE26B103780D9A31" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[393,575,1310,1333]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555BFFADFE26B103780D9A31" box="[393,575,1310,1333]" italics="true" pageId="4" pageNumber="291">Pearsonia avana</emphasis>
</taxonomicName>
, which is vertical. The sutural tube of both
<taxonomicName id="4C2C25D0555BFFADFE19B12378E09A51" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[438,722,1341,1365]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555BFFADFE19B12378E09A51" box="[438,722,1341,1365]" italics="true" pageId="4" pageNumber="291">Pearsonia lamphunensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C25D0555BFFADFF27B1437B4C9A71" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[136,382,1373,1397]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555BFFADFF27B1437B4C9A71" box="[136,382,1373,1397]" italics="true" pageId="4" pageNumber="291">Pearsonia bhamoensis</emphasis>
</taxonomicName>
is curved upward and backward in the suture, whereas in
<taxonomicName id="4C2C25D0555BFFADFED8B163781E9A91" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[375,556,1406,1429]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555BFFADFED8B163781E9A91" box="[375,556,1406,1429]" italics="true" pageId="4" pageNumber="291">Pearsonia avana</emphasis>
</taxonomicName>
it is curved upward. The colour pattern of
<taxonomicName id="4C2C25D0555BFFADFEDAB183784C9AB1" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2015" box="[373,638,1437,1461]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="lamphunensis">
<emphasis id="B9588241555BFFADFEDAB183784C9AB1" box="[373,638,1437,1461]" italics="true" pageId="4" pageNumber="291">Pearsonia lamphunensis</emphasis>
</taxonomicName>
differs from these two geographically close species by the presence of dark brown zig-zag stripes on both dorsal and ventral sides of the shell, whereas stripes are only present on the dorsal side in
<taxonomicName id="4C2C25D0555BFFADFF7EB2037BB09931" baseAuthorityName="Blanford" baseAuthorityYear="1863" box="[209,386,1566,1589]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="avana">
<emphasis id="B9588241555BFFADFF7EB2037BB09931" box="[209,386,1566,1589]" italics="true" pageId="4" pageNumber="291">Pearsonia avana</emphasis>
</taxonomicName>
and absent in
<taxonomicName id="4C2C25D0555BFFADFDBCB20379329931" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[531,768,1565,1589]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="291" phylum="Tracheophyta" rank="species" species="bhamoensis">
<emphasis id="B9588241555BFFADFDBCB20379329931" box="[531,768,1565,1589]" italics="true" pageId="4" pageNumber="291">Pearsonia bhamoensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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