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<document id="B0915152067EDC0F5C3CEF1B06D3DAF4" ID-CLB-Dataset="9574" ID-DOI="10.5281/zenodo.4272127" ID-GBIF-Dataset="724cb358-6f18-4816-afc7-bb42cb9b6942" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298182318" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2010" docId="0385915EFF5509EA609EFB5DCD84F9EA" docLanguage="en" docName="ActaEntMusNatPra.50.suppl.1-254.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 50" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Saprinillus Kryzhanovskij 1974" docType="treatment" docVersion="16" lastPageNumber="188" masterDocId="FFBCE926FF9709296157FFE7CE2AFFFE" masterDocTitle="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)" masterLastPageNumber="254" masterPageNumber="1" pageNumber="191" updateTime="1698846217483" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4C507B321123DE59FCF9066F8674AA9C">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<mods:date id="DA5DD71357CD3B71F67A5893BFE6F67B">2010</mods:date>
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<mods:title id="461512E9E94A74A35D3CB0D6C690C732">suppl.</mods:title>
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<subSubSection id="C33673C3FF5509EB609EFB5DCD1EFB2D" box="[457,820,1210,1235]" pageId="194" pageNumber="191" type="nomenclature">
<paragraph id="8B932048FF5509EB609EFB5DCD1EFB2D" blockId="194.[457,820,1210,1235]" box="[457,820,1210,1235]" pageId="194" pageNumber="191">
<heading id="D0DB9724FF5509EB609EFB5DCD1EFB2D" bold="true" box="[457,820,1210,1235]" centered="true" fontSize="10" level="2" pageId="194" pageNumber="191" reason="2">
<taxonomicName id="4C2C5BCBFF5509EB609EFB5DCD1EFB2D" ID-CoL="7CBX" authority="Kryzhanovskij, 1974" authorityName="Kryzhanovskij" authorityYear="1974" box="[457,820,1210,1235]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5509EB609EFB5DCD1EFB2D" bold="true" box="[457,820,1210,1235]" pageId="194" pageNumber="191">
<emphasis id="B958FC5AFF5509EB609EFB5DCC69FB2D" bold="true" box="[457,579,1210,1235]" italics="true" pageId="194" pageNumber="191">Saprinillus</emphasis>
<bibRefCitation id="EFBD5DB9FF5509EB631EFB5CCD1EFB2D" author="KRYZHANOVSKIJ O. L." box="[585,820,1211,1235]" pageId="194" pageNumber="191" refId="ref116877" refString="KRYZHANOVSKIJ O. L. 1974: K faune Histeridae (Coleoptera) Mongolskoy Narodnoy Respubliky II. (On the histerid fauna of the Mongolian Peoples' Republic (Coleoptera )). Pp. 102 - 109. In: Nasekomye Mongolii. Vyp. 2, tom. IV. [Insects of Mongolia. Vol. 2 (4)]. Nauka, Leningrad, xxx pp (in Russian,)." type="book" year="1974">Kryzhanovskij, 1974</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitationGroup id="AB3C0766FF5509EB61C0FB05CF35FAF1" pageId="194" pageNumber="191">
<subSubSection id="C33673C3FF5509EB61C0FB05CFC7FB0A" box="[151,493,1249,1269]" pageId="194" pageNumber="191" type="reference_group">
<treatmentCitation id="0A8D0659FF5509EB61C0FB05CFC3FB0A" author="KRYZHANOVSKIJ O. L." box="[151,489,1249,1269]" page="106" pageId="194" pageNumber="191" year="1974">
<taxonomicName id="4C2C5BCBFF5509EB61C0FB05CFC3FB0A" ID-CoL="7CBX" authority="Kryzhanovskij, 1974: 106" authorityName="Kryzhanovskij" authorityPageNumber="106" authorityYear="1974" box="[151,489,1249,1269]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5509EB61C0FB05CEDCFB0A" box="[151,246,1250,1268]" italics="true" pageId="194" pageNumber="191">Saprinillus</emphasis>
<bibRefCitation id="EFBD5DB9FF5509EB61A8FB06CFC3FB0A" author="KRYZHANOVSKIJ O. L." box="[255,489,1249,1269]" pageId="194" pageNumber="191" refId="ref116877" refString="KRYZHANOVSKIJ O. L. 1974: K faune Histeridae (Coleoptera) Mongolskoy Narodnoy Respubliky II. (On the histerid fauna of the Mongolian Peoples' Republic (Coleoptera )). Pp. 102 - 109. In: Nasekomye Mongolii. Vyp. 2, tom. IV. [Insects of Mongolia. Vol. 2 (4)]. Nauka, Leningrad, xxx pp (in Russian,)." type="book" year="1974">Kryzhanovskij, 1974: 106</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</subSubSection>
<subSubSection id="C33673C3FF5509EB60A3FB05CF35FAF1" pageId="194" pageNumber="191" type="type_taxon">
<typeStatus id="54979EEAFF5509EB60A3FB05CC0AFB0B" box="[500,544,1250,1269]" pageId="194" pageNumber="191">Type</typeStatus>
species:
<taxonomicName id="4C2C5BCBFF5509EB6321FB05CA3DFB0B" ID-CoL="79L9D" authority="Kryzhanovskij, 1974" authorityName="Kryzhanovskij" authorityYear="1974" box="[630,1047,1249,1269]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="species" species="paromaloides">
<emphasis id="B958FC5AFF5509EB6321FB05CD7EFB0A" box="[630,852,1250,1268]" italics="true" pageId="194" pageNumber="191">Saprinillus paromaloides</emphasis>
<bibRefCitation id="EFBD5DB9FF5509EB620AFB06CA3DFB0B" author="KRYZHANOVSKIJ O. L." box="[861,1047,1249,1269]" pageId="194" pageNumber="191" refId="ref116877" refString="KRYZHANOVSKIJ O. L. 1974: K faune Histeridae (Coleoptera) Mongolskoy Narodnoy Respubliky II. (On the histerid fauna of the Mongolian Peoples' Republic (Coleoptera )). Pp. 102 - 109. In: Nasekomye Mongolii. Vyp. 2, tom. IV. [Insects of Mongolia. Vol. 2 (4)]. Nauka, Leningrad, xxx pp (in Russian,)." type="book" year="1974">Kryzhanovskij, 1974</bibRefCitation>
</taxonomicName>
, original designation.
</subSubSection>
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</paragraph>
<subSubSection id="C33673C3FF5509EB61C0FAFFCC35FABB" pageId="194" pageNumber="191" type="reference_group">
<paragraph id="8B932048FF5509EB61C0FAFFCC35FABB" blockId="194.[151,1127,1249,1349]" pageId="194" pageNumber="191">
<treatmentCitationGroup id="AB3C0766FF5509EB61C0FAFFCC35FABB" pageId="194" pageNumber="191">
<treatmentCitation id="0A8D0659FF5509EB61C0FAFFCD2FFAD4" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[151,773,1303,1322]" page="216" pageId="194" pageNumber="191" year="1976">
<taxonomicName id="4C2C5BCBFF5509EB61C0FAFFCCF2FAD4" ID-CoL="7CBX" authority=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT (1976)" authorityName=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[151,728,1303,1322]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5509EB61C0FAFFCEDFFAD4" box="[151,245,1304,1322]" italics="true" pageId="194" pageNumber="191">Saprinillus</emphasis>
: Kryzhanovskij in
<bibRefCitation id="EFBD5DB9FF5509EB60CAFAF0CCF2FAD4" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[413,728,1303,1322]" pageId="194" pageNumber="191" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
</taxonomicName>
: 216
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF5509EB6258FAF0CD85FAD4" author="MAZUR S." box="[783,943,1303,1322]" page="83" pageId="194" pageNumber="191" year="1984">
<bibRefCitation id="EFBD5DB9FF5509EB6258FAF0CDA7FAD4" author="MAZUR S." box="[783,909,1303,1322]" pageId="194" pageNumber="191" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 83
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF5509EB62EEFAF0CA49FAD4" author="MAZUR S." box="[953,1123,1303,1322]" page="249" pageId="194" pageNumber="191" year="1997">
<bibRefCitation id="EFBD5DB9FF5509EB62EEFAF0CA1DFAD4" author="MAZUR S." box="[953,1079,1303,1322]" pageId="194" pageNumber="191" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 249
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF5509EB61E2FAD5CF79FABA" author="MAZUR S." box="[181,339,1329,1349]" page="96" pageId="194" pageNumber="191" year="2004">
<bibRefCitation id="EFBD5DB9FF5509EB61E2FAD5CF18FABA" author="MAZUR S." box="[181,306,1329,1349]" pageId="194" pageNumber="191" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
: 96
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF5509EB600CFAD5CC31FABB" author="LACKNER T." box="[347,539,1329,1349]" page="108" pageId="194" pageNumber="191" year="2009">
<bibRefCitation id="EFBD5DB9FF5509EB600CFAD5CFDAFABA" author="LACKNER T." box="[347,496,1329,1349]" pageId="194" pageNumber="191" pagination="107 - 118" refId="ref117251" refString="LACKNER T. 2009 a: Revision of the genus Saprinillus, with description of a new species (Coleoptera: Histeridae). Folia Heyrovskyana, Series A 16: 107 - 118." type="journal article" year="2009">LACKNER (2009a)</bibRefCitation>
: 108
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF5509EA61C0FABACDF5FBCA" lastPageId="195" lastPageNumber="188" pageId="194" pageNumber="191" type="diagnosis">
<paragraph id="8B932048FF5509EA61C0FABACF85FEEA" blockId="194.[151,1127,1373,1589]" lastBlockId="195.[113,1089,219,1556]" lastPageId="195" lastPageNumber="188" pageId="194" pageNumber="191">
<emphasis id="B958FC5AFF5509EB61C0FABACF26FA8B" bold="true" box="[151,268,1373,1397]" pageId="194" pageNumber="191">Diagnosis.</emphasis>
Body very small for the subfamily (
<quantity id="4CD48DADFF5509EB63DBFABACD01FA8B" box="[652,811,1373,1397]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="194" pageNumber="191" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.40–1.80 mm</quantity>
), cuticle without metallic luster, body sub-cylindrical to elongate oval, convex; clypeus and anterior third of frontal disc rugulose-lacunose, rest of frontal disc with sparse but coarse punctures; frontal stria largely interrupted, forming acute angles behind eyes; pronotal foveae absent; pronotal hypomeron glabrous (in case of
<taxonomicName id="4C2C5BCBFF5509EB6038FA39CC0BFA0B" authorityName="Lackner" authorityYear="2009" box="[367,545,1502,1525]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="species" species="kryzhanovskyi">
<emphasis id="B958FC5AFF5509EB6038FA39CC0BFA0B" box="[367,545,1502,1525]" italics="true" pageId="194" pageNumber="191">S. kryzhanovskyi</emphasis>
</taxonomicName>
with extremely short sparse setae). Apical elytral stria absent, sutural stria complete; pre-apical foveae present, well impressed; prosternal process narrowed; both sets of prosternal striae well developed (in the case of
<emphasis id="B958FC5AFF5509EA6222F9F9CEB0FF0A" italics="true" lastPageId="195" lastPageNumber="188" pageId="194" pageNumber="191">
<taxonomicName id="4C2C5BCBFF5509EB6222F9F9CDC0F9CB" authorityName=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[885,1002,1566,1589]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="194" pageNumber="191" phylum="Arthropoda" rank="genus">Saprinillus</taxonomicName>
kryzhanov- skyi
</emphasis>
lateral prosternal striae interrupted by antennal cavity); protibia with 5–6 short denticles on outer margin, teeth absent.
</paragraph>
<paragraph id="8B932048FF5409EA61C7FEFBCD1FFE6A" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
This genus has been recently revised by
<bibRefCitation id="EFBD5DB9FF5409EA636AFEFBCCDDFECA" author="LACKNER T." box="[573,759,284,308]" pageId="195" pageNumber="188" pagination="107 - 118" refId="ref117251" refString="LACKNER T. 2009 a: Revision of the genus Saprinillus, with description of a new species (Coleoptera: Histeridae). Folia Heyrovskyana, Series A 16: 107 - 118." type="journal article" year="2009">LACKNER (2009a)</bibRefCitation>
. For the sake of consistency its diagnosis, biology and distribution are repeated here, if slightly altered to fit the style used in this publication. Likewise, since this paper introduces some new terminology, this has also been taken into the account and the relevant parts are duly altered.
</paragraph>
<paragraph id="8B932048FF5409EA6126FE7CCDF5FBCA" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
<emphasis id="B958FC5AFF5409EA6126FE7CCF44FE4D" bold="true" box="[113,366,411,435]" pageId="195" pageNumber="188">Differential diagnosis.</emphasis>
Genus
<taxonomicName id="4C2C5BCBFF5409EA6094FE7ACC12FE4A" authorityName=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[451,568,413,436]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6094FE7ACC12FE4A" box="[451,568,413,436]" italics="true" pageId="195" pageNumber="188">Saprinillus</emphasis>
</taxonomicName>
stands out from the other genera of Palaearctic
<taxonomicName id="4C2C5BCBFF5409EA6126FE5BCEC9FE2A" authorityName="Blanchard" authorityYear="1845" box="[113,227,444,468]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
by its small body size, sub-cylindrical body, coarsely and rugosely granulate clypeus and anterior part of frontal disc, and by the shape of protibia which is without typical articulated triangular teeth and instead is adorned with denticles, growing out directly from the outer margin of protibia. Only few other genera of the Palaearctic
<taxonomicName id="4C2C5BCBFF5409EA6211FDFBCD92FDCA" authorityName="Blanchard" authorityYear="1845" box="[838,952,540,564]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
possess denticles (as opposed to articulated teeth) on the outer protibial margin, namely
<taxonomicName id="4C2C5BCBFF5409EA62CAFDDACA10FDAA" authorityName=": MAZUR" authorityYear="1984" box="[925,1082,573,596]" class="Insecta" family="Histeridae" genus="Microsaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA62CAFDDACA10FDAA" box="[925,1082,573,596]" italics="true" pageId="195" pageNumber="188">Microsaprinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFF5409EA6126FDBACF3EFD8A" authorityName=": ROSS" authorityYear="1940" box="[113,276,605,628]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6126FDBACF3EFD8A" box="[113,276,605,628]" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFF5409EA6073FDBACFA7FD8A" authorityName=": THOMSON" authorityYear="1867" box="[292,397,605,628]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6073FDBACFA7FD8A" box="[292,397,605,628]" italics="true" pageId="195" pageNumber="188">Myrmetes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C5BCBFF5409EA6092FDBACC44FD8A" authorityName="Tishechkin" authorityYear="2005" box="[453,622,605,628]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6092FDBACC44FD8A" box="[453,622,605,628]" italics="true" pageId="195" pageNumber="188">Turanostyphrus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C2C5BCBFF5409EA63B8FDBACD4EFD8A" authorityName=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[751,868,605,628]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA63B8FDBACD4EFD8A" box="[751,868,605,628]" italics="true" pageId="195" pageNumber="188">Saprinillus</emphasis>
</taxonomicName>
differs significantly from all the above-mentioned genera: from the genus
<taxonomicName id="4C2C5BCBFF5409EA63CBFD9ACD12FD6A" authorityName=": MAZUR" authorityYear="1984" box="[668,824,637,660]" class="Insecta" family="Histeridae" genus="Microsaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA63CBFD9ACD12FD6A" box="[668,824,637,660]" italics="true" pageId="195" pageNumber="188">Microsaprinus</emphasis>
</taxonomicName>
by the rugulose-lacunose clypeus and anterior third of frontal disc, anteriorly united carinal and lateral prosternal striae (anteriorly ‘open’ in
<taxonomicName id="4C2C5BCBFF5409EA6000FD5ACFDEFD2A" authorityName=": MAZUR" authorityYear="1984" box="[343,500,701,724]" class="Insecta" family="Histeridae" genus="Microsaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6000FD5ACFDEFD2A" box="[343,500,701,724]" italics="true" pageId="195" pageNumber="188">Microsaprinus</emphasis>
</taxonomicName>
), present pre-apical foveae (absent in
<taxonomicName id="4C2C5BCBFF5409EA62CCFD5ACA12FD2A" authorityName=": MAZUR" authorityYear="1984" box="[923,1080,701,724]" class="Insecta" family="Histeridae" genus="Microsaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA62CCFD5ACA12FD2A" box="[923,1080,701,724]" italics="true" pageId="195" pageNumber="188">Microsaprinus</emphasis>
</taxonomicName>
) and by the more elongate, almost cylindrical body; from
<taxonomicName id="4C2C5BCBFF5409EA63B6FD3ACDAEFD0A" authorityName=": ROSS" authorityYear="1940" box="[737,900,733,756]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA63B6FD3ACDAEFD0A" box="[737,900,733,756]" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
it also differs by the rugulose-lacunose clypeus and anterior third of frontal disc (finely punctate in
<taxonomicName id="4C2C5BCBFF5409EA62A6FD1ACEE7FCCA" authorityName=": ROSS" authorityYear="1940" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA62A6FD1ACEE7FCCA" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
), present pre-apical foveae (absent in
<taxonomicName id="4C2C5BCBFF5409EA6333FCFACD2DFCCA" authorityName=": ROSS" authorityYear="1940" box="[612,775,797,820]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6333FCFACD2DFCCA" box="[612,775,797,820]" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
), by the sub-cylindrical body shape (roundly oval in
<taxonomicName id="4C2C5BCBFF5409EA6039FCDACC3BFCAA" authorityName=": ROSS" authorityYear="1940" box="[366,529,829,852]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6039FCDACC3BFCAA" box="[366,529,829,852]" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
), and the much shorter meso- and metatibiae; from
<taxonomicName id="4C2C5BCBFF5409EA6126FCBACEF0FC8A" authorityName=": THOMSON" authorityYear="1867" box="[113,218,861,884]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6126FCBACEF0FC8A" box="[113,218,861,884]" italics="true" pageId="195" pageNumber="188">Myrmetes</emphasis>
</taxonomicName>
it differs by the same characters as from
<taxonomicName id="4C2C5BCBFF5409EA63CFFCBACD11FC8A" authorityName=": ROSS" authorityYear="1940" box="[664,827,861,884]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA63CFFCBACD11FC8A" box="[664,827,861,884]" italics="true" pageId="195" pageNumber="188">Eremosaprinus</emphasis>
</taxonomicName>
; furthermore,
<taxonomicName id="4C2C5BCBFF5409EA6280FCBACA6AFC8A" authorityName=": THOMSON" authorityYear="1867" box="[983,1088,861,884]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6280FCBACA6AFC8A" box="[983,1088,861,884]" italics="true" pageId="195" pageNumber="188">Myrmetes</emphasis>
</taxonomicName>
has almost impunctate dorsal surface; and from the species of the genus
<taxonomicName id="4C2C5BCBFF5409EA62D6FC9ACA00FC6A" authorityName="Tishechkin" authorityYear="2005" box="[897,1066,893,916]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA62D6FC9ACA00FC6A" box="[897,1066,893,916]" italics="true" pageId="195" pageNumber="188">Turanostyphrus</emphasis>
</taxonomicName>
it differs by the above-mentioned characters, plus asetose elytral epipleuron, bearing setae in
<taxonomicName id="4C2C5BCBFF5409EA6126FC5ACF30FC2A" authorityName="Tishechkin" authorityYear="2005" box="[113,282,957,980]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6126FC5ACF30FC2A" box="[113,282,957,980]" italics="true" pageId="195" pageNumber="188">Turanostyphrus</emphasis>
</taxonomicName>
. On the other hand, it superficially resembles the genus
<taxonomicName id="4C2C5BCBFF5409EA623AFC5ACDE3FC2A" authorityName=": MAZUR" authorityYear="1984" box="[877,969,957,980]" class="Insecta" family="Histeridae" genus="Axelinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA623AFC5ACDE3FC2A" box="[877,969,957,980]" italics="true" pageId="195" pageNumber="188">Axelinus</emphasis>
</taxonomicName>
, especially by the rugulose-lacunose anterior part of frontal disc, with largely interrupted frontal stria, but differing from it chiefly by the shape of protibia (bearing three large teeth topped with denticles in
<taxonomicName id="4C2C5BCBFF5409EA61A5FBFACF64FBCA" authorityName=": MAZUR" authorityYear="1984" box="[242,334,1053,1076]" class="Insecta" family="Histeridae" genus="Axelinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA61A5FBFACF64FBCA" box="[242,334,1053,1076]" italics="true" pageId="195" pageNumber="188">Axelinus</emphasis>
</taxonomicName>
), and by the more slender metatibiae (thickened in
<taxonomicName id="4C2C5BCBFF5409EA6224FBFACDE5FBCA" authorityName=": MAZUR" authorityYear="1984" box="[883,975,1053,1076]" class="Insecta" family="Histeridae" genus="Axelinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6224FBFACDE5FBCA" box="[883,975,1053,1076]" italics="true" pageId="195" pageNumber="188">Axelinus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF5409EA6126FBDCCFA5FAEA" pageId="195" pageNumber="188" type="biology_ecology">
<paragraph id="8B932048FF5409EA6126FBDCCFA5FAEA" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
<emphasis id="B958FC5AFF5409EA6126FBDCCEE4FBAD" bold="true" box="[113,206,1083,1107]" pageId="195" pageNumber="188">Biology.</emphasis>
The biology of this genus is poorly documented. The
<typeStatus id="54979EEAFF5409EA6275FBDBCD7AFBAA" box="[802,848,1084,1108]" pageId="195" pageNumber="188">type</typeStatus>
series of both species were collected in debris under plant
<taxonomicName id="4C2C5BCBFF5409EA60BFFBBACD4EFB8A" authorityName="Fenzl" authorityYear="1851" box="[488,868,1116,1140]" class="Magnoliopsida" family="Amaranthaceae" genus="Kalidium" kingdom="Plantae" order="Caryophyllales" pageId="195" pageNumber="188" phylum="Tracheophyta" rank="species" species="gracile">
<emphasis id="B958FC5AFF5409EA60BFFBBACCB1FB8A" box="[488,667,1117,1140]" italics="true" pageId="195" pageNumber="188">Kalidium gracile</emphasis>
(Chenopodiaceae)
</taxonomicName>
. The specimens from
<collectingCountry id="F33B60D8FF5409EA6126FB9BCF2FFB6A" box="[113,261,1148,1172]" name="Turkmenistan" pageId="195" pageNumber="188">Turkmenistan</collectingCountry>
and
<collectingCountry id="F33B60D8FF5409EA6061FB9BCF98FB6A" box="[310,434,1148,1172]" name="Kazakhstan" pageId="195" pageNumber="188">Kazakhstan</collectingCountry>
were found in a burrow of a Giant Gerbil (
<taxonomicName id="4C2C5BCBFF5409EA623AFB9ACF64FB4A" authority="(Lichtenstein, 1823))" authorityName="" baseAuthorityName="Lichtenstein" baseAuthorityYear="1823" class="Mammalia" family="Muridae" genus="Rhombomys" kingdom="Animalia" order="Rodentia" pageId="195" pageNumber="188" phylum="Chordata" rank="species" species="opimus">
<emphasis id="B958FC5AFF5409EA623AFB9ACA6BFB6A" box="[877,1089,1149,1172]" italics="true" pageId="195" pageNumber="188">Rhombomys opimus</emphasis>
(Lichtenstein, 1823))
</taxonomicName>
or trapped by a pitfall trap, respectively (
<bibRefCitation id="EFBD5DB9FF5409EA63A1FB7BCDFEFB4A" author="KRYZHANOVSKIJ O. L." box="[758,980,1180,1204]" pageId="195" pageNumber="188" pagination="24 - 38" refId="ref117012" refString="KRYZHANOVSKIJ O. L. 1987: Novye i maloizvestnye palearkticheskie taksony podsemeystva Saprininae (Coleoptera Histeridae). [New and poorly known Palaearctic taxa of the subfamily Saprininae (Coleoptera Histeridae)]. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 164: 24 - 38 (in Russian)." type="journal article" year="1987">KRYZHANOVSKIJ 1987</bibRefCitation>
).
<bibRefCitation id="EFBD5DB9FF5409EA62B3FB7BCEEEFB2A" author="LACKNER T." pageId="195" pageNumber="188" pagination="107 - 118" refId="ref117251" refString="LACKNER T. 2009 a: Revision of the genus Saprinillus, with description of a new species (Coleoptera: Histeridae). Folia Heyrovskyana, Series A 16: 107 - 118." type="journal article" year="2009">LACKNER (2009a)</bibRefCitation>
speculated that this taxon might rank amongst inqulines rather than psammophiles, in accordance with
<bibRefCitation id="EFBD5DB9FF5409EA6011FB3BCC1CFB0A" author="KRYZHANOVSKIJ O. L." box="[326,566,1244,1268]" pageId="195" pageNumber="188" pagination="24 - 38" refId="ref117012" refString="KRYZHANOVSKIJ O. L. 1987: Novye i maloizvestnye palearkticheskie taksony podsemeystva Saprininae (Coleoptera Histeridae). [New and poorly known Palaearctic taxa of the subfamily Saprininae (Coleoptera Histeridae)]. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 164: 24 - 38 (in Russian)." type="journal article" year="1987">KRYZHANOVSKIJ (1987)</bibRefCitation>
who discussed the shape of protibia as unfit for the psammophilous habits.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF5409EA6126FAFCCEC8FA6A" pageId="195" pageNumber="188" type="distribution">
<paragraph id="8B932048FF5409EA6126FAFCCF0CFAAA" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
<emphasis id="B958FC5AFF5409EA6126FAFCCF2EFACD" bold="true" box="[113,260,1307,1331]" pageId="195" pageNumber="188">Distribution.</emphasis>
<collectingCountry id="F33B60D8FF5409EA605CFAFBCF5DFACA" box="[267,375,1308,1332]" name="Mongolia" pageId="195" pageNumber="188">Mongolia</collectingCountry>
: South Gobi Aimak,
<collectingCountry id="F33B60D8FF5409EA630BFAFBCCDAFACA" box="[604,752,1308,1332]" name="Turkmenistan" pageId="195" pageNumber="188">Turkmenistan</collectingCountry>
and
<collectingCountry id="F33B60D8FF5409EA6272FAFBCD8BFACA" box="[805,929,1308,1332]" name="Kazakhstan" pageId="195" pageNumber="188">Kazakhstan</collectingCountry>
(
<bibRefCitation id="EFBD5DB9FF5409EA62E6FAFBCA17FACA" author="MAZUR S." box="[945,1085,1308,1332]" pageId="195" pageNumber="188" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR 1997</bibRefCitation>
,
<bibRefCitation id="EFBD5DB9FF5409EA6126FADBCF31FAAA" author="LACKNER T." box="[113,283,1340,1364]" pageId="195" pageNumber="188" pagination="107 - 118" refId="ref117251" refString="LACKNER T. 2009 a: Revision of the genus Saprinillus, with description of a new species (Coleoptera: Histeridae). Folia Heyrovskyana, Series A 16: 107 - 118." type="journal article" year="2009">LACKNER 2009a</bibRefCitation>
).
</paragraph>
<paragraph id="8B932048FF5409EA6126FABCCEC8FA6A" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
<emphasis id="B958FC5AFF5409EA6126FABCCF6AFA8D" bold="true" box="[113,320,1371,1395]" pageId="195" pageNumber="188">Species examined.</emphasis>
<taxonomicName id="4C2C5BCBFF5409EA601FFABACD2AFA8A" authority="Lackner, 2009" authorityName="Lackner" authorityYear="2009" box="[328,768,1372,1396]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="species" species="kryzhanovskyi">
<emphasis id="B958FC5AFF5409EA601FFABACC71FA8A" box="[328,603,1373,1396]" italics="true" pageId="195" pageNumber="188">Saprinillus kryzhanovskyi</emphasis>
Lackner, 2009
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFF5409EA625BFABACEF4FA6A" authority="Kryzhanovskij, 1974" authorityName="Kryzhanovskij" authorityYear="1974" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="species" species="paromaloides">
<emphasis id="B958FC5AFF5409EA625BFABACD90FA8A" box="[780,954,1373,1396]" italics="true" pageId="195" pageNumber="188">S. paromaloides</emphasis>
<bibRefCitation id="EFBD5DB9FF5409EA6296FABBCEF4FA6A" author="KRYZHANOVSKIJ O. L." pageId="195" pageNumber="188" refId="ref116877" refString="KRYZHANOVSKIJ O. L. 1974: K faune Histeridae (Coleoptera) Mongolskoy Narodnoy Respubliky II. (On the histerid fauna of the Mongolian Peoples' Republic (Coleoptera )). Pp. 102 - 109. In: Nasekomye Mongolii. Vyp. 2, tom. IV. [Insects of Mongolia. Vol. 2 (4)]. Nauka, Leningrad, xxx pp (in Russian,)." type="book" year="1974">Kryzhanovskij, 1974</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF5409EA6126FA7CCD84F9EA" pageId="195" pageNumber="188" type="discussion">
<paragraph id="8B932048FF5409EA6126FA7CCD84F9EA" blockId="195.[113,1089,219,1556]" pageId="195" pageNumber="188">
<emphasis id="B958FC5AFF5409EA6126FA7CCEDBFA4D" bold="true" box="[113,241,1435,1459]" pageId="195" pageNumber="188">Discussion.</emphasis>
<taxonomicName id="4C2C5BCBFF5409EA61A1FA7ACF41FA4A" authorityName=": Kryzhanovskij in KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[246,363,1437,1460]" class="Insecta" family="Histeridae" genus="Saprinillus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA61A1FA7ACF41FA4A" box="[246,363,1437,1460]" italics="true" pageId="195" pageNumber="188">Saprinillus</emphasis>
</taxonomicName>
belongs probably to the group of the genera (together with
<taxonomicName id="4C2C5BCBFF5409EA6289FA7ACA10FA4A" authorityName=": MAZUR" authorityYear="1984" box="[990,1082,1437,1460]" class="Insecta" family="Histeridae" genus="Axelinus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6289FA7ACA10FA4A" box="[990,1082,1437,1460]" italics="true" pageId="195" pageNumber="188">Axelinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFF5409EA6126FA5ACF3AFA2A" authorityName="Kanaar" authorityYear="2008" box="[113,272,1469,1492]" class="Insecta" family="Histeridae" genus="Alienocacculus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6126FA5ACF3AFA2A" box="[113,272,1469,1492]" italics="true" pageId="195" pageNumber="188">Alienocacculus</emphasis>
</taxonomicName>
, etc.) that are probably deeply nested within the paraphyletic genus
<taxonomicName id="4C2C5BCBFF5409EA6295FA5ACEBAFA0A" authorityName="Bickhardt" authorityYear="1914" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="195" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF5409EA6295FA5ACEBAFA0A" italics="true" pageId="195" pageNumber="188">Hypocacculus</emphasis>
</taxonomicName>
. It is supported by several ‘weak’ synapomorphies e.g. cylindrical body, rugulose-lacunose frons or interrupted frontal stria. Its monophyly should be tested in the future.
</paragraph>
</subSubSection>
</treatment>
</document>
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