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<mods:title id="92947259A7F6CB2DE7194DCA392A6A39">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<figureCitation id="13173CCDFF4D09F360A1FAFCCC14FAD0" box="[502,574,1307,1326]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="218" pageNumber="215">Figs. 12</figureCitation>
, 
<figureCitation id="13173CCDFF4D09F36310FAFCCC74FAD0" box="[583,606,1307,1326]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="218" pageNumber="215">71</figureCitation>
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<figureCitation id="13173CCDFF4D09F36330FAFCCCA3FAD0" box="[615,649,1307,1326]" captionStart="Figs" captionStartId="36.[151,190,1161,1180]" captionTargetBox="[151,1126,219,1130]" captionTargetId="figure-100@36.[151,1126,219,1132]" captionTargetPageId="36" captionText="Figs.99–107.Saprininae, mandibles,ventral view:99 – Philothis (Farabius) hexeris Reichardt, 1930; 100 – Pholioxenus phoenix (Reichardt,1930); 101 – Reichardtiolus duriculus (Reitter, 1904); 102 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 103 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 104 – Saprinus (Phaonius) pharao (Marseul, 1855); 105 – Styphrus corpulentus Motschulsky, 1845; 106 – Xenonychus tridens (Jacquelin-Duval, 1852); 107 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4272155" httpUri="https://zenodo.org/record/4272155/files/figure.png" pageId="218" pageNumber="215">105</figureCitation>
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<figureCitation id="13173CCDFF4D09F363C6FAFCCC99FAD0" box="[657,691,1307,1326]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="218" pageNumber="215">136</figureCitation>
, 
<figureCitation id="13173CCDFF4D09F363EBFAFCCD22FAD0" box="[700,776,1307,1326]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="220.[151,190,1516,1535]" captionStartId-1="221.[113,152,1087,1106]" captionTargetBox-0="[231,1047,219,1496]" captionTargetBox-1="[259,950,225,1062]" captionTargetId-0="figure-11@220.[231,1047,219,1497]" captionTargetId-1="figure-12@221.[229,974,216,1068]" captionTargetPageId-0="220" captionTargetPageId-1="221" captionText-0="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." captionText-1="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272302" figureDoi-1="http://doi.org/10.5281/zenodo.4272304" httpUri-0="https://zenodo.org/record/4272302/files/figure.png" httpUri-1="https://zenodo.org/record/4272304/files/figure.png" pageId="218" pageNumber="215">677–693</figureCitation>
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<emphasis id="B958FC5AFF4D09F361C0FAA2CF66FAA9" box="[151,332,1349,1367]" italics="true" pageId="218" pageNumber="215">Styphrus corpulentus</emphasis>
<bibRefCitation id="EFBD5DB9FF4D09F36006FAA3CC32FAA9" author="MOTSCHULSKY V." box="[337,536,1348,1367]" pageId="218" pageNumber="215" pagination="3 - 127" refId="ref118985" refString="MOTSCHULSKY V. 1845: Remarques sur la collection de Coleopteres Russes. 1 er Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou 18 (I): 3 - 127." type="journal article" year="1845">Motschulsky, 1845: 54</bibRefCitation>
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<treatmentCitation id="0A8D0659FF4D09F361C0FA87CC30FA8C" author="JAKOBSON G. G." box="[151,538,1375,1395]" page="651" pageId="218" pageNumber="215" year="1911">
<taxonomicName id="4C2C5BCBFF4D09F361C0FA87CFC4FA8C" ID-CoL="53BWP" authority=": JAKOBSON (1911)" authorityName=": JAKOBSON" authorityYear="1911" box="[151,494,1375,1395]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="218" pageNumber="215" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4D09F361C0FA87CF64FA8C" box="[151,334,1376,1394]" italics="true" pageId="218" pageNumber="215">Styphrus corpulentus</emphasis>
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<bibRefCitation id="EFBD5DB9FF4D09F3600CFA87CFC4FA8C" author="JAKOBSON G. G." box="[347,494,1375,1395]" pageId="218" pageNumber="215" pagination="641 - 720" refId="ref116036" refString="JAKOBSON G. G. 1911: Zhuki Rosii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. 9. [Beetles of Russia and western Europe. Manual for the beetles' determination. 9]. A. F. Devrien, St. Petersburg, pp. 1 + 641 - 720 (in Russian)." type="journal article" year="1911">JAKOBSON (1911)</bibRefCitation>
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<treatmentCitation id="0A8D0659FF4D09F36372FA87CCC7FA8C" author="REICHARDT A." box="[549,749,1375,1395]" page="174" pageId="218" pageNumber="215" year="1941">
<bibRefCitation id="EFBD5DB9FF4D09F36372FA87CCE8FA8C" author="REICHARDT A." box="[549,706,1375,1395]" pageId="218" pageNumber="215" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
: 174
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<figureCitation id="13173CCDFF4D09F363AFFAB8CD49FA8C" box="[760,867,1375,1394]" captionStart="Figs" captionStartId="34.[153,192,1459,1478]" captionTargetBox="[153,1128,576,1437]" captionTargetId="figure-144@34.[151,1128,576,1439]" captionTargetPageId="34" captionText="Figs. 81–89. Saprininae, mandibles, ventral view: 81 –Alienocacculus neftensis (Olexa, 1984); 82 – Ammostyphrus cerberus Reichardt, 1924; 83 – Chalcionellus amoenus (Erichson, 1834); 84 – Chivaenius kryzhanovskii Olexa, 1980; 85 – Ctenophilotis chobauti (Théry, 1900); 86 – Eopachylopus ripae (Lewis, 1885); 87 – Eremosaprinus vlasovi (Reichardt, 1941); 88 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 89 – Exaesiopus grossipes grossipes (Marseul, 1855)." figureDoi="http://doi.org/10.5281/zenodo.4272151" httpUri="https://zenodo.org/record/4272151/files/figure.png" pageId="218" pageNumber="215">Figs. 83, 84</figureCitation>
; 
<treatmentCitation id="0A8D0659FF4D09F3623AFA87CF36FA73" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." page="187" pageId="218" pageNumber="215" year="1976">
<bibRefCitation id="EFBD5DB9FF4D09F3623AFA87CEDBFA73" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." pageId="218" pageNumber="215" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
: 187
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, 
<figureCitation id="13173CCDFF4D09F36073FA9DCF8BFA73" box="[292,417,1402,1421]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="122.[151,190,1491,1510]" captionStartId-1="123.[113,152,1215,1234]" captionTargetBox-0="[234,1042,219,1469]" captionTargetBox-1="[169,1031,223,1190]" captionTargetId-0="figure-11@122.[234,1043,219,1469]" captionTargetId-1="figure-100@123.[157,1044,219,1199]" captionTargetPageId-0="122" captionTargetPageId-1="123" captionText-0="Figs. 360–370.Gnathoncus rotundatus (Kugelann, 1792), SEM micrographs:360 – habitus, dorsal view; 361 – ditto, ventral view; 362 – antennal club, ventral view; 363 – head, dorsal view; 364 – ditto, ventral view; 365 – mentum, ventral view; 366 – prosternum; 367 – mesoventrite and metaventrite; 368 – lateral disc of metaventrite, metepisternum and fused metepimeron; 369 – protibia, ventral view; 370 – mesotibia, ventral view." captionText-1="Figs. 371–378. Gnathoncus rotundatus (Kugelann, 1792), male terminalia (after ÔHARA 1994): 371 – 8th sternite and tergite, ventral view; 372 – ditto, dorsal view; 373 – ditto, lateral view; 374 – 9th tergite, 10th tergite, dorsal view; 375 – ditto, lateral view; 376 – spiculum gastrale, ventral view; 377 – aedeagus, dorsal view; 378 – ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272228" figureDoi-1="http://doi.org/10.5281/zenodo.4272230" httpUri-0="https://zenodo.org/record/4272228/files/figure.png" httpUri-1="https://zenodo.org/record/4272230/files/figure.png" pageId="218" pageNumber="215">Figs. 367–372</figureCitation>
; 
<treatmentCitation id="0A8D0659FF4D09F360FDFA9DCC6DFA73" author="MAZUR S." box="[426,583,1402,1421]" page="78" pageId="218" pageNumber="215" year="1984">
<bibRefCitation id="EFBD5DB9FF4D09F360FDFA9DCC0CFA73" author="MAZUR S." box="[426,550,1402,1421]" pageId="218" pageNumber="215" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 78
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF4D09F36318FA9DCCDDFA73" author="MAZUR S." box="[591,759,1402,1421]" page="245" pageId="218" pageNumber="215" year="1997">
<bibRefCitation id="EFBD5DB9FF4D09F36318FA9DCCE7FA73" author="MAZUR S." box="[591,717,1402,1421]" pageId="218" pageNumber="215" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 245
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF4D09F36257FA9DCD8DFA73" author="MAZUR S." box="[768,935,1402,1421]" page="101" pageId="218" pageNumber="215" year="2004">
<bibRefCitation id="EFBD5DB9FF4D09F36257FA9DCD56FA73" author="MAZUR S." box="[768,892,1402,1421]" pageId="218" pageNumber="215" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
: 101
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4D09F361C0FA72CD0DFA59" blockId="218.[151,1126,1348,1528]" box="[151,807,1428,1448]" pageId="218" pageNumber="215">
<treatmentCitationGroup id="AB3C0766FF4D09F361C0FA72CD0DFA59" box="[151,807,1428,1448]" pageId="218" pageNumber="215">
<treatmentCitation id="0A8D0659FF4D09F361C0FA72CFE3FA56" author="SCHMIDT J." box="[151,457,1428,1448]" page="19" pageId="218" pageNumber="215" year="1890">
<taxonomicName id="4C2C5BCBFF4D09F361C0FA72CFE3FA56" ID-CoL="9TJCF" authority="Schmidt, 1890: 19" authorityName="Schmidt" authorityPageNumber="19" authorityYear="1890" box="[151,457,1428,1448]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="218" pageNumber="215" phylum="Arthropoda" rank="species" species="akinini">
<emphasis id="B958FC5AFF4D09F361C0FA72CF09FA59" box="[151,291,1429,1447]" italics="true" pageId="218" pageNumber="215">Saprinus akinini</emphasis>
<bibRefCitation id="EFBD5DB9FF4D09F3607EFA73CFE3FA56" author="SCHMIDT J." box="[297,457,1428,1448]" pageId="218" pageNumber="215" pagination="81 - 87" refId="ref121039" refString="SCHMIDT J. 1890: Neun neue Saprinus aus der Gruppe des metallescens und rufipes. Deutsche Entomologische Zeitschrift 1890: 81 - 87." type="journal article" year="1890">Schmidt, 1890: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by 
<treatmentCitation id="0A8D0659FF4D09F36331FA72CD09FA59" author="JAKOBSON G." box="[614,803,1428,1448]" page="264" pageId="218" pageNumber="215" year="1910">
<bibRefCitation id="EFBD5DB9FF4D09F36331FA72CCDDFA56" author="JAKOBSON G." box="[614,759,1429,1448]" pageId="218" pageNumber="215" pagination="263 - 264" refId="ref116006" refString="JAKOBSON G. 1910: Note synonymique sur quelques Coleopteres palearctiques [Histeridae et Malacodermata]. Bulletin de la Societe Entomologique de France 1910: 263 - 264." type="journal article" year="1910">JAKOBSON (1910)</bibRefCitation>
: 264
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4D09F361C0FA57CD0DFA3C" blockId="218.[151,1126,1348,1528]" box="[151,807,1455,1475]" pageId="218" pageNumber="215">
<treatmentCitationGroup id="AB3C0766FF4D09F361C0FA57CD0DFA3C" box="[151,807,1455,1475]" pageId="218" pageNumber="215">
<treatmentCitation id="0A8D0659FF4D09F361C0FA57CFE3FA3C" author="REITTER E." box="[151,457,1455,1474]" page="29" pageId="218" pageNumber="215" year="1904">
<taxonomicName id="4C2C5BCBFF4D09F361C0FA57CFE3FA3C" ID-CoL="9TJCV" authority="Reitter, 1904: 29" authorityName="Reitter" authorityPageNumber="29" authorityYear="1904" box="[151,457,1455,1474]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="218" pageNumber="215" phylum="Arthropoda" rank="species" species="mimulus">
<emphasis id="B958FC5AFF4D09F361C0FA57CF1AFA3C" box="[151,304,1456,1474]" italics="true" pageId="218" pageNumber="215">Saprinus mimulus</emphasis>
<bibRefCitation id="EFBD5DB9FF4D09F36061FA48CFE3FA3C" author="REITTER E." box="[310,457,1455,1474]" pageId="218" pageNumber="215" pagination="29 - 36" refId="ref120480" refString="REITTER E. 1904: Ueber neue und wenig bekannte Histeriden (Coleoptera). Wiener Entomologische Zeitung 23: 29 - 36." type="journal article" year="1904">Reitter, 1904: 29</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by 
<treatmentCitation id="0A8D0659FF4D09F36331FA57CD09FA3C" author="JAKOBSON G." box="[614,803,1455,1475]" page="263" pageId="218" pageNumber="215" year="1910">
<bibRefCitation id="EFBD5DB9FF4D09F36331FA57CCDDFA3C" author="JAKOBSON G." box="[614,759,1455,1475]" pageId="218" pageNumber="215" pagination="263 - 264" refId="ref116006" refString="JAKOBSON G. 1910: Note synonymique sur quelques Coleopteres palearctiques [Histeridae et Malacodermata]. Bulletin de la Societe Entomologique de France 1910: 263 - 264." type="journal article" year="1910">JAKOBSON (1910)</bibRefCitation>
: 263
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4D09F361C0FA2CCFCBFA23" blockId="218.[151,1126,1348,1528]" box="[151,481,1482,1501]" pageId="218" pageNumber="215">
<treatmentCitationGroup id="AB3C0766FF4D09F361C0FA2CCFCBFA23" box="[151,481,1482,1501]" pageId="218" pageNumber="215">
<treatmentCitation id="0A8D0659FF4D09F361C0FA2CCFF7FA23" author="LEWIS G." box="[151,477,1482,1501]" page="78" pageId="218" pageNumber="215" year="1905">
<taxonomicName id="4C2C5BCBFF4D09F361C0FA2CCF97FA23" authority=": LEWIS (1905)" authorityName="LEWIS" authorityYear="1905" box="[151,445,1482,1501]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="218" pageNumber="215" phylum="Arthropoda" rank="species" species="akinini">
<emphasis id="B958FC5AFF4D09F361C0FA2CCF15FA23" box="[151,319,1483,1501]" italics="true" pageId="218" pageNumber="215">Xenonychus akinini</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF4D09F3601DFA2DCF97FA23" author="LEWIS G." box="[330,445,1482,1501]" pageId="218" pageNumber="215" refId="ref117719" refString="LEWIS G. 1905: A systematic catalogue of Histeridae. Kessinger, London, vi + 81 pp." type="book" year="1905">LEWIS (1905)</bibRefCitation>
</taxonomicName>
: 78
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4D09F361C0FA02CFD2FA09" blockId="218.[151,1126,1348,1528]" box="[151,504,1508,1528]" pageId="218" pageNumber="215">
<treatmentCitationGroup id="AB3C0766FF4D09F361C0FA02CFD2FA09" box="[151,504,1508,1528]" pageId="218" pageNumber="215">
<treatmentCitation id="0A8D0659FF4D09F361C0FA02CFDFFA09" author="REITTER E." box="[151,501,1508,1528]" page="268" pageId="218" pageNumber="215" year="1906">
<taxonomicName id="4C2C5BCBFF4D09F361C0FA02CFE3FA09" authority=": REITTER (1906)" authorityName="REITTER" authorityYear="1906" box="[151,457,1508,1528]" class="Insecta" family="Histeridae" genus="Pachylopus" kingdom="Animalia" order="Coleoptera" pageId="218" pageNumber="215" phylum="Arthropoda" rank="species" species="akinini">
<emphasis id="B958FC5AFF4D09F361C0FA02CF11FA09" box="[151,315,1509,1527]" italics="true" pageId="218" pageNumber="215">Pachylopus akinini</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF4D09F36010FA02CFE3FA09" author="REITTER E." box="[327,457,1508,1528]" pageId="218" pageNumber="215" refId="ref120504" refString="REITTER E. 1906: Histeridae. In: HEYDEN L. V., REITTER E. &amp; WEISE J. (eds.): Catalogus Coleopterorum Europae, Caucasi et Armeniae Rossicae. Editio secunda. Berlin, Paskau, Caen, vi + 775 pp." type="book" year="1906">REITTER (1906)</bibRefCitation>
</taxonomicName>
: 268
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4C09F26126FF3CCD16FF11" blockId="219.[113,1088,219,319]" box="[113,828,219,239]" pageId="219" pageNumber="212">
<treatmentCitationGroup id="AB3C0766FF4C09F26126FF3CCD16FF11" box="[113,828,219,239]" pageId="219" pageNumber="212">
<treatmentCitation id="0A8D0659FF4C09F26126FF3CCFF8FF11" author="REITTER E." box="[113,466,219,239]" page="13" pageId="219" pageNumber="212" year="1910">
<taxonomicName id="4C2C5BCBFF4C09F26126FF3CCFF8FF11" ID-CoL="9TJDW" authority="Reitter, 1910: 13" authorityName="Reitter" authorityPageNumber="13" authorityYear="1910" box="[113,466,219,239]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="laevidorsis">
<emphasis id="B958FC5AFF4C09F26126FF3CCF16FF10" box="[113,316,219,238]" italics="true" pageId="219" pageNumber="212">Xenonychus laevidorsis</emphasis>
<bibRefCitation id="EFBD5DB9FF4C09F26016FF3CCFF8FF11" author="REITTER E." box="[321,466,219,239]" pageId="219" pageNumber="212" pagination="13" refId="ref120599" refString="REITTER E. 1910: Uebersicht der Arten der Coleopterengattung Xenonychus Woll. Entomologische Blatter 6: 13." type="journal article" year="1910">Reitter, 1910: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by 
<treatmentCitation id="0A8D0659FF4C09F26326FF3BCD12FF11" author="BICKHARDT H." box="[625,824,219,239]" page="103" pageId="219" pageNumber="212" year="1916">
<bibRefCitation id="EFBD5DB9FF4C09F26326FF3BCD27FF10" author="BICKHARDT H." box="[625,781,219,239]" pageId="219" pageNumber="212" refId="ref112724" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum, Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
: 103
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4C09F26126FF11CFDCFEF7" blockId="219.[113,1088,219,319]" box="[113,502,246,266]" pageId="219" pageNumber="212">
<treatmentCitationGroup id="AB3C0766FF4C09F26126FF11CFDCFEF7" box="[113,502,246,266]" pageId="219" pageNumber="212">
<treatmentCitation id="0A8D0659FF4C09F26126FF11CFDBFEF7" author="BICKHARDT H." box="[113,497,246,266]" page="107" pageId="219" pageNumber="212" year="1910">
<taxonomicName id="4C2C5BCBFF4C09F26126FF11CFECFEF7" authority=": BICKHARDT (1910)" authorityName="BICKHARDT" authorityYear="1910" box="[113,454,246,266]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="laevidorsis">
<emphasis id="B958FC5AFF4C09F26126FF11CF35FEF7" box="[113,287,246,265]" italics="true" pageId="219" pageNumber="212">Styphrus laevidorsis</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF4C09F2607DFF10CFECFEF7" author="BICKHARDT H." box="[298,454,246,266]" pageId="219" pageNumber="212" refId="ref112602" refString="BICKHARDT H. 1910: Histeridae. In: JUNK W. &amp; SCHENKLING S. (eds.): Coleopterorum Catalogus, volume 24. W. Junk, Berlin, 137 pp." type="book" year="1910">BICKHARDT (1910)</bibRefCitation>
</taxonomicName>
: 107
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF4C09F26126FEF6CEDCFEC1" blockId="219.[113,1088,219,319]" pageId="219" pageNumber="212">
<treatmentCitationGroup id="AB3C0766FF4C09F26126FEF6CEDCFEC1" pageId="219" pageNumber="212">
<taxonomicName id="4C2C5BCBFF4C09F26126FEF6CFD8FEDA" ID-CoL="9TJD4" authority="Dahlgren, 1971: 50" authorityName="Dahlgren" authorityPageNumber="50" authorityYear="1971" box="[113,498,273,292]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="rugosipennis">
<emphasis id="B958FC5AFF4C09F26126FEF6CF12FEDD" box="[113,312,273,292]" italics="true" pageId="219" pageNumber="212">Saprinus rugosipennis</emphasis>
Dahlgren, 1971: 50
</taxonomicName>
. Synonymized by Kryzhanovskij in 
<treatmentCitation id="0A8D0659FF4C09F26268FEF6CEDBFEC1" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." page="187" pageId="219" pageNumber="212" year="1976">
<bibRefCitation id="EFBD5DB9FF4C09F26268FEF6CEE3FEC0" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." pageId="219" pageNumber="212" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
: 187
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF4C09F26126FEB2CE80FE10" pageId="219" pageNumber="212" type="materials_examined">
<paragraph id="8B932048FF4C09F26126FEB2CF80FE96" blockId="219.[113,1089,340,707]" box="[113,426,340,360]" pageId="219" pageNumber="212">
<emphasis id="B958FC5AFF4C09F26126FEB2CEC7FE96" bold="true" box="[113,237,340,360]" pageId="219" pageNumber="212">
<typeStatus id="54979EEAFF4C09F26126FEB2CEB5FE96" box="[113,159,341,360]" pageId="219" pageNumber="212">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B60D8FF4C09F261A5FEB2CF7EFE96" box="[242,340,341,360]" name="Uzbekistan" pageId="219" pageNumber="212">Uzbekistan</collectingCountry>
, Buchara.
</paragraph>
<paragraph id="8B932048FF4C09F26126FE97CCE9FE46" blockId="219.[113,1089,340,707]" pageId="219" pageNumber="212">
<emphasis id="B958FC5AFF4C09F26126FE97CF79FE7D" bold="true" box="[113,339,367,387]" pageId="219" pageNumber="212">Type material examined.</emphasis>
<taxonomicName id="4C2C5BCBFF4C09F26000FE97CC21FE7D" authorityName=": JAKOBSON" authorityYear="1911" box="[343,523,368,387]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4C09F26000FE97CC21FE7D" box="[343,523,368,387]" italics="true" pageId="219" pageNumber="212">Styphrus corpulentus</emphasis>
</taxonomicName>
. 
<materialsCitation id="3B442A15FF4C09F26343FE97CCCEFE60" ID-GBIF-Occurrence="2981920385" country="Uzbekistan" location="Buchara" municipality="Kyzyl Kum" pageId="219" pageNumber="212" specimenCount="1" typeStatus="neotype">
<typeStatus id="54979EEAFF4C09F26343FE97CC76FE7C" box="[532,604,368,387]" pageId="219" pageNumber="212" type="neotype">NEOTYPE</typeStatus>
(here designated):♁, ‘ 
<location id="8EF37693FF4C09F2624DFE97CD4DFE7D" LSID="urn:lsid:plazi:treatment:0385915EFF4D09F660CBFB1CCD34FE2D:8EF37693FF4C09F2624DFE97CD4DFE7D" box="[794,871,368,387]" country="Uzbekistan" municipality="Kyzyl Kum" name="Buchara" pageId="219" pageNumber="212">Buchara</location>
, 
<collectingMunicipality id="6BF7BA32FF4C09F26238FE97CDF8FE7D" box="[879,978,368,387]" pageId="219" pageNumber="212">Kyzyl Kum</collectingMunicipality>
/ 
<collectingCountry id="F33B60D8FF4C09F262B6FE97CA6BFE7D" box="[993,1089,368,387]" name="Uzbekistan" pageId="219" pageNumber="212">Uzbekistan</collectingCountry>
28.–30.IV. / A. 
<bibRefCitation id="EFBD5DB9FF4C09F261A9FE6CCF49FE63" author="OLEXA A." box="[254,355,394,414]" pageId="219" pageNumber="212" pagination="21" refId="ref119438" refString="OLEXA A. 1975: Sur la variabilite de Saprinus perrisi Marseul, 1871 (Col. Histeridae). Bulletin de la Societe Entomologique de Mulhouse 31: 21." type="journal article" year="1975">Olexa 1975</bibRefCitation>
[printed] // D07-026 [pink label, written] //
</materialsCitation>
<typeStatus id="54979EEAFF4C09F263BCFE6CCD70FE60" box="[747,858,395,414]" pageId="219" pageNumber="212" type="neotype">NEOTYPUS</typeStatus>
/ 
<taxonomicName id="4C2C5BCBFF4C09F26238FE6CCD90FE60" authorityName=": JAKOBSON" authorityYear="1911" box="[879,954,395,414]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF4C09F26238FE6CCD90FE60" box="[879,954,395,414]" italics="true" pageId="219" pageNumber="212">Styphrus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C2C5BCBFF4C09F26298FE6CCA1EFE60" authorityName=": JAKOBSON" authorityYear="1911" box="[975,1076,395,414]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4C09F26298FE6CCA1EFE60" box="[975,1076,395,414]" italics="true" pageId="219" pageNumber="212">corpulentus</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFBD5DB9FF4C09F26126FE42CF3AFE46" author="MOTSCHULSKY V." box="[113,272,421,440]" pageId="219" pageNumber="212" pagination="3 - 127" refId="ref118985" refString="MOTSCHULSKY V. 1845: Remarques sur la collection de Coleopteres Russes. 1 er Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou 18 (I): 3 - 127." type="journal article" year="1845">Motschulsky 1845</bibRefCitation>
/ Des. T. Lackner 2009 [red label, written]’ (
<collectionCode id="ED3DB88DFF4C09F263C6FE42CC93FE46" box="[657,697,421,440]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="219" pageNumber="212" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B932048FF4C09F261C7FE27CE80FE10" blockId="219.[113,1089,340,707]" pageId="219" pageNumber="212">
<taxonomicName id="4C2C5BCBFF4C09F261C7FE27CF31FE2D" authorityName="Schmidt" authorityYear="1890" box="[144,283,448,467]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="akinini">
<emphasis id="B958FC5AFF4C09F261C7FE27CF31FE2D" box="[144,283,448,467]" italics="true" pageId="219" pageNumber="212">Saprinus akinini</emphasis>
</taxonomicName>
. Par at ype: 
<specimenCount id="9D2AEBC1FF4C09F260D7FE27CF97FE2D" box="[384,445,448,467]" pageId="219" pageNumber="212" type="generic">1 spec.</specimenCount>
, ‘ 
<taxonomicName id="4C2C5BCBFF4C09F2609CFE27CC3DFE2D" authorityName=": LACORDAIRE" authorityYear="1854" box="[459,535,448,467]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF4C09F2609CFE27CC3DFE2D" box="[459,535,448,467]" italics="true" pageId="219" pageNumber="212">Saprinus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C2C5BCBFF4C09F26372FE27CC4AFE2D" authorityName="Schmidt" authorityYear="1890" box="[549,608,448,467]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="akinini">
<emphasis id="B958FC5AFF4C09F26372FE27CC4AFE2D" box="[549,608,448,467]" italics="true" pageId="219" pageNumber="212">akinini</emphasis>
</taxonomicName>
/ Schmidt / typ [written] // 
<typeStatus id="54979EEAFF4C09F26213FE27CDB2FE2D" box="[836,920,448,467]" pageId="219" pageNumber="212" type="paratype">Paratypus</typeStatus>
[red label, written]’ (
<collectionCode id="ED3DB88DFF4C09F2612FFE3CCE8AFE10" box="[120,160,475,494]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="219" pageNumber="212" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF4C09F26126FE13CFB3FD3D" pageId="219" pageNumber="212" type="discussion">
<paragraph id="8B932048FF4C09F26126FE13CC24FD8D" blockId="219.[113,1089,340,707]" pageId="219" pageNumber="212">
<emphasis id="B958FC5AFF4C09F26126FE13CEE5FDF6" bold="true" box="[113,207,500,520]" pageId="219" pageNumber="212">Comment.</emphasis>
Type specimen(s) of this species were not found in Motschulsky’s collection,housed at ZMUM.According to N. Nikitsky (ZMUM), a part of the type specimens described by Motschulsky in 1845 has been destroyed and lost. Perhaps the type specimen(s) of 
<taxonomicName id="4C2C5BCBFF4C09F260E3FDCCCC40FDC0" authorityName=": JAKOBSON" authorityYear="1911" box="[436,618,555,574]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4C09F260E3FDCCCC40FDC0" box="[436,618,555,574]" italics="true" pageId="219" pageNumber="212">Styphrus corpulentus</emphasis>
</taxonomicName>
were among those. The Motschulsky’s collection is in a very poor state. Therefore, it is necessary to designate a 
<typeStatus id="54979EEAFF4C09F26337FDA1CC8EFDA7" box="[608,676,582,601]" pageId="219" pageNumber="212" type="neotype">neotype</typeStatus>
for this species to fix its identity as type species of 
<taxonomicName id="4C2C5BCBFF4C09F261DFFD87CEF9FD8D" authorityName=": JAKOBSON" authorityYear="1911" box="[136,211,608,627]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="219" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF4C09F261DFFD87CEF9FD8D" box="[136,211,608,627]" italics="true" pageId="219" pageNumber="212">Styphrus</emphasis>
</taxonomicName>
. The 
<typeStatus id="54979EEAFF4C09F26053FD87CF62FD8D" box="[260,328,608,627]" pageId="219" pageNumber="212" type="neotype">neotype</typeStatus>
will be housed at ZIN.
</paragraph>
<paragraph id="8B932048FF4C09F26126FD9DCFB3FD3D" blockId="219.[113,1089,340,707]" pageId="219" pageNumber="212">
<emphasis id="B958FC5AFF4C09F26126FD9DCFA3FD70" bold="true" box="[113,393,634,654]" pageId="219" pageNumber="212">Additional material examined.</emphasis>
<materialsCitation id="3B442A15FF4C09F260D8FD9DCA23FD70" ID-GBIF-Occurrence="2981920368" box="[399,1033,632,655]" collectingDate="1975-04-28" collectingDateMax="1975-04-30" collectingDateMin="1975-04-28" collectorName="A. Olexa" country="Uzbekistan" location="Kyzylkum" municipality="Buchara" pageId="219" pageNumber="212" specimenCount="1">
<emphasis id="B958FC5AFF4C09F260D8FD9DCC09FD70" bold="true" box="[399,547,634,654]" pageId="219" pageNumber="212">
<collectingCountry id="F33B60D8FF4C09F260D8FD9DCC34FD70" box="[399,542,634,654]" name="Uzbekistan" pageId="219" pageNumber="212">UZBEKISTAN</collectingCountry>
:
</emphasis>
<location id="8EF37693FF4C09F2637EFD9CCCAFFD70" LSID="urn:lsid:plazi:treatment:0385915EFF4D09F660CBFB1CCD34FE2D:8EF37693FF4C09F2637EFD9CCCAFFD70" box="[553,645,635,654]" country="Uzbekistan" municipality="Buchara" name="Kyzylkum" pageId="219" pageNumber="212">Kyzylkum</location>
, 
<collectingMunicipality id="6BF7BA32FF4C09F263D9FD9CCCFDFD70" box="[654,727,635,654]" pageId="219" pageNumber="212">Buchara</collectingMunicipality>
, 
<date id="FF920688FF4C09F263B6FD9DCD4EFD73" box="[737,868,634,653]" pageId="219" pageNumber="212" value="1975-04-28" valueMax="1975-04-30" valueMin="1975-04-28">
<collectingDate id="EFD6FF60FF4C09F263B6FD9DCD4EFD73" box="[737,868,634,653]" pageId="219" pageNumber="212" value="1975-04-28" valueMax="1975-04-30" valueMin="1975-04-28">28.–30.iv.1975</collectingDate>
</date>
, 1 ♁, 
<collectorName id="26D9459EFF4C09F262CDFD9CCDC2FD70" box="[922,1000,635,654]" pageId="219" pageNumber="212">A. Olexa</collectorName>
lgt
</materialsCitation>
.; Kyzyk-Kum, Chiva [= Khiva], 
<date id="FF920688FF4C09F2600AFD72CF90FD56" box="[349,442,661,680]" pageId="219" pageNumber="212" value="1972-04-25">25.iv.1972</date>
, 1 ♁ 
<specimenCount id="9D2AEBC1FF4C09F260BDFD72CC21FD57" box="[490,523,658,681]" pageId="219" pageNumber="212" type="female">1 ♀</specimenCount>
, A. Olexa lgt.; 
<materialsCitation id="3B442A15FF4C09F263C5FD72CDCCFD57" ID-GBIF-Occurrence="2981920356" box="[658,998,658,681]" collectingDate="1978-05-03" collectingDateMax="1975-04-30" collectingDateMin="1975-04-28" collectorName="Olexa" country="Uzbekistan" location="Kyzylkum" municipality="Buchara" pageId="219" pageNumber="212" specimenCount="3" specimenCount-female="3">
ditto, but 
<date id="FF920688FF4C09F263B2FD72CD18FD56" box="[741,818,661,680]" pageId="219" pageNumber="212" value="1978-05-03">
<collectingDate id="EFD6FF60FF4C09F263B2FD72CD18FD56" box="[741,818,661,680]" pageId="219" pageNumber="212" value="1978-05-03">3.v.1978</collectingDate>
</date>
, A. 
<collectorName id="26D9459EFF4C09F26201FD72CDA3FD56" box="[854,905,661,680]" pageId="219" pageNumber="212">Olexa</collectorName>
lgt., 
<specimenCount id="9D2AEBC1FF4C09F262E0FD72CDCCFD57" box="[951,998,658,681]" pageId="219" pageNumber="212" type="female">3 ♀♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B442A15FF4C09F262A4FD72CFBFFD3D" ID-GBIF-Occurrence="2981920387" collectingDate="1972-04-23" collectingDateMax="1975-04-30" collectingDateMin="1975-04-28" collectionCode="TLAN" collectorName="A. Olexa" country="Uzbekistan" location="Kyzylkum" municipality="Buchara" pageId="219" pageNumber="212" specimenCount="1">
ditto, but 
<date id="FF920688FF4C09F26126FD57CEE5FD3D" box="[113,207,688,707]" pageId="219" pageNumber="212" value="1972-04-23">
<collectingDate id="EFD6FF60FF4C09F26126FD57CEE5FD3D" box="[113,207,688,707]" pageId="219" pageNumber="212" value="1972-04-23">23.iv.1972</collectingDate>
</date>
, 
<collectorName id="26D9459EFF4C09F26180FD57CF0EFD3D" box="[215,292,688,707]" pageId="219" pageNumber="212">A. Olexa</collectorName>
lgt. (
<collectionCode id="ED3DB88DFF4C09F26006FD57CFA5FD3D" box="[337,399,688,707]" pageId="219" pageNumber="212">TLAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF4C09F66126FD3CCD34FE2D" lastPageId="223" lastPageNumber="216" pageId="219" pageNumber="212" type="description">
<paragraph id="8B932048FF4C09F26126FD3CCC70FCED" blockId="219.[113,1089,731,1620]" pageId="219" pageNumber="212">
<emphasis id="B958FC5AFF4C09F26126FD3CCF3DFD0D" bold="true" box="[113,279,731,755]" pageId="219" pageNumber="212">Redescription.</emphasis>
Body length: PEL: 
<quantity id="4CD48DADFF4C09F260BCFD3CCCA3FD0D" box="[491,649,731,755]" metricMagnitude="-3" metricUnit="m" metricValue="2.975" metricValueMax="3.45" metricValueMin="2.5" pageId="219" pageNumber="212" unit="mm" value="2.975" valueMax="3.45" valueMin="2.5">2.50–3.45 mm</quantity>
; APW: 
<quantity id="4CD48DADFF4C09F2638BFD3CCD50FD0D" box="[732,890,731,755]" metricMagnitude="-3" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="219" pageNumber="212" unit="mm" value="1.025" valueMax="1.2" valueMin="0.85">0.85–1.20 mm</quantity>
; PPW: 
<quantity id="4CD48DADFF4C09F2629EFD3CCEB4FCED" metricMagnitude="-3" metricUnit="m" metricValue="2.3875" metricValueMax="2.8" metricValueMin="1.975" pageId="219" pageNumber="212" unit="mm" value="2.3875" valueMax="2.8" valueMin="1.975">1.975–2.80 mm</quantity>
; EL: 
<quantity id="4CD48DADFF4C09F2618FFD1CCF5CFCED" box="[216,374,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.25" metricValueMin="1.55" pageId="219" pageNumber="212" unit="mm" value="1.9" valueMax="2.25" valueMin="1.55">1.55–2.25 mm</quantity>
; EW: 
<quantity id="4CD48DADFF4C09F260EFFD1BCC7CFCED" box="[440,598,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.1" metricValueMin="2.2" pageId="219" pageNumber="212" unit="mm" value="2.65" valueMax="3.1" valueMin="2.2">2.20–3.10 mm</quantity>
.
</paragraph>
<paragraph id="8B932048FF4C09F261C7FCFCCF76FC8D" blockId="219.[113,1089,731,1620]" pageId="219" pageNumber="212">
Body (
<figureCitation id="13173CCDFF4C09F2618EFCFCCF5CFCCD" box="[217,374,795,819]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Figs. 677–678</figureCitation>
) roundly oval, ventrally strongly convex, moderately convex dorsally; cuticle dark brown with slight metallic tinge, elytral apex rufous; legs, mouthparts and antennae rufopiceous.
</paragraph>
<paragraph id="8B932048FF4C09F261C7FC9CCF8EFBAD" blockId="219.[113,1089,731,1620]" pageId="219" pageNumber="212">
Antennal scape (
<figureCitation id="13173CCDFF4C09F26014FC9CCFB4FC6D" box="[323,414,891,915]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Fig. 681</figureCitation>
) substrigulate, with numerous well sclerotized long setae; club (
<figureCitation id="13173CCDFF4C09F2612EFC7CCF3CFC4D" box="[121,278,923,947]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Figs. 679–680</figureCitation>
) large, flattened dorso-ventrally, round, without visible articulation, dorsal surface with large apical sensory area with dense short sensilla, intermingled with longer, much sparser erect sensilla; surface of club otherwise imbricate, with sparse long erect sensilla (except for basal fourth); sensory structures of antennal club (
<figureCitation id="13173CCDFF4C09F2625DFC1CCD72FBEA" box="[778,856,1019,1044]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="219" pageNumber="212">Fig. 12</figureCitation>
) in form of one large apical sensory area and four rather large ventral sensory areas and one vesicle on internal distal margin of ventral side.
</paragraph>
<paragraph id="8B932048FF4C09F261C7FBBCCC92FA8D" blockId="219.[113,1089,731,1620]" pageId="219" pageNumber="212">
Mouthparts. Mandibles (
<figureCitation id="13173CCDFF4C09F260C5FBBCCFC1FB8A" box="[402,491,1115,1140]" captionStart="Figs" captionStartId="36.[151,190,1161,1180]" captionTargetBox="[151,1126,219,1130]" captionTargetId="figure-100@36.[151,1126,219,1132]" captionTargetPageId="36" captionText="Figs.99–107.Saprininae, mandibles,ventral view:99 – Philothis (Farabius) hexeris Reichardt, 1930; 100 – Pholioxenus phoenix (Reichardt,1930); 101 – Reichardtiolus duriculus (Reitter, 1904); 102 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 103 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 104 – Saprinus (Phaonius) pharao (Marseul, 1855); 105 – Styphrus corpulentus Motschulsky, 1845; 106 – Xenonychus tridens (Jacquelin-Duval, 1852); 107 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4272155" httpUri="https://zenodo.org/record/4272155/files/figure.png" pageId="219" pageNumber="212">Fig. 104</figureCitation>
) with rounded outer margin, mandibular apex acute; subapical tooth of left mandible moderately large; labrum (
<figureCitation id="13173CCDFF4C09F26392FB9CCD3EFB6A" box="[709,788,1147,1172]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="219" pageNumber="212">Fig. 70</figureCitation>
) with scattered microscopic punctation, convex, with a slight median depression; labral pits with two weakly sclerotized setae; terminal labial palpomere thickened, its width about half its length; mentum (
<figureCitation id="13173CCDFF4C09F26288FB5CCA13FB2D" box="[991,1081,1211,1235]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Fig. 683</figureCitation>
) square-shaped, imbricate, anterior margin medially with conspicuous notch (
<figureCitation id="13173CCDFF4C09F262DDFB3CCDCEFB0D" box="[906,996,1243,1267]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="219" pageNumber="212">Fig. 136</figureCitation>
), fringed with numerous long setae, lateral margins as well as disc of mentum with numerous much shorter setae; cardo of maxilla on lateral margin with several short setae; stipes triangular, with three much longer setae; terminal maxillary palpomere thickened, its width about half its length, approximately twice as long as penultimate.
</paragraph>
<paragraph id="8B932048FF4C09F261C7FA9CCD9BFA0D" blockId="219.[113,1089,731,1620]" pageId="219" pageNumber="212">
Clypeus (
<figureCitation id="13173CCDFF4C09F261A2FA9CCF64FA6D" box="[245,334,1403,1427]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Fig. 681</figureCitation>
) rectangular, basally constricted, depressed laterally, with shallow irregular punctures; frontal stria well impressed, usually interrupted medially, shortly prolonged onto clypeus; supraorbital stria well impressed, carinate; frontal disc (
<figureCitation id="13173CCDFF4C09F26218FA5CCD83FA2D" box="[847,937,1467,1491]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Fig. 681</figureCitation>
) smooth, only with scattered microscopic punctation; eyes convex, conspicuous from above.
</paragraph>
<paragraph id="8B932048FF4C09F461C7FA1CCC6FFAAB" blockId="219.[113,1089,731,1620]" lastBlockId="221.[113,1089,1309,1621]" lastPageId="221" lastPageNumber="214" pageId="219" pageNumber="212">
Pronotal sides (
<figureCitation id="13173CCDFF4C09F26061FA1CCFBBF9ED" box="[310,401,1531,1555]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="219" pageNumber="212">Fig. 677</figureCitation>
) moderately narrowed anteriorly, apical angles obtuse; pronotal foveae vaguely impressed, often absent; marginal pronotal stria complete, slightly weakened behind head; pronotal disc (except for inconspicuous scattered punctation along lateral margins and a double row of small ellipsoid punctures along pronotal base) entirely smooth; pronotal hypomeron setose; scutellum small, visible.
</paragraph>
<caption id="DF5370C0FF4B09F561C0FA0BCFA0F9B1" ID-DOI="http://doi.org/10.5281/zenodo.4272302" ID-Zenodo-Dep="4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="220" pageNumber="217" startId="220.[151,190,1516,1535]" targetBox="[231,1047,219,1496]" targetPageId="220">
<paragraph id="8B932048FF4B09F561C0FA0BCFA0F9B1" blockId="220.[151,1129,1516,1615]" pageId="220" pageNumber="217">
Figs.677–686. 
<taxonomicName id="4C2C5BCBFF4B09F5604DFA0ACC5CFA01" authority="Motschulsky, 1845" authorityName="Motschulsky" authorityYear="1845" box="[282,630,1516,1535]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="217" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4B09F5604DFA0ACFE4FA01" box="[282,462,1517,1535]" italics="true" pageId="220" pageNumber="217">Styphrus corpulentus</emphasis>
<bibRefCitation id="EFBD5DB9FF4B09F56086FA0BCC5CFA01" author="MOTSCHULSKY V." box="[465,630,1516,1535]" pageId="220" pageNumber="217" pagination="3 - 127" refId="ref118985" refString="MOTSCHULSKY V. 1845: Remarques sur la collection de Coleopteres Russes. 1 er Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou 18 (I): 3 - 127." type="journal article" year="1845">Motschulsky, 1845</bibRefCitation>
</taxonomicName>
, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view.
</paragraph>
</caption>
<caption id="DF5370C0FF4A09F46126FBD8CF34FB5C" ID-DOI="http://doi.org/10.5281/zenodo.4272304" ID-Zenodo-Dep="4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="221" pageNumber="214" startId="221.[113,152,1087,1106]" targetBox="[259,950,225,1062]" targetPageId="221">
<paragraph id="8B932048FF4A09F46126FBD8CF34FB5C" blockId="221.[113,1089,1085,1186]" pageId="221" pageNumber="214">
Figs.687–693. 
<taxonomicName id="4C2C5BCBFF4A09F461A4FBA7CC64FBAC" authority="Motschulsky, 1845" authorityName="Motschulsky" authorityYear="1845" box="[243,590,1087,1106]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="221" pageNumber="214" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B958FC5AFF4A09F461A4FBA7CF8CFBAC" box="[243,422,1088,1106]" italics="true" pageId="221" pageNumber="214">Styphrus corpulentus</emphasis>
<bibRefCitation id="EFBD5DB9FF4A09F460FDFBD8CC64FBAC" author="MOTSCHULSKY V." box="[426,590,1087,1106]" pageId="221" pageNumber="214" pagination="3 - 127" refId="ref118985" refString="MOTSCHULSKY V. 1845: Remarques sur la collection de Coleopteres Russes. 1 er Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou 18 (I): 3 - 127." type="journal article" year="1845">Motschulsky, 1845</bibRefCitation>
</taxonomicName>
, male terminalia: 687 – 8 
<superScript id="7C598D00FF4A09F46277FBDACD00FBB7" attach="left" box="[800,810,1085,1097]" fontSize="5" pageId="221" pageNumber="214">th</superScript>
sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9 
<superScript id="7C598D00FF4A09F46308FBBFCC43FB9A" attach="left" box="[607,617,1112,1124]" fontSize="5" pageId="221" pageNumber="214">th</superScript>
tergite, 10 
<superScript id="7C598D00FF4A09F46393FBBFCCE4FB9A" attach="left" box="[708,718,1112,1124]" fontSize="5" pageId="221" pageNumber="214">th</superScript>
tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9 
<superScript id="7C598D00FF4A09F46060FB94CF6BFB81" attach="left" box="[311,321,1139,1151]" fontSize="5" pageId="221" pageNumber="214">th</superScript>
tergite, 10 
<superScript id="7C598D00FF4A09F460C9FB94CF82FB81" attach="left" box="[414,424,1139,1151]" fontSize="5" pageId="221" pageNumber="214">th</superScript>
tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8B932048FF4A09F761C7FABACD31FEED" blockId="221.[113,1089,1309,1621]" lastBlockId="222.[151,1127,219,1619]" lastPageId="222" lastPageNumber="219" pageId="221" pageNumber="214">Elytral epipleura shiny, almost smooth, with scattered microscopic punctures; marginal epipleural stria weakly impressed, complete; marginal elytral stria well impressed, continued as weakly impressed complete apical elytral stria; inner subhumeral stria well impressed, rather long, apically reaching as far as four-fifths of elytral length, basally connected with weakly impressed humeral elytral stria; elytra with four rather thin dorsal elytral striae 1–4, apically reaching about elytral half, occasionally second dorsal elytral stria slightly longer; fourth dorsal elytral stria basally connected with complete sutural elytral stria; sutural elytral stria well impressed, complete, apically connected with apical elytral stria. Elytral disc on apical half with dense, shallow, often confluent punctures, basal half with much finer and sparser punctation; elytral flanks and elytral humeri smooth.</paragraph>
<paragraph id="8B932048FF4909F761E2FEFCCD48FEAD" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">Propygidium almost completely exposed; punctation of propygidium and pygidium shallow but dense, punctures confluent, becoming more granulose apically.</paragraph>
<paragraph id="8B932048FF4909F761E2FEBCCDF5FDEA" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">
Anterior margin of median portion of prosternum (
<figureCitation id="13173CCDFF4909F763B7FEBCCD17FE8D" box="[736,829,347,371]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="222" pageNumber="219">Fig. 685</figureCitation>
) almost straight, pre-apical foveae absent; marginal prosternal stria present laterally and as thin apical fragment; prosternal process flattened; surface between carinal prosternal striae smooth; laterally substrigulate, with sparse punctures fringed with setae; carinal prosternal striae on prosternal apophysis slightly divergent, approximate, sub-parallel, slightly divergent apically, not united; lateral prosternal striae well impressed, shortened, slightly carinate, anteriorly ‘open’.
</paragraph>
<paragraph id="8B932048FF4909F761E2FDFCCD48FCED" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">Anterior margin of mesoventrite straight; discal marginal mesoventral stria well impressed, slightly carinate; disc of mesoventrite flat, with shallow scattered punctation, few punctures with microscopic setae; meso-metaventral sutural stria well impressed, undulate; intercoxal disc of metaventrite convex, almost smooth, along posterior margin with irregular sparse fine punctures. Lateral metaventral stria well impressed, slightly carinate, almost straight, shortened apically; lateral disc of metaventrite concave, with scattered shallow punctures fringed with long setae; metepisternum on basal three-fourths with similar punctation and setae, on apical fourth + fused metepimeron punctation and setae sparser and finer.</paragraph>
<paragraph id="8B932048FF4909F761E2FCFCCCE2FC8D" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">Intercoxal disc of the first abdominal sternite completely striate laterally; surface of disc with sparse scattered punctation, interspaces imbricate, apically punctation sparser and finer; lateral sides of all visible abdominal sternites setose.</paragraph>
<paragraph id="8B932048FF4909F761E2FC9CCF7BFB2D" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">
Protibia (
<figureCitation id="13173CCDFF4909F76040FC9CCF5BFC6D" box="[279,369,891,915]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="222" pageNumber="219">Fig. 686</figureCitation>
) dilated, outer margin with three low teeth topped with rather long acute denticle, second and third teeth larger than first, followed by six short denticles diminishing in size in proximal direction; setae of outer row regular, rather sparse; protarsal groove deep; anterior protibial stria absent; setae of median row much shorter and sparser than those of outer row, present only on basal half; single tiny tarsal denticle present near tarsal insertion; protibial spur well developed, bent, growing out from apical margin of protibia; apical margin of protibia posteriorly with three short denticles; outer part of posterior surface of protibia (
<figureCitation id="13173CCDFF4909F761ABFBBCCF71FB8D" box="[252,347,1115,1139]" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="222" pageNumber="219">Fig. 686</figureCitation>
) broadly strigate, interspaces smooth, separated from glabrous median part of posterior surface by clear-cut boundary; posterior protibial stria complete, apically terminating in two tiny inner ventral denticles; inner row of setae lamelliform, strongly sclerotized, long.
</paragraph>
<paragraph id="8B932048FF4909F761E2FB3CCFF6F9AD" blockId="222.[151,1127,219,1619]" pageId="222" pageNumber="219">
Mesotibia slender, outer margin with single row of short thin denticles, only slightly growing in size in proximal direction; setae of outer row situated approximately in middle of posterior mesotibial surface, setae regular, dense, about as long as denticles themselves; setae of median row absent; posterior mesotibial stria shortened apically; anterior surface of mesotibia microscopically punctate; anterior mesotibial stria almost complete; inner anterior denticles absent; mesotibial spur rather long and stout; apical margin of mesotibia anteriorly with two tiny denticles; inner margin of mesotibia with long dense well sclerotized setae; claws of apical tarsomere almost straight, thin, almost as long as apical-most tarsomere itself; each mesotarsomere anteriorly with single long well sclerotized seta; metatibia (
<figureCitation id="13173CCDFF4909F7656CFA3CCEEAF9ED" captionStart="Figs" captionStartId="220.[151,190,1516,1535]" captionTargetBox="[231,1047,219,1496]" captionTargetId="figure-11@220.[231,1047,219,1497]" captionTargetPageId="220" captionText="Figs.677–686. Styphrus corpulentus Motschulsky, 1845, SEM micrographs:677 – habitus, dorsal view; 678 – ditto, ventral view; 679 – antennal club, dorsal view; 680 – ditto, ventral view; 681 – head, dorsal view; 682 – ditto, ventral view; 683 – mentum, cardines and stipites of maxilla, ventral view; 684 – metatibia, dorsal view; 685 – prosternum; 686 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272302" httpUri="https://zenodo.org/record/4272302/files/figure.png" pageId="222" pageNumber="219">Fig. 684</figureCitation>
) similar to mesotibia, but slightly more thickened and another row of four sparse longer denticles appears on apical half of outer margin and posterior metatibial stria even shorter than posterior mesotibial stria.
</paragraph>
<paragraph id="8B932048FF4809F661C7FF3CCD34FE2D" blockId="223.[113,1089,219,467]" pageId="223" pageNumber="216">
Male genitalia. Eighth sternite (
<figureCitation id="13173CCDFF4809F660B6FF3CCC58FF0D" box="[481,626,219,243]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Fig. 687–688</figureCitation>
) widely separated along their entire length, apically with large inflatable membrane (velum) mesally and laterally with several rows of long close-set setae creating a conspicuous apical brush; eighth tergite and eighth sternite fused laterally (
<figureCitation id="13173CCDFF4809F6604FFEDCCF5CFEAD" box="[280,374,315,339]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Fig. 689</figureCitation>
). Ninth tergite (
<figureCitation id="13173CCDFF4809F66348FEDCCC91FEAD" box="[543,699,315,339]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Figs. 690–691</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="13173CCDFF4809F66183FEBCCF1BFE8D" box="[212,305,347,371]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Fig. 690</figureCitation>
) expanded on both ends. Aedeagus (
<figureCitation id="13173CCDFF4809F66396FEBCCD74FE8D" box="[705,862,347,371]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Figs. 692–693</figureCitation>
) slender; basal piece of aedeagus rather short, ratio of its length: length of parameres 1: 3.50; parameres fused almost along their apical length, on apical tenth with opening for the median lobe (best seen from lateral view); aedeagus strongly curved ventrad (
<figureCitation id="13173CCDFF4809F663E3FE5CCD38FE2D" box="[692,786,443,467]" captionStart="Figs" captionStartId="221.[113,152,1087,1106]" captionTargetBox="[259,950,225,1062]" captionTargetId="figure-12@221.[229,974,216,1068]" captionTargetPageId="221" captionText="Figs.687–693.Styphrus corpulentus Motschulsky, 1845, male terminalia: 687 – 8th sternite and tergite, ventral view; 688 – ditto, dorsal view; 689 – ditto, lateral view; 690 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 691 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 692 – aedeagus, dorsal view; 693 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272304" httpUri="https://zenodo.org/record/4272304/files/figure.png" pageId="223" pageNumber="216">Fig. 693</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>