167 lines
22 KiB
XML
167 lines
22 KiB
XML
<document id="8E124E57B295D61BF8AEFC60FCE74574" ID-CLB-Dataset="9574" ID-DOI="10.5281/zenodo.4272127" ID-GBIF-Dataset="724cb358-6f18-4816-afc7-bb42cb9b6942" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298182318" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2010" docId="0385915EFF4809C860CAFE1DCDDDFB06" docLanguage="en" docName="ActaEntMusNatPra.50.suppl.1-254.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 50" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Turanostyphrus Tishechkin 2005" docType="treatment" docVersion="17" lastPageNumber="216" masterDocId="FFBCE926FF9709296157FFE7CE2AFFFE" masterDocTitle="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)" masterLastPageNumber="254" masterPageNumber="1" pageNumber="216" updateTime="1698846217483" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8C840AF698DC73547B268A0158314E27">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="475DAC6FF808613D1EEE29648D803CC5">Lackner, Tomáš</mods:namePart>
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<mods:title id="025C4B46D3F3D5F37548CE3DD5F77BD9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E9367816F30BE2097C1AB3D8BFC0E690">2010</mods:date>
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<mods:title id="D52024DF0001F357971DDEA84CE828AF">suppl.</mods:title>
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<mods:number id="8EF033F19AF6E0EFB495A5EC7E95B8F8">2010-10-20</mods:number>
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<treatment id="0385915EFF4809C860CAFE1DCDDDFB06" ID-DOI="http://doi.org/10.5281/zenodo.4342071" ID-GBIF-Taxon="169092635" ID-Zenodo-Dep="4342071" LSID="urn:lsid:plazi:treatment:0385915EFF4809C860CAFE1DCDDDFB06" httpUri="http://treatment.plazi.org/id/0385915EFF4809C860CAFE1DCDDDFB06" lastPageId="225" lastPageNumber="216" pageId="223" pageNumber="216">
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<subSubSection id="C33673C3FF4809F660CAFE1DCD3EFDED" box="[413,788,506,531]" pageId="223" pageNumber="216" type="nomenclature">
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<paragraph id="8B932048FF4809F660CAFE1DCD3EFDED" blockId="223.[413,788,506,531]" box="[413,788,506,531]" pageId="223" pageNumber="216">
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<heading id="D0DB9724FF4809F660CAFE1DCD3EFDED" bold="true" box="[413,788,506,531]" centered="true" fontSize="10" level="2" pageId="223" pageNumber="216" reason="2">
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<taxonomicName id="4C2C5BCBFF4809F660CAFE1DCD3EFDED" ID-CoL="83SF" authority="Tishechkin, 2005" authorityName="Tishechkin" authorityYear="2005" box="[413,788,506,531]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="genus">
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<emphasis id="B958FC5AFF4809F660CAFE1DCD3EFDED" bold="true" box="[413,788,506,531]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F660CAFE1DCC67FDED" bold="true" box="[413,589,506,531]" italics="true" pageId="223" pageNumber="216">Turanostyphrus</emphasis>
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<bibRefCitation id="EFBD5DB9FF4809F66303FE1CCD3EFDED" author="TISHECHKIN A. K." box="[596,788,507,531]" pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">Tishechkin, 2005</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<paragraph id="8B932048FF4809F66126FDC5CEE6FDB1" blockId="223.[113,1088,545,591]" pageId="223" pageNumber="216">
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<treatmentCitationGroup id="AB3C0766FF4809F66126FDC5CEE6FDB1" pageId="223" pageNumber="216">
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<subSubSection id="C33673C3FF4809F66126FDC5CFEEFDCB" box="[113,452,545,565]" pageId="223" pageNumber="216" type="reference_group">
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<treatmentCitation id="0A8D0659FF4809F66126FDC5CFEAFDCB" author="TISHECHKIN A. K." box="[113,448,545,565]" page="348" pageId="223" pageNumber="216" year="2005">
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<taxonomicName id="4C2C5BCBFF4809F66126FDC5CFEAFDCB" ID-CoL="83SF" authority="Tishechkin, 2005: 348" authorityName="Tishechkin" authorityPageNumber="348" authorityYear="2005" box="[113,448,545,565]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="genus">
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<emphasis id="B958FC5AFF4809F66126FDC5CED2FDCA" box="[113,248,546,564]" italics="true" pageId="223" pageNumber="216">Turanostyphrus</emphasis>
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<bibRefCitation id="EFBD5DB9FF4809F661A9FDC6CFEAFDCB" author="TISHECHKIN A. K." box="[254,448,545,565]" pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">Tishechkin, 2005: 348</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</subSubSection>
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<subSubSection id="C33673C3FF4809F6609FFDC5CEE6FDB1" pageId="223" pageNumber="216" type="type_taxon">
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<typeStatus id="54979EEAFF4809F6609FFDC5CFDEFDCB" box="[456,500,546,565]" pageId="223" pageNumber="216">Type</typeStatus>
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species:
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<taxonomicName id="4C2C5BCBFF4809F66315FDC5CD96FDCA" ID-CoL="59MJL" authority="Tishechkin, 2005" authorityName="Tishechkin" authorityYear="2005" box="[578,956,545,564]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="ignoratus">
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<emphasis id="B958FC5AFF4809F66315FDC5CD0BFDCA" box="[578,801,546,564]" italics="true" pageId="223" pageNumber="216">Turanostyphrus ignoratus</emphasis>
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<bibRefCitation id="EFBD5DB9FF4809F66271FDC6CD96FDCA" author="TISHECHKIN A. K." box="[806,956,545,564]" pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">Tishechkin, 2005</bibRefCitation>
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</taxonomicName>
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, original designation.
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</subSubSection>
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C33673C3FF4809F66126FD80CD8DFB61" pageId="223" pageNumber="216" type="diagnosis">
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<paragraph id="8B932048FF4809F66126FD80CA3CFD01" blockId="223.[113,1089,615,1599]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FD80CECCFD81" bold="true" box="[113,230,615,639]" pageId="223" pageNumber="216">Diagnosis.</emphasis>
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Cuticle without prominent metallic luster; frontal and supraorbital striae absent; pronotal hypomeron setose; elytral epipleuron with short setae; both sets of prosternal striae present; pre-apical foveae absent. Protibia dilated, outer margin explanate, concealing numerous denticles (similar to
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<taxonomicName id="4C2C5BCBFF4809F66026FD2ECC74FD1E" authorityName="Kryzhanovskij" authorityYear="1966" box="[369,606,713,736]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="vlasovi">
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<emphasis id="B958FC5AFF4809F66026FD2ECC74FD1E" box="[369,606,713,736]" italics="true" pageId="223" pageNumber="216">Eremosaprinus vlasovi</emphasis>
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</taxonomicName>
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); metatibia with explanate outer margin almost entirely concealing two rows of numerous denticles observable only from ventral view.
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</paragraph>
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<paragraph id="8B932048FF4809F661C7FCE0CC84FC81" blockId="223.[113,1089,615,1599]" pageId="223" pageNumber="216">This genus has been erected only recently and TISHECHKIN’ s (2005) diagnosis serves the purpose well. However, for the sake of consistency and clarity concerning the morphology used in this work it is repeated here. The concerning parts are duly modified and characters omitted by Tishechkin in his paper are supplemented.</paragraph>
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<paragraph id="8B932048FF4809F66126FC60CD8DFB61" blockId="223.[113,1089,615,1599]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FC60CF49FC61" bold="true" box="[113,355,903,927]" pageId="223" pageNumber="216">Differential diagnosis.</emphasis>
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This genus is one of the most peculiar among the Palaearctic
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<taxonomicName id="4C2C5BCBFF4809F66286FC60CA6BFC61" authorityName="Blanchard" authorityYear="1845" box="[977,1089,903,927]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
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genera. The structure of protibia, and especially metatibia (with explanate outer margins almost completely concealing one or two rows of short denticles, respectively) and the setose elytral epipleuron does not allow it to be confused with any currently known taxon. By the absent frontal and supraorbital striae and pre-apical foveae, as well as the explanate outer margin of protibia it can be compared to the species
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<taxonomicName id="4C2C5BCBFF4809F66373FBCECD3DFBBE" authorityName="Kryzhanovskij" authorityYear="1966" box="[548,791,1065,1088]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="vlasovi">
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<emphasis id="B958FC5AFF4809F66373FBCECD3DFBBE" box="[548,791,1065,1088]" italics="true" pageId="223" pageNumber="216">Eremosaprinus vlasovi</emphasis>
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</taxonomicName>
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, but clearly differing from it by differently structured metatibia (long and slender, almost without denticles
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<emphasis id="B958FC5AFF4809F662F9FBAECDD8FB9E" box="[942,1010,1097,1120]" italics="true" pageId="223" pageNumber="216">versus</emphasis>
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dilated and explanate concealing two rows of numerous denticles) as well as by the setose pronotal hypomeron and elytral epipleuron
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<emphasis id="B958FC5AFF4809F660B5FB6ECC0CFB5E" box="[482,550,1161,1184]" italics="true" pageId="223" pageNumber="216">versus</emphasis>
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glabrous in
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<taxonomicName id="4C2C5BCBFF4809F663FCFB6ECD8AFB5E" authorityName="Kryzhanovskij" authorityYear="1966" box="[683,928,1161,1184]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="vlasovi">
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<emphasis id="B958FC5AFF4809F663FCFB6ECD8AFB5E" box="[683,928,1161,1184]" italics="true" pageId="223" pageNumber="216">Eremosaprinus vlasovi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33673C3FF4809F66126FB40CF72FAE1" pageId="223" pageNumber="216" type="biology_ecology">
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<paragraph id="8B932048FF4809F66126FB40CF72FAE1" blockId="223.[113,1089,615,1599]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FB40CEE4FB41" bold="true" box="[113,206,1191,1215]" pageId="223" pageNumber="216">Biology.</emphasis>
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Virtually unknown; the only two described species (each known only from a
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<typeStatus id="54979EEAFF4809F6655FFB40CE8AFB1E" pageId="223" pageNumber="216" type="holotype">holotype</typeStatus>
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) were collected in a rodent burrow and trapped by a pitfall trap, respectively.
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<bibRefCitation id="EFBD5DB9FF4809F6629BFB2FCE93FB01" author="TISHECHKIN A. K." pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">TISHECHKIN (2005)</bibRefCitation>
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hypothesized that this taxon probably leads psammophilous way of life, based on its external morphology.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33673C3FF4809F66126FAC0CE84FA81" pageId="223" pageNumber="216" type="distribution">
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<paragraph id="8B932048FF4809F66126FAC0CD90FAC1" blockId="223.[113,1089,615,1599]" box="[113,954,1319,1343]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FAC0CF2EFAC1" bold="true" box="[113,260,1319,1343]" pageId="223" pageNumber="216">Distribution.</emphasis>
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Central Asian province of Turan:
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<collectingCountry id="F33B60D8FF4809F66324FAC0CD2DFAC1" box="[627,775,1319,1343]" name="Turkmenistan" pageId="223" pageNumber="216">Turkmenistan</collectingCountry>
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and
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<collectingCountry id="F33B60D8FF4809F6626CFAC0CD9DFAC1" box="[827,951,1319,1343]" name="Uzbekistan" pageId="223" pageNumber="216">Uzbekistan</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B932048FF4809F66126FAA0CE84FA81" blockId="223.[113,1089,615,1599]" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FAA0CF6BFAA1" bold="true" box="[113,321,1351,1375]" pageId="223" pageNumber="216">Species examined.</emphasis>
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<taxonomicName id="4C2C5BCBFF4809F6601EFAAECD0CFAA1" authority="Tishechkin, 2005" authorityName="Tishechkin" authorityYear="2005" box="[329,806,1351,1376]" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="ignoratus">
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<emphasis id="B958FC5AFF4809F6601EFAAECC4AFA9E" box="[329,608,1353,1376]" italics="true" pageId="223" pageNumber="216">Turanostyphrus ignoratus</emphasis>
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<bibRefCitation id="EFBD5DB9FF4809F6633EFAA0CD0CFAA1" author="TISHECHKIN A. K." box="[617,806,1351,1375]" pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">Tishechkin, 2005</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C2C5BCBFF4809F66264FAAECE80FA81" authority="Tishechkin, 2005" authorityName="Tishechkin" authorityYear="2005" class="Insecta" family="Histeridae" genus="Turanostyphrus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="species" species="kizilkumis">
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<emphasis id="B958FC5AFF4809F66264FAAECD91FA9E" box="[819,955,1353,1376]" italics="true" pageId="223" pageNumber="216">T. kizilkumis</emphasis>
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<bibRefCitation id="EFBD5DB9FF4809F66294FAA0CE80FA81" author="TISHECHKIN A. K." pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">Tishechkin, 2005</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33673C3FF4809C86126FA60CDDDFB06" lastPageId="225" lastPageNumber="218" pageId="223" pageNumber="216" type="discussion">
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<paragraph id="8B932048FF4809C86126FA60CDDDFB06" blockId="223.[113,1089,615,1599]" lastBlockId="225.[113,1089,1120,1272]" lastPageId="225" lastPageNumber="218" pageId="223" pageNumber="216">
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<emphasis id="B958FC5AFF4809F66126FA60CEDBFA61" bold="true" box="[113,241,1415,1439]" pageId="223" pageNumber="216">Discussion.</emphasis>
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Resolving the exact taxomonic position within the Palaearctic
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<taxonomicName id="4C2C5BCBFF4809F662D0FA60CDD3FA61" authorityName="Blanchard" authorityYear="1845" box="[903,1017,1415,1439]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
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of this genus is a puzzling task, which needs to be examined by further studies. In the preliminary studies on the phylogeny of
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<taxonomicName id="4C2C5BCBFF4809F660FEFA20CC31FA21" authorityName="Blanchard" authorityYear="1845" box="[425,539,1479,1503]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
|
||
(LACKNER, in prep.) it came out in several equally parsimonious trees on the bottom, close to the group ‘
|
||
<emphasis id="B958FC5AFF4809F663F7FA0ECEBFF9DE" italics="true" pageId="223" pageNumber="216">
|
||
<taxonomicName id="4C2C5BCBFF4809F663F7FA0ECD69F9FE" authorityName=": ROSS" authorityYear="1940" box="[672,835,1513,1536]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="genus">Eremosaprinus</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C2C5BCBFF4809F6620DFA0ECDE9F9FE" authorityName=": THOMSON" authorityYear="1867" box="[858,963,1513,1536]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="genus">Myrmetes</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C2C5BCBFF4809F6628DFA0ECEBFF9DE" authorityName="Lackner" authorityYear="2010" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="216" phylum="Arthropoda" rank="genus">Gnathoncus</taxonomicName>
|
||
</emphasis>
|
||
’; weighted analysis, however, placed it near the most derived psammophilous taxa, as originally also indicated by
|
||
<bibRefCitation id="EFBD5DB9FF4809F660F7F9CFCC4DF9C1" author="TISHECHKIN A. K." box="[416,615,1575,1599]" pageId="223" pageNumber="216" refId="ref122089" refString="TISHECHKIN A. K. 2005: Descriptions of new taxa of the Palaearctic Saprininae (Coleoptera: Histeridae). Pp. 347 - 355. In: KONSTANTINOV A., TISHECHKIN A. & PENEV L. (eds.): Contributions to Systematics and Biology of Beetles, Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin. Series Faunistica 43. Pensoft, Sofia-Moscow, 450 pp." type="book" year="2005">TISHECHKIN (2005)</bibRefCitation>
|
||
. It is probably a well-defined monophyletic taxon, supported by several ‘strong’ autapomorphies, e.g. explanate outer margin of protibia and metatibia, concealing denticles. Absent frontal and supraorbital striae can be regarded as putative synapomorphies, shared also with several other taxa, e.g. the ‘
|
||
<emphasis id="B958FC5AFF7609C862C9FB46CFBBFB26" italics="true" pageId="225" pageNumber="218">
|
||
<taxonomicName id="4C2C5BCBFF7609C862C9FB46CA6BFB46" authorityName=": ROSS" authorityYear="1940" box="[926,1089,1185,1208]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="218" phylum="Arthropoda" rank="genus">Eremosaprinus</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C2C5BCBFF7609C861DFFB26CEDBFB26" authorityName=": THOMSON" authorityYear="1867" box="[136,241,1217,1240]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="218" phylum="Arthropoda" rank="genus">Myrmetes</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C2C5BCBFF7609C86047FB27CFBBFB26" authorityName="Lackner" authorityYear="2010" box="[272,401,1216,1240]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="218" phylum="Arthropoda" rank="genus">Gnathoncus</taxonomicName>
|
||
</emphasis>
|
||
’ complex. Especially puzzling is the ciliate elytral epipleuron, which possibly represents an evolutionary reversal, or, alternatively a plesiomorphy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |