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<mods:title id="AB01ACF88F508DEBF0C83549451B13D1">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<taxonomicName id="4C2C5BCBFF29099760F6FBCDCD77FBBC" ID-CoL="794HH" authority="(Reitter, 1904)" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" baseAuthorityName="Reitter" baseAuthorityYear="1904" box="[417,861,1066,1091]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">
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<emphasis id="B958FC5AFF29099760F6FBCDCC98FBBD" bold="true" box="[417,690,1066,1091]" italics="true" pageId="190" pageNumber="187">Reichardtiolus duriculus</emphasis>
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<bibRefCitation id="EFBD5DB9FF2909976397FBCDCD7CFBBC" author="REITTER E." box="[704,854,1066,1090]" pageId="190" pageNumber="187" pagination="29 - 36" refId="ref120480" refString="REITTER E. 1904: Ueber neue und wenig bekannte Histeriden (Coleoptera). Wiener Entomologische Zeitung 23: 29 - 36." type="journal article" year="1904">Reitter, 1904</bibRefCitation>
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<figureCitation id="13173CCDFF290997635BFBAECC79FBA2" box="[524,595,1097,1116]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="190" pageNumber="187">Figs. 27</figureCitation>
, 
<figureCitation id="13173CCDFF290997630BFBAECC59FBA2" box="[604,627,1097,1116]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="190" pageNumber="187">67</figureCitation>
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<figureCitation id="13173CCDFF290997632BFBAECCB4FBA2" box="[636,670,1097,1116]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="190" pageNumber="187">132</figureCitation>
, 
<figureCitation id="13173CCDFF29099763F0FBAECCD9FBA2" box="[679,755,1097,1116]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="192.[151,190,1521,1540]" captionStartId-1="193.[113,152,1016,1035]" captionTargetBox-0="[240,1040,219,1499]" captionTargetBox-1="[252,954,227,997]" captionTargetId-0="figure-11@192.[240,1040,219,1499]" captionTargetId-1="figure-139@193.[221,980,217,1010]" captionTargetPageId-0="192" captionTargetPageId-1="193" captionText-0="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." captionText-1="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272282" figureDoi-1="http://doi.org/10.5281/zenodo.4272284" httpUri-0="https://zenodo.org/record/4272282/files/figure.png" httpUri-1="https://zenodo.org/record/4272284/files/figure.png" pageId="190" pageNumber="187">593–610</figureCitation>
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<treatmentCitation id="0A8D0659FF29099761C0FB94CFFBFB78" author="REITTER E." box="[151,465,1139,1158]" page="31" pageId="190" pageNumber="187" year="1904">
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<emphasis id="B958FC5AFF29099761C0FB94CF13FB78" box="[151,313,1139,1158]" italics="true" pageId="190" pageNumber="187">Saprinus duriculus</emphasis>
<bibRefCitation id="EFBD5DB9FF2909976069FB94CFFBFB78" author="REITTER E." box="[318,465,1139,1158]" pageId="190" pageNumber="187" pagination="29 - 36" refId="ref120480" refString="REITTER E. 1904: Ueber neue und wenig bekannte Histeriden (Coleoptera). Wiener Entomologische Zeitung 23: 29 - 36." type="journal article" year="1904">Reitter, 1904: 31</bibRefCitation>
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<treatmentCitationGroup id="AB3C0766FF29099761C0FB6ACC2EFB5E" box="[151,516,1165,1185]" pageId="190" pageNumber="187">
<treatmentCitation id="0A8D0659FF29099761C0FB6ACC2AFB5E" author="JAKOBSON G. G." box="[151,512,1165,1185]" page="651" pageId="190" pageNumber="187" year="1911">
<taxonomicName id="4C2C5BCBFF29099761C0FB6ACFFFFB5F" authority=": JAKOBSON (1911)" authorityName="JAKOBSON" authorityYear="1911" box="[151,469,1165,1185]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF29099761C0FB6ACF12FB5E" box="[151,312,1165,1184]" italics="true" pageId="190" pageNumber="187">Styphrus duriculus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF2909976014FB69CFFFFB5F" author="JAKOBSON G. G." box="[323,469,1166,1185]" pageId="190" pageNumber="187" pagination="641 - 720" refId="ref116036" refString="JAKOBSON G. G. 1911: Zhuki Rosii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. 9. [Beetles of Russia and western Europe. Manual for the beetles' determination. 9]. A. F. Devrien, St. Petersburg, pp. 1 + 641 - 720 (in Russian)." type="journal article" year="1911">JAKOBSON (1911)</bibRefCitation>
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: 651
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<paragraph id="8B932048FF29099761C0FB4FCC18FB45" blockId="190.[151,1127,1139,1292]" box="[151,562,1192,1212]" pageId="190" pageNumber="187">
<treatmentCitationGroup id="AB3C0766FF29099761C0FB4FCC18FB45" box="[151,562,1192,1212]" pageId="190" pageNumber="187">
<treatmentCitation id="0A8D0659FF29099761C0FB4FCC04FB45" author="BICKHARDT H." box="[151,558,1192,1212]" page="97" pageId="190" pageNumber="187" year="1916">
<taxonomicName id="4C2C5BCBFF29099761C0FB4FCC27FB45" authority=": BICKHARDT (1916)" authorityName="BICKHARDT" authorityYear="1916" box="[151,525,1192,1212]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF29099761C0FB4FCF4CFB45" box="[151,358,1192,1211]" italics="true" pageId="190" pageNumber="187">Hypocacculus duriculus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF2909976026FB4ECC27FB45" author="BICKHARDT H." box="[369,525,1192,1212]" pageId="190" pageNumber="187" refId="ref112724" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum, Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
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: 97
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<treatmentCitationGroup id="AB3C0766FF29099761C0FB24CD49FB28" box="[151,867,1219,1238]" pageId="190" pageNumber="187">
<treatmentCitation id="0A8D0659FF29099761C0FB24CC3EFB28" author="REICHARDT A." box="[151,532,1219,1238]" page="17" pageId="190" pageNumber="187" year="1926">
<taxonomicName id="4C2C5BCBFF29099761C0FB24CFDEFB28" authority=": REICHARDT (1926)" authorityName="REICHARDT" authorityYear="1926" box="[151,500,1219,1238]" class="Insecta" family="Histeridae" genus="Exaesiopus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF29099761C0FB24CF65FB28" box="[151,335,1219,1238]" italics="true" pageId="190" pageNumber="187">Exaesiopus duriculus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF290997600DFB24CFDEFB28" author="REICHARDT A." box="[346,500,1219,1238]" pageId="190" pageNumber="187" pagination="12 - 18" refId="ref120278" refString="REICHARDT A. 1926: Uber die mit Pachylopus verwandten Arten. Entomologische Blatter 22: 12 - 18." type="journal article" year="1926">REICHARDT (1926)</bibRefCitation>
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: 17
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; 
<treatmentCitation id="0A8D0659FF290997634AFB24CCC9FB28" author="REICHARDT A." box="[541,739,1219,1238]" page="330" pageId="190" pageNumber="187" year="1941">
<bibRefCitation id="EFBD5DB9FF290997634AFB24CC93FB28" author="REICHARDT A." box="[541,697,1219,1238]" pageId="190" pageNumber="187" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
: 330
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, 333, 
<figureCitation id="13173CCDFF2909976241FB24CD4AFB28" box="[790,864,1219,1238]" captionStart="Figs" captionStartId="58.[151,190,1544,1563]" captionTargetBox="[178,1096,461,1510]" captionTargetId="figure-11@58.[178,1096,459,1512]" captionTargetPageId="58" captionText="Figs. 168–173. Saprininae, male terminalia, schematic: 168 – 8th sternite, ventral view; 169 – 8th sternite and tergite, dorsal view; 170 – ditto, lateral view; 171 – ninth tergites and tenth tergite, dorsal view; 172 – spiculum gastrale (10th sternite), ventral view; 173 – aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272183" httpUri="https://zenodo.org/record/4272183/files/figure.png" pageId="190" pageNumber="187">Fig. 172</figureCitation>
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</paragraph>
<paragraph id="8B932048FF29099761C0FB3ACFECFAF5" blockId="190.[151,1127,1139,1292]" pageId="190" pageNumber="187">
<treatmentCitationGroup id="AB3C0766FF29099761C0FB3ACFECFAF5" pageId="190" pageNumber="187">
<treatmentCitation id="0A8D0659FF29099761C0FB3ACCE4FB0F" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[151,718,1245,1265]" page="239" pageId="190" pageNumber="187" year="1976">
<taxonomicName id="4C2C5BCBFF29099761C0FB3ACC8FFB0E" ID-CoL="794HH" authority=": KRYZHANOVSKIJ &amp; REICHARDT (1976)" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" baseAuthorityName="Reitter" baseAuthorityYear="1904" box="[151,677,1245,1265]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF29099761C0FB3ACF42FB0E" box="[151,360,1245,1264]" italics="true" pageId="190" pageNumber="187">Reichardtiolus duriculus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF2909976024FB39CC8FFB0E" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[371,677,1245,1265]" pageId="190" pageNumber="187" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
</taxonomicName>
: 239
</treatmentCitation>
, 
<figureCitation id="13173CCDFF2909976382FB3ACD52FB0E" box="[725,888,1245,1264]" captionStart="Figs" captionStartId="151.[113,152,1018,1037]" captionTargetBox="[213,993,222,986]" captionTargetId="figure-165@151.[150,1051,206,1005]" captionTargetPageId="151" captionText="Figs. 462–470. Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807), male terminalia: 462 – 8th sternite and tergite, ventral view; 463 – ditto, dorsal view; 464 – ditto, lateral view; 465 – 9th tergite, 10th tergite, dorsal view; 466 – ditto, lateral view; 467 – aedeagus, dorsal view; 468 – ditto, lateral view; 469 – spiculum gastrale, ventral view; 470 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272250" httpUri="https://zenodo.org/record/4272250/files/figure.png" pageId="190" pageNumber="187">Figs. 465, 466, 468</figureCitation>
; 
<treatmentCitation id="0A8D0659FF2909976228FB39CA09FB0F" author="MAZUR S." box="[895,1059,1246,1265]" page="103" pageId="190" pageNumber="187" year="1984">
<bibRefCitation id="EFBD5DB9FF2909976228FB39CDD0FB0F" author="MAZUR S." box="[895,1018,1246,1265]" pageId="190" pageNumber="187" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 103
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF290997657CFB39CF37FAF5" author="MAZUR S." page="265" pageId="190" pageNumber="187" year="1997">
<bibRefCitation id="EFBD5DB9FF290997657CFB39CED8FAF5" author="MAZUR S." pageId="190" pageNumber="187" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 265
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF2909976072FB1ECFE8FAF5" author="MAZUR S." box="[293,450,1272,1292]" page="96" pageId="190" pageNumber="187" year="2004">
<bibRefCitation id="EFBD5DB9FF2909976072FB1ECF88FAF5" author="MAZUR S." box="[293,418,1272,1292]" pageId="190" pageNumber="187" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
: 96
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF29099761C0FAC5CFEAFA2B" pageId="190" pageNumber="187" type="materials_examined">
<paragraph id="8B932048FF29099761C0FAC5CFE4FACB" blockId="190.[151,1127,1313,1493]" box="[151,462,1313,1333]" pageId="190" pageNumber="187">
<emphasis id="B958FC5AFF29099761C0FAC5CF39FACB" bold="true" box="[151,275,1313,1333]" pageId="190" pageNumber="187">
<typeStatus id="54979EEAFF29099761C0FAC5CEEFFACB" box="[151,197,1314,1333]" pageId="190" pageNumber="187">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B60D8FF290997604FFAC5CFBBFACB" box="[280,401,1314,1333]" name="Turkmenistan" pageId="190" pageNumber="187">Turkmenistan</collectingCountry>
, 
<collectingRegion id="49E8EEAAFF29099760CEFAC5CFE0FACB" box="[409,458,1314,1333]" country="Turkmenistan" name="Mary Welayaty" pageId="190" pageNumber="187">Mary</collectingRegion>
.
</paragraph>
<paragraph id="8B932048FF29099761C0FADACA7EFA7B" blockId="190.[151,1127,1313,1493]" pageId="190" pageNumber="187">
<emphasis id="B958FC5AFF29099761C0FADACF5EFAAE" bold="true" box="[151,372,1340,1360]" pageId="190" pageNumber="187">Type material examined.</emphasis>
<typeStatus id="54979EEAFF2909976020FADACFE3FAB0" box="[375,457,1341,1360]" pageId="190" pageNumber="187" type="holotype">HOLOTYPE</typeStatus>
: 
<specimenCount id="9D2AEBC1FF2909976085FADACC24FAAE" box="[466,526,1341,1360]" pageId="190" pageNumber="187" type="generic">1 spec.</specimenCount>
, last two metatarsomeres of right hind tibia broken off:‘Merw [printed] // Ahnger [printed] // coll. Reitter [printed] // 
<typeStatus id="54979EEAFF29099760AFFAB0CC7AFA94" box="[504,592,1367,1386]" pageId="190" pageNumber="187" type="holotype">Holotypus</typeStatus>
1904 / 
<taxonomicName id="4C2C5BCBFF29099763DCFAB0CCFFFA94" authorityName="Erichson" authorityYear="1834" box="[651,725,1367,1386]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ duriculus / Reitter [red-framed printed-written] // 1960 / 
<taxonomicName id="4C2C5BCBFF29099761B3FA95CF6FFA7B" authorityName=", MAZUR" authorityYear="1984" box="[228,325,1394,1413]" class="Insecta" family="Histeridae" genus="Exaesiopus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
/ (
<taxonomicName id="4C2C5BCBFF290997600CFA95CFF0FA7B" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[347,474,1394,1413]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="genus">Reichardtiolus</taxonomicName>
) / 
<taxonomicName id="4C2C5BCBFF29099760A6FA95CC55FA7B" authority="Rchdt." authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" baseAuthorityName="Reitter" baseAuthorityYear="1904" box="[497,639,1394,1413]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="187" phylum="Arthropoda" rank="species" species="duriculus">duriculus Rchdt.</taxonomicName>
[sic!] / Kryzhanovskij det.[printed-written]’ (
<collectionCode id="ED3DB88DFF2909976553FA95CA60FA7B" box="[1028,1098,1394,1413]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="190" pageNumber="187">HNHM</collectionCode>
).
</paragraph>
<paragraph id="8B932048FF29099761C0FA6BCFEAFA2B" blockId="190.[151,1127,1313,1493]" pageId="190" pageNumber="187">
<emphasis id="B958FC5AFF29099761C0FA6BCF87FA5E" bold="true" box="[151,429,1420,1440]" pageId="190" pageNumber="187">Additional material examined.</emphasis>
<materialsCitation id="3B442A15FF29099760E5FA6BCF43FA44" ID-GBIF-Occurrence="2981920337" collectingDate="1994-04-05" collectorName="Becvar J." country="Jordan" elevation="1000" latitude="29.333334" location="El Mudawwara" longLatPrecision="1227" longitude="35.533333" pageId="190" pageNumber="187" specimenCount="3">
<emphasis id="B958FC5AFF29099760E5FA6BCC39FA5E" bold="true" box="[434,531,1420,1440]" pageId="190" pageNumber="187">
<collectingCountry id="F33B60D8FF29099760E5FA6BCC24FA5E" box="[434,526,1420,1440]" name="Jordan" pageId="190" pageNumber="187">JORDAN</collectingCountry>
:
</emphasis>
<quantity id="4CD48DADFF290997634FFA6ACC67FA5E" box="[536,589,1421,1440]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="190" pageNumber="187" unit="km" value="60.0">60 km</quantity>
N 
<location id="8EF37693FF2909976330FA6ACCC7FA5E" LSID="urn:lsid:plazi:treatment:0385915EFF2909EB60F6FBCDCDBDFB6D:8EF37693FF2909976330FA6ACCC7FA5E" box="[615,749,1421,1440]" country="Jordan" latitude="29.333334" longLatPrecision="1227" longitude="35.533333" name="El Mudawwara" pageId="190" pageNumber="187">El Mudawwara</location>
, 
<quantity id="4CD48DADFF29099763A2FA6ACD12FA5E" box="[757,824,1421,1440]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="190" pageNumber="187" unit="m" value="1000.0">
<elevation id="0001C77BFF29099763A2FA6ACD12FA5E" box="[757,824,1421,1440]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="190" pageNumber="187" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, 
<geoCoordinate id="EE18468FFF2909976217FA6ACDADFA5E" box="[832,903,1421,1440]" degrees="29" direction="north" minutes="20" orientation="latitude" pageId="190" pageNumber="187" precision="925" value="29.333334">29º20'N</geoCoordinate>
, 
<geoCoordinate id="EE18468FFF29099762D8FA6BCDF9FA61" box="[911,979,1420,1439]" degrees="35" direction="east" minutes="32" orientation="longitude" pageId="190" pageNumber="187" precision="925" value="35.533333">35º32'E</geoCoordinate>
, 
<date id="FF920688FF290997628BFA6BCA05FA61" box="[988,1071,1420,1439]" pageId="190" pageNumber="187" value="1994-04-05">
<collectingDate id="EFD6FF60FF290997628BFA6BCA05FA61" box="[988,1071,1420,1439]" pageId="190" pageNumber="187" value="1994-04-05">5.iv.1994</collectingDate>
</date>
, 2 ♁♁ 
<specimenCount id="9D2AEBC1FF29099761C0FA40CEF9FA45" box="[151,211,1447,1467]" pageId="190" pageNumber="187" type="generic">3 spec.</specimenCount>
, 
<collectorName id="26D9459EFF290997618EFA4FCF0CFA45" box="[217,294,1447,1467]" pageId="190" pageNumber="187">Bečvář J.</collectorName>
&amp; S. lgt
</materialsCitation>
. 
<materialsCitation id="3B442A15FF2909976027FA41CD9AFA44" ID-GBIF-Occurrence="2981920313" box="[368,944,1444,1467]" collectingDate="1981-04-21" collectorName="A. Olexa" country="Turkmenistan" location="Annau" municipality="Karakum" pageId="190" pageNumber="187" specimenCount="1">
<emphasis id="B958FC5AFF2909976027FA41CC0FFA44" bold="true" box="[368,549,1446,1466]" pageId="190" pageNumber="187">
<collectingCountry id="F33B60D8FF2909976027FA41CC0BFA44" box="[368,545,1446,1466]" name="Turkmenistan" pageId="190" pageNumber="187">TURKMENISTAN</collectingCountry>
:
</emphasis>
<location id="8EF37693FF290997637EFA40CC49FA44" LSID="urn:lsid:plazi:treatment:0385915EFF2909EB60F6FBCDCDBDFB6D:8EF37693FF290997637EFA40CC49FA44" box="[553,611,1447,1466]" country="Turkmenistan" municipality="Karakum" name="Annau" pageId="190" pageNumber="187">Annau</location>
, 
<collectingMunicipality id="6BF7BA32FF290997633EFA40CC93FA44" box="[617,697,1447,1466]" pageId="190" pageNumber="187">Karakum</collectingMunicipality>
, 
<date id="FF920688FF2909976397FA40CD31FA44" box="[704,795,1447,1466]" pageId="190" pageNumber="187" value="1981-04-21">
<collectingDate id="EFD6FF60FF2909976397FA40CD31FA44" box="[704,795,1447,1466]" pageId="190" pageNumber="187" value="1981-04-21">21.iv.1981</collectingDate>
</date>
, 1♁, 
<collectorName id="26D9459EFF290997621EFA40CDB9FA44" box="[841,915,1447,1466]" pageId="190" pageNumber="187">A. Olexa</collectorName>
lgt
</materialsCitation>
.; 
<materialsCitation id="3B442A15FF29099762ECFA40CF96FA2B" ID-GBIF-Occurrence="2981920366" collectingDate="1989-04-12" collectionCode="TLAN" collectorName="M. Nikodym" country="Turkmenistan" location="Repetek" pageId="190" pageNumber="187" specimenCount="1">
<location id="8EF37693FF29099762ECFA40CA28FA44" LSID="urn:lsid:plazi:treatment:0385915EFF2909EB60F6FBCDCDBDFB6D:8EF37693FF29099762ECFA40CA28FA44" box="[955,1026,1447,1466]" country="Turkmenistan" name="Repetek" pageId="190" pageNumber="187">Repetek</location>
, 
<date id="FF920688FF290997655FFA40CA49FA44" box="[1032,1123,1447,1466]" pageId="190" pageNumber="187" value="1989-04-12">
<collectingDate id="EFD6FF60FF290997655FFA40CA49FA44" box="[1032,1123,1447,1466]" pageId="190" pageNumber="187" value="1989-04-12">12.iv.1989</collectingDate>
</date>
, 
<specimenCount id="9D2AEBC1FF29099761C0FA25CEFCFA2B" box="[151,214,1474,1493]" pageId="190" pageNumber="187" type="generic">1 spec.</specimenCount>
, 
<collectorName id="26D9459EFF290997618AFA25CF66FA2B" box="[221,332,1474,1493]" pageId="190" pageNumber="187">M. Nikodým</collectorName>
lgt. (
<collectionCode id="ED3DB88DFF290997602FFA25CF9CFA2B" box="[376,438,1474,1493]" pageId="190" pageNumber="187">TLAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF2909EB61C0FA0ACDC4FBAD" lastPageId="194" lastPageNumber="191" pageId="190" pageNumber="187" type="description">
<paragraph id="8B932048FF29099761C0FA0ACCAAF9DB" blockId="190.[151,1127,1517,1573]" pageId="190" pageNumber="187">
<emphasis id="B958FC5AFF29099761C0FA0ACF17F9FB" bold="true" box="[151,317,1517,1541]" pageId="190" pageNumber="187">Redescription.</emphasis>
Body length: PEL: 2.00– 
<quantity id="4CD48DADFF290997631AFA0ACC84F9FB" box="[589,686,1517,1541]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="190" pageNumber="187" unit="mm" value="3.4">3.40 mm</quantity>
; APW: 
<quantity id="4CD48DADFF2909976255FA0ACD8AF9FB" box="[770,928,1517,1541]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.5" metricValueMin="6.5" pageId="190" pageNumber="187" unit="mm" value="0.85" valueMax="1.05" valueMin="0.65">0.65–1.05 mm</quantity>
; PPW: 
<quantity id="4CD48DADFF29099762B9FA0ACEEEF9DB" metricMagnitude="-3" metricUnit="m" metricValue="1.8875" metricValueMax="2.4" metricValueMin="1.375" pageId="190" pageNumber="187" unit="mm" value="1.8875" valueMax="2.4" valueMin="1.375">1.375–2.40 mm</quantity>
; EL: 
<quantity id="4CD48DADFF29099761AAF9EACFB6F9DB" box="[253,412,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.25" metricValueMin="1.25" pageId="190" pageNumber="187" unit="mm" value="1.75" valueMax="2.25" valueMin="1.25">1.25–2.25 mm</quantity>
; EW: 
<quantity id="4CD48DADFF2909976089F9EACC56F9DB" box="[478,636,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="190" pageNumber="187" unit="mm" value="2.1" valueMax="2.7" valueMin="1.5">1.50–2.70 mm</quantity>
.
</paragraph>
<paragraph id="8B932048FF28099661C7FF3CCF22FE4D" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">
Body (
<figureCitation id="13173CCDFF280996618CFF3CCF50FF0D" box="[219,378,219,243]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Figs. 593–594</figureCitation>
) elongate oval, strongly convex, cuticle dark brown with feeble metallic luster, legs, antennae and mouthparts rufous. Antennal scape (
<figureCitation id="13173CCDFF28099662D6FF1CCDF5FEED" box="[897,991,251,275]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 596</figureCitation>
) slightly thickened, with several short setae; club (
<figureCitation id="13173CCDFF280996637DFEFCCCAFFECD" box="[554,645,283,307]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 595</figureCitation>
) rather large, without visible articulation, apical four-fifths covered with short sensilla intermingled with longer sparse erect sensilla, basal fifth glabrous; sensory structures of antennal club (
<figureCitation id="13173CCDFF28099663B6FEBCCD1BFE8A" box="[737,817,347,372]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="191" pageNumber="184">Fig. 27</figureCitation>
) in form of stipe-shaped vesicle situated under circular sensory area on internal distal margin of the ventral side of antennal club.
</paragraph>
<paragraph id="8B932048FF28099661C7FE5CCFC3FD2D" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">
Mouthparts. Mandibles (
<figureCitation id="13173CCDFF28099660CFFE5CCFD9FE2A" box="[408,499,443,468]" captionStart="Figs" captionStartId="36.[151,190,1161,1180]" captionTargetBox="[151,1126,219,1130]" captionTargetId="figure-100@36.[151,1126,219,1132]" captionTargetPageId="36" captionText="Figs.99–107.Saprininae, mandibles,ventral view:99 – Philothis (Farabius) hexeris Reichardt, 1930; 100 – Pholioxenus phoenix (Reichardt,1930); 101 – Reichardtiolus duriculus (Reitter, 1904); 102 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 103 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 104 – Saprinus (Phaonius) pharao (Marseul, 1855); 105 – Styphrus corpulentus Motschulsky, 1845; 106 – Xenonychus tridens (Jacquelin-Duval, 1852); 107 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4272155" httpUri="https://zenodo.org/record/4272155/files/figure.png" pageId="191" pageNumber="184">Fig. 101</figureCitation>
) with rounded outer margin, strongly curved inwardly, mandibular apex acutely pointed; sub-apical tooth on inner margin of left mandible blunt; labrum (
<figureCitation id="13173CCDFF280996619CFE1CCF33FDED" box="[203,281,507,531]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="191" pageNumber="184">Fig. 67</figureCitation>
) convex, coarsely punctate; with two labral pits, each with two well-sclerotized setae; terminal labial palpomere thickened, its width about half its length; mentum subtrapezoid, anterior margin shallowly emarginate medially (
<figureCitation id="13173CCDFF28099663BDFDDCCD63FDAA" box="[746,841,571,596]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="191" pageNumber="184">Fig. 132</figureCitation>
); antero-lateral corners with few short setae, lateral margins with a single row of short ramose setae; disc of mentum imbricate, asetose; cardo of maxilla with few short setae on lateral margin; stipes triangular, with three short setae; terminal maxillary palpomere thickened, its width about half its length, about twice as long as penultimate.
</paragraph>
<paragraph id="8B932048FF28099661C7FD3CCFD8FCAD" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">
Clypeus (
<figureCitation id="13173CCDFF28099661A4FD3CCF66FD0D" box="[243,332,731,755]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 596</figureCitation>
) slightly concave medially, rounded laterally, rugulose-lacunose; frontal stria well impressed, carinate, almost straight, somewhat weakened medially, continued as well-impressed, carinate supraorbital stria; frontal disc (
<figureCitation id="13173CCDFF2809966385FCFCCD05FCCD" box="[722,815,795,819]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 596</figureCitation>
) rugulose-lacunose; eyes slightly convex, visible from above.
</paragraph>
<paragraph id="8B932048FF28099661C7FCBBCCC6FC2D" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">
Pronotum (
<figureCitation id="13173CCDFF2809966052FCBCCF77FC8D" box="[261,349,859,883]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 593</figureCitation>
) convex, pronotal sides rounded, strongly convergent anteriorly, apical angles inconspicuous; marginal pronotal stria complete, carinate; disc with very deep, dense and coarse punctures, laterally rugulose-lacunose, medially punctation weakens and becomes sparser; pronotal hypomeron with sparse short amber setae.
</paragraph>
<paragraph id="8B932048FF28099661C7FC3CCF4EFAED" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">Elytral epipleura with a row of deep punctures; marginal epipleural stria well impressed, complete; marginal elytral stria complete, deeply impressed, carinate, continued as complete apical elytral stria. Humeral elytral stria weakly impressed on basal third, often doubled; inner subhumeral stria inconspicuous, present as tiny median fragment; elytra with four dorsal striae 1–4, bearing large punctures; first, second and third dorsal striae about the same length, reaching approximately elytral half apically, fourth dorsal elytral stria weakly impressed on basal third (occasionally longer apically), connected to complete sutural elytral stria. Elytral disc with deep round punctation, punctures separated by 2–4 times their diameter, becoming finer apically and laterally; between sutural elytral stria and elytral suture a row of regular fine punctures present.</paragraph>
<paragraph id="8B932048FF28099661C7FAFCCEF4FA8D" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">Propygidium transverse, almost completely exposed, coarsely and densely punctate; pygidium almost as long as broad, with similar punctation; interspaces in both cases finely imbricate.</paragraph>
<paragraph id="8B932048FF28099661C7FA9CCF3EF9AD" blockId="191.[113,1089,219,1619]" pageId="191" pageNumber="184">
Anterior margin of median portion of prosternum (
<figureCitation id="13173CCDFF28099663F1FA9CCD2AFA6D" box="[678,768,1403,1427]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="191" pageNumber="184">Fig. 598</figureCitation>
) rounded; marginal prosternal stria present laterally and as vague anterior fragment; pre-apical foveae rather small; prosternal process rather narrow, slightly concave; carinal prosternal striae slightly carinate, almost parallel, united in front of strongly carinate shortened lateral prosternal striae. Surface between carinal prosternal striae almost smooth, prosternal apophysis with several microscopic setae; lateral parts of prosternal process strigulate with scattered microscopic punctures fringed with tiny setae.
</paragraph>
<caption id="DF5370C0FF5709E961C0FA16CCDCF9AA" ID-DOI="http://doi.org/10.5281/zenodo.4272282" ID-Zenodo-Dep="4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="192" pageNumber="189" startId="192.[151,190,1521,1540]" targetBox="[240,1040,219,1499]" targetPageId="192">
<paragraph id="8B932048FF5709E961C0FA16CCDCF9AA" blockId="192.[151,1129,1521,1621]" pageId="192" pageNumber="189">
Figs. 593–603. 
<taxonomicName id="4C2C5BCBFF5709E9604CFA14CC5BF9FB" authority="(Reitter, 1904)" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" baseAuthorityName="Reitter" baseAuthorityYear="1904" box="[283,625,1521,1541]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="192" pageNumber="189" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF5709E9604CFA14CFC4F9FB" box="[283,494,1523,1541]" italics="true" pageId="192" pageNumber="189">Reichardtiolus duriculus</emphasis>
(
<bibRefCitation id="EFBD5DB9FF5709E960AFFA16CC40F9FB" author="REITTER E." box="[504,618,1521,1541]" pageId="192" pageNumber="189" pagination="29 - 36" refId="ref120480" refString="REITTER E. 1904: Ueber neue und wenig bekannte Histeriden (Coleoptera). Wiener Entomologische Zeitung 23: 29 - 36." type="journal article" year="1904">Reitter, 1904</bibRefCitation>
)
</taxonomicName>
, SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view.
</paragraph>
</caption>
<caption id="DF5370C0FF5609E86126FC1FCF34FBA5" ID-DOI="http://doi.org/10.5281/zenodo.4272284" ID-Zenodo-Dep="4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="193" pageNumber="186" startId="193.[113,152,1016,1035]" targetBox="[252,954,227,997]" targetPageId="193">
<paragraph id="8B932048FF5609E86126FC1FCF34FBA5" blockId="193.[113,1089,1014,1115]" pageId="193" pageNumber="186">
Figs. 604–610. 
<taxonomicName id="4C2C5BCBFF5609E861A3FC1ECC63FBF5" authority="(Reitter, 1904)" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" baseAuthorityName="Reitter" baseAuthorityYear="1904" box="[244,585,1016,1035]" class="Insecta" family="Histeridae" genus="Reichardtiolus" kingdom="Animalia" order="Coleoptera" pageId="193" pageNumber="186" phylum="Arthropoda" rank="species" species="duriculus">
<emphasis id="B958FC5AFF5609E861A3FC1ECFEDFBF5" box="[244,455,1017,1035]" italics="true" pageId="193" pageNumber="186">Reichardtiolus duriculus</emphasis>
(
<bibRefCitation id="EFBD5DB9FF5609E86086FC1FCC69FBF5" author="REITTER E." box="[465,579,1016,1035]" pageId="193" pageNumber="186" pagination="29 - 36" refId="ref120480" refString="REITTER E. 1904: Ueber neue und wenig bekannte Histeriden (Coleoptera). Wiener Entomologische Zeitung 23: 29 - 36." type="journal article" year="1904">Reitter, 1904</bibRefCitation>
)
</taxonomicName>
, male terminalia: 604 – 8 
<superScript id="7C598D00FF5609E8624AFC11CD0DFBFC" attach="left" box="[797,807,1014,1026]" fontSize="5" pageId="193" pageNumber="186">th</superScript>
sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9 
<superScript id="7C598D00FF5609E86308FBF6CC43FBE3" attach="left" box="[607,617,1041,1053]" fontSize="5" pageId="193" pageNumber="186">th</superScript>
tergite, 10 
<superScript id="7C598D00FF5609E86393FBF6CCE4FBE3" attach="left" box="[708,718,1041,1053]" fontSize="5" pageId="193" pageNumber="186">th</superScript>
tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9 
<superScript id="7C598D00FF5609E86060FBCBCF6BFBC6" attach="left" box="[311,321,1068,1080]" fontSize="5" pageId="193" pageNumber="186">th</superScript>
tergite, 10 
<superScript id="7C598D00FF5609E860C9FBCBCF82FBC6" attach="left" box="[414,424,1068,1080]" fontSize="5" pageId="193" pageNumber="186">th</superScript>
tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8B932048FF5609E861C7FB7CCD2CFAED" blockId="193.[113,1089,1179,1587]" pageId="193" pageNumber="186">
Anterior margin of mesoventrite (
<figureCitation id="13173CCDFF5609E860ABFB7CCC72FB4D" box="[508,600,1179,1203]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="193" pageNumber="186">Fig. 599</figureCitation>
) feebly emarginate medially; discal marginal mesoventral stria well-impressed, carinate, slightly weakened anteriorly; disc of mesoventrite with scattered deep, round punctures, fringed with microscopic setae; meso-metaventral sutural stria absent; meso-metaventral suture well discernible.
</paragraph>
<paragraph id="8B932048FF5609E861C7FAFCCF62FA0D" blockId="193.[113,1089,1179,1587]" pageId="193" pageNumber="186">
Intercoxal disc of metaventrite longitudinally concave in male, with coarse scattered punctures, area around lateral metaventral stria smooth; lateral metaventral stria (
<figureCitation id="13173CCDFF5609E862C4FADCCDC7FAAD" box="[915,1005,1339,1363]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="193" pageNumber="186">Fig. 600</figureCitation>
) deeply impressed, carinate, extending obliquely and shortened apically; lateral disc of metaventrite (
<figureCitation id="13173CCDFF5609E8612EFA9CCEFFFA6D" box="[121,213,1403,1427]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="193" pageNumber="186">Fig. 600</figureCitation>
) with shallow large setiferous punctures; metepisternum on basal half with similar punctation, apical half of metepisternum (
<figureCitation id="13173CCDFF5609E8637AFA7CCCA3FA4D" box="[557,649,1435,1459]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="193" pageNumber="186">Fig. 600</figureCitation>
) almost smooth, fused metepimeron with few punctures; metepisternal stria present along entire fused metepimeron and metepisternum, basally intermittent.
</paragraph>
<paragraph id="8B932048FF5609E861C7FA1CCF1EF9CD" blockId="193.[113,1089,1179,1587]" pageId="193" pageNumber="186">Intercoxal disc of the first abdominal sternite completely striate laterally, with sparse coarse punctation.</paragraph>
<paragraph id="8B932048FF5509EB61E2FF3CCCF6FD8D" blockId="194.[151,1127,219,1172]" pageId="194" pageNumber="191">
Protibia (
<figureCitation id="13173CCDFF5509EB604CFF3CCF52FF0D" box="[283,376,219,243]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="194" pageNumber="191">Fig. 603</figureCitation>
) flattened and somewhat dilated, anterior margin formed by anterior margin of large sub-triangular distal-most tooth topped with large triangular denticle, outer margin apart from this tooth with another similar tooth topped with large triangular denticle, followed by another, much lower tooth topped by much smaller triangular denticle and another microscopic denticle entombed in outer margin of protibia; setae of outer row sparse, regular and short; setae of median row similarly sparse and regular, much shorter than those of outer row; protarsal groove moderately deep; anterior protibial stria present only on basal third; tarsal denticles absent; protibial spur tiny, bent, growing out from apical protibial margin; apical margin of protibia posteriorly without denticles; outer part of posterior surface of protibia sparsely punctate, distinctly separated from glabrous median part of posterior surface by irregular costiform stria fringed with sparse microscopic setae; posterior protibial stria complete, deeply impressed, with sparse microscopic setae; inner-ventral denticles absent; inner margin with single row of well sclerotized setae.
</paragraph>
<paragraph id="8B932048FF5509EB61E2FD9CCD28FC6D" blockId="194.[151,1127,219,1172]" pageId="194" pageNumber="191">
Mesotibia (
<figureCitation id="13173CCDFF5509EB6079FD9CCFA2FD6D" box="[302,392,635,659]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="194" pageNumber="191">Fig. 601</figureCitation>
) slightly thickened, outer margin with two sparse rows of thin denticles growing in size apically; setae of outer row rather dense, strongly sclerotized and longer than denticles of outer margin; setae of median row sparse, microscopic; posterior mesotibial stria inconspicuous; anterior surface of mesotibia imbricate, with scattered minuscule punctures with microscopic setae; anterior mesotibial stria shortened apically, almost complete; mesotibial spur stout, rather short; apical margin with several tiny denticles; claws of apical tarsomere longer than half its length; metatibia basically similar to mesotibia, but much more thickened and dilated, rows of denticles of outer margin widely separated, outer row of denticles (
<figureCitation id="13173CCDFF5509EB6076FC9CCF56FC6D" box="[289,380,891,915]" captionStart="Figs" captionStartId="192.[151,190,1521,1540]" captionTargetBox="[240,1040,219,1499]" captionTargetId="figure-11@192.[240,1040,219,1499]" captionTargetPageId="192" captionText="Figs. 593–603. Reichardtiolus duriculus (Reitter, 1904), SEM micrographs: 593 – habitus, dorsal view; 594 – ditto, ventral view; 595 – antennal club, dorsal view; 596 – head, dorsal view; 597 – ditto, ventral view; 598 – prosternum; 599 – mesoventrite; 600 – lateral disc of metaventrite, metepisternum and fused metepimeron; 601 – mesotibia, dorsal view; 602 – metatibia, ventral view; 603 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272282" httpUri="https://zenodo.org/record/4272282/files/figure.png" pageId="194" pageNumber="191">Fig. 602</figureCitation>
) observable only from ventral view.
</paragraph>
<paragraph id="8B932048FF5509EB61E2FC7CCDC4FBAD" blockId="194.[151,1127,219,1172]" pageId="194" pageNumber="191">
Male genitalia. Eighth sternite (
<figureCitation id="13173CCDFF5509EB6357FC7CCCB3FC4D" box="[512,665,923,947]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Figs. 604–605</figureCitation>
) longitudinally separated medially, apically with large setose inflatable membrane (velum), with numerous dense short setae; eighth tergite and eighth sternite fused laterally (
<figureCitation id="13173CCDFF5509EB636FFC3CCCBCFC0D" box="[568,662,987,1011]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Fig. 606</figureCitation>
). Morphology of 9 
<superScript id="7C598D00FF5509EB6209FC3DCD40FC17" attach="left" box="[862,874,986,1001]" fontSize="6" pageId="194" pageNumber="191">th</superScript>
tergite (
<figureCitation id="13173CCDFF5509EB6293FC3CCA75FC0D" box="[964,1119,987,1011]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Figs. 607–608</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="13173CCDFF5509EB632CFC1CCCFDFBED" box="[635,727,1019,1043]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Fig. 607</figureCitation>
) expanded on both ends. Basal piece of aedeagus (
<figureCitation id="13173CCDFF5509EB6072FBFCCFEBFBCD" box="[293,449,1051,1075]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Figs. 609–610</figureCitation>
) short, ratio of its length: length of parameres 1: 5; parameres fused along their basal third; aedeagus conspicuously curved ventrad (
<figureCitation id="13173CCDFF5509EB62D3FBDCCDC8FBAD" box="[900,994,1083,1107]" captionStart="Figs" captionStartId="193.[113,152,1016,1035]" captionTargetBox="[252,954,227,997]" captionTargetId="figure-139@193.[221,980,217,1010]" captionTargetPageId="193" captionText="Figs. 604–610. Reichardtiolus duriculus (Reitter, 1904), male terminalia: 604 – 8th sternite and tergite, ventral view; 605 – ditto, dorsal view; 606 – ditto, lateral view; 607 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 608 – 9th tergite, 10th tergite and spiculum gastrale, lateral view; 609 – aedeagus, dorsal view; 610 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272284" httpUri="https://zenodo.org/record/4272284/files/figure.png" pageId="194" pageNumber="191">Fig. 610</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF5509EB61C0FBBBCDBDFB6D" pageId="194" pageNumber="191" type="discussion">
<paragraph id="8B932048FF5509EB61C0FBBBCDBDFB6D" blockId="194.[151,1127,219,1172]" pageId="194" pageNumber="191">
<emphasis id="B958FC5AFF5509EB61C0FBBBCF23FB8A" bold="true" box="[151,265,1116,1140]" pageId="194" pageNumber="191">Remarks.</emphasis>
<bibRefCitation id="EFBD5DB9FF5509EB6043FBBBCFDAFB8D" author="MAZUR S." box="[276,496,1115,1140]" pageId="194" pageNumber="191" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997: 265)</bibRefCitation>
erroneously stated the 
<typeStatus id="54979EEAFF5509EB6257FBBBCD05FB8A" box="[768,815,1116,1140]" pageId="194" pageNumber="191">type</typeStatus>
locality for this species as ‘Kazakhstan’. In fact, it is the city of 
<collectingRegion id="49E8EEAAFF5509EB6371FB9BCC48FB6A" box="[550,610,1148,1172]" country="Turkmenistan" name="Mary Welayaty" pageId="194" pageNumber="191">Mary</collectingRegion>
, which lies in 
<collectingCountry id="F33B60D8FF5509EB63ABFB9CCDBEFB6D" box="[764,916,1147,1171]" name="Turkmenistan" pageId="194" pageNumber="191">Turkmenistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>