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<document id="75841CA39AF1F97792F3DB50918F40BE" ID-DOI="10.26028/cybium/2023-034" ID-ISSN="2101-0315" ID-Zenodo-Dep="12713757" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720651148950" checkinUser="felipe" docAuthor="Keith, Philippe &amp; Mennesson, Marion I." docDate="2023" docId="03EA87D826190D11FEFDF88EF5D2F863" docLanguage="en" docName="Cybium.47.4.431-466.pdf" docOrigin="Cybium 47 (4)" docSource="http://dx.doi.org/10.26028/cybium/2023-034" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Butis butis" docType="treatment" docVersion="3" lastPageNumber="445" masterDocId="FFD3FFA026170D1FFFD5F857F53CFF10" masterDocTitle="Review of Butis (Teleostei: Butidae) from Indo-Pacific islands with description of three new species" masterLastPageNumber="466" masterPageNumber="431" pageNumber="445" updateTime="1720659241642" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DFDAB8EA369E38324AC2CCE02A416FA8">Review of Butis (Teleostei: Butidae) from Indo-Pacific islands with description of three new species</mods:title>
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<mods:namePart id="EA2B7A49B69777979F98FB0ED172569A">Keith, Philippe</mods:namePart>
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<mods:namePart id="2E8B57E943B2DFEEBFC2C102BE2E763B">Mennesson, Marion I.</mods:namePart>
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<treatment id="03EA87D826190D11FEFDF88EF5D2F863" ID-DOI="http://doi.org/10.5281/zenodo.12714196" ID-Zenodo-Dep="12714196" LSID="urn:lsid:plazi:treatment:03EA87D826190D11FEFDF88EF5D2F863" httpUri="http://treatment.plazi.org/id/03EA87D826190D11FEFDF88EF5D2F863" lastPageNumber="445" pageId="14" pageNumber="445">
<subSubSection id="C359654526190D11FEFDF88EF75FFFE1" box="[296,611,217,241]" pageId="14" pageNumber="445" type="nomenclature">
<paragraph id="8BFC36CE26190D11FEFDF88EF75FFFE1" blockId="14.[296,611,217,241]" box="[296,611,217,241]" pageId="14" pageNumber="445">
<heading id="D0B481A226190D11FEFDF88EF75FFFE1" bold="true" box="[296,611,217,241]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="445" reason="2">
<taxonomicName id="4C434D4D26190D11FEFDF88EF75FFFE1" ID-CoL="NZ3X" authority="(Hamilton, 1822)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[296,611,217,241]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FEFDF88EF75FFFE1" bold="true" box="[296,611,217,241]" pageId="14" pageNumber="445">
<emphasis id="B937EADC26190D11FEFDF88EF4A1FFE1" bold="true" box="[296,413,217,241]" italics="true" pageId="14" pageNumber="445">Butis butis</emphasis>
(
<bibRefCitation id="EFD24B3F26190D11FE7FF88EF760FFE1" author="HAMILTON F." box="[426,604,217,241]" pageId="14" pageNumber="445" refId="ref26231" refString="HAMILTON F., 1822. - An Account of the Fishes Found in the River Ganges and its Branches. Edinburgh &amp; London: 405 p." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359654526190D11FEF9F8AEF724F9E3" pageId="14" pageNumber="445" type="description">
<paragraph id="8BFC36CE26190D11FEF9F8AEF763FE01" blockId="14.[300,607,249,273]" box="[300,607,249,273]" pageId="14" pageNumber="445">
(
<figureCitation id="13782A4B26190D11FEE1F8AEF49CFE01" box="[308,416,249,273]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[1077,1141,1813,1835]" captionStartId-1="5.[126,189,1574,1596]" captionStartId-2="7.[132,195,1245,1267]" captionTargetBox-0="[138,1058,221,1894]" captionTargetBox-1="[132,1425,407,1517]" captionTargetBox-2="[133,1454,216,1229]" captionTargetId-0="figure-54@2.[132,1060,215,1900]" captionTargetId-1="figure-154@5.[132,1455,401,1558]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="Figure 1. A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." captionText-1="Figure 2 (see also following page). A: Bayesian tree of the cytochrome c oxidase subunit (COI; 452 bp) for sequenced specimens of Butis. Numbers at each node represent posterior probabilities. B: Barcoding NJ K2P tree of the COI gene (196 bp) for sequenced specimens of Butis including the holotype of Butis humeralis from the MNHN collection. Numbers at each node represent bootstrap values. C, D: Barcoding NJ K2P trees of the 12S and the 16S genes (129 and 270 bp respectively) for sequenced specimens of Butis collected during the 19th century from the MNHN collection including a paratype of B. audebertae (MNHN A-1503). Numbers at each node represent bootstrap values. [A high resolution version is available online.]" captionText-2="Figure 3. A, B: Distribution of Butis species." figureDoi-0="http://doi.org/10.5281/zenodo.12713759" figureDoi-1="http://doi.org/10.5281/zenodo.12713761" figureDoi-2="http://doi.org/10.5281/zenodo.12713763" httpUri-0="https://zenodo.org/record/12713759/files/figure.png" httpUri-1="https://zenodo.org/record/12713761/files/figure.png" httpUri-2="https://zenodo.org/record/12713763/files/figure.png" pageId="14" pageNumber="445">Figs 1-3B</figureCitation>
,
<figureCitation id="13782A4B26190D11FE7EF8AEF484FE01" box="[427,440,249,273]" captionStart="Figure 5" captionStartId="15.[132,199,1556,1578]" captionTargetBox="[132,775,215,1539]" captionTargetId="figure-567@15.[132,775,439,1012]" captionTargetPageId="15" captionText="Figure 5. A: Butis butis, MNHN 2022-0233, neotype (male, 55.2 mm SL), India (Photo P. Keith); B: Butis butis, Hamiltons original, unpublished illustration (from Britz, 2019); C: Butis butis, Fly River, Papua New Guinea (Photo G. Allen); D: Butis butis, MNHN 2022-0213 (male, 95 mm SL), Solomon, tag 17612 (Photo P. Keith)." figureDoi="http://doi.org/10.5281/zenodo.12713767" httpUri="https://zenodo.org/record/12713767/files/figure.png" pageId="14" pageNumber="445">5</figureCitation>
; Tables III-IV)
</paragraph>
<paragraph id="8BFC36CE26190D11FF7FF96EF7DAFD61" blockId="14.[132,776,313,1779]" pageId="14" pageNumber="445">
The species of the genus
<taxonomicName id="4C434D4D26190D11FE0DF96DF72FFE41" authorityName="Bleeker" authorityYear="1856" box="[472,531,314,337]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="genus">
<emphasis id="B937EADC26190D11FE0DF96DF72FFE41" box="[472,531,314,337]" italics="true" pageId="14" pageNumber="445">Butis</emphasis>
</taxonomicName>
are often fished and numerous papers refer to these species. As it exists several cryptic species (this paper) and at least 6 different species in
<collectingCountry id="F354765E26190D11FF51F9CEF587FEA1" box="[132,187,409,433]" name="India" pageId="14" pageNumber="445">India</collectingCountry>
(this paper), it is necessary to clarify the status of
<taxonomicName id="4C434D4D26190D11FD05F9CDF4BCFEC2" authority="(Hamilton, 1822)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FD05F9CDF585FEC1" italics="true" pageId="14" pageNumber="445">Butis butis</emphasis>
(
<bibRefCitation id="EFD24B3F26190D11FF1FF9EEF44BFEC2" author="HAMILTON F." box="[202,375,441,466]" pageId="14" pageNumber="445" refId="ref26231" refString="HAMILTON F., 1822. - An Account of the Fishes Found in the River Ganges and its Branches. Edinburgh &amp; London: 405 p." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F354765E26190D11FE17F9EEF4C7FEC1" box="[450,507,441,465]" name="India" pageId="14" pageNumber="445">India</collectingCountry>
(type locality), which is one of the most cited species. This name has been used for a species of
<taxonomicName id="4C434D4D26190D11FF26F9ADF416FD01" authorityName="Bleeker" authorityYear="1856" box="[243,298,506,529]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="genus">
<emphasis id="B937EADC26190D11FF26F9ADF416FD01" box="[243,298,506,529]" italics="true" pageId="14" pageNumber="445">Butis</emphasis>
</taxonomicName>
since 1822 and it seems reasonable to decide to settle it. As the type of this species is not objectively identifiable, the name
<taxonomicName id="4C434D4D26190D11FE86FA6DF495FD42" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[339,425,570,594]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FE86FA6DF495FD42" box="[339,425,570,594]" italics="true" pageId="14" pageNumber="445">B. butis</emphasis>
</taxonomicName>
can only be linked to a known taxon by a
<typeStatus id="54F8886C26190D11FF2CFA0DF471FD62" box="[249,333,602,626]" pageId="14" pageNumber="445" type="neotype">neotype</typeStatus>
designation (
<bibRefCitation id="EFD24B3F26190D11FE09FA0DF760FD62" author="ICZN" box="[476,604,602,626]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.1).
</paragraph>
<paragraph id="8BFC36CE26190D11FF7FFA2DF5EBFB62" blockId="14.[132,776,313,1779]" pageId="14" pageNumber="445">
It is currently considered that there are no
<typeStatus id="54F8886C26190D11FDA9FA2DF78AFD82" box="[636,694,634,658]" pageId="14" pageNumber="445">types</typeStatus>
known (
<bibRefCitation id="EFD24B3F26190D11FF58FACDF46FFDA2" author="FRICKE R. &amp; ESCHMEYER W. N. &amp; VAN DER LAAN R. &amp; EDS" box="[141,339,666,690]" pageId="14" pageNumber="445" refId="ref26140" refString="FRICKE R., ESCHMEYER W. N. &amp; VAN DER LAAN R. (EDS), 2022. - Eschmeyer's catalog of fishes: genera, species, references. Electronic version accessed 5 March 2023. http: // researcharchive. calacademy. org / research / ichthyology / catalog / fish- catmain. asp" type="url" year="2022">
Fricke
<emphasis id="B937EADC26190D11FF0CFACDF435FDA2" box="[217,265,666,690]" italics="true" pageId="14" pageNumber="445">et al</emphasis>
., 2022
</bibRefCitation>
) and
<bibRefCitation id="EFD24B3F26190D11FE45FACDF729FDA2" author="BRITZ R." box="[400,533,666,690]" pageId="14" pageNumber="445" refId="ref25815" refString="BRITZ R., 2019. - Francis Hamilton's Gangetic Fishes in colour. A new edition of the 1822 monograph, with reproductions of unpublished coloured illustrations. Ray Society, London: 405 p." type="book" year="2019">Britz (2019)</bibRefCitation>
, expert on Hamiltons work, wrote that there is no evidence at all that Hamilton preserved any fish when making his descriptions.
<bibRefCitation id="EFD24B3F26190D11FD78FA8DF5F7FC02" author="GUNTHER A." pageId="14" pageNumber="445" refId="ref26199" refString="GUNTHER A., 1861. - Catalogue of the Acanthopterygian Fishes in the Collection of the British Museum. Vol. 3. British Museum, London: 586 p." type="book" year="1861">Günther (1861)</bibRefCitation>
listed
<specimenCount id="9D45FD4726190D11FEC4FAADF49FFC02" box="[273,419,762,786]" count="1" pageId="14" pageNumber="445" type="generic">one specimen</specimenCount>
presented by G.R. Waterhouse as a probable
<typeStatus id="54F8886C26190D11FF28FB4DF410FC22" box="[253,300,794,818]" pageId="14" pageNumber="445">type</typeStatus>
(
<emphasis id="B937EADC26190D11FEE9FB4CF459FC22" box="[316,357,795,818]" italics="true" pageId="14" pageNumber="445">i.e.,</emphasis>
possible
<typeStatus id="54F8886C26190D11FE19FB4DF4C7FC22" box="[460,507,794,818]" pageId="14" pageNumber="445">type</typeStatus>
BMNH 1858.8.15.142).
<bibRefCitation id="EFD24B3F26190D11FF51FB6DF437FC42" author="HORA S. L." box="[132,267,826,850]" pageId="14" pageNumber="445" pagination="169 - 192" refId="ref26334" refString="HORA S. L., 1929. - An aid to the study of Hamilton Buchanan's &quot; Gangetic Fishes &quot;. Mem. Indian Mus., 9 (4): 169 - 192." type="journal article" year="1929">Hora (1929)</bibRefCitation>
indicated no Hamilton
<typeStatus id="54F8886C26190D11FDDAFB6DF702FC42" box="[527,574,826,850]" pageId="14" pageNumber="445">type</typeStatus>
material retained, arguing that none of the material presented by Waterhouse could be
<typeStatus id="54F8886C26190D11FF3CFB2DF41BFC82" box="[233,295,890,914]" pageId="14" pageNumber="445">types</typeStatus>
. After examination of the specimen BMNH 1858.8.15.142 (this study;
<figureCitation id="13782A4B26190D11FE61FBCDF731FCA2" box="[436,525,922,946]" captionStart="Figure 7" captionStartId="18.[813,880,1837,1859]" captionTargetBox="[813,1456,1219,1820]" captionTargetId="figure-780@18.[813,1456,1219,1529]" captionTargetPageId="18" captionText="Figure 7. A: Butis humeralis, BMNH 1858.8.15.142 (male, 83.5 mm SL), India (Photo L. Goodayle, BMNH); B: Butis humeralis, Thailand (Photo D.D. Tran; http://ffish.asia)." figureDoi="http://doi.org/10.5281/zenodo.12713771" httpUri="https://zenodo.org/record/12713771/files/figure.png" pageId="14" pageNumber="445">Fig. 7A</figureCitation>
;
<tableCitation id="C6C1037526190D11FDCCFBCDF75BFCA1" box="[537,615,922,946]" captionStart="Table I" captionStartId="30.[126,179,421,447]" captionText="Table I. Continued." pageId="14" pageNumber="445">Table I</tableCitation>
), we consider that it is a
<taxonomicName id="4C434D4D26190D11FF27FBECF447FCC2" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[242,379,954,978]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="humeralis">
<emphasis id="B937EADC26190D11FF27FBECF447FCC2" box="[242,379,954,978]" italics="true" pageId="14" pageNumber="445">B. humeralis</emphasis>
</taxonomicName>
and there is no evidence that it could be the
<typeStatus id="54F8886C26190D11FF1BFB8CF5C1FCE3" box="[206,253,987,1011]" pageId="14" pageNumber="445">type</typeStatus>
<taxonomicName id="4C434D4D26190D11FED1FB8CF465FCE2" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[260,345,986,1010]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FED1FB8CF465FCE2" box="[260,345,986,1010]" italics="true" pageId="14" pageNumber="445">B. butis</emphasis>
</taxonomicName>
according to the description of Hamilton. Finally, J. Maclaine (BMNH, Senior Curator, fish section; pers. comm.) confirmed that there is no
<typeStatus id="54F8886C26190D11FDB2FC4CF7A9FB23" box="[615,661,1051,1075]" pageId="14" pageNumber="445">type</typeStatus>
in BMNH considered to be the
<taxonomicName id="4C434D4D26190D11FEB0FC6CF484FB42" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[357,440,1082,1106]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FEB0FC6CF484FB42" box="[357,440,1082,1106]" italics="true" pageId="14" pageNumber="445">B. butis</emphasis>
</taxonomicName>
of Hamilton (
<bibRefCitation id="EFD24B3F26190D11FD87FC6DF7E9FB42" author="ICZN" box="[594,725,1082,1106]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.4).
</paragraph>
<paragraph id="8BFC36CE26190D11FF7FFC2CF724F9E3" blockId="14.[132,776,313,1779]" pageId="14" pageNumber="445">
For these reasons, to avoid confusion we designate here the specimen MNHN 2022-0233 (male,
<quantity id="4CBB9B2B26190D11FDEEFCCDF7A0FBA2" box="[571,668,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="5.52" pageId="14" pageNumber="445" unit="mm" value="55.2">55.2 mm</quantity>
SL) from
<collectingCountry id="F354765E26190D11FF51FCEDF580FBC2" box="[132,188,1210,1234]" name="India" pageId="14" pageNumber="445">India</collectingCountry>
as
<typeStatus id="54F8886C26190D11FF34FCECF40BFBC3" box="[225,311,1211,1235]" pageId="14" pageNumber="445" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C434D4D26190D11FE89FCECF7A8FBC2" authority="(Hamilton, 1822)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[348,660,1210,1234]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="445" phylum="Chordata" rank="species" species="butis">
<emphasis id="B937EADC26190D11FE89FCECF4ECFBC2" box="[348,464,1210,1234]" italics="true" pageId="14" pageNumber="445">Butis butis</emphasis>
(
<bibRefCitation id="EFD24B3F26190D11FE0AFCEDF7B0FBC2" author="HAMILTON F." box="[479,652,1210,1234]" pageId="14" pageNumber="445" refId="ref26231" refString="HAMILTON F., 1822. - An Account of the Fishes Found in the River Ganges and its Branches. Edinburgh &amp; London: 405 p." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="13782A4B26190D11FD76FCEDF7C6FBC2" box="[675,762,1210,1234]" captionStart="Figure 5" captionStartId="15.[132,199,1556,1578]" captionTargetBox="[132,775,215,1539]" captionTargetId="figure-567@15.[132,775,439,1012]" captionTargetPageId="15" captionText="Figure 5. A: Butis butis, MNHN 2022-0233, neotype (male, 55.2 mm SL), India (Photo P. Keith); B: Butis butis, Hamiltons original, unpublished illustration (from Britz, 2019); C: Butis butis, Fly River, Papua New Guinea (Photo G. Allen); D: Butis butis, MNHN 2022-0213 (male, 95 mm SL), Solomon, tag 17612 (Photo P. Keith)." figureDoi="http://doi.org/10.5281/zenodo.12713767" httpUri="https://zenodo.org/record/12713767/files/figure.png" pageId="14" pageNumber="445">Fig. 5A</figureCitation>
), which comes from the original type locality (
<bibRefCitation id="EFD24B3F26190D11FDAFFC8DF7C3FBE2" author="ICZN" box="[634,767,1242,1266]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.6). The MNHN is a recognized scientific institution that maintains a research collection, with proper facilities for preserving name-bearing types, and that makes them accessible for study (
<bibRefCitation id="EFD24B3F26190D11FEFBFD0CF48DFA63" author="ICZN" box="[302,433,1371,1395]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.7). Data and diagnostic characters differentiating from other taxa are given below (in the description and comparison paragraphs) to ensure recognition of the specimen designated (
<bibRefCitation id="EFD24B3F26190D11FD99FDECF7EEFAC3" author="ICZN" box="[588,722,1467,1491]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.2 and 75.3.3). The description of
<bibRefCitation id="EFD24B3F26190D11FDF2FD8CF7E1FAE3" author="HAMILTON F." box="[551,733,1499,1523]" pageId="14" pageNumber="445" refId="ref26231" refString="HAMILTON F., 1822. - An Account of the Fishes Found in the River Ganges and its Branches. Edinburgh &amp; London: 405 p." type="book" year="1822">Hamilton (1822)</bibRefCitation>
did not give sufficient or diagnostic data to state on the species (several species have the two red spots at the base of the pectoral fins as cited by
<bibRefCitation id="EFD24B3F26190D11FEB9FE6CF719F943" author="HAMILTON F." box="[364,549,1595,1619]" pageId="14" pageNumber="445" refId="ref26231" refString="HAMILTON F., 1822. - An Account of the Fishes Found in the River Ganges and its Branches. Edinburgh &amp; London: 405 p." type="book" year="1822">Hamilton (1822)</bibRefCitation>
and as shown in the unpublished drawing (
<figureCitation id="13782A4B26190D11FEA0FE0CF4F4F963" box="[373,456,1627,1651]" captionStart="Figure 5" captionStartId="15.[132,199,1556,1578]" captionTargetBox="[132,775,215,1539]" captionTargetId="figure-567@15.[132,775,439,1012]" captionTargetPageId="15" captionText="Figure 5. A: Butis butis, MNHN 2022-0233, neotype (male, 55.2 mm SL), India (Photo P. Keith); B: Butis butis, Hamiltons original, unpublished illustration (from Britz, 2019); C: Butis butis, Fly River, Papua New Guinea (Photo G. Allen); D: Butis butis, MNHN 2022-0213 (male, 95 mm SL), Solomon, tag 17612 (Photo P. Keith)." figureDoi="http://doi.org/10.5281/zenodo.12713767" httpUri="https://zenodo.org/record/12713767/files/figure.png" pageId="14" pageNumber="445">Fig. 5B</figureCitation>
) of Hamilton and reproduced by
<bibRefCitation id="EFD24B3F26190D11FF7CFE2CF40FF983" author="BRITZ R." box="[169,307,1659,1683]" pageId="14" pageNumber="445" refId="ref25815" refString="BRITZ R., 2019. - Francis Hamilton's Gangetic Fishes in colour. A new edition of the 1822 monograph, with reproductions of unpublished coloured illustrations. Ray Society, London: 405 p." type="book" year="2019">Britz (2019)</bibRefCitation>
, or have 18 rays in pectoral fins). Thus a
<typeStatus id="54F8886C26190D11FF51FECCF5E5F9A3" box="[132,217,1691,1715]" pageId="14" pageNumber="445" type="neotype">neotype</typeStatus>
designation is needed to clarify the identity, and the
<typeStatus id="54F8886C26190D11FF51FEECF5E5F9C3" box="[132,217,1723,1747]" pageId="14" pageNumber="445" type="neotype">neotype</typeStatus>
here is consistent with the poor original description (see below) (
<bibRefCitation id="EFD24B3F26190D11FEDBFE8CF4B2F9E3" author="ICZN" box="[270,398,1755,1779]" pageId="14" pageNumber="445" refId="ref26371" refString="ICZN, 1999. - International Code of Zoological Nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN, 1999</bibRefCitation>
: art. 75.3.5).
</paragraph>
</subSubSection>
<subSubSection id="C359654526190D11FF51FF4CF5D2F863" pageId="14" pageNumber="445" type="materials_examined">
<paragraph id="8BFC36CE26190D11FF51FF4CF467F823" blockId="14.[132,775,1819,1907]" box="[132,347,1819,1843]" pageId="14" pageNumber="445">
<emphasis id="B937EADC26190D11FF51FF4CF467F823" bold="true" box="[132,347,1819,1843]" pageId="14" pageNumber="445">Material examined</emphasis>
</paragraph>
<paragraph id="8BFC36CE26190D11FF7FFF6CF5D2F863" blockId="14.[132,775,1819,1907]" pageId="14" pageNumber="445">
<materialsCitation id="3B2B3C9326190D11FF7FFF6CF5D5F863" collectionCode="MNHN" country="India" location="India" pageId="14" pageNumber="445" specimenCode="MNHN 2022-0233" specimenCount="1" typeStatus="neotype">
<specimenCode id="DBE59EB526190D11FF7FFF6CF440F843" box="[170,380,1851,1875]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="445">MNHN 2022-0233</specimenCode>
,
<typeStatus id="54F8886C26190D11FE5DFF6CF4DEF843" box="[392,482,1851,1875]" pageId="14" pageNumber="445" type="neotype">neotype</typeStatus>
, male (
<quantity id="4CBB9B2B26190D11FDFAFF6CF7ADF842" box="[559,657,1851,1875]" metricMagnitude="-2" metricUnit="m" metricValue="5.52" pageId="14" pageNumber="445" unit="mm" value="55.2">55.2 mm</quantity>
SL),
<collectingCountry id="F354765E26190D11FD1AFF6CF63BF843" box="[719,775,1851,1875]" name="India" pageId="14" pageNumber="445">India</collectingCountry>
(
<figureCitation id="13782A4B26190D11FF59FF0CF5DEF863" box="[140,226,1883,1907]" captionStart="Figure 5" captionStartId="15.[132,199,1556,1578]" captionTargetBox="[132,775,215,1539]" captionTargetId="figure-567@15.[132,775,439,1012]" captionTargetPageId="15" captionText="Figure 5. A: Butis butis, MNHN 2022-0233, neotype (male, 55.2 mm SL), India (Photo P. Keith); B: Butis butis, Hamiltons original, unpublished illustration (from Britz, 2019); C: Butis butis, Fly River, Papua New Guinea (Photo G. Allen); D: Butis butis, MNHN 2022-0213 (male, 95 mm SL), Solomon, tag 17612 (Photo P. Keith)." figureDoi="http://doi.org/10.5281/zenodo.12713767" httpUri="https://zenodo.org/record/12713767/files/figure.png" pageId="14" pageNumber="445">Fig. 5A</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>