219 lines
29 KiB
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219 lines
29 KiB
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<mods:title id="21F37932B363EBEAD1452777B9060ED9">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="F669BCF83151DFC6BE488AF57D3EBB33">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="3723E76F837320F28BCD63D90637BBE6">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:affiliation id="9DE36082783B3DEEA222D10535BEFCA1">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<taxonomicName id="4C634D67FFCFFFF0FDA1477EBB593FAA" ID-CoL="4FSK3" authority="Moctezuma & Halffter, 2017" authorityName="Moctezuma & Halffter" authorityYear="2017" box="[521,1067,1397,1424]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zoque">
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<emphasis id="B917EAF6FFCFFFF0FDA1477EBDBD3FAA" bold="true" box="[521,719,1397,1424]" italics="true" pageId="49" pageNumber="50">Phanaeus zoque</emphasis>
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<bibRefCitation id="EFF24B15FFCFFFF0FD7E477DBB593FAA" author="Moctezuma V. & Halffter G." box="[726,1067,1398,1424]" pageId="49" pageNumber="50" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter, 2017</bibRefCitation>
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<paragraph id="8BDC36E4FFCFFFF0FD3D4793BCEC3F88" blockId="49.[521,1067,1397,1459]" box="[661,926,1432,1459]" pageId="49" pageNumber="50">
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<figureCitation id="13582A61FFCFFFF0FD3D4793BD863F88" box="[661,756,1432,1458]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="49" pageNumber="50">Figs 1Q</figureCitation>
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,
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<figureCitation id="13582A61FFCFFFF0FCAA4793BC533F89" box="[770,801,1432,1459]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="49" pageNumber="50">15</figureCitation>
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,
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<figureCitation id="13582A61FFCFFFF0FC864793BC133F88" box="[814,865,1432,1458]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="49" pageNumber="50">18R</figureCitation>
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,
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<figureCitation id="13582A61FFCFFFF0FCC54793BCEC3F88" box="[877,926,1432,1458]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="49" pageNumber="50">19R</figureCitation>
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<taxonomicName id="4C634D67FFCFFFF0FF1547D2BC7A3FC9" ID-CoL="4FSK3" authority="Moctezuma & Halffter, 2017: 47" authorityName="Moctezuma & Halffter" authorityPageNumber="47" authorityYear="2017" box="[189,776,1497,1524]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zoque">
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<emphasis id="B917EAF6FFCFFFF0FF1547D2BE0E3FC9" box="[189,380,1497,1523]" italics="true" pageId="49" pageNumber="50">Phanaeus zoque</emphasis>
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<bibRefCitation id="EFF24B15FFCFFFF0FE2C47D1BC7A3FC9" author="Moctezuma V. & Halffter G." box="[388,776,1497,1524]" pageId="49" pageNumber="50" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter, 2017: 47</bibRefCitation>
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, 48, 52, 55–56, figs 1–15, 23.
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<emphasis id="B917EAF6FFCFFFF0FF154411BEF03C0E" box="[189,386,1562,1588]" italics="true" pageId="49" pageNumber="50">Phanaeus zoque</emphasis>
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– Moctezuma
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<emphasis id="B917EAF6FFCFFFF0FDEA4410BD093C0E" box="[578,635,1562,1588]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
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. 2017: 113–114, 126, 128, 131–132, fig. 19; 2019: 252–253, fig. 5. —
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<treatmentCitation id="0AC210F5FFCFFFF0FEF24436BDFB3C6D" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[346,649,1597,1623]" page="83" pageId="49" pageNumber="50" year="2018">
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<bibRefCitation id="EFF24B15FFCFFFF0FEF24436BDFB3C6D" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[346,649,1597,1623]" pageId="49" pageNumber="50" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
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Kohlmann
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<emphasis id="B917EAF6FFCFFFF0FE754435BD633C6D" box="[477,529,1597,1623]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
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. 2018: 83
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, 88.
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<emphasis id="B917EAF6FFCFFFF0FF154475BE1E3CA2" bold="true" box="[189,364,1662,1688]" pageId="49" pageNumber="50">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDC36E4FFCFFFF0FF4244A7BE2E3CFC" blockId="49.[189,1398,1708,1803]" box="[234,348,1708,1734]" pageId="49" pageNumber="50">
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<emphasis id="B917EAF6FFCFFFF0FF4244A7BE2E3CFC" bold="true" box="[234,348,1708,1734]" pageId="49" pageNumber="50">
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<typeStatus id="54D88846FFCFFFF0FF4244A7BE2E3CFC" box="[234,348,1708,1734]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
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|
||
<collectingCountry id="F3747674FFCFFFF0FF1544C4BE433CD3" box="[189,305,1743,1769]" name="Mexico" pageId="49" pageNumber="50">MEXICO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D65FD6DFFCFFFF0FEC944C4BE0B3CD3" box="[353,377,1743,1769]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
|
||
,
|
||
<bibRefCitation id="EFF24B15FFCFFFF0FE3944C4BC483CD3" author="Moctezuma V. & Halffter G." box="[401,826,1742,1769]" pageId="49" pageNumber="50" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017: 48</bibRefCitation>
|
||
(revised);
|
||
<collectingRegion id="49A7F806FFCFFFF0FC7944C4BB5E3CD3" box="[977,1068,1743,1769]" country="Mexico" name="Oaxaca" pageId="49" pageNumber="50">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BB8AC9EFFCFFFF0FBEB44C4BA003CD2" box="[1091,1394,1742,1769]" pageId="49" pageNumber="50">Santa María Chimalapa</collectingMunicipality>
|
||
,
|
||
<location id="8EBC603FFFCFFFF0FF1544FABE0B3D31" LSID="urn:lsid:plazi:treatment:03CA87F2FFCFFFF3FDA1477EBDC13913:8EBC603FFFCFFFF0FF1544FABE0B3D31" box="[189,377,1777,1803]" country="Mexico" municipality="Santa Maria Chimalapa" name="Chocomanatlan" pageId="49" pageNumber="50" stateProvince="Oaxaca">Chocomanatlán</location>
|
||
;
|
||
<collectionCode id="ED72AE21FFCFFFF0FE2D44FABEBD3D31" box="[389,463,1777,1803]" pageId="49" pageNumber="50">IEXA</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCFFFF0FF154539BE2F3D76" blockId="49.[189,349,1842,1868]" box="[189,349,1842,1868]" pageId="49" pageNumber="50">
|
||
<heading id="D0948188FFCFFFF0FF154539BE2F3D76" bold="true" box="[189,349,1842,1868]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
|
||
<emphasis id="B917EAF6FFCFFFF0FF154539BE2F3D76" bold="true" box="[189,349,1842,1868]" pageId="49" pageNumber="50">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCFFFF0FF15456ABB1F3D46" blockId="49.[189,1133,1889,1916]" box="[189,1133,1889,1916]" pageId="49" pageNumber="50">
|
||
<collectingCountry id="F3747674FFCFFFF0FF15456ABE683D41" box="[189,282,1889,1915]" name="Mexico" pageId="49" pageNumber="50">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A7F806FFCFFFF0FE8D456ABEF23D41" box="[293,384,1889,1915]" country="Mexico" name="Oaxaca" pageId="49" pageNumber="50">Oaxaca</collectingRegion>
|
||
, Santa María Chimalapa, Chocomanatlán (José López Portillo).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFCFFFF0FF1545AABBBD3DD3" box="[189,1231,1953,2025]" pageId="49" pageNumber="50" type="distribution">
|
||
<paragraph id="8BDC36E4FFCFFFF0FF1545AABE243D81" blockId="49.[189,342,1953,1979]" box="[189,342,1953,1979]" pageId="49" pageNumber="50">
|
||
<heading id="D0948188FFCFFFF0FF1545AABE243D81" bold="true" box="[189,342,1953,1979]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
|
||
<emphasis id="B917EAF6FFCFFFF0FF1545AABE243D81" bold="true" box="[189,342,1953,1979]" pageId="49" pageNumber="50">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCFFFF0FF1545C4BBBD3DD3" blockId="49.[189,1231,1998,2025]" box="[189,1231,1998,2025]" pageId="49" pageNumber="50">
|
||
Temperate mountains of Los Chimalapas, eastern
|
||
<collectingRegion id="49A7F806FFCFFFF0FCA045C5BC123DD2" box="[776,864,1998,2024]" country="Mexico" name="Oaxaca" pageId="49" pageNumber="50">Oaxaca</collectingRegion>
|
||
and western
|
||
<collectingRegion id="49A7F806FFCFFFF0FC5345C5BB283DD2" box="[1019,1114,1998,2024]" country="Mexico" name="Chiapas" pageId="49" pageNumber="50">Chiapas</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCFFFF0FBC245C5BBB33DD3" box="[1130,1217,1998,2025]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="49" pageNumber="50">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFCCFFF3FF154302BDC13913" pageId="50" pageNumber="51" type="discussion">
|
||
<paragraph id="8BDC36E4FFCCFFF3FF154302BE5D3B19" blockId="50.[189,303,265,291]" box="[189,303,265,291]" pageId="50" pageNumber="51">
|
||
<heading id="D0948188FFCCFFF3FF154302BE5D3B19" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="B917EAF6FFCCFFF3FF154302BE5D3B19" bold="true" box="[189,303,265,291]" pageId="50" pageNumber="51">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCCFFF3FF15433DBE6B38B8" blockId="50.[188,1400,310,642]" pageId="50" pageNumber="51">
|
||
Mean length
|
||
<quantity id="4C9B9B01FFCCFFF3FEF2433DBEB13B6A" box="[346,451,310,337]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="50" pageNumber="51" unit="mm" value="15.8">15.8 mm</quantity>
|
||
(
|
||
<quantity id="4C9B9B01FFCCFFF3FE7C433DBDF43B6A" box="[468,646,310,336]" metricMagnitude="-2" metricUnit="m" metricValue="1.5799999999999998" metricValueMax="1.8399999999999999" metricValueMin="1.3199999999999998" pageId="50" pageNumber="51" unit="mm" value="15.799999999999999" valueMax="18.4" valueMin="13.2">13.2–18.4 mm</quantity>
|
||
). To the original description we add the elytral striae impressed basally as distinct fossae; right and left lobes of endophallite copulatrix similar in size; right lobe obtusely triangular in shape, distinctly rounded superiorly; left lobe strongly developed, obtusely lobed; central ridge less developed than central column (
|
||
<figureCitation id="13582A61FFCCFFF3FCA94397BC2C3B8C" box="[769,862,412,438]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="50" pageNumber="51">Fig. 1Q</figureCitation>
|
||
). For the female, the head showing a cephalic trituberculate carina; with conical, similar in size tubercles; middle tubercle slightly more frontally projected than lateral tubercles; frons with almost effaced to effaced punctures; pronotal sculpture completely smooth, with almost effaced to effaced punctures; pronotal process trituberculate, with posterior concavity; with dentiform middle tubercle, strongly more developed and projected frontally than lateral tubercles; carinate or rounded lateral tubercles; posterior pronotal midline almost completely effaced.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCCFFF3FF1540A3BDC13913" blockId="50.[189,1399,680,810]" pageId="50" pageNumber="51">
|
||
After having described
|
||
<taxonomicName id="4C634D67FFCCFFF3FE7940A2BCEF38F8" authority="(Moctezuma & Halffter 2017)" baseAuthorityName="Moctezuma & Halffter" baseAuthorityYear="2017" box="[465,925,680,706]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="zoque">
|
||
<emphasis id="B917EAF6FFCCFFF3FE7940A2BD4038F8" box="[465,562,681,706]" italics="true" pageId="50" pageNumber="51">P. zoque</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B15FFCCFFF3FDE940A3BCE438F8" author="Moctezuma V. & Halffter G." box="[577,918,680,706]" pageId="50" pageNumber="51" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the following
|
||
<typeStatus id="54D88846FFCCFFF3FBE640A2BBCC38F9" box="[1102,1214,681,707]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
were deposited by us in additional collections: CEMT:
|
||
<specimenCount id="9D65FD6DFFCCFFF3FD3240C1BD9238DF" box="[666,736,714,741]" pageId="50" pageNumber="51" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFCCFFF3FD4740C1BC4738DF" box="[751,821,714,741]" pageId="50" pageNumber="51" type="female">5 ♀♀</specimenCount>
|
||
; CDC:
|
||
<specimenCount id="9D65FD6DFFCCFFF3FC3C40C1BCA838DF" box="[916,986,714,741]" pageId="50" pageNumber="51" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFCCFFF3FC4140C1BB5C38DF" box="[1001,1070,714,741]" pageId="50" pageNumber="51" type="female">5 ♀♀</specimenCount>
|
||
; CMNC:
|
||
<specimenCount id="9D65FD6DFFCCFFF3FB0F40C1BBAA38DF" box="[1191,1240,714,741]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFCCFFF3FB4F40C1BA6438DF" box="[1255,1302,714,741]" pageId="50" pageNumber="51" type="female">1 ♀</specimenCount>
|
||
; CPFA:
|
||
<specimenCount id="9D65FD6DFFCCFFF3FF1540E7BE71393D" box="[189,259,748,775]" pageId="50" pageNumber="51" type="male">6 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFCCFFF3FEB940E7BE24393D" box="[273,342,748,775]" pageId="50" pageNumber="51" type="female">6 ♀♀</specimenCount>
|
||
; UVGC:
|
||
<specimenCount id="9D65FD6DFFCCFFF3FE6240E7BE88393D" box="[458,506,748,775]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFCCFFF3FDA040E7BD45393D" box="[520,567,748,775]" pageId="50" pageNumber="51" type="female">1 ♀</specimenCount>
|
||
. These
|
||
<typeStatus id="54D88846FFCCFFF3FD3D40E6BC77393D" box="[661,773,749,775]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
were previously housed at VMC (see the full list of
|
||
<typeStatus id="54D88846FFCCFFF3FF15411BBE5F3910" box="[189,301,784,810]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<bibRefCitation id="EFF24B15FFCCFFF3FEFB411BBDD43913" author="Moctezuma V. & Halffter G." box="[339,678,783,810]" pageId="50" pageNumber="51" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |