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<document ID-DOI="http://doi.org/10.5281/zenodo.3789754" ID-GBIF-Dataset="7a68e13d-3c05-49d5-86a3-7bdb5c4af1ce" ID-GBIF-Taxon="163755349" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789754" ID-ZooBank="553C91CE-FEA1-4867-929B-2865647E4B2E" approvalRequired="14" approvalRequired_for_textStreams="14" checkinTime="1588002532591" checkinUser="plazi" docAuthor="Ballerio, Alberto &amp; Maruyama, Munetoshi" docDate="2010" docId="03CA87D50455FFF465FE49D34D7C2C27" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="ZooKeys 34 (34)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5da20b76" docStyleName="zookeys.2008.journal_article" docTitle="Madrasostes malayanum Paulian 1979" docType="treatment" docVersion="4" lastPageNumber="85" masterDocId="FFF3FFAD045DFFFC65724A5A4E182815" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="8" pageNumber="85" updateTime="1643557208252" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ballerio, Alberto</mods:namePart>
<mods:affiliation>, Brescia, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maruyama, Munetoshi</mods:namePart>
<mods:affiliation>The Kyushu University Museum, Fukuoka, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-01-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>34</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.34.268</mods:identifier>
<mods:identifier type="GBIF-Dataset">7a68e13d-3c05-49d5-86a3-7bdb5c4af1ce</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576606</mods:identifier>
<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789754" ID-GBIF-Taxon="163755349" ID-Zenodo-Dep="3789754" LSID="urn:lsid:plazi:treatment:03CA87D50455FFF465FE49D34D7C2C27" httpUri="http://treatment.plazi.org/id/03CA87D50455FFF465FE49D34D7C2C27" lastPageNumber="85" pageId="8" pageNumber="85">
<subSubSection pageId="8" pageNumber="85" type="nomenclature">
<paragraph blockId="8.[140,1107,905,1074]" box="[140,716,905,932]" pageId="8" pageNumber="85">
<heading box="[140,716,905,932]" fontSize="11" level="2" pageId="8" pageNumber="85" reason="3">
<taxonomicName authority="Paulian, 1979" authorityName="Paulian" authorityYear="1979" box="[140,608,905,932]" class="Insecta" family="Hybosoridae" genus="Madrasostes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="malayanum">
<emphasis bold="true" box="[140,608,905,932]" pageId="8" pageNumber="85">
<emphasis bold="true" box="[140,434,905,932]" italics="true" pageId="8" pageNumber="85">
<emphasis box="[140,286,905,932]" inLineHeading="true" pageId="8" pageNumber="85" reason="1">Madrasostes</emphasis>
malayanum
</emphasis>
<bibRefCitation author="Paulian R" box="[442,608,905,931]" pageId="8" pageNumber="85" pagination="171 - 174" refId="ref11483" refString="Paulian R (1979) Sur quelques ceratocanthides de Malaisie (Col. Scarabaeoidea). Bulletin de la Societe Entomologique de France 84: 171 - 174." type="journal article" year="1979">Paulian, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[623,708,906,932]" captionStart="Figure 2" captionStartId="15.[140,212,1558,1580]" captionText="Figure 2. Ceratocanthinae of Ulu Gombak. A Madrasostes depressum habitus dorsal B Madrasostes boucomonti habitus dorsal C Madrasostes agostii sp. n. habitus dorsal D Madrasostes clypeale male habitus dorsal E Madrasostes malyanum habitus dorsal F Madrasostes mirificum sp. n. habitus dorsal G habitus lateral of male H fore body of female I male protibia dorsal view Į male protibia lateral view." pageId="8" pageNumber="85">Fig. 2E</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="8.[140,1107,905,1074]" box="[185,703,941,968]" pageId="8" pageNumber="85">
Known distribution: West
<collectingCountry box="[488,587,941,967]" name="Malaysia" pageId="8" pageNumber="85">Malaysia</collectingCountry>
(
<collectingRegion box="[602,691,941,968]" country="Malaysia" name="Pahang" pageId="8" pageNumber="85">Pahang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="discussion">
<paragraph blockId="8.[140,1107,905,1074]" pageId="8" pageNumber="85">
Remarks. First record for
<collectingRegion box="[475,575,976,1002]" country="Malaysia" name="Selangor" pageId="8" pageNumber="85">Selangor</collectingRegion>
. This species was known only by its
<typeStatus box="[985,1033,977,1003]" pageId="8" pageNumber="85">type</typeStatus>
series, which consists only on females. The overall morphology suggests a close similarity to
<taxonomicName authority="Paulian, 1989" authorityName="Paulian" authorityYear="1989" box="[140,614,1047,1073]" class="Insecta" family="Hybosoridae" genus="Madrasostes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="parcepunctatum">
<emphasis box="[140,449,1047,1073]" italics="true" pageId="8" pageNumber="85">Madrasostes parcepunctatum</emphasis>
<bibRefCitation author="Paulian R" box="[456,614,1047,1073]" pageId="8" pageNumber="85" pagination="297 - 318" refId="ref11512" refString="Paulian R (1989) Notes sur les coleopteres Scarabaeoidea du Museum de Geneve. IV. Revue Suisse de Zoologie 96: 297 - 318." type="journal article" year="1989">Paulian, 1989</bibRefCitation>
</taxonomicName>
from Borneo (
<collectingRegion box="[782,856,1047,1074]" country="Malaysia" name="Sabah" pageId="8" pageNumber="85">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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