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<document id="214E08D9CB778B909E29A10FB3B8B820" ID-DOI="10.11646/zootaxa.5433.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954420" ID-ZooBank="7ADF6A54-E31C-453A-AA25-44DEDBCF2752" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712752719575" checkinUser="plazi" docAuthor="Souza, Leonardo Santos De, Araújo, Tarciso Almeida De &amp; Pimenta, Alexandre Dias" docDate="2024" docId="03CA878ABA7FDE2FFF4FF9627503FDA2" docLanguage="en" docName="zootaxa.5433.4.1.pdf" docOrigin="Zootaxa 5433 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aclis pulchra Souza, Araujo &amp; Pimenta" docType="treatment" docVersion="2" lastPageNumber="469" masterDocId="FFF3FFF2BA6EDE3DFFD8FFD77042FF9F" masterDocTitle="Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena" masterLastPageNumber="499" masterPageNumber="451" pageNumber="468" updateTime="1712753971016" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0BC312760279D17B23E4DC3AF7C3EA8A">Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena</mods:title>
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<mods:name id="2BED0E027D4AA2962F91941FA6C5A89D" type="personal">
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<mods:namePart id="BFF12FD3140222BA019722952A74765B">Souza, Leonardo Santos De</mods:namePart>
<mods:nameIdentifier id="6E503BADDB9A7F22630DDF035ABB19BD" type="ORCID">0000-0002-8242-010X</mods:nameIdentifier>
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<mods:namePart id="359D38AE7406D886373F71616648847C">Araújo, Tarciso Almeida De</mods:namePart>
<mods:nameIdentifier id="86719C9315814123D7EE8525A4F25056" type="ORCID">0000-0002-8242-010X</mods:nameIdentifier>
<mods:affiliation id="423CA48663BA51A993CAE41E1F6EF657">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista., São Cristóvão, 20940 - 040, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="55ACAD16A559E44C6F32F0028A9FB6AB">Pimenta, Alexandre Dias</mods:namePart>
<mods:nameIdentifier id="BE90B6889E35E92D21D74A65CA2746DB" type="ORCID">0000-0002-8242-010X</mods:nameIdentifier>
<mods:affiliation id="F69488DE60B4B77095226C0653A74DC3">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista., São Cristóvão, 20940 - 040, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:title id="6ED87CCCD0362C0FB1146814D27605BE">Zootaxa</mods:title>
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<mods:date id="0495AC4281334ACB1C2635ABD523A8FD">2024</mods:date>
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<mods:number id="686EE2AF64A93B7DD0B5A28E086A6248">2024-04-09</mods:number>
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<mods:detail id="58DCA9263158D628A3F8E3AC2497ACE4" type="volume">
<mods:number id="D2C6FB2DC03882B8CEB0174D8403CB58">5433</mods:number>
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<mods:number id="1263D7DEC8ABDCCD42FE2DC26C8F2BE3">4</mods:number>
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<mods:identifier id="7367807FF63CCF16E495B41D957E6DFA" type="DOI">10.11646/zootaxa.5433.4.1</mods:identifier>
<mods:identifier id="4829AA1FCA6F3C3A686FB25719875505" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2D7AA2487774C93C391C252D980B1DFD" type="Zenodo-Dep">10954420</mods:identifier>
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<treatment id="03CA878ABA7FDE2FFF4FF9627503FDA2" ID-DOI="http://doi.org/10.5281/zenodo.10954585" ID-Zenodo-Dep="10954585" LSID="urn:lsid:plazi:treatment:03CA878ABA7FDE2FFF4FF9627503FDA2" httpUri="http://treatment.plazi.org/id/03CA878ABA7FDE2FFF4FF9627503FDA2" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<subSubSection id="C3796517BA7FDE2CFF4FF9627328F950" box="[151,874,1717,1744]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8BDC369CBA7FDE2CFF4FF9627328F950" blockId="17.[151,874,1717,1780]" box="[151,874,1717,1744]" pageId="17" pageNumber="468">
<heading id="D09481F0BA7FDE2CFF4FF9627328F950" bold="true" box="[151,874,1717,1744]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="B917EA8EBA7FDE2CFF4FF9627328F950" bold="true" box="[151,874,1717,1744]" pageId="17" pageNumber="468">
<taxonomicName id="4C634D1FBA7FDE2CFF4FF96272E5F950" authority="Souza, Araujo &amp; Pimenta" authorityName="Souza, Araujo &amp; Pimenta" box="[151,679,1717,1744]" class="Gastropoda" family="Eulimidae" genus="Aclis" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="468" phylum="Mollusca" rank="species" species="pulchra">
<emphasis id="B917EA8EBA7FDE2CFF4FF9627091F94F" bold="true" box="[151,211,1717,1744]" italics="true" pageId="17" pageNumber="468">Aclis</emphasis>
cf.
<emphasis id="B917EA8EBA7FDE2CFED8F962711DF94F" bold="true" box="[256,351,1717,1744]" italics="true" pageId="17" pageNumber="468">pulchra</emphasis>
Souza, Araújo &amp; Pimenta
</taxonomicName>
(present study)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA7FDE2CFF4FF90E709BF96B" box="[151,217,1753,1780]" pageId="17" pageNumber="468" type="description">
<paragraph id="8BDC369CBA7FDE2CFF4FF90E709BF96B" blockId="17.[151,874,1717,1780]" box="[151,217,1753,1780]" pageId="17" pageNumber="468">
<figureCitation id="13582A19BA7FDE2CFF4FF90E709BF96B" box="[151,217,1753,1780]" captionStart="FIGURE 9" captionStartId="17.[151,250,1539,1563]" captionTargetBox="[151,1435,273,1515]" captionTargetId="figure-67@17.[151,1435,273,1515]" captionTargetPageId="17" captionText="FIGURE 9. Aclis cf. pulchra: MNRJ 34072. A, C. Shell in frontal view. B. Shell in lateral view. D. Detail of teleoconch surface. E. Detail of apical whorls. F–G. Detail of body whorl in frontal view. Scale bars: A–C. 1 mm; D. 20 μm; E. 200 μm; F–G. 500 μm." figureDoi="http://doi.org/10.5281/zenodo.10954447" httpUri="https://zenodo.org/record/10954447/files/figure.png" pageId="17" pageNumber="468">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA7FDE2FFF4FF8F67174FF66" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="8BDC369CBA7FDE2CFF4FF8F6715BF8FF" blockId="17.[151,1437,1825,2032]" pageId="17" pageNumber="468">
<emphasis id="B917EA8EBA7FDE2CFF4FF8F671C1F8A4" bold="true" box="[151,387,1825,1851]" pageId="17" pageNumber="468">Material examined.</emphasis>
<collectingCountry id="F374760CBA7FDE2CFE54F8F67197F8A4" box="[396,469,1825,1851]" name="Brazil" pageId="17" pageNumber="468">Brazil</collectingCountry>
:
<collectingRegion id="49A7F87EBA7FDE2CFE3AF8F67288F8A3" box="[482,714,1825,1852]" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="468">Rio de Janeiro state</collectingRegion>
: HAB 13 sta. H3;
<collectionCode id="ED72AE59BA7FDE2CFC7BF8F573ACF8A3" box="[931,1006,1826,1852]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="17" pageNumber="468">MNRJ</collectionCode>
16811 [1†]; HAB 16 sta. H3;
<collectionCode id="ED72AE59BA7FDE2CFA89F8F575DEF8A3" box="[1361,1436,1826,1852]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="17" pageNumber="468">MNRJ</collectionCode>
34072 [2†].
</paragraph>
<paragraph id="8BDC369CBA7FDE2FFF1FF8BE7174FF66" blockId="17.[151,1437,1825,2032]" lastBlockId="18.[151,1436,151,573]" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<emphasis id="B917EA8EBA7FDE2CFF1FF8BE71D8F81C" bold="true" box="[199,410,1897,1923]" pageId="17" pageNumber="468">Characterization.</emphasis>
Shell tall, conical, spire angle 15°, reaching 3.0 mm long, 1.0 mm wide, eight whorls, colorless. Protoconch subcylindrical, dome-shaped, about 2.0 whorls, 320 μm high, surface smooth. Teleoconch with strongly convex whorls, surface glossy presenting faint microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl about 40% of the total length, base rounded; aperture high, about 67% of body whorl length, strongly expanded laterally, rounded anteriorly, acute posteriorly; outer lip thin, strongly orthocline, sinuous, moderately advanced close to distal area; inner lip thin, reflected. Wide umbilicus present, bordered by a rounded edge.
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA7CDE2FFF1FFED4739CFEFA" pageId="18" pageNumber="469" type="description">
<paragraph id="8BDC369CBA7CDE2FFF1FFED47088FEDE" blockId="18.[151,1436,151,573]" pageId="18" pageNumber="469">
<emphasis id="B917EA8EBA7CDE2FFF1FFED4713CFE82" bold="true" box="[199,382,259,285]" pageId="18" pageNumber="469">Measurements.</emphasis>
MNRJ 34072: 6 whorls; SL= 3.0 mm; BWL=
<quantity id="4C9B9B79BA7CDE2FFC55FED473A5FE82" box="[909,999,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="469" unit="mm" value="1.2">1.2 mm</quantity>
; AL=
<quantity id="4C9B9B79BA7CDE2FFBF5FED474C4FE82" box="[1069,1158,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="469" unit="mm" value="0.8">0.8 mm</quantity>
; SW= 1.0 mm; AW=
<quantity id="4C9B9B79BA7CDE2FFAA1FED47084FEDE" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="469" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="8BDC369CBA7CDE2FFF1FFE9D739CFEFA" blockId="18.[151,1436,151,573]" box="[199,990,330,357]" pageId="18" pageNumber="469">
<emphasis id="B917EA8EBA7CDE2FFF1FFE9D71B6FEFB" bold="true" box="[199,500,330,356]" pageId="18" pageNumber="469">Geographic Distribution.</emphasis>
<materialsCitation id="3B0B3CC1BA7CDE2FFE22FE9C7338FEFA" box="[506,890,331,357]" country="Brazil" elevation="71" location="From" pageId="18" pageNumber="469" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="F374760CBA7CDE2FFE22FE9C7201FEFA" box="[506,579,331,357]" name="Brazil" pageId="18" pageNumber="469">Brazil</collectingCountry>
:
<collectingRegion id="49A7F87EBA7CDE2FFD96FE9C72B6FEFA" box="[590,756,331,357]" country="Brazil" name="Rio de Janeiro" pageId="18" pageNumber="469">Rio de Janeiro</collectingRegion>
.
<location id="8EBC6047BA7CDE2FFD25FE9C7378FEFA" LSID="urn:lsid:plazi:treatment:03CA878ABA7FDE2FFF4FF9627503FDA2:8EBC6047BA7CDE2FFD25FE9C7378FEFA" box="[765,826,331,357]" country="Brazil" name="From" pageId="18" pageNumber="469" stateProvince="Rio de Janeiro">From</location>
<quantity id="4C9B9B79BA7CDE2FFC99FE9C7338FEFA" box="[833,890,331,357]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="18" pageNumber="469" unit="m" value="71.0">
<elevation id="004ED1AFBA7CDE2FFC99FE9C7338FEFA" box="[833,890,331,357]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="18" pageNumber="469" unit="m" value="71.0">71 m</elevation>
</quantity>
</materialsCitation>
to
<specimenCount id="9D65FD15BA7CDE2FFC46FE9C739CFEFA" box="[926,990,331,357]" count="73" pageId="18" pageNumber="469" type="generic">
<quantity id="4C9B9B79BA7CDE2FFC46FE9C7398FEFA" box="[926,986,331,357]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="18" pageNumber="469" unit="m" value="73.0">73 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA7CDE2FFF1FFEB87503FDA2" pageId="18" pageNumber="469" type="discussion">
<paragraph id="8BDC369CBA7CDE2FFF1FFEB87503FDA2" blockId="18.[151,1436,151,573]" pageId="18" pageNumber="469">
<emphasis id="B917EA8EBA7CDE2FFF1FFEB87179FE16" bold="true" box="[199,315,367,393]" pageId="18" pageNumber="469">Remarks.</emphasis>
These shells (
<figureCitation id="13582A19BA7CDE2FFE3BFEB87268FE16" box="[483,554,367,393]" captionStart="FIGURE 9" captionStartId="17.[151,250,1539,1563]" captionTargetBox="[151,1435,273,1515]" captionTargetId="figure-67@17.[151,1435,273,1515]" captionTargetPageId="17" captionText="FIGURE 9. Aclis cf. pulchra: MNRJ 34072. A, C. Shell in frontal view. B. Shell in lateral view. D. Detail of teleoconch surface. E. Detail of apical whorls. F–G. Detail of body whorl in frontal view. Scale bars: A–C. 1 mm; D. 20 μm; E. 200 μm; F–G. 500 μm." figureDoi="http://doi.org/10.5281/zenodo.10954447" httpUri="https://zenodo.org/record/10954447/files/figure.png" pageId="18" pageNumber="469">Fig. 9</figureCitation>
) are similar to
<taxonomicName id="4C634D1FBA7CDE2FFD07FEB8731AFE16" authority="Souza &amp; Araújo &amp; Pimenta, 2024" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" box="[735,856,367,393]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="pulchra" status="sp. nov">
<emphasis id="B917EA8EBA7CDE2FFD07FEB8731AFE16" box="[735,856,367,393]" italics="true" pageId="18" pageNumber="469">A. pulchra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457F5BA7CDE2FFCBAFEB873F7FE16" box="[866,949,367,393]" pageId="18" pageNumber="469" rank="species">sp. nov</taxonomicNameLabel>
(
<figureCitation id="13582A19BA7CDE2FFC1FFEB87451FE16" box="[967,1043,367,393]" captionStart="FIGURE 7" captionStartId="15.[151,246,1943,1967]" captionTargetBox="[162,1425,181,1918]" captionTargetId="figure-17@15.[162,1425,181,1918]" captionTargetPageId="15" captionText="FIGURE 7. Aclis pulchra sp. nov. A–C, F–I. Holotype, MNRJ 16825. D–E, J. paratype, MNRJ 26601. A, C–D. Shell in frontal view. B, E. Shell in lateral view. F–G. Detail of teleoconch surface; square in F indicates detail in G. H. Detail of body whorl in frontal view. I. Detail of apical whorls. J. Protoconch in apical view. Scale bars: A–E. 1 mm; F–G. 50 μm; H. 500 μm; I–J. 200 μm." figureDoi="http://doi.org/10.5281/zenodo.10954440" httpUri="https://zenodo.org/record/10954440/files/figure.png" pageId="18" pageNumber="469">Figs 7</figureCitation>
,
<figureCitation id="13582A19BA7CDE2FFBF9FEB8746DFE16" box="[1057,1071,367,393]" captionStart="FIGURE 8" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[162,1425,181,1919]" captionTargetId="figure-23@16.[162,1425,181,1919]" captionTargetPageId="16" captionText="FIGURE 8. Aclis pulchra sp. nov. A. Paratype, MNRJ 16323. B, F, H. Paratype, MNRJ 17713. C. Paratype, MNRJ 30872. D, G, I. paratype, MORG 39469. E. Paratype, MNRJ 26601. A–D. Shell in frontal view. E–G. Detail of apical whorls. H–I. Detail of body whorl in frontal view. Scale bars: A–D. 1 mm; E–G. 200 μm; H–I. 500 μm." figureDoi="http://doi.org/10.5281/zenodo.10954443" httpUri="https://zenodo.org/record/10954443/files/figure.png" pageId="18" pageNumber="469">8</figureCitation>
) described herein in relation to shell shape, dimensions of the protoconch and convexity of teleoconch whorls. However, shells with this tentative identification (‘cf.’) reach a higher number of whorls, has a more regular increase in diameter, which results in a more turriform shape, besides a less advanced outer lip, a slightly smaller aperture compared to the total length (0.25
<emphasis id="B917EA8EBA7CDE2FFF01FE2870B5FD87" box="[217,247,511,536]" italics="true" pageId="18" pageNumber="469">vs.</emphasis>
0.30) and a slightly narrower umbilicus. We avoid describing these specimens as a new species due to the scarcity of studied material that hampers a proper comparison of shape variability with
<taxonomicName id="4C634D1FBA7CDE2FFBB2FDF474A2FDA2" authority="Souza &amp; Araújo &amp; Pimenta, 2024" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" box="[1130,1248,547,573]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="pulchra" status="sp. nov.">
<emphasis id="B917EA8EBA7CDE2FFBB2FDF474A2FDA2" box="[1130,1248,547,573]" italics="true" pageId="18" pageNumber="469">A. pulchra</emphasis>
</taxonomicName>
<emphasis id="B917EA8EBA7CDE2FFB3FFDF47503FDA2" bold="true" box="[1255,1345,547,573]" pageId="18" pageNumber="469">
<taxonomicNameLabel id="A22457F5BA7CDE2FFB3FFDF47503FDA2" box="[1255,1345,547,573]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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